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<document id="6C9067EA46EB93A443AA1B2EFC3B88C8" ID-DOI="10.5281/zenodo.5176618" ID-ISSN="0374-1036" ID-Zenodo-Dep="5176618" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628601973277" checkinUser="carolina" docAuthor="Hájek, Jiří &amp; Šťastný, Jaroslav" docDate="2005" docId="FE0987CFFFFDE93EFE06EF87FEC11550" docLanguage="en" docName="ActaEntMusNatPra.45.51-58.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 45" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Laccophilus uncletan Hájek &amp; Šťastný 2005, sp. nov." docType="treatment" docVersion="3" lastPageNumber="54" masterDocId="0230FFB7FFFFE93DFFDCEB58FFFA1347" masterDocTitle="Two new species of Laccophilus Leach, 1815 (Coleoptera: Dytiscidae) from India and Borneo with notes on other species of the genus" masterLastPageNumber="58" masterPageNumber="51" pageNumber="53" updateTime="1699047269640" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="275C2DD4A9D85E87A2058C5DE9A5D35C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0FD6C0F0C1099698DB699ED2F5C714D8">
<mods:title id="3C98FD8625F68B9EB68D348E83A769B9">Two new species of Laccophilus Leach, 1815 (Coleoptera: Dytiscidae) from India and Borneo with notes on other species of the genus</mods:title>
</mods:titleInfo>
<mods:name id="74439810FFC6312312B7D37CC19A3E4B" type="personal">
<mods:role id="0D1160121A3348A2B4185424225224BA">
<mods:roleTerm id="AF0AA320DF1A51F47B198082FB9E7891">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3041B689C3FC015328834F9957C6F209">Hájek, Jiří</mods:namePart>
<mods:affiliation id="A1913C77CE2CD28463A29CB7EBC967E1">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: jiri _ hajek @ nm. cz</mods:affiliation>
</mods:name>
<mods:name id="29A1CF71C960F0FE72D8737571F662B4" type="personal">
<mods:role id="7B525D9CADD41515FA494BEA5D92D0F6">
<mods:roleTerm id="8BBDA3247C5B268A1F7BDCA3F98F1F12">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB6AE19C87E66D01A42F7A4989843357">Šťastný, Jaroslav</mods:namePart>
<mods:affiliation id="4D7C2856BD6FD740E5C25720F16D51A4">Kosmonautů 359, CZ- 460 05 Liberec, Czech Republic; e-mail: stastnyj @ jergym. hiedu. cz</mods:affiliation>
</mods:name>
<mods:typeOfResource id="96DA3FF7FC586A4031D556A8BDDDB712">text</mods:typeOfResource>
<mods:relatedItem id="46A691BDFA93395B8C12941D23A9D1F9" type="host">
<mods:titleInfo id="A8853146813E613ED44D2CAAB3435712">
<mods:title id="088E85DA07765F59375D5873093618A1">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="8FD8EE6ED293C7EEF8585FD5BA8F70E0">
<mods:date id="87FF21EECC15CD10EB25848919184514">2005</mods:date>
<mods:detail id="C32AFB8A2EDE411F6C7E887F30AE2300" type="pubDate">
<mods:number id="4D50D90A06A6BF8F3EE516998C29BEAE">2005-12-31</mods:number>
</mods:detail>
<mods:detail id="9AD4AF6C3DA36F1E7715F3AAE58C22BD" type="volume">
<mods:number id="1FB880499DDA60B48DFA8856E27839A1">45</mods:number>
</mods:detail>
<mods:extent id="99F7E8CABCABE693C14006BC631FF28B" unit="page">
<mods:start id="60CC192A406A5F9726F528FFB73328F1">51</mods:start>
