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<document id="FD55BBD7E9AD0FFD984440B4F97D4F2C" ID-CLB-Dataset="198299" ID-DOI="10.11646/palaeoentomology.5.5.3" ID-GBIF-Dataset="57587277-64dd-4e57-9a2b-8d8401b7f3ca" ID-ISSN="2624-2834" ID-Zenodo-Dep="7333681" ID-ZooBank="DDE5EFC6-DD26-4B92-836B-7C91D4CBBE32" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668766693217" checkinUser="plazi" docAuthor="SHCHERBAKOV, DMITRY E." docDate="2022" docId="038E7975217EA259D8D0FAA1FC2C943D" docLanguage="en" docName="palaeoentomology.5.5.3.pdf" docOrigin="Palaeoentomology 5 (5)" docSource="http://dx.doi.org/10.11646/palaeoentomology.5.5.3" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Strivicia davidi SHCHERBAKOV 2022, sp. nov." docType="treatment" docUuid="A093D1D3-16B8-4F13-8A29-570E7C3EDD58" docUuidSource="ZooBank" docVersion="8" lastPageNumber="435" masterDocId="FFB7010D217FA258D858FF9FFF9A9C11" masterDocTitle="A new genus of Ipsviciidae (Hemiptera, Cicadomorpha) with a tegminal strigil from the Triassic of Madygen" masterLastPageNumber="438" masterPageNumber="434" pageNumber="435" updateTime="1699411749870" updateUser="ExternalLinkService">
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<mods:title id="86AD7EA5CF74E9D9AEDCCC357BFE4582">A new genus of Ipsviciidae (Hemiptera, Cicadomorpha) with a tegminal strigil from the Triassic of Madygen</mods:title>
</mods:titleInfo>
<mods:name id="30A78381075C2B3DC4E9CDD706630C78" type="personal">
<mods:role id="EADD77E22564C840CA21F7887B0AB33C">
<mods:roleTerm id="FDEF6C939742C50475BC0401C4107B0F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="21FB89FB09E2597654FDDD289AC5061B">SHCHERBAKOV, DMITRY E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="66F7B01067659F02BAA3B8492AC5E7BE">text</mods:typeOfResource>
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<mods:title id="CCEADBF85B5EC9154F17FA4EE6BE2756">Palaeoentomology</mods:title>
</mods:titleInfo>
<mods:part id="7C9EF19257951352A8877A2E2AEFD109">
<mods:date id="8BCA206357FFF333A05B62B8B727A56B">2022</mods:date>
<mods:detail id="30003E5AD688B5AC5D8DD8341FE58F72" type="pubDate">
<mods:number id="D9A55103C98F4002C9D02879FC22D116">2022-09-22</mods:number>
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<mods:number id="4573F2C1F1A80FFF41E1D506DBB65418">5</mods:number>
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<mods:number id="F5459ACC5BC690E6B517C5AECC575768">5</mods:number>
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<mods:url id="124631494A6BC30CFCCF476EB5083309">http://dx.doi.org/10.11646/palaeoentomology.5.5.3</mods:url>
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<mods:classification id="78F00FE687FD3444ECACBCAA64D2F76D">journal article</mods:classification>
<mods:identifier id="34777DB0548D3F42DC7B313534F6BBB4" type="CLB-Dataset">198299</mods:identifier>
<mods:identifier id="ABBD2C58F24099E5B1ACDFB607D7A073" type="DOI">10.11646/palaeoentomology.5.5.3</mods:identifier>
<mods:identifier id="8A118611A061510C2085501D2E90B23B" type="GBIF-Dataset">57587277-64dd-4e57-9a2b-8d8401b7f3ca</mods:identifier>
<mods:identifier id="D9A1F9AEFE7CB2F4C7290F0C4DF61B3B" type="ISSN">2624-2834</mods:identifier>
<mods:identifier id="EC23A6363A40F5CCB7E004CCCE948530" type="Zenodo-Dep">7333681</mods:identifier>
<mods:identifier id="CA34C14DC9909680F127E2AA1C0F3C23" type="ZooBank">DDE5EFC6-DD26-4B92-836B-7C91D4CBBE32</mods:identifier>
</mods:mods>
<treatment id="038E7975217EA259D8D0FAA1FC2C943D" ID-DOI="http://doi.org/10.5281/zenodo.7383823" ID-GBIF-Taxon="205255164" ID-Zenodo-Dep="7383823" LSID="urn:lsid:zoobank.org:act:A093D1D3-16B8-4F13-8A29-570E7C3EDD58" httpUri="http://treatment.plazi.org/id/038E7975217EA259D8D0FAA1FC2C943D" lastPageNumber="435" pageId="1" pageNumber="435">
