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257 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.889.39090" ID-GBIF-Dataset="f50f6008-500c-492f-84b8-295ba991ec7b" ID-PMC="PMC6872603" ID-Pensoft-Pub="1313-2970-889-81" ID-Pensoft-UUID="0C546DE0BA425539879C1421D8C25A63" ID-PubMed="31777439" ID-ZooBank="7E7A3D196D7040398C087B248A27EB33" ModsDocID="1313-2970-889-81" checkinTime="1574358895821" checkinUser="pensoft" docAuthor="Hendrich, Lars, Shaverdo, Helena, Hajek, Jiri & Balke, Michael" docDate="2019" docId="4BC8C219282D5B35AA34375993E0145A" docLanguage="en" docName="ZooKeys 889: 81-152" docOrigin="ZooKeys 889" docSource="http://dx.doi.org/10.3897/zookeys.889.39090" docTitle="Copelatus martinbaehri Hendrich, Shaverdo, Hajek & Balke, 2019, sp. nov." docType="treatment" docVersion="4" id="0C546DE0BA425539879C1421D8C25A63" lastPageNumber="81" masterDocId="0C546DE0BA425539879C1421D8C25A63" masterDocTitle="Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="152" masterPageNumber="81" pageNumber="81" updateTime="1668168070667" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hendrich, Lars</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hajek, Jiri</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>889</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>81</mods:start>
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<mods:end>152</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.889.39090</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.889.39090</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-889-81</mods:identifier>
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<mods:identifier type="ZooBank">7E7A3D196D7040398C087B248A27EB33</mods:identifier>
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<mods:identifier type="Pensoft-UUID">0C546DE0BA425539879C1421D8C25A63</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="160511459" LSID="urn:lsid:plazi:treatment:4BC8C219282D5B35AA34375993E0145A" httpUri="http://treatment.plazi.org/id/4BC8C219282D5B35AA34375993E0145A" lastPageNumber="81" pageId="0" pageNumber="81">
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<subSubSection pageId="0" pageNumber="81" type="nomenclature">
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<paragraph pageId="0" pageNumber="81">
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<taxonomicName LSID="4BC8C219-282D-5B35-AA34-375993E0145A" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus martinbaehri" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="martinbaehri">Copelatus martinbaehri</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="81">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus and colouration of Copelatus martinbaehri sp. nov., paratype (Mount Tozer), male. Total length 6.7 mm." figureDoi="10.3897/zookeys.889.39090.figure7" httpUri="https://binary.pensoft.net/fig/356476" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure7">Figures 7</figureCitation>
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,
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Copelatus martinbaehri sp. nov., paratype from Mount Tozer A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure16" httpUri="https://binary.pensoft.net/fig/356485" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure16">16</figureCitation>
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,
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<figureCitation captionStart="Figures 27, 28" captionStartId="F26" captionText="Figures 27, 28. Distribution of 27 Copelatus tenebrosus 28 C. martinbaehri sp. nov." figureDoi="10.3897/zookeys.889.39090.figures27-28" httpUri="https://binary.pensoft.net/fig/356495" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figures27-28">28</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="type locality">
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<paragraph pageId="0" pageNumber="81">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="81">
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Moreguina (St
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<normalizedToken originalValue="Stephen’s">Stephen's</normalizedToken>
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Mission), Central Province, Papua New Guinea Central, 50 m,
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<geoCoordinate degrees="10.018104" direction="south" orientation="latitude" precision="1" value="-10.018104">10.018104S</geoCoordinate>
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,
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<geoCoordinate degrees="148.467793" direction="east" orientation="longitude" precision="1" value="148.46779">148.467793E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="type material">
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<paragraph pageId="0" pageNumber="81">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Holotype:</emphasis>
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Male, "Papua New Guinea Central, Moreguina, 18.viii.2008 Posman (PNG184)" [printed label]; "Holotype
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus martinbaehri" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="martinbaehri">
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<emphasis italics="true" pageId="0" pageNumber="81">Copelatus martinbaehri</emphasis>
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</taxonomicName>
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sp.n. Hendrich, Shaverdo, Hajek & Balke des. 2019 [red printed label] (ZSM).