<mods:end id="B9F7A20B5127DAA83BDF48224180A1DC">58</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8D3A2B154F01A6A83F6C54C9B034E676">journal article</mods:classification>
<mods:identifier id="CDAF54B819995B6D63760DCCF5167F89" type="DOI">10.5281/zenodo.5176618</mods:identifier>
<mods:identifier id="97ADBCF84D7A28A1B133AAB7960F19DE" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="51525D96DDA03F8C8806129CC3DB67B4" type="Zenodo-Dep">5176618</mods:identifier>
</mods:mods>
<treatment id="FE0987CFFFFDE93EFE06EF87FEC11550" LSID="urn:lsid:plazi:treatment:FE0987CFFFFDE93EFE06EF87FEC11550" httpUri="http://treatment.plazi.org/id/FE0987CFFFFDE93EFE06EF87FEC11550" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53">
<subSubSection id="3EBA6552FFFDE93FFE06EF87FCDE17B0" box="[474,804,1247,1271]" pageId="2" pageNumber="53" type="nomenclature">
<paragraph id="761F36D9FFFDE93FFE06EF87FCDE17B0" blockId="2.[474,804,1247,1271]" box="[474,804,1247,1271]" pageId="2" pageNumber="53">
<heading id="2D5781B5FFFDE93FFE06EF87FCDE17B0" bold="true" box="[474,804,1247,1271]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="53" reason="3">
<emphasis id="44D4EACBFFFDE93FFE06EF87FCDE17B0" bold="true" box="[474,804,1247,1271]" pageId="2" pageNumber="53">
<taxonomicName id="B1A04D5AFFFDE93FFE06EF87FD3C17B0" ID-CoL="3RMCH" authority="Hájek &amp; Šťastný, 2005" authorityName="Hájek &amp; Šťastný" authorityYear="2005" box="[474,710,1247,1271]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="uncletan" status="sp. nov.">
<emphasis id="44D4EACBFFFDE93FFE06EF87FD3C17B0" bold="true" box="[474,710,1247,1271]" italics="true" pageId="2" pageNumber="53">Laccophilus uncletan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FE757B0FFFDE93FFD12EF87FCDE17B0" box="[718,804,1247,1271]" pageId="2" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFDE93FFD8CEFA7FD541655" box="[592,686,1279,1298]" pageId="2" pageNumber="53" type="description">
<paragraph id="761F36D9FFFDE93FFD8CEFA7FD541655" blockId="2.[151,742,1279,1333]" box="[592,686,1279,1298]" pageId="2" pageNumber="53">
(
<figureCitation id="EE9B2A5CFFFDE93FFD8AEFA7FD5D1655" box="[598,679,1279,1298]" captionStart="Figs 1-4. 1, 3" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[167,1112,215,1451]" captionTargetId="figure-9@4.[167,1112,215,1451]" captionTargetPageId="4" captionText="Figs 1-4. 1, 3 – Laccophilus boukali sp. nov. 1 – habitus; 3 – median lobe in lateral view. 2, 4 – L. uncletan sp. nov. 2 – habitus; 4 – aedeagus (a – median lobe in lateral view; b – apex of median lobe in dorsal view; c – left paramere; d – right paramere). Scale bars: 2 mm (Figs 1-2); 0.5 mm (Figs 3-4)." figureDoi="http://doi.org/10.5281/zenodo.5176620" httpUri="https://zenodo.org/record/5176620/files/figure.png" pageId="2" pageNumber="53">Figs. 2, 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFDE93FFF4BEE7AFD1C1672" box="[151,742,1314,1333]" pageId="2" pageNumber="53" type="reference_group">
<paragraph id="761F36D9FFFDE93FFF4BEE7AFD1C1672" blockId="2.[151,742,1279,1333]" box="[151,742,1314,1333]" pageId="2" pageNumber="53">
<treatmentCitationGroup id="56B011F7FFFDE93FFF4BEE7AFD1C1672" box="[151,742,1314,1333]" pageId="2" pageNumber="53">
?