<subSubSection id="C33D9BE8217EA259D8D0FAA1FEA4998F" pageId="1" pageNumber="435" type="nomenclature">
<paragraph id="8B98C863217EA259D8D0FAA1FE139947" blockId="1.[136,778,1341,1475]" box="[136,393,1341,1366]" pageId="1" pageNumber="435">
<heading id="D0D07F0F217EA259D8D0FAA1FE139947" bold="true" box="[136,393,1341,1366]" fontSize="10" level="2" pageId="1" pageNumber="435" reason="3">
<emphasis id="B9531471217EA259D8D0FAA1FE139947" bold="true" box="[136,393,1341,1366]" pageId="1" pageNumber="435">
<taxonomicName id="4C27B3E0217EA259D8D0FAA1FEB49944" authority="SHCHERBAKOV, 2022" authorityName="SHCHERBAKOV" authorityYear="2022" box="[136,302,1341,1366]" class="Insecta" family="Ipsviciidae" genus="Strivicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="435" phylum="Arthropoda" rank="species" species="davidi" status="sp. nov.">
<emphasis id="B9531471217EA259D8D0FAA1FEB49944" bold="true" box="[136,302,1341,1366]" italics="true" pageId="1" pageNumber="435">Strivicia davidi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260A90A217EA259D96CFAA1FE139947" box="[308,393,1342,1366]" pageId="1" pageNumber="435" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B98C863217EA259D8D0FAFDFEA4998F" blockId="1.[136,778,1341,1475]" pageId="1" pageNumber="435">
<uri id="FFB6C461217EA259D8D0FAFDFEA4998F" pageId="1" pageNumber="435">
urn:lsid:zoobank.org:act:
<uuid id="FF81F2B6217EA259D9F4FAFDFEA4998F" pageId="1" pageNumber="435">A093D1D3-16B8-4F13-8A29- 570E7C3EDD58</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C33D9BE8217EA259D8D0FA34FF4299D2" box="[136,216,1451,1475]" pageId="1" pageNumber="435" type="description">
<paragraph id="8B98C863217EA259D8D0FA34FF4299D2" blockId="1.[136,778,1341,1475]" box="[136,216,1451,1475]" pageId="1" pageNumber="435">
(
<figureCitation id="131CD4E6217EA259D8C8FA34FF4A99D2" box="[144,208,1451,1475]" captionStart="FIGURE 2" captionStartId="2.[136,235,1886,1910]" captionTargetBox="[151,1436,189,1862]" captionTargetId="figure-15@2.[151,1436,189,1862]" captionTargetPageId="2" captionText="FIGURE 2. Strivicia davidi gen. et sp. nov. A–D, Holotype, left tegmen, A, B, Counterpart, whole tegmen. C, D, Part (images flipped). C, Proximal part of tegmen with strigil. D, Proximal half of strigil. E, F, Paratype PIN 3288/388, right tegmen (part). G, H, Paratype PIN 2785/2432, left tegmen (counterpart). A, F, G, Photographs. B–E, H, SEM (BSE). I, Line drawing of tegmen (based on the holotype and paratypes; dashed line, costal fracture; dotted contour, strigil). Scale bars = 1 mm (A, B, E–I), 0.2 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.7333685" httpUri="https://zenodo.org/record/7333685/files/figure.png" pageId="1" pageNumber="435">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33D9BE8217EA259D8D0FA6CFEFE9AF7" pageId="1" pageNumber="435" type="materials_examined">
<paragraph id="8B98C863217EA259D8D0FA6CFEFE9AF7" blockId="1.[136,778,1523,2093]" pageId="1" pageNumber="435">
<emphasis id="B9531471217EA259D8D0FA6CFF689A1A" bold="true" box="[136,242,1523,1547]" pageId="1" pageNumber="435">Material.</emphasis>
<materialsCitation id="3B4FC23E217EA259D8A6FA6CFE219A21" ID-GBIF-Occurrence="3978668301" collectionCode="PIN" country="Kyrgyzstan" county="Batken Region" location="southwestern Madygen" municipality="Leilek District" pageId="1" pageNumber="435" specimenCode="PIN 3288, PIN 2785" specimenCount="3" typeStatus="holotype">
<typeStatus id="549C76C1217EA259D8A6FA6CFEFB9A1A" box="[254,353,1523,1547]" pageId="1" pageNumber="435" type="holotype">Holotype</typeStatus>
<specimenCode id="DB816018217EA259D935FA6BFE439A1A" box="[365,473,1523,1547]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="1" pageNumber="435">PIN 3288</specimenCode>
/316±, finely preserved left tegmen (clavus missing) and
</materialsCitation>
