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<mods:title id="086355C220612E0EFFC5054ECEB03505">A revised circumscription for the Blakeeae (Melastomataceae) with associated nomenclatural adjustments</mods:title>
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<mods:namePart id="99AF46ED0705FE5B38FA21327D11CDD8">Penneys, Darin S.</mods:namePart>
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<subSubSection id="DBD77AD9B0C8DBC5CFB4BB796BD78C6B" pageId="3" pageNumber="20" type="nomenclature">
<paragraph id="FF7F3C43FB4A645FA557D387DF7EB6C5" pageId="3" pageNumber="20">
<taxonomicName id="A03659B56590A02F6C754A49D7A89036" LSID="F154FE85-2D66-5A52-8248-A3CE122988AE" authority="Naudin, Ann. Sci. Nat., Bot. ser. 3 16: 99. 1851." class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Chalybea" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
Chalybea Naudin, Ann. Sci. Nat., Bot. 
<normalizedToken id="8DA2076221A9A932F3C028701B58F4F9" originalValue="sér">ser</normalizedToken>
. 3 16: 99. 1851.
</taxonomicName>
</paragraph>
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<subSubSection id="52C05E10C7A1766985B38A829637866E" pageId="3" pageNumber="20" type="reference_group">
<paragraph id="5A40D41D513EFD555C26E29A0E6BDBAB" pageId="3" pageNumber="20">
<taxonomicName id="C84C2D7003F04C9C1E6FF34237E464F6" authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">Huilaea</taxonomicName>
Wurdack, Brittonia 9: 106. 1957. Type: 
<taxonomicName id="9F8590DBCBC97D9497E0703778B12065" authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="species" species="penduliflora">
<emphasis id="FC2C17CE092875B88EFD6C7ECF128DF7" italics="true" pageId="3" pageNumber="20">Huilaea penduliflora</emphasis>
</taxonomicName>
Wurdack, Brittonia 9: 106. 1957. 
<emphasis id="9A3B0F4C3293AA24658B986606A90234" bold="true" pageId="3" pageNumber="20">TYPE. COLOMBIA.</emphasis>
<emphasis id="B4170B5B78B8906A59A7037D0B41A8A6" bold="true" pageId="3" pageNumber="20">Huila:</emphasis>
Cordillera Oriental, wet temperate forest of deep moist canyon near camp on north side of 
<normalizedToken id="6A1009D6128244D02C46F1434B899CA1" originalValue="Río">Rio</normalizedToken>
Venadito, 25 km SE of La Bodega, 2450 m, 1 Dec 1944, E.Little 9101 (holotype: US!; isotypes: COL!, NY!).
</paragraph>
</subSubSection>
<subSubSection id="79348F6D09E77A5D560A0E88C53C681C" pageId="3" pageNumber="20" type="type">
<paragraph id="45574A8EB9447E1B922CCB3DDB29E09A" pageId="3" pageNumber="20">Type.</paragraph>
<paragraph id="60BDB060B6E65AA99F3D5AA3360BE708" pageId="3" pageNumber="20">
<taxonomicName id="AE3AC34CAA5DC8B9298A5559DAEF6476" class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="species" species="corymbifera">
<emphasis id="F307F9FE6D02AEE7E4BAB975C863F115" italics="true" pageId="3" pageNumber="20">Chalybea corymbifera</emphasis>
</taxonomicName>
Naudin, Ann. Sci. Nat., Bot. 
<normalizedToken id="9AC86D816377DE86840B9C855808BDF1" originalValue="sér">ser</normalizedToken>
. 3 16: 100. 1851. TYPE. COLOMBIA. Santander: Pamplona, La Baja, ca. 3000 m, Dec 1846, Funck &amp; Schlim 1312 (holotype: P!, photos at NY!, US!; isotypes: BM!, BR! (2 sheets), G!, photos of G sheet at NY!, US!).
