229 lines
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229 lines
33 KiB
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<document id="8E54A1DD1867287B516CCFB100D2752E" ID-CLB-Dataset="8104" ID-DOI="10.37520/aemnp.2020.031" ID-GBIF-Dataset="c1898d54-cd34-4727-96c9-fd832a13e196" ID-ISSN="1804-6487" ID-Zenodo-Dep="4489548" ID-ZooBank="CC5352E8-213D-40E0-B340-AFB35A17665D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612221854605" checkinUser="felipe" docAuthor="Murakami, Hiroyuki" docDate="2020" docId="E30F87CAA749925CFEBB7B8F18A113F6" docLanguage="en" docName="ActaEntMusNatPrag.60.2.475-492.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 60 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Cladiscus liaoi Murakami 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="489" masterDocId="1F36FFB2A7479252FFF37E711C3A1523" masterDocTitle="Review of the genus Cladiscus (Coleoptera: Cleridae) from Japan and Taiwan, with descriptions of two new species" masterLastPageNumber="492" masterPageNumber="475" pageNumber="489" updateTime="1698926216820" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="43BD98F3086BA501E4DFDA706AED0670">Review of the genus Cladiscus (Coleoptera: Cleridae) from Japan and Taiwan, with descriptions of two new species</mods:title>
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<mods:namePart id="863FDECFC0904A81103010431C58799D">Murakami, Hiroyuki</mods:namePart>
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<mods:typeOfResource id="8E3B792F4BD35CE8BD54FFFA38B2521A">text</mods:typeOfResource>
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<mods:title id="D8BB55B5547D2BFB9F0197188C368951">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="86F7865A4EFBDF7BE20C31D5BCBD356D">
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<mods:date id="FC3FD22D56A9CF17E71DC29D376E6EFC">2020</mods:date>
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<mods:number id="94F0F5AF3E6FA9EFDCEAF359B1A50547">2020-08-31</mods:number>
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<mods:number id="48CC011486980FB03D61C28C06BAB2C8">2</mods:number>
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<mods:identifier id="65EA1C56E245FC5107AE918F285CEDEE" type="CLB-Dataset">8104</mods:identifier>
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<mods:identifier id="92018B685818CF6E7D86B447AB13EE3E" type="DOI">10.37520/aemnp.2020.031</mods:identifier>
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<treatment id="E30F87CAA749925CFEBB7B8F18A113F6" ID-DOI="http://doi.org/10.5281/zenodo.4551189" ID-GBIF-Taxon="177428805" ID-Zenodo-Dep="4551189" LSID="urn:lsid:plazi:treatment:E30F87CAA749925CFEBB7B8F18A113F6" httpUri="http://treatment.plazi.org/id/E30F87CAA749925CFEBB7B8F18A113F6" lastPageNumber="489" pageId="14" pageNumber="489">
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<subSubSection id="23BC6557A749925CFEBB7B8F1E7D1336" box="[328,583,1534,1557]" pageId="14" pageNumber="489" type="nomenclature">
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<paragraph id="6B1936DCA749925CFEBB7B8F1E7D1336" blockId="14.[328,583,1534,1557]" box="[328,583,1534,1557]" pageId="14" pageNumber="489">
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<heading id="305181B0A749925CFEBB7B8F1E7D1336" box="[328,583,1534,1557]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="489" reason="3">
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<emphasis id="59D2EACEA749925CFEBB7B8F1E7D1336" bold="true" box="[328,583,1534,1557]" pageId="14" pageNumber="489">
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<taxonomicName id="ACA64D5FA749925CFEBB7B8F1DD11336" authority="Murakami, 2020" authorityName="Murakami" authorityYear="2020" box="[328,491,1534,1557]" class="Insecta" family="Cleridae" genus="Cladiscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="489" phylum="Arthropoda" rank="species" species="liaoi" status="sp. nov.">