<materialsCitation id="3B4FC23E217EA259D99AF987FEA79A69" ID-GBIF-Occurrence="3978668303" collectionCode="PIN" country="Kyrgyzstan" county="Batken Region" location="southwestern Madygen" municipality="Leilek District" pageId="1" pageNumber="435" specimenCode="PIN 3288, PIN 2785" specimenCount="3" typeStatus="paratype">
<typeStatus id="549C76C1217EA259D99AF987FD849A21" box="[450,542,1560,1584]" pageId="1" pageNumber="435" type="paratype">paratype</typeStatus>
<specimenCode id="DB816018217EA259DA7CF987FD179A21" box="[548,653,1560,1584]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="1" pageNumber="435">PIN 3288</specimenCode>
/388±, right tegmen (clavus except apex missing) from
<location id="8EF89EB8217EA259DA25F9A3FF779A69" LSID="urn:lsid:plazi:treatment:038E7975217EA259D8D0FAA1FC2C943D:8EF89EB8217EA259DA25F9A3FF779A69" country="Kyrgyzstan" county="Batken Region" municipality="Leilek District" name="southwestern Madygen" pageId="1" pageNumber="435">
southwestern
<location id="8EF89EB8217EA259D8D0F9FFFF779A69" LSID="urn:lsid:plazi:treatment:038E7975217EA259D8D0FAA1FC2C943D:8EF89EB8217EA259D8D0F9FFFF779A69" box="[136,237,1632,1656]" country="Kyrgyzstan" county="Batken Region" municipality="Leilek District" name="Madygen area" pageId="1" pageNumber="435">Madygen</location>
</location>
area;
</materialsCitation>
<materialsCitation id="3B4FC23E217EA259D901F9FEFEFE9AF7" ID-GBIF-Occurrence="3978668302" collectionCode="PIN" country="Kyrgyzstan" county="Batken Region" location="southwestern Madygen" municipality="Leilek District" pageId="1" pageNumber="435" specimenCode="PIN 3288, PIN 2785" specimenCount="3" typeStatus="paratype">
<typeStatus id="549C76C1217EA259D901F9FEFE2F9A68" box="[345,437,1633,1657]" pageId="1" pageNumber="435" type="paratype">paratype</typeStatus>
<specimenCode id="DB816018217EA259D989F9FEFDD69A69" box="[465,588,1632,1656]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="1" pageNumber="435">PIN 2785</specimenCode>
/2432 (negative impression), complete left tegmen from
<location id="8EF89EB8217EA259DA22F91AFC9C9A8C" LSID="urn:lsid:plazi:treatment:038E7975217EA259D8D0FAA1FC2C943D:8EF89EB8217EA259DA22F91AFC9C9A8C" box="[634,774,1669,1693]" country="Kyrgyzstan" county="Batken Region" municipality="Leilek District" name="Dzhailoucho" pageId="1" pageNumber="435">Dzhailoucho</location>
;
<location id="8EF89EB8217EA259D8D0F936FF779AD0" LSID="urn:lsid:plazi:treatment:038E7975217EA259D8D0FAA1FC2C943D:8EF89EB8217EA259D8D0F936FF779AD0" box="[136,237,1705,1729]" country="Kyrgyzstan" county="Batken Region" municipality="Leilek District" name="Madygen" pageId="1" pageNumber="435">Madygen</location>
(
<locationDeviation id="9A2AD141217EA259D95BF936FE779AD0" box="[259,493,1705,1729]" location="Isfara" pageId="1" pageNumber="435">
<quantity id="4CDF6586217EA259D95BF936FED59AD0" box="[259,335,1705,1729]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="435" unit="km" value="35.0">35 km</quantity>
WSW Isfara
</locationDeviation>
),
<collectingMunicipality id="6BFC5219217EA259DA5FF936FD369AD0" box="[519,684,1705,1729]" pageId="1" pageNumber="435">Leilek District</collectingMunicipality>
,
<collectingCounty id="62F9B0EF217EA259DAE6F936FF429AF4" pageId="1" pageNumber="435">Batken Region</collectingCounty>
,
<collectingCountry id="F33088F3217EA259D8BAF951FEFA9AF7" box="[226,352,1742,1766]" name="Kyrgyzstan" pageId="1" pageNumber="435">Kyrgyzstan</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D9BE8217EA259D8E5F96DFDA39B3F" pageId="1" pageNumber="435" type="etymology">
<paragraph id="8B98C863217EA259D8E5F96DFDA39B3F" blockId="1.[136,778,1523,2093]" pageId="1" pageNumber="435">
<emphasis id="B9531471217EA259D8E5F96DFEA49B1B" bold="true" box="[189,318,1778,1802]" pageId="1" pageNumber="435">Etymology.</emphasis>
In honour of our colleague David Grimaldi, for his excellent studies of fossil insects.