<treatmentCitation id="F70110C8FFFDE93FFF7DEE7AFD691672" author="BALKE M. &amp; MAZZOLDI P. &amp; HENDRICH L." box="[161,659,1314,1333]" page="72" pageId="2" pageNumber="53" year="1998">
<taxonomicName id="B1A04D5AFFFDE93FFF7DEE7AFD691672" ID-CoL="3RM63" authority="Balke, Mazzoldi &amp; Hendrich, 1998: 72" authorityName="Balke, Mazzoldi &amp; Hendrich" authorityPageNumber="72" authorityYear="1998" box="[161,659,1314,1333]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="chini">
<emphasis id="44D4EACBFFFDE93FFF7DEE7AFEC31672" box="[161,313,1314,1333]" italics="true" pageId="2" pageNumber="53">Laccophilus chini</emphasis>
<bibRefCitation id="12314B28FFFDE93FFE9CEE7AFD691672" author="BALKE M. &amp; MAZZOLDI P. &amp; HENDRICH L." box="[320,659,1314,1333]" pageId="2" pageNumber="53" pagination="71 - 77" refId="ref3715" refString="BALKE M., MAZZOLDI P. &amp; HENDRICH L. 1998: Two new Laccophilus species (Coleoptera: Dytiscidae) from Southeast Asia, and notes on other species of the genus. Raffles Bulletin of Zoology 46: 71 - 77." type="journal article" year="1998">Balke, Mazzoldi &amp; Hendrich, 1998: 72</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(partim).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFDE93FFF4BEE0FFD9C1568" pageId="2" pageNumber="53" type="materials_examined">
<paragraph id="761F36D9FFFDE93FFF4BEE0FFC6A1629" blockId="2.[151,1127,1367,1583]" box="[151,912,1367,1391]" pageId="2" pageNumber="53">
<emphasis id="44D4EACBFFFDE93FFF4BEE0FFED41628" bold="true" box="[151,302,1367,1391]" pageId="2" pageNumber="53">
<typeStatus id="A91B887BFFFDE93FFF4BEE0FFF341628" box="[151,206,1367,1391]" pageId="2" pageNumber="53">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="0EB77649FFFDE93FFEE8EE0FFE621628" box="[308,408,1367,1391]" name="Malaysia" pageId="2" pageNumber="53">Malaysia</collectingCountry>
,
<collectingRegion id="B464F83BFFFDE93FFE7FEE0FFE1C1628" box="[419,486,1367,1391]" country="Malaysia" name="Sabah" pageId="2" pageNumber="53">Sabah</collectingRegion>
, Kinabatangan river, Uncle Tan’s camp.
</paragraph>
<paragraph id="761F36D9FFFDE93FFF4BEE2FFD9C1568" blockId="2.[151,1127,1367,1583]" pageId="2" pageNumber="53">
<emphasis id="44D4EACBFFFDE93FFF4BEE2FFEC616C8" bold="true" box="[151,316,1399,1423]" pageId="2" pageNumber="53">Type material.</emphasis>
<materialsCitation id="C6C83C84FFFDE93FFE9EEE2FFD9116E8" collectorName="J. Stastny" country="Malaysia" location="Sabah" pageId="2" pageNumber="53" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="A91B887BFFFDE93FFE9EEE2FFE5316CA" box="[322,425,1399,1422]" pageId="2" pageNumber="53" type="holotype">HOLOTYPE</typeStatus>
: J, ‘
<collectingCountry id="0EB77649FFFDE93FFE06EE2FFDB816C8" box="[474,578,1399,1423]" name="Malaysia" pageId="2" pageNumber="53">Malaysia</collectingCountry>
,
<collectingRegion id="B464F83BFFFDE93FFD91EE2FFD7416C8" box="[589,654,1399,1423]" country="Malaysia" name="Sabah" pageId="2" pageNumber="53">Sabah</collectingRegion>
/ Kinabatangan riv.[er] / 8.-15.6.[20]03, Uncle Tan’s / camp,
<collectorName id="DB55530FFFFDE93FFE84EECFFE3916E8" box="[344,451,1425,1455]" pageId="2" pageNumber="53">J. Šťastný</collectorName>
lgt. [printed] //
</materialsCitation>
<materialsCitation id="C6C83C84FFFDE93FFDAFEECFFC961688" collectionCode="NMPC" country="Malaysia" determinerName="J. Hajek &amp; J. Stastny" pageId="2" pageNumber="53" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="A91B887BFFFDE93FFDAFEECFFD0516E8" box="[627,767,1431,1455]" pageId="2" pageNumber="53" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="B1A04D5AFFFDE93FFCCBEECFFC3416E8" authorityName="Leach" authorityYear="1815" box="[791,974,1431,1455]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">LACCOPHILUS</taxonomicName>
