135 lines
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135 lines
18 KiB
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<document id="D489FC5B5724744E75627866C84B950C" ID="10.11646/zootaxa.4138.3.2" ID-DOI="10.11646/zootaxa.4138.3.2" ID-GBIF-Dataset="c785c9ac-b71c-4487-b770-35ebfe797c8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="258464" ID-ZooBank="DDF68118-EA96-49A5-B316-47814E19ED3D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468823189128" checkinUser="plazi" docAuthor="Leschen, Richard A. B., Escalona, Hermes E. & Elgueta, Mario" docDate="2016" docId="746C8797FFFFD003ACDDF997FEA895DB" docLanguage="en" docName="zootaxa.4138.3.2.pdf" docOrigin="Zootaxa 4138 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ulobostrichus Leschen and Escalona, gen. n." docType="treatment" docUuid="E02B4845-66BF-43EB-990F-02EF1F83185C" docUuidSource="ZooBank" docVersion="9" lastPageNumber="451" masterDocId="8855FFEFFFF6D009AC4AFF96FF8B9636" masterDocTitle="Phylogeny of the Gondwanan beetle family Ulodidae (Tenebrionoidea)" masterLastPageNumber="473" masterPageNumber="441" pageNumber="450" updateTime="1698365200336" updateUser="plazi">
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<mods:mods id="8541E930ACA7F54A9CA101A9DE0B954F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0DC62F13763168CD33149C6C68428540">
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<mods:title id="A85DBE395E7D359713148A2D85C0A622">Phylogeny of the Gondwanan beetle family Ulodidae (Tenebrionoidea)</mods:title>
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</mods:titleInfo>
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<mods:name id="007476D1FB8FDBC99ABB9A755172E55A" type="personal">
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<mods:role id="EE0759AB11F0435FA2992D62A2EA1CBD">
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<mods:roleTerm id="42CC0310F6351A83B86B7EF096D8762E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DBE624F01F1E89480224D1518C84CE43">Leschen, Richard A. B.</mods:namePart>
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</mods:name>
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<mods:name id="080F69E239C5EE8A9EE9C4DC385A03C3" type="personal">
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</mods:role>
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<mods:namePart id="D4B55779B8272E55BDE1CA8B3EAF77C8">Escalona, Hermes E.</mods:namePart>
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</mods:name>
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<mods:name id="AF8BB4E285608DD4DA05F07CF8E0751E" type="personal">
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<mods:roleTerm id="5903A7DE12DB9BCED16474EE200A372F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="68ECE6A3F40DB3DD495A7713976A052B">Elgueta, Mario</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="0694CB46840924031D5FC51D91B315FD">text</mods:typeOfResource>
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<mods:title id="CC19610AAF2C5141AE86EE0BE305A8E2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6389C8565C3AAC8DE85DF645B5B660D0">
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<mods:date id="8F2D4BB2BC6E864C0A86B8D2AC4F6802">2016</mods:date>
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<mods:detail id="6A6B79DCB4A1B11EA5FACEC89405BA10" type="volume">
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<mods:number id="0A314B8B0C5740E6EEDB2A1972206A44">4138</mods:number>
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</mods:detail>
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<mods:detail id="83B643E0310FE50412547106FA736D05" type="issue">
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<mods:number id="3A923C769B23E7D210E9E52586FF4A5B">3</mods:number>
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<mods:start id="878A3AB5B65FA3F848EA07F329D34365">441</mods:start>
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<mods:end id="508E67E64B0D27196961DACF837E272A">473</mods:end>
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<mods:classification id="4D646886D6A86C0679B606DBD77DA032">journal article</mods:classification>
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<mods:identifier id="E913508566C06D0D8560C980194A7AA6" type="DOI">10.11646/zootaxa.4138.3.2</mods:identifier>