</paragraph>
</subSubSection>
<subSubSection id="9FEB21C44B94476ED4E3F80D57B52B06" pageId="3" pageNumber="20" type="remarks">
<paragraph id="5D8F433902F85065065F8AE99343421B" pageId="3" pageNumber="20">Remarks.</paragraph>
<paragraph id="74F81F4420F7E534E6B6C5BC4B8AC66B" pageId="3" pageNumber="20">
<emphasis id="A1D975034CB406BBA0B6686C00DAC4ED" bold="true" pageId="3" pageNumber="20">Phylogenetic analyses</emphasis>
(
<bibRefCitation id="C18A340829C8BB61EA2AC5CC5E5F430E" author="Morales-P, ME" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="12" pageNumber="29" refId="B12" refString="Morales-P, ME, 2010. Analisis filogenetico de Huilaea Wurdack (Melastomataceae) basado en datos morfologicos y moleculares. PhD Thesis, Universidad Nacional de Colombia, Colombia." title="Analisis filogenetico de Huilaea Wurdack (Melastomataceae) basado en datos morfologicos y moleculares. PhD Thesis, Universidad Nacional de Colombia, Colombia." year="2010">Morales-P 2010</bibRefCitation>
, 
<bibRefCitation id="77D120C0F2938BC46B086A673A26A881" author="Morales-P, ME" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="13" pageNumber="30" publicationUrl="10.1007/s12228-009-9092-x" refId="B15" refString="Morales-P, ME, Penneys, DS, Judd, W, Gonzalez, F, submitted. Revision taxonomica del clado Chalybea/Huilaea (Melastomataceae). Caldasia., 10.1007/s12228-009-9092-x" title="Revision taxonomica del clado Chalybea / Huilaea (Melastomataceae). Caldasia" url="10.1007/s12228-009-9092-x" year="submitted">Morales-P et al. submitted</bibRefCitation>
, 
<bibRefCitation id="822E498B8CE54A6923BE2D241356469C" author="Penneys, DS" journalOrPublisher="Caldasia" pageId="13" pageNumber="30" refId="B19" refString="Penneys, DS, Judd, WS, 2010. Phylogeny and character evolution in the Blakeeae (Melastomataceae). Abstract ID no. 699. Botany 2010. July 31 -August 4. Providence, Rhode Island, USA. http://2010.botanyconference.org/engine/search/index.php?func=detail&amp;aid=699." title="Phylogeny and character evolution in the Blakeeae (Melastomataceae). Abstract ID no. 699. Botany 2010. July 31 - August 4. Providence, Rhode Island, USA. http: // 2010. botanyconference. org / engine / search / index. php? func = detail &amp; aid = 699" year="2010">Penneys and Judd 2010</bibRefCitation>
) indicate that 
<taxonomicName id="9B8F395E3D7E4BD6957FCAD2BBC86E76" class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
<emphasis id="2BE792FCCEE5B34819FF73C177B3FCC5" italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
</taxonomicName>
and 
<taxonomicName id="D548B73AFB6C9CDAA880522738E9A148" authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis id="BC34054A228391D508594C778995F086" italics="true" pageId="3" pageNumber="20">Huilaea</emphasis>
</taxonomicName>
form a strongly-supported clade. However, as 
<taxonomicName id="74F1C38B347F11B912CB6D1775AD713C" class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
<emphasis id="E05046EAB50CCE4059A05FB0D7003779" italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
</taxonomicName>
is nested within 
<taxonomicName id="5767F2BFD414D1DF529DF9E7AD667934" authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis id="37B668CA1CC133EFA04D597B03AB791D" italics="true" pageId="3" pageNumber="20">Huilaea</emphasis>
</taxonomicName>
, species belonging to the latter genus must be transferred because 
<taxonomicName id="45D5DA0EC92764FB496348F16C67107E" class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
<emphasis id="51ECE7302F854193167383936E15C3C7" italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
</taxonomicName>
has nomenclatural priority. 
<taxonomicName id="E9D7837D88CB8697C444941A437E9F89" class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
<emphasis id="FD60B3C2F3C706FD44522BD30202EE99" italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
</taxonomicName>
has been distinguished from 
<taxonomicName id="4DB31E0C49E745197F4085048B3DBA81" authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis id="456829F6CF216769A444C634E0FCB334" italics="true" pageId="3" pageNumber="20">Huilaea</emphasis>
</taxonomicName>
by having inflorescences with 21-39 flowers (vs. 3-17 in 
<taxonomicName id="159C5F1FF74F31436FE4DEE7D42166E5" authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis id="30662AC6EBC8C53F6514D76869D4D41A" italics="true" pageId="3" pageNumber="20">Huilaea</emphasis>
</taxonomicName>
), the flowers 13-20 mm long (vs. 45-55 mm long), the hypanthium 7-10 mm long and 8-10 mm wide (vs. 13-31 mm long x 11-19 mm wide), the petals 12-14 mm long and 5-7 mm wide (vs. 26-44 mm long x 10-22 mm wide), and white to cream or green, sometimes 
<normalizedToken id="BC0772A9D9CF33E46B2BA26D886A201A" originalValue="pink–tinged">pink-tinged</normalizedToken>
apically (vs. pink to red, paler inside), the anthers 3-4 mm long (vs. 7-10 mm long), and the style 10-12 mm long (vs. 22-33 mm long).
</paragraph>
<paragraph id="DF174A151530A3216493A12AE0803113" pageId="3" pageNumber="20">
<taxonomicName id="7368531C16C9596D456137AC4558F6C2" class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
<emphasis id="EF0BC742591323FD964AD0B514B6DCF9" italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
</taxonomicName>
includes ten described species of small, South American trees: seven endemic to Colombia, two to Ecuador, and one to Peru. All are found in the Andes, except one species that is restricted to the Sierra Nevada de Santa Marta, Colombia.