/ uncletan
<taxonomicNameLabel id="5FE757B0FFFDE93FFB94EECFFF3F1689" pageId="2" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
/
<determinerName id="83C3DA1DFFFDE93FFF07EEEFFEC91688" box="[219,307,1463,1487]" pageId="2" pageNumber="53">J. Hájek</determinerName>
&amp;
<determinerName id="83C3DA1DFFFDE93FFE84EEEFFE381688" box="[344,450,1457,1487]" pageId="2" pageNumber="53">J. Šťastný</determinerName>
det. 2005’ [printed, red label] (
<collectionCode id="10B1AE1CFFFDE93FFCC8EEEFFC9F1688" box="[788,869,1463,1487]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="53">NMPC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="A91B887BFFFDE93FFCA4EEEFFC1E168A" box="[888,996,1463,1486]" pageId="2" pageNumber="53" type="paratype">PARATYPES</typeStatus>
: 2JJ
<specimenCount id="60A6FD50FFFDE93FFBF1EEEFFBA21697" box="[1069,1112,1460,1488]" pageId="2" pageNumber="53" type="female">5♀♀</specimenCount>
, same label data as
<typeStatus id="A91B887BFFFDE93FFEBAEE8FFE3916A8" box="[358,451,1495,1519]" pageId="2" pageNumber="53" type="holotype">holotype</typeStatus>
(
<collectionCode id="10B1AE1CFFFDE93FFE0FEE8FFDE816A8" box="[467,530,1495,1519]" pageId="2" pageNumber="53">JSCL</collectionCode>
,
<collectionCode id="10B1AE1CFFFDE93FFDC3EE8FFD9716A8" box="[543,621,1495,1519]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="53">NMPC</collectionCode>
,
<collectionCode id="10B1AE1CFFFDE93FFDA5EE8FFD3816A8" box="[633,706,1495,1519]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="53">NMW</collectionCode>
). Each
<typeStatus id="A91B887BFFFDE93FFCCFEE8FFC9416A8" box="[787,878,1495,1519]" pageId="2" pageNumber="53" type="paratype">paratype</typeStatus>
is provided with a red label similar to that of the
<typeStatus id="A91B887BFFFDE93FFE68EEAFFDE91548" box="[436,531,1527,1551]" pageId="2" pageNumber="53" type="holotype">holotype</typeStatus>
, except ‘PARATYPE’ instead of ‘HOLOTYPE’, the respective sex symbol and collection number.
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFCE93EFFADEB87FE951710" pageId="3" pageNumber="54" type="description">
<paragraph id="761F36D9FFFCE93EFFADEB87FDDE1270" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">
<emphasis id="44D4EACBFFFCE93EFFADEB87FF0113B0" bold="true" box="[113,251,223,247]" pageId="3" pageNumber="54">Description.</emphasis>
Oval and oblong; head testaceous, pronotum testaceous with blackish basal band and two subbasal spots, elytra blackish with testaceous pattern as depicted in
<figureCitation id="EE9B2A5CFFFCE93EFC26EBA7FBC71250" box="[1018,1085,255,279]" captionStart="Figs 1-4. 1, 3" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[167,1112,215,1451]" captionTargetId="figure-9@4.[167,1112,215,1451]" captionTargetPageId="4" captionText="Figs 1-4. 1, 3 – Laccophilus boukali sp. nov. 1 – habitus; 3 – median lobe in lateral view. 2, 4 – L. uncletan sp. nov. 2 – habitus; 4 – aedeagus (a – median lobe in lateral view; b – apex of median lobe in dorsal view; c – left paramere; d – right paramere). Scale bars: 2 mm (Figs 1-2); 0.5 mm (Figs 3-4)." figureDoi="http://doi.org/10.5281/zenodo.5176620" httpUri="https://zenodo.org/record/5176620/files/figure.png" pageId="3" pageNumber="54">Fig. 2</figureCitation>
. Ventral parts and appendages testaceous.