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<mods:identifier id="05D68D3CAEA06F2952F6F5F670DED7FC" type="GBIF-Dataset">c785c9ac-b71c-4487-b770-35ebfe797c8a</mods:identifier>
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<mods:identifier id="C9E6C7566DEC6B9EC644E143AAE44211" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="7A6EF6E4241EEFF82A9639493A5B7975" type="Zenodo-Dep">258464</mods:identifier>
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<mods:identifier id="851F199E26DDD29CD3610417F856BF47" type="ZooBank">DDF68118-EA96-49A5-B316-47814E19ED3D</mods:identifier>
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</mods:mods>
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<treatment id="746C8797FFFFD003ACDDF997FEA895DB" ID-DOI="http://doi.org/10.5281/zenodo.5668195" ID-GBIF-Taxon="123195953" ID-Zenodo-Dep="5668195" LSID="urn:lsid:zoobank.org:act:E02B4845-66BF-43EB-990F-02EF1F83185C" httpUri="http://treatment.plazi.org/id/746C8797FFFFD003ACDDF997FEA895DB" lastPageId="10" lastPageNumber="451" pageId="9" pageNumber="450">
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<subSubSection id="B4DF650AFFFFD000ACDDF997FE569008" pageId="9" pageNumber="450" type="nomenclature">
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<paragraph id="FC7A3681FFFFD000ACDDF997FD2E902C" blockId="9.[151,677,1537,1598]" box="[151,677,1537,1562]" pageId="9" pageNumber="450">
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<heading id="A73281EDFFFFD000ACDDF997FD2E902C" bold="true" box="[151,677,1537,1562]" fontSize="10" level="2" pageId="9" pageNumber="450" reason="2">
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<emphasis id="CEB1EA93FFFFD000ACDDF997FD2E902C" bold="true" box="[151,677,1537,1562]" pageId="9" pageNumber="450">
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<taxonomicName id="3BC54D02FFFFD000ACDDF997FDC1902C" authority="Leschen and Escalona" authorityName="Leschen and Escalona" box="[151,586,1537,1562]" class="Insecta" family="Ulodidae" genus="Ulobostrichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="450" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="CEB1EA93FFFFD000ACDDF997FEB2902C" bold="true" box="[151,313,1537,1562]" italics="true" pageId="9" pageNumber="450">Ulobostrichus</emphasis>
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Leschen and Escalona
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</taxonomicName>
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,
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<taxonomicNameLabel id="D58257E8FFFFD000AE1CF997FD2E902C" box="[598,677,1537,1562]" pageId="9" pageNumber="450" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="FC7A3681FFFFD000ACDDF9B3FE569008" blockId="9.[151,677,1537,1598]" box="[151,477,1573,1598]" pageId="9" pageNumber="450">
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(
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<figureCitation id="64FE2A04FFFFD000ACD5F9B3FE939008" box="[159,280,1573,1598]" captionStart="FIGURES 4 A – J. A" captionStartId="17.[151,264,1786,1808]" captionTargetBox="[151,1434,194,1758]" captionTargetId="figure@17.[151,1436,193,1765]" captionTargetPageId="17" captionText="FIGURES 4 A – J. A, Trachyderas costulatum Fairmaire & Germain, length 7.3 mm; B, Trachyderastes dipsaconiae Kaszab, length 7.5 mm; C, Ulodes sp., length 4.8 mm (Australia, Queensland); D, Ulodes hispidus Pascoe, length 7.2 mm. E – G, Ulobostrichus monteithi sp. n., dorsal, lateral and anterior view, length 7.0 mm; H – J, Waitomophylax worthyi (Leschen & Rhode), head, dorsal, lateral and ventral." httpUri="https://zenodo.org/record/258469/files/figure.png" pageId="9" pageNumber="450">Figs 4E–G</figureCitation>
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,