</paragraph>
</subSubSection>
<subSubSection id="C33D9BE8217EA259D8E5F8A5FE649B8A" pageId="1" pageNumber="435" type="diagnosis">
<paragraph id="8B98C863217EA259D8E5F8A5FE529B42" blockId="1.[136,778,1523,2093]" box="[189,456,1850,1875]" pageId="1" pageNumber="435">
<emphasis id="B9531471217EA259D8E5F8A5FEA89B43" bold="true" box="[189,306,1850,1874]" pageId="1" pageNumber="435">Diagnosis.</emphasis>
As for genus.
</paragraph>
<paragraph id="8B98C863217EA259D8E5F8C1FE649B8A" blockId="1.[136,778,1523,2093]" pageId="1" pageNumber="435">
<emphasis id="B9531471217EA259D8E5F8C1FE2B9B67" bold="true" box="[189,433,1886,1911]" pageId="1" pageNumber="435">Locality and horizon.</emphasis>
Madygen Formation, Ladinian– Carnian, Middle or Upper Triassic.
</paragraph>
</subSubSection>
<subSubSection id="C33D9BE8217EA259D8E5F838FC2C943D" pageId="1" pageNumber="435" type="description">
<paragraph id="8B98C863217EA259D8E5F838FC2C943D" blockId="1.[136,778,1523,2093]" lastBlockId="1.[810,1452,615,2092]" pageId="1" pageNumber="435">
<emphasis id="B9531471217EA259D8E5F838FED29BAE" bold="true" box="[189,328,1959,1983]" pageId="1" pageNumber="435">Description.</emphasis>
Tegmen length
<quantity id="4CDF6586217EA259D9A7F838FD1F9BAE" box="[511,645,1959,1983]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.4" metricValueMin="4.2" pageId="1" pageNumber="435" unit="mm" value="4.3" valueMax="4.4" valueMin="4.2">4.2–4.4 mm</quantity>
, width
<quantity id="4CDF6586217EA259DA83F838FF449BF2" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="1" pageNumber="435" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.6– 1.8 mm</quantity>
, elongate (length/width ratio 2.5–2.6:1), as seen in the
<typeStatus id="549C76C1217EA259D8E8F86FFE979419" box="[176,269,2032,2056]" pageId="1" pageNumber="435" type="holotype">holotype</typeStatus>
and
<typeStatus id="549C76C1217EA259D91BF86FFE059419" box="[323,415,2032,2056]" pageId="1" pageNumber="435" type="paratype">paratype</typeStatus>
from the SW Madygen area. The
<typeStatus id="549C76C1217EA259D8D0F78AFF7E943C" box="[136,228,2069,2093]" pageId="1" pageNumber="435" type="paratype">paratype</typeStatus>
from Dzhailoucho is stretched transversely (its surface punctures are transformed into short subtransverse wrinkles) and appears much shorter and broader, but in fact it is of nearly the same size (length
<quantity id="4CDF6586217EA259DCF5FD2FFA989ED6" box="[1197,1282,688,712]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="435" unit="mm" value="3.4">3.4 mm</quantity>
, width
<quantity id="4CDF6586217EA259DD0AFD2FFA3D9ED6" box="[1362,1447,688,712]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="435" unit="mm" value="1.8">1.8 mm</quantity>
, √LW
<quantity id="4CDF6586217EA259DB3DFD4BFC2C9EFD" box="[869,950,724,748]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="435" unit="mm" value="2.5">2.5 mm</quantity>
) as the two former (√LW
<quantity id="4CDF6586217EA259DC9BFD4BFADC9EFD" box="[1219,1350,724,748]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="1" pageNumber="435" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">2.6–2.8 mm</quantity>