</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CEA18FCD61230" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">
Measurements. Body length
<quantity id="B1589B3CFFFCE93EFE1FEA67FDA21210" box="[451,600,319,343]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="3" pageNumber="54" unit="mm" value="3.65" valueMax="3.8" valueMin="3.5">3.50-3.80 mm</quantity>
(mean value =
<quantity id="B1589B3CFFFCE93EFCDDEA67FC991210" box="[769,867,319,343]" metricMagnitude="-3" metricUnit="m" metricValue="3.62" pageId="3" pageNumber="54" unit="mm" value="3.62">3.62 mm</quantity>
,
<typeStatus id="A91B887BFFFCE93EFCB2EA67FC311210" box="[878,971,319,343]" pageId="3" pageNumber="54" type="holotype">holotype</typeStatus>
<quantity id="B1589B3CFFFCE93EFC0EEA67FBCF1210" box="[978,1077,319,343]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" pageId="3" pageNumber="54" unit="mm" value="3.65">3.65 mm</quantity>
), width
<quantity id="B1589B3CFFFCE93EFF68EA07FEB31230" box="[180,329,351,375]" metricMagnitude="-3" metricUnit="m" metricValue="1.89" metricValueMax="1.95" metricValueMin="1.83" pageId="3" pageNumber="54" unit="mm" value="1.89" valueMax="1.95" valueMin="1.83">1.83-1.95 mm</quantity>
(mean value =
<quantity id="B1589B3CFFFCE93EFE33EA07FDAA1230" box="[495,592,351,375]" metricMagnitude="-3" metricUnit="m" metricValue="1.88" pageId="3" pageNumber="54" unit="mm" value="1.88">1.88 mm</quantity>
,
<typeStatus id="A91B887BFFFCE93EFD86EA07FD4D1230" box="[602,695,351,375]" pageId="3" pageNumber="54" type="holotype">holotype</typeStatus>
<quantity id="B1589B3CFFFCE93EFD61EA07FCDA1230" box="[701,800,351,375]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="3" pageNumber="54" unit="mm" value="1.85">1.85 mm</quantity>
).
</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CEA27FBE71290" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">Head. Coloration testaceous with indistinct darker spots. Reticulation double, composed of almost imperceptible polygonal meshes, and microreticulation of fine isodiametric meshes. Frons between eyes and clypeus with several large punctures. Mouthparts testaceous.</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CEAB8FCD91110" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">Pronotum. Coloration testaceous with thin basal blackish band and two subbasal blackish spots. Reticulation double, composed of slightly impressed large polygonal meshes and microreticulation of fine isodiametric meshes similar to that of head. Anterior margin, basal margin and posterior angles with numerous setigerous punctures.</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CE907FCD011B0" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">Elytra. Coloration blackish with testaceous markings comprising subbasal transverse sinuous band, mediolateral spot, subapical transverse sinuous band and apical spot. Sutural part of elytra blackish; in some specimens all testaceous markings connected in epipleural part. Reticulation on whole elytra distinctly double, similar to that of pronotum. Each elytron with sutural, discal and lateral row of large serial setigerous punctures.</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CE9A7FF341030" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">Ventral side. Testaceous and microreticulated. Prosternal process long and acute. Abdominal sternites laterally with oblique grooves and several large punctures. Posterior margin of anal sternite prominent medially with cluster of setigerous punctures, and moderately incised laterally.</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CE8D8FF481090" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">Legs. Completely testaceous. Male tarsomeres 1-3 of fore and middle legs only weakly dilated with microtrichia on ventral side. Claws of fore as well as middle tarsi simple and equal.</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CE887FCBB10B0" blockId="3.[113,1090,223,1559]" box="[144,833,991,1015]" pageId="3" pageNumber="54">
Male genitalia. Shape of median lobe and parameres as in
<figureCitation id="EE9B2A5CFFFCE93EFD20E887FCC710B0" box="[764,829,991,1015]" captionStart="Figs 1-4. 1, 3" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[167,1112,215,1451]" captionTargetId="figure-9@4.[167,1112,215,1451]" captionTargetPageId="4" captionText="Figs 1-4. 1, 3 – Laccophilus boukali sp. nov. 1 – habitus; 3 – median lobe in lateral view. 2, 4 – L. uncletan sp. nov. 2 – habitus; 4 – aedeagus (a – median lobe in lateral view; b – apex of median lobe in dorsal view; c – left paramere; d – right paramere). Scale bars: 2 mm (Figs 1-2); 0.5 mm (Figs 3-4)." figureDoi="http://doi.org/10.5281/zenodo.5176620" httpUri="https://zenodo.org/record/5176620/files/figure.png" pageId="3" pageNumber="54">Fig. 4</figureCitation>
.