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<figureCitation id="64FE2A04FFFFD000AD69F9B3FED19008" box="[291,346,1573,1598]" captionStart="FIGURES 5 A – J. A – D" captionStartId="18.[151,264,1725,1747]" captionTargetBox="[193,1397,210,1689]" captionTargetId="figure@18.[180,1408,193,1703]" captionTargetPageId="18" captionText="FIGURES 5 A – J. A – D, Head and prothorax, ventral view: A, Ulocyphaleus valdivianus (Philippi & Philippi); B, Trachyderas costulatum Fairmaire & Germain; C, Syrphetodes marginatus Pascoe; D, Arthopus brouni Sharp. E, Trachyderastes dipsaconiae Kaszab, antenna detail; F – I, Ulobostrichus monteithi sp. n. F, antenna, G, elytral apex, H, body lateral view, I, body latero-ventral view; J, Meryx rugosa Latreille, wing." httpUri="https://zenodo.org/record/258470/files/figure.png" pageId="9" pageNumber="450">5F–I</figureCitation>
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,
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<figureCitation id="64FE2A04FFFFD000AD2CF9B0FE5F9008" box="[358,468,1574,1598]" captionStart="FIGURES 8 A – P. A – B" captionStartId="21.[151,264,1802,1824]" captionTargetBox="[167,1433,205,1780]" captionTargetId="figure@21.[151,1436,193,1780]" captionTargetPageId="21" captionText="FIGURES 8 A – P. A – B, Exohadrus volutithorax (Broun), prothorax, dorsal and ventral; C, Ulodes hispidus Pascoe, pronotum; D, Trachyderastes dipsaconiae Kaszab, abdomen; E, Ulobostrichus monteithi sp. n., female reproductive track; F – I, Arthopus brouni Sharp, F – G, tegmen, H – I penis, lateral view and apex detail; J, Dipsaconia sp., K – M, Ulocyphaleus valdivianus (Philippi & Philippi), penis, ventral, dorsal and lateral; N – P, Ulobostrichus monteithi sp. n., N – O, genitalia (N, penis, lateral view; O, ventral view of tegmen); P, female genitalia." httpUri="https://zenodo.org/record/258473/files/figure.png" pageId="9" pageNumber="450">8E, 8N–P</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B4DF650AFFFFD000ACDDF9FBFCC590B5" box="[151,846,1645,1667]" pageId="9" pageNumber="450" type="materials_examined">
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<paragraph id="FC7A3681FFFFD000ACDDF9FBFCC590B5" blockId="9.[151,846,1645,1667]" box="[151,846,1645,1667]" pageId="9" pageNumber="450">
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<emphasis id="CEB1EA93FFFFD000ACDDF9FBFEAD90B5" bold="true" box="[151,294,1645,1667]" pageId="9" pageNumber="450">
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<typeStatus id="237E8823FFFFD000ACDDF9FBFF4790B5" box="[151,204,1645,1667]" pageId="9" pageNumber="450">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="3BC54D02FFFFD000AD61F9FBFC8190B5" authority="Leschen and Escalona" authorityName="Leschen and Escalona" box="[299,778,1645,1667]" class="Insecta" family="Ulodidae" genus="Ulobostrichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="450" phylum="Arthropoda" rank="species" species="monteithi" status="sp. nov.">
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<emphasis id="CEB1EA93FFFFD000AD61F9FBFDAA90B5" box="[299,545,1645,1667]" italics="true" pageId="9" pageNumber="450">Ulobostrichus monteithi</emphasis>
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Leschen and Escalona
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</taxonomicName>
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,
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<emphasis id="CEB1EA93FFFFD000AF5EF9FBFCC590B5" bold="true" box="[788,846,1645,1667]" pageId="9" pageNumber="450">
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<taxonomicNameLabel id="D58257E8FFFFD000AF5EF9FBFCC590B5" box="[788,846,1645,1667]" pageId="9" pageNumber="450" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="B4DF650AFFFFD000ACDDF939FAE59126" pageId="9" pageNumber="450" type="diagnosis">
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<paragraph id="FC7A3681FFFFD000ACDDF939FAE59126" blockId="9.[151,1437,1711,2025]" pageId="9" pageNumber="450">
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<emphasis id="CEB1EA93FFFFD000ACDDF939FE9A90FE" bold="true" box="[151,273,1711,1736]" pageId="9" pageNumber="450">Diagnosis.</emphasis>
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<taxonomicName id="3BC54D02FFFFD000AD50F939FE3290FE" box="[282,441,1711,1736]" class="Insecta" family="Ulodidae" genus="Ulobostrichus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="441" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="CEB1EA93FFFFD000AD50F939FE3290FE" box="[282,441,1711,1736]" italics="true" pageId="9" pageNumber="450">Ulobostrichus</emphasis>
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</taxonomicName>