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<emphasis bold="true" pageId="0" pageNumber="81">
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<emphasis italics="true" pageId="0" pageNumber="81">Paratypes (23 specimens)</emphasis>
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:
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<emphasis italics="true" pageId="0" pageNumber="81">Australia</emphasis>
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</emphasis>
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: 7 exs., "
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<geoCoordinate degrees="12.445" direction="south" orientation="latitude" precision="55" value="-12.445">12.445S</geoCoordinate>
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,
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<geoCoordinate degrees="143.14" direction="east" orientation="longitude" precision="555" value="143.14">143.14E</geoCoordinate>
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QLD 3 km ENE Mt. Tozer 28
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<normalizedToken originalValue="June–">June-</normalizedToken>
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4July 1986 D.H.Colless Malaise Trap" "ANIC Database No. 25 019356" [printed label]; "Aust. Nat. Ins. Coll." [green printed label] (ANIC, ZSM); 1 male, "N. Queensland IRON RANGE Gordon's CK. [Creek,
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<geoCoordinate degrees="12.715780" direction="south" orientation="latitude" precision="1" value="-12.71578">-12.715780</geoCoordinate>
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,
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<geoCoordinate degrees="143.302092" direction="east" orientation="longitude" precision="1" value="143.3021">143.302092</geoCoordinate>
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] 10.5.71 at light leg: J.G. Brooks" [hw on both label sides], "COLL. HENDRICH BERLIN" (NMW); 1 male, 1 female, "Gordon's CK [Creek,
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<geoCoordinate degrees="12.715780" direction="south" orientation="latitude" precision="1" value="-12.71578">-12.715780</geoCoordinate>
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,
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<geoCoordinate degrees="143.302092" direction="east" orientation="longitude" precision="1" value="143.3021">143.302092</geoCoordinate>
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] Iron RA [Range] N.Q. [North Queensland] 100' [100 m] 10.5.71 J.G. Brooks "at light" (CLH); 1 female, "Iron RA N.Q. 4.5.71 J.G. Brooks "at light" (CLH); 2 females, "Iron Range, Cape York Pen., N.-Qld 26
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<normalizedToken originalValue="May–">May-</normalizedToken>
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2 June 1971 B.K. Cantrell" (QM); 2 males, 2 females, "Iron Range, Cape York Pen. N.Q. 13.-20.V.1975 K.J. Houston At light" (QDPC, ZSM).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Papua New Guinea</emphasis>
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: 1 male, 1 female, same data as holotype, one male additionally with a green label "DNA M.Balke 3803" (ZSM). 2 females, "Papua New Guinea: Central, Moreguina, 16.viii.2008 Posman (PNG183)" [printed label] (ZSM). 1 male, "Papua New Guinea: Northern Kokoda, 410 m, i.2008, 53.4[?]81S 147 43.648E, Posman, (PNG 174)" (ZSM). All paratypes with our red printed labels.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="description">
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<paragraph pageId="0" pageNumber="81">Description of male holotype.</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Body shape:</emphasis>
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In dorsal view, oblong-oval, broadest in basal third of elytra, moderately convex. Body outline without discontinuity between pronotum and elytra. Head relatively broad; anterior margin of clypeus not bordered. Pronotum broadest between posterior angles, lateral margins moderately curved. Base of elytra as broad as pronotal base; lateral margins of elytra moderately curved (
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus and colouration of Copelatus martinbaehri sp. nov., paratype (Mount Tozer), male. Total length 6.7 mm." figureDoi="10.3897/zookeys.889.39090.figure7" httpUri="https://binary.pensoft.net/fig/356476" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure7">Fig. 7</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.889.39090.figure7" httpUri="https://binary.pensoft.net/fig/356476" pageId="0" pageNumber="81" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" pageId="0" pageNumber="81">Figure 7.</emphasis>
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Habitus and colouration of
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus martinbaehri" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="martinbaehri">
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<emphasis italics="true" pageId="0" pageNumber="81">Copelatus martinbaehri</emphasis>
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</taxonomicName>
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sp. nov., paratype (Mount Tozer), male. Total length 6.7 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Colouration:</emphasis>
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Body black, most of clypeus, anterior angles of pronotum, base and tip of elytra, appendages and much of ventral surface testaceous.