</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CE8A7FE951710" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">Female. Generally similar to male. Posterior margin of anal sternite laterally only slightly incised, medially more prominent than in male, with truncate apex. Tarsomeres of fore and middle tarsi not dilated.</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFCE93EFFADEF07FE8C1650" pageId="3" pageNumber="54" type="diagnosis">
<paragraph id="761F36D9FFFCE93EFFADEF07FE8C1650" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">
<emphasis id="44D4EACBFFFCE93EFFADEF07FE931730" bold="true" box="[113,361,1119,1143]" pageId="3" pageNumber="54">Differential diagnosis.</emphasis>
Based on the double elytral reticulation and the long and acute prosternal process, the new species is classified into the
<taxonomicName id="B1A04D5AFFFCE93EFD54EF27FC8F17D0" authorityName="Sharp" authorityYear="1882" box="[648,885,1151,1175]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="44D4EACBFFFCE93EFD54EF27FC8F17D0" box="[648,885,1151,1175]" italics="true" pageId="3" pageNumber="54">Laccophilus siamensis</emphasis>
</taxonomicName>
-group sensu BRAN- CUCCI (1983). Its sister species is
<taxonomicName id="B1A04D5AFFFCE93EFE13EFF8FC6517F0" authority="Balke, Mazzoldi &amp; Hendrich, 1998" authorityName="Balke, Mazzoldi &amp; Hendrich" authorityYear="1998" box="[463,927,1183,1207]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="chini">
<emphasis id="44D4EACBFFFCE93EFE13EFF8FDDA17F0" box="[463,544,1183,1207]" italics="true" pageId="3" pageNumber="54">L. chini</emphasis>
<bibRefCitation id="12314B28FFFCE93EFDFBEFC7FC6517F0" author="BALKE M. &amp; MAZZOLDI P. &amp; HENDRICH L." box="[551,927,1183,1207]" pageId="3" pageNumber="54" pagination="71 - 77" refId="ref3715" refString="BALKE M., MAZZOLDI P. &amp; HENDRICH L. 1998: Two new Laccophilus species (Coleoptera: Dytiscidae) from Southeast Asia, and notes on other species of the genus. Raffles Bulletin of Zoology 46: 71 - 77." type="journal article" year="1998">Balke, Mazzoldi &amp; Hendrich, 1998</bibRefCitation>
</taxonomicName>
, from which
<taxonomicName id="B1A04D5AFFFCE93EFBF7EFF8FF361790" authority="Hájek &amp; Šťastný, 2005" authorityName="Hájek &amp; Šťastný" authorityYear="2005" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="uncletan" status="sp. nov.">
<emphasis id="44D4EACBFFFCE93EFBF7EFF8FF361790" italics="true" pageId="3" pageNumber="54">L. uncletan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FE757B0FFFCE93EFF0FEF98FEDC1790" box="[211,294,1216,1239]" pageId="3" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
can be reliably distinguished only by the male genitalia. In
<taxonomicName id="B1A04D5AFFFCE93EFC78EF98FBE11790" authority="Hájek &amp; Šťastný, 2005" authorityName="Hájek &amp; Šťastný" authorityYear="2005" box="[932,1051,1215,1239]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="uncletan" status="sp. nov.">
<emphasis id="44D4EACBFFFCE93EFC78EF98FBE11790" box="[932,1051,1215,1239]" italics="true" pageId="3" pageNumber="54">L. uncletan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FE757B0FFFCE93EFBFEEF98FF6517B0" pageId="3" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
the median lobe is more slender, ventrally without a subbasal rectangular spur, and has an almost rounded apex.