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can easily be distinguished from all other ulodids by the tubulate body form and head with the vertex strongly declined. Distinctive characters for the genus are the 10-segmented antennae with 2- segmented club, which does not extend beyond the posterior margin of the pronotum, and the declivous elytra.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4DF650AFFFFD003AC8DF88AFA99956B" lastPageId="10" lastPageNumber="451" pageId="9" pageNumber="450" type="description">
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<paragraph id="FC7A3681FFFFD003AC8DF88AFA99956B" blockId="9.[151,1437,1711,2025]" lastBlockId="10.[151,1437,151,1005]" lastPageId="10" lastPageNumber="451" pageId="9" pageNumber="450">
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<emphasis id="CEB1EA93FFFFD000AC8DF88AFED29103" bold="true" box="[199,345,1820,1845]" pageId="9" pageNumber="450">Description.</emphasis>
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Body about 2.2 times as long as wide, tubulate with sides parallel. Body surfaces rugose and subglabrous; unicolorous; vestiture consisting of scales, obscured in some specimens by encrustation. Head with vertex declined, mouthparts more or less directed ventrally. Antenna 10 segmented with 2-segmented club; antenna not reaching posterior edge of pronotum, length of antennomere 3 greater than 1 and 2 combined; antennomeres 9 and 10 with sensory areas demarked by a rim. Eye ovoid and entire, interfacetal setae absent. Temples absent. Frons impressed and lacking anterolateral horns in male, frontoclypeal suture absent. Incisor edge of right mandible with single tooth, but not greatly enlarged. Last maxillary palpomere not expanded, width more or less equal to preceding palpomeres; uncus present on lacinea. Prementum sclerotized. Postmental groove absent; gular sutures short or absent (posterior tentorial pits well-developed). Subocular tubercle absent. Prothorax narrowly explanate, almost as wide as elytra. Prothorax widest near middle; anterior angles angulate and produced, but short; posterior angles present. Pronotal disc uneven, paired basal pits and bead absents. Prothoracic carinae sharp and dentate. Anterior portion of prosternum longer than width of procoxae; lacking pits. Notosternal sutures complete. Protrochanteral notch absent or weakly indicated. Procoxal cavities closed behind and separated by more than their diameters. Scutellar shield present and elevated. Elytral punctation seriate and lacking scutellary striole; punctures opened. Elytral surface convex with declivous apex; bearing large tubercles and carinate. Epipleuron welldeveloped with elytral carina simple. Mesoventrite lacking prosternal rest; width of the apex of mesoventrite greater than the width of mesoventral process. Discrimen absent. Metacoxal separation narrow, mesometaventral junction less than mesocoxal width. Metendosternite lacking laminae and anterior tendons. Hind wings absent. Abdominal ventrites with abdominal flanges; glandular fields absent; apical groove of ventrite 5 absent. Tarsal formula 5-5-4; protarsomere 2 quadrate; length of tarsomere 2 of middle leg equal to 1 and 3. Sternite VIII in male with short anterior strut (spiculum relictum), weakly emarginate at apex; tergite VIII lightly sclerotized and broadly rounded. Segment IX in male with broad spiculum gastrale and moderately sclerotized laterotergites; segment X membranous. Aedeagus (
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<figureCitation id="64FE2A04FFFCD003ADF9FD41FDA794C6" box="[435,556,727,752]" captionStart="FIGURES 8 A – P. A – B" captionStartId="21.[151,264,1802,1824]" captionTargetBox="[167,1433,205,1780]" captionTargetId="figure@21.[151,1436,193,1780]" captionTargetPageId="21" captionText="FIGURES 8 A – P. A – B, Exohadrus volutithorax (Broun), prothorax, dorsal and ventral; C, Ulodes hispidus Pascoe, pronotum; D, Trachyderastes dipsaconiae Kaszab, abdomen; E, Ulobostrichus monteithi sp. n., female reproductive track; F – I, Arthopus brouni Sharp, F – G, tegmen, H – I penis, lateral view and apex detail; J, Dipsaconia sp., K – M, Ulocyphaleus valdivianus (Philippi & Philippi), penis, ventral, dorsal and lateral; N – P, Ulobostrichus monteithi sp. n., N – O, genitalia (N, penis, lateral view; O, ventral view of tegmen); P, female genitalia." httpUri="https://zenodo.org/record/258473/files/figure.png" pageId="10" pageNumber="451">Fig. 8N–O</figureCitation>