). Tegmen widest just before midlength, costal margin arched there, nearly straight more proximally and arched at base. Precostal carina wider proximally, deflected dorsad; hypocostal carina narrow; basicostal projection low; bSc large arch; costal fracture transverse; costal area moderately wide, its part between costal fracture and dSc sloped and occupied with oblong strigil. Basal cell short, triangular, closed with short M+CuA anastomosis. R(+dSc) stem convex, short, oblique; dSc continuing its direction, convex, sigmoidal, moderately long (1/5–1/3 tegmen length, much longer than R stem), joining margin before or at (Dzhailoucho
<typeStatus id="549C76C1217EA259DCA8FBFBFAD4986D" box="[1264,1358,1124,1148]" pageId="1" pageNumber="435" type="paratype">paratype</typeStatus>
) tegmen midlength. R distal to dSc longitudinal, with 3 anterior branches, bent at origin of third branch; apex of tegmen at narrow apical cell between R and M. M and CuA stems arched in opposite directions, enclosing lanceolate medial area, CuA arched close to CuP. Branches of CuA and M forming common posterior pecten with 4 terminations, CuA
<subScript id="17A3CA26217EA259DDFCFAD1FA36994D" attach="right" box="[1444,1452,1358,1372]" fontSize="6" pageId="1" pageNumber="435">1</subScript>
apparently fused to posterior M branch. Crossvein
<emphasis id="B9531471217EA259DD1BFAFCFAF3996B" box="[1347,1385,1379,1402]" italics="true" pageId="1" pageNumber="435">r-m</emphasis>
short, much more distal than free basal section of CuA
<subScript id="17A3CA26217EA259DD18FA09FAD299B5" attach="left" box="[1344,1352,1430,1444]" fontSize="6" pageId="1" pageNumber="435">1</subScript>
(looking like
<emphasis id="B9531471217EA259DB38FA34FC0F99D3" box="[864,917,1451,1474]" italics="true" pageId="1" pageNumber="435">m-cu</emphasis>
). Clavus occupying 2/3 tegmen length, claval veins (Pcu and 1A) united at 2/3 clavus length, their common stalk entering commissural margin much before claval apex. Upper surface of tegmen covered with rasp-like punctures, coarser and denser in proximal half, especially on basicostal piece, finer and sparser in distal half. Stridulatory area on underside of tegmen 1.1 ×
<quantity id="4CDF6586217EA259DC1EF91AFB3D9A8D" box="[1094,1191,1669,1693]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="435" unit="mm" value="0.23">0.23 mm</quantity>
, consisting of more than 40 oblique rows of microtrichia; rows 4 times more dense at base of strigil (14 per 100 μm) than in its distal part (3.5 per 100 μm); microtrichia (except near strigil base) transformed into short ridgelets up to 15 μm long, oriented across rows,
<emphasis id="B9531471217EA259DB72F8A4FCAB9B43" box="[810,817,1851,1874]" italics="true" pageId="1" pageNumber="435">i</emphasis>
.
<emphasis id="B9531471217EA259DB60F8A4FCDE9B43" box="[824,836,1851,1874]" italics="true" pageId="1" pageNumber="435">e</emphasis>
. almost longitudinally. Large pale ocellate spot margined with dark in widest part of medial area, similar smaller spot more distally in radial area, and probably one more at 1A; vein terminations margined with dark, CuA
<subScript id="17A3CA26217EA259DCAEF828FB649BD4" attach="left" box="[1270,1278,1975,1989]" fontSize="6" pageId="1" pageNumber="435">2</subScript>
with dark mark, dark specks and spots (more developed in Dzhailoucho
<typeStatus id="549C76C1217EA259DB72F86FFC129419" box="[810,904,2032,2056]" pageId="1" pageNumber="435" type="paratype">paratype</typeStatus>
) along veins and in anterior area; ground colour pale fuscous.
</paragraph>
</subSubSection>
</treatment>
</document>
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