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</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Dorsal surface sculpture:</emphasis>
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Whole surface shiny (
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus and colouration of Copelatus martinbaehri sp. nov., paratype (Mount Tozer), male. Total length 6.7 mm." figureDoi="10.3897/zookeys.889.39090.figure7" httpUri="https://binary.pensoft.net/fig/356476" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure7">Fig. 7</figureCitation>
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). Head uniformly microreticulated, reticulation composed of moderately deeply impressed isodiametric very small meshes. Punctation composed of very small punctures spread sparsely on surface; rows of deep and coarse punctures present around inner margin of eyes and in small depression anterolaterally of eyes. Pronotum with some weak and short striae laterally. Microreticulation and punctation similar to that of head; row of coarse setigerous punctures present along anterior margin, basal margin (except for basomedially), and laterally close to sides. Elytra with microreticulation similar to that of head and pronotum, but less impressed. Punctation consisting of very fine sparse punctures. Each elytron with six strongly impressed discal and one submarginal longitudinal striae, intervals subequal, striae 1 and 5 tending to be shorter basally; submarginal stria reaching from little behind midlength of elytron almost to end of stria 6. Serial punctures on elytron untraceable.
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</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Antennae and legs:</emphasis>
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Antenna with antennomeres long and slender. Protibia modified, distinctly broadened anteriorly (2/3rd) and strongly narrowed basally (1/3rd). Pro- and mesotarsomeres 1-3 slightly broadened, with adhesive discs on their ventral side; claws simple.
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</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Ventral part:</emphasis>
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Finely microreticulated, with intermixed, sparsely distributed, very small punctures. Prosternal process rather flat, distinctly bordered laterally, weakly and bluntly pointed. Lateral parts of metaventrite tongue-shaped, very slender. Metacoxal lines close, well-marked and moderately divergent anteriorly. Metacoxae with long and deep striae, abdominal ventrite I with numerous striae and ventrites 2-3 with a few longitudinal striae.
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</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Male genitalia:</emphasis>
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Median lobe sickle-shaped, narrow, simple; in ventral view, appearing somewhat flattened; in lateral view, apically tapering and strongly curved downwards (
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Copelatus martinbaehri sp. nov., paratype from Mount Tozer A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure16" httpUri="https://binary.pensoft.net/fig/356485" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure16">Fig. 16B, C</figureCitation>
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). Shape of paramere broadly triangular, with weak, relatively short setae, mainly along dorsal margin of subdistal part (
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Copelatus martinbaehri sp. nov., paratype from Mount Tozer A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure16" httpUri="https://binary.pensoft.net/fig/356485" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure16">Fig. 16D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="female">
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<paragraph pageId="0" pageNumber="81">Female.</paragraph>
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<paragraph pageId="0" pageNumber="81">Similar to male in habitus. Protibia not modified. Pro- and mesotarsomeres not broadened, without adhesive setae.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="description">
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<paragraph pageId="0" pageNumber="81">Measurements.</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Holotype:</emphasis>
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TL = 6.35 mm; TL-H = 5.85 mm; MW = 3.1 mm.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Paratypes</emphasis>
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: TL = 6.2-6.75 mm; TL-H = 5.85-6.1 mm; MW = 3.1-3.2 mm.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="variability">
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<paragraph pageId="0" pageNumber="81">Variability.</paragraph>
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<paragraph pageId="0" pageNumber="81">All specimens studied are rather uniform in shape and size but vary a bit in the extension of the testaceous elytral markings.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="81">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="81">
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Based on the characteristic sickle-shaped median lobe, the new species belongs to a difficult complex of species distributed in the Sunda Islands and New Guinea, including
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<taxonomicName lsidName="C. geniculatus" pageId="0" pageNumber="81" rank="species" species="geniculatus">
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<emphasis italics="true" pageId="0" pageNumber="81">C. geniculatus</emphasis>
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</taxonomicName>
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Sharp, 1882,
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<taxonomicName lsidName="C. gentilis" pageId="0" pageNumber="81" rank="species" species="gentilis">
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<emphasis italics="true" pageId="0" pageNumber="81">C. gentilis</emphasis>