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) with phallobase slightly larger than apicale. Penis about as long as phallobase and apicale combined, very slender and parallel-sided. Sternite VIII in female with long, slender, slightly curved anterior strut (spiculum ventrale). Ovipositor slightly longer than wide, lightly sclerotized, bursa saccular with a subacute apex and a narrow spermathecal gland (
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<figureCitation id="64FE2A04FFFCD003AEF7FCD2FC9F956A" box="[701,788,836,861]" captionStart="FIGURES 8 A – P. A – B" captionStartId="21.[151,264,1802,1824]" captionTargetBox="[167,1433,205,1780]" captionTargetId="figure@21.[151,1436,193,1780]" captionTargetPageId="21" captionText="FIGURES 8 A – P. A – B, Exohadrus volutithorax (Broun), prothorax, dorsal and ventral; C, Ulodes hispidus Pascoe, pronotum; D, Trachyderastes dipsaconiae Kaszab, abdomen; E, Ulobostrichus monteithi sp. n., female reproductive track; F – I, Arthopus brouni Sharp, F – G, tegmen, H – I penis, lateral view and apex detail; J, Dipsaconia sp., K – M, Ulocyphaleus valdivianus (Philippi & Philippi), penis, ventral, dorsal and lateral; N – P, Ulobostrichus monteithi sp. n., N – O, genitalia (N, penis, lateral view; O, ventral view of tegmen); P, female genitalia." httpUri="https://zenodo.org/record/258473/files/figure.png" pageId="10" pageNumber="451">Fig. 8E</figureCitation>
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). Coxites with gonostyli inserted subapically.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4DF650AFFFCD003AC8DFCF1FB9D9593" pageId="10" pageNumber="451" type="etymology">
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<paragraph id="FC7A3681FFFCD003AC8DFCF1FB9D9593" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="451">
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<emphasis id="CEB1EA93FFFCD003AC8DFCF1FEC595B6" bold="true" box="[199,334,871,896]" pageId="10" pageNumber="451">Etymology.</emphasis>
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The generic name combines the prefix
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<emphasis id="CEB1EA93FFFCD003AF4FFCF1FCB995B6" box="[773,818,871,896]" italics="true" pageId="10" pageNumber="451">ulo-</emphasis>
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derived from
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<taxonomicName id="3BC54D02FFFCD003AF9AFCF1FBBC95B6" box="[976,1079,871,896]" class="Insecta" family="Ulodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="451" phylum="Arthropoda" rank="family">Ulodidae</taxonomicName>
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with the suffix -
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<emphasis id="CEB1EA93FFFCD003A8A5FCF1FAE895B6" box="[1263,1379,871,896]" italics="true" pageId="10" pageNumber="451">bostrichus</emphasis>
|
|||
|
|
with reference to the similar body form found in many tubulate wood-boring beetles.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="B4DF650AFFFCD003AC8DFC39FEA895DB" pageId="10" pageNumber="451" type="discussion">
|
|||
|
|
<paragraph id="FC7A3681FFFCD003AC8DFC39FEA895DB" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="451">
|
|||
|
|
<emphasis id="CEB1EA93FFFCD003AC8DFC39FEC495FE" bold="true" box="[199,335,943,968]" pageId="10" pageNumber="451">Comments.</emphasis>
|
|||
|
|
This genus is represented by a single species found in subtropical forests of
|
|||
|
|
<collectingRegion id="3E01F863FFFCD003A8EFFC26FAE695FE" box="[1189,1389,943,968]" country="Australia" name="New South Wales" pageId="10" pageNumber="451">
|
|||
|
|
New South
|
|||
|
|
<collectingCountry id="84D27611FFFCD003A963FC39FAE695FE" box="[1321,1389,943,968]" name="United Kingdom" pageId="10" pageNumber="451">Wales</collectingCountry>
|
|||
|
|
</collectingRegion>
|
|||
|
|
and
|
|||
|
|
<collectingRegion id="3E01F863FFFCD003ACDDFC42FE9495DB" box="[151,287,980,1005]" country="Australia" name="Queensland" pageId="10" pageNumber="451">Queensland</collectingRegion>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|