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFCE93EFFADEE47FC3316F0" pageId="3" pageNumber="54" type="distribution">
<paragraph id="761F36D9FFFCE93EFFADEE47FCDC1670" blockId="3.[113,1090,223,1559]" box="[113,806,1311,1335]" pageId="3" pageNumber="54">
<emphasis id="44D4EACBFFFCE93EFFADEE47FEF81670" bold="true" box="[113,258,1311,1335]" pageId="3" pageNumber="54">Distribution.</emphasis>
Known only from northern
<collectingRegion id="B464F83BFFFCE93EFDF2EE47FD5C1670" box="[558,678,1311,1335]" country="Malaysia" name="Sabah" pageId="3" pageNumber="54">Sabah state</collectingRegion>
(
<collectingCountry id="0EB77649FFFCE93EFD68EE47FCE11670" box="[692,795,1311,1335]" name="Malaysia" pageId="3" pageNumber="54">Malaysia</collectingCountry>
).
</paragraph>
<paragraph id="761F36D9FFFCE93EFFADEE67FC3316F0" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">
<emphasis id="44D4EACBFFFCE93EFFADEE67FF261610" bold="true" box="[113,220,1343,1367]" pageId="3" pageNumber="54">Bionomy.</emphasis>
The specimens were collected in primary lowland forest in a shallow oxbow of the river Kinabatangan together with
<taxonomicName id="B1A04D5AFFFCE93EFE0AEE38FCFC1630" authority="Brancucci, 1983" authorityName="Brancucci" authorityYear="1983" box="[470,774,1375,1399]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="bacchusi">
<emphasis id="44D4EACBFFFCE93EFE0AEE38FDAA1630" box="[470,592,1375,1399]" italics="true" pageId="3" pageNumber="54">L. bacchusi</emphasis>
<bibRefCitation id="12314B28FFFCE93EFD8BEE07FCFC1630" author="BRANCUCCI M." box="[599,774,1375,1399]" pageId="3" pageNumber="54" pagination="241 - 426" refId="ref3760" refString="BRANCUCCI M. 1983: Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey 31 / 32: 241 - 426." type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B1A04D5AFFFCE93EFCCDEE38FBBB1630" authority="Regimbart, 1880" authorityName="Regimbart" authorityYear="1880" box="[785,1089,1375,1399]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="ritsemae">
<emphasis id="44D4EACBFFFCE93EFCCDEE38FC721630" box="[785,904,1376,1399]" italics="true" pageId="3" pageNumber="54">L. ritsemae</emphasis>
Régimbart, 1880
</taxonomicName>
and
<emphasis id="44D4EACBFFFCE93EFF42EED8FF3116D0" box="[158,203,1408,1431]" italics="true" pageId="3" pageNumber="54">L. s.</emphasis>
<taxonomicName id="B1A04D5AFFFCE93EFF0EEED8FE3816D0" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[210,450,1407,1431]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="44D4EACBFFFCE93EFF0EEED8FEC316D0" box="[210,313,1408,1431]" italics="true" pageId="3" pageNumber="54">siamensis</emphasis>
Sharp, 1882
</taxonomicName>
. They inhabited muddy water near the banks of small drains flowing to the oxbow. The substrate was silty with accumulated decaying leaves.
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFCE93EFFADEEE7FEC11550" pageId="3" pageNumber="54" type="etymology">
<paragraph id="761F36D9FFFCE93EFFADEEE7FEC11550" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">
<emphasis id="44D4EACBFFFCE93EFFADEEE7FF0B1690" bold="true" box="[113,241,1471,1495]" pageId="3" pageNumber="54">Etymology.</emphasis>
This species is dedicated to Mr. Tan Su Lim known as ‘Uncle Tan’, an enthusiastic supporter of ecology popularisation and a leading environmentalist in
<collectingRegion id="B464F83BFFFCE93EFCABEE87FC4116B0" box="[887,955,1503,1527]" country="Malaysia" name="Sabah" pageId="3" pageNumber="54">Sabah</collectingRegion>
. The noun is used in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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