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<emphasis id="59D2EACEA749925CFEBB7B8F1DD11336" bold="true" box="[328,491,1534,1557]" italics="true" pageId="14" pageNumber="489">Cladiscus liaoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="42E157B5A749925CFE027B8F1E7D1336" box="[497,583,1534,1557]" pageId="14" pageNumber="489" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="23BC6557A749925CFEFF786D1EB9130C" box="[268,643,1564,1583]" pageId="14" pageNumber="489" type="description">
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<paragraph id="6B1936DCA749925CFEFF786D1EB9130C" blockId="14.[151,761,1564,1705]" box="[268,643,1564,1583]" pageId="14" pageNumber="489">
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(
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<figureCitation id="F39D2A59A749925CFEE1786D1DA4130C" box="[274,414,1564,1583]" captionStart="Figs 1–13" captionStartId="2.[151,188,2001,2020]" captionTargetBox="[174,1418,197,1920]" captionTargetId="figure-13@2.[166,1421,189,1930]" captionTargetPageId="2" captionText="Figs 1–13. Habitus and head of Cladiscus spp. (1–5, 6, 8, 10, 12, male; 7, 9, 11, 13, female). 1 – Cladiscus obeliscus Lewis, 1892; 2 – C. weyersi Kraatz, 1899; 3 – C. sauteri Schenkling, 1912; 4 – C. terminalis Schenkling, 1912; 5 – C. pallidicornis Corporaal & van der Wiel, 1949; 6, 7, 10, 11 – C. hachijoensis sp. nov. (6, 10, holotype; 7, 11, paratype); 8, 9, 12, 13 – C. liaoi sp. nov. (8, 12, holotype; 9, 13, paratype). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4489552" httpUri="https://zenodo.org/record/4489552/files/figure.png" pageId="14" pageNumber="489">Figs 8, 9, 12, 13</figureCitation>
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,
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<figureCitation id="F39D2A59A749925CFE54786D1DE4130C" box="[423,478,1564,1583]" captionStart="Figs 14–26" captionStartId="3.[150,187,900,919]" captionTargetBox="[151,1434,200,863]" captionTargetId="figure-472@3.[151,1437,200,871]" captionText="Figs 14–26. Antennae (14, 16, 17, 19, 21, 23, 25 – male; 15, 18, 20, 22, 24, 26 – female). 14, 15 – Cladiscus obeliscus Lewis, 1892; 16 – C. weyersi Kraatz, 1899; 17, 18 – C. sauteri Schenkling, 1912; 19, 20 – C. terminalis Schenkling, 1912; 21, 22 – C. pallidicornis Corporaal & van der Wiel, 1949; 23, 24 – C. hachijoensis sp. nov.; 25, 26 – C. liaoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4551445" httpUri="https://zenodo.org/record/4551445/files/figure.png" pageId="14" pageNumber="489">25, 26</figureCitation>
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, 33, 107–118, 123)
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</paragraph>
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</subSubSection>
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<subSubSection id="23BC6557A749925CFF6478371E4A138A" pageId="14" pageNumber="489" type="materials_examined">
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<paragraph id="6B1936DCA749925CFF6478371E4A138A" blockId="14.[151,761,1564,1705]" pageId="14" pageNumber="489">
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<emphasis id="59D2EACEA749925CFF6478371D27137A" bold="true" box="[151,285,1606,1625]" pageId="14" pageNumber="489">
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<typeStatus id="B41D887EA749925CFF6478371CFE137A" box="[151,196,1606,1625]" pageId="14" pageNumber="489">Type</typeStatus>
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material.
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</emphasis>
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HOLOΤYΡΕ: 1 ♁, ‘[Taiwan] Dahanshan (Mt.). | Alt. ca.