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</taxonomicName>
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Sharp, 1882,
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<taxonomicName lsidName="C. lineatus" pageId="0" pageNumber="81" rank="species" species="lineatus">
|
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<emphasis italics="true" pageId="0" pageNumber="81">C. lineatus</emphasis>
|
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</taxonomicName>
|
|||
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(
|
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|
|
<normalizedToken originalValue="Guérin-Méneville">Guerin-Meneville</normalizedToken>
|
|||
|
|
, 1838) and
|
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|
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<taxonomicName lsidName="C. subterraneus" pageId="0" pageNumber="81" rank="species" species="subterraneus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="81">C. subterraneus</emphasis>
|
|||
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</taxonomicName>
|
|||
|
|
<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
|
|||
|
|
, 1978 (of the
|
|||
|
|
<taxonomicName lsidName="C. irinus" pageId="0" pageNumber="81" rank="species" species="irinus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="81">C. irinus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species group) and several additional undescribed species, both from the
|
|||
|
|
<taxonomicName lsidName="C. irinus" pageId="0" pageNumber="81" rank="species" species="irinus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="81">C. irinus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
group (i.e., with six dorsal striae on elytra) and the
|
|||
|
|
<taxonomicName lsidName="C. trilobatus" pageId="0" pageNumber="81" rank="species" species="trilobatus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="81">C. trilobatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
group (with 11 dorsal striae). All those species are rather uniform in body shape and colouration; they differ in elytral striation (which may be, however, variable) and less so in the shape of the median lobe in lateral view, especially in the width of the medial part and in length and curvature of the apical part.
|
|||
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus martinbaehri" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="martinbaehri">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="81">Copelatus martinbaehri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. nov. differs from the other species of this complex by the shape of the median lobe, which has the central part in lateral view broader, but without distinct tubercle on the ventral side; additionally, the apical part is shorter and almost straight (the angle between central and apical part of median lobe in lateral view is nearly rectangular).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="81">
|
|||
|
|
Within other Australian species with six elytral striae,
|
|||
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus martinbaehri" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="martinbaehri">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="81">Copelatus martinbaehri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. nov. can be easily distinguished by its larger size (
|
|||
|
|
<taxonomicName lsidName="C. tenebrosus" pageId="0" pageNumber="81" rank="species" species="tenebrosus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="81">C. tenebrosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is always less than 5 mm), more elongate habitus (
|
|||
|
|
<taxonomicName lsidName="C. portior" pageId="0" pageNumber="81" rank="species" species="portior">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="81">C. portior</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
more ovoid, oval), elytral colouration (
|
|||
|
|
<taxonomicName lsidName="C. marginatus" pageId="0" pageNumber="81" rank="species" species="marginatus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="81">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName lsidName="C. tenebrosus" pageId="0" pageNumber="81" rank="species" species="tenebrosus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="81">C. tenebrosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
with almost black dorsal surface), and the shape of the median lobe.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="81" type="etymology">
|
|||
|
|
<paragraph pageId="0" pageNumber="81">Etymology.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="81">This species is dedicated in honour of our late colleague Dr Martin Baehr (*10.3.1943, †17.4.2019, Munich, Germany), coleopterist, arachnologist, and others as well as the most knowledgeable authority for Australian ground beetles. The specific epithet is a substantive in the genitive case.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="81" type="distribution">
|
|||
|
|
<paragraph pageId="0" pageNumber="81">Distribution.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="81">
|
|||
|
|
Northern Queensland (Iron Range National Park at Cape York Peninsula) and south-eastern Papua New Guinea (Central Province) (
|
|||
|
|
<figureCitation captionStart="Figures 27, 28" captionStartId="F26" captionText="Figures 27, 28. Distribution of 27 Copelatus tenebrosus 28 C. martinbaehri sp. nov." figureDoi="10.3897/zookeys.889.39090.figures27-28" httpUri="https://binary.pensoft.net/fig/356495" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figures27-28">Fig. 28</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="81" type="habitat">
|
|||
|
|
<paragraph pageId="0" pageNumber="81">Habitat.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="81">Unknown. Most probably, the new species is an inhabitant of temporary lowland rainforest pools. The type specimens were collected in a Malaise Trap and at light.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|