</paragraph>
<paragraph id="BA1A122CE188F86483838DBC0EE78B75" pageId="3" pageNumber="20">
For more information on the systematics of 
<taxonomicName id="989AFB457A652D14845218CD061E9BDF" class="Magnoliopsida" family="Melastomataceae" genus="Chalybea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="genus">
<emphasis id="F981711387B3AE357B1F4066D1333CFB" italics="true" pageId="3" pageNumber="20">Chalybea</emphasis>
</taxonomicName>
(and 
<taxonomicName id="2BE0D0D31AA77A1A14E42FD973E87540" authorityName="Wurdack" authorityYear="1957" class="Magnoliopsida" family="Melastomataceae" genus="Huilaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis id="E7DA3D9037FBF9A9B797E16C43330511" italics="true" pageId="3" pageNumber="20">Huilaea</emphasis>
</taxonomicName>
), see 
<bibRefCitation id="0936C54F28EDF2D6333844797F3C95AE" author="Lozano-Contreras, G" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="12" pageNumber="29" refId="B10" refString="Lozano-Contreras, G, Ruiz-R, N, 1996. El genero Huilaea Wurdack (Melastomataceae). Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 20: 237-242. http://www.accefyn.org.co/revista/Vol_20/77/237-242.pdf." title="El genero Huilaea Wurdack (Melastomataceae). Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 20: 237 - 242. http: // www. accefyn. org. co / revista / Vol _ 20 / 77 / 237 - 242. pdf" year="1996">Lozano-Contreras and Ruiz-R (1996)</bibRefCitation>
, 
<bibRefCitation id="730A8112D3916A10B9F4D2B483122A58" author="Morales-P, ME" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="13" pageNumber="30" refId="B13" refString="Morales-P, ME, Gonzalez, F, 2005. Redescubrimiento de Chalybea Naudin e implicaciones en la delimitacion generica de Huilaea Wurdack (Melastomataceae). Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 29: 171-178. http://www.accefyn.org.co/revista/Vol_29/111/03_171_178.pdf." title="Redescubrimiento de Chalybea Naudin e implicaciones en la delimitacion generica de Huilaea Wurdack (Melastomataceae). Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 29: 171 - 178. http: // www. accefyn. org. co / revista / Vol _ 29 / 111 / 03 _ 171 _ 178. pdf" year="2005">
Morales-P and 
<normalizedToken id="9E16C0248E6B38EB2FBFFF142EBE97F5" originalValue="González">Gonzalez</normalizedToken>
(2005)
</bibRefCitation>
, 
<bibRefCitation id="AEDD815A4962D916BBBB1C7AF91FE0BB" author="Morales-P, ME" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="12" pageNumber="29" refId="B12" refString="Morales-P, ME, 2010. Analisis filogenetico de Huilaea Wurdack (Melastomataceae) basado en datos morfologicos y moleculares. PhD Thesis, Universidad Nacional de Colombia, Colombia." title="Analisis filogenetico de Huilaea Wurdack (Melastomataceae) basado en datos morfologicos y moleculares. PhD Thesis, Universidad Nacional de Colombia, Colombia." year="2010">Morales-P (2010)</bibRefCitation>
, 
<bibRefCitation id="E9398585CDAD044C9B1205DDD66AAFCC" author="Morales-P, ME" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="13" pageNumber="30" refId="B14" refString="Morales-P, ME, Penneys, DS, 2010. New species of Chalybea Naudin and Huilaea Wurdack. Brittonia 62: 26-34. http://www.springerlink.com/content/47873303k2k84836/." title="New species of Chalybea Naudin and Huilaea Wurdack. Brittonia 62: 26 - 34. http: // www. springerlink. com / content / 47873303 k 2 k 84836 /" year="2010">Morales-P and Penneys (2010)</bibRefCitation>
, and 
<bibRefCitation id="54CBFF7F249BA22D2BDFA6F876B67E15" author="Morales-P, ME" journalOrPublisher="Bulletin of the Florida Museum of Natural History, Biological Sciences" pageId="13" pageNumber="30" publicationUrl="10.1007/s12228-009-9092-x" refId="B15" refString="Morales-P, ME, Penneys, DS, Judd, W, Gonzalez, F, submitted. Revision taxonomica del clado Chalybea/Huilaea (Melastomataceae). Caldasia., 10.1007/s12228-009-9092-x" title="Revision taxonomica del clado Chalybea / Huilaea (Melastomataceae). Caldasia" url="10.1007/s12228-009-9092-x" year="submitted">Morales-P et al. (submitted)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
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