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<quantity id="AC5E9B39A749925CFF6478101CE51357" box="[151,223,1633,1652]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="489" unit="m" value="1200.0">1,200 m</quantity>
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, Chunrih | Township,
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<collectingRegion id="A962F83EA749925CFE6A78101E101357" box="[409,554,1633,1652]" country="Taiwan" name="Pingtung" pageId="14" pageNumber="489">Pingtung County</collectingRegion>
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<emphasis id="59D2EACEA749925CFDCB78111ECD1350" bold="true" box="[568,759,1632,1651]" pageId="14" pageNumber="489">DZAEḆĦHṞ†Oiffi</emphasis>
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<date id="1F18101CA749925CFF51780A1D3313AD" box="[162,265,1659,1678]" pageId="14" pageNumber="489" value="2015-05-28">28. V. 2015</date>
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, W. C. Liao leg.’ (
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<collectionCode id="0DB7AE19A749925CFE5A780A1DDC13AD" box="[425,486,1659,1678]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="14" pageNumber="489">EUMJ</collectionCode>
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). PΑRΑΤYΡΕ:
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<specimenCount id="7DA0FD55A749925CFDA0780A1E4E13AC" box="[595,628,1656,1679]" pageId="14" pageNumber="489" type="female">1 ♀</specimenCount>
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, ‘FORMOSA |
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<collectingCountry id="13B1764CA749925CFF6478E71CD3138A" box="[151,233,1686,1705]" name="Taiwan" pageId="14" pageNumber="489">TAIWAN</collectingCountry>
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| 1968.....1971’, unknown collector. (
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<collectionCode id="0DB7AE19A749925CFDDA78E71E5C138A" box="[553,614,1686,1705]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="14" pageNumber="489">EUMJ</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="23BC6557A749925CFF6478B3197E10B6" pageId="14" pageNumber="489" type="description">
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<paragraph id="6B1936DCA749925CFF6478B31EFD13DA" blockId="14.[151,760,1730,2073]" pageId="14" pageNumber="489">
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<emphasis id="59D2EACEA749925CFF6478B31D5913FA" bold="true" box="[151,355,1730,1753]" pageId="14" pageNumber="489">
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Description.
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<emphasis id="59D2EACEA749925CFED878B31D5913FA" bold="true" box="[299,355,1730,1753]" italics="true" pageId="14" pageNumber="489">Male</emphasis>
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</emphasis>
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(
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<figureCitation id="F39D2A59A749925CFE8078B31DDD13FA" box="[371,487,1730,1753]" captionStart="Figs 1–13" captionStartId="2.[151,188,2001,2020]" captionTargetBox="[174,1418,197,1920]" captionTargetId="figure-13@2.[166,1421,189,1930]" captionTargetPageId="2" captionText="Figs 1–13. Habitus and head of Cladiscus spp. (1–5, 6, 8, 10, 12, male; 7, 9, 11, 13, female). 1 – Cladiscus obeliscus Lewis, 1892; 2 – C. weyersi Kraatz, 1899; 3 – C. sauteri Schenkling, 1912; 4 – C. terminalis Schenkling, 1912; 5 – C. pallidicornis Corporaal & van der Wiel, 1949; 6, 7, 10, 11 – C. hachijoensis sp. nov. (6, 10, holotype; 7, 11, paratype); 8, 9, 12, 13 – C. liaoi sp. nov. (8, 12, holotype; 9, 13, paratype). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4489552" httpUri="https://zenodo.org/record/4489552/files/figure.png" pageId="14" pageNumber="489">Figs 8, 12</figureCitation>
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). Head, pronotum, elytra yellowish brown; antennae and legs brownish black.
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</paragraph>
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<paragraph id="6B1936DCA749925CFF4679731DD4127A" blockId="14.[151,760,1730,2073]" pageId="14" pageNumber="489">
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Antennae (
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<figureCitation id="F39D2A59A749925CFEDA79731D4C123A" box="[297,374,1794,1817]" captionStart="Figs 14–26" captionStartId="3.[150,187,900,919]" captionTargetBox="[151,1434,200,863]" captionTargetId="figure-472@3.[151,1437,200,871]" captionText="Figs 14–26. Antennae (14, 16, 17, 19, 21, 23, 25 – male; 15, 18, 20, 22, 24, 26 – female). 14, 15 – Cladiscus obeliscus Lewis, 1892; 16 – C. weyersi Kraatz, 1899; 17, 18 – C. sauteri Schenkling, 1912; 19, 20 – C. terminalis Schenkling, 1912; 21, 22 – C. pallidicornis Corporaal & van der Wiel, 1949; 23, 24 – C. hachijoensis sp. nov.; 25, 26 – C. liaoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4551445" httpUri="https://zenodo.org/record/4551445/files/figure.png" pageId="14" pageNumber="489">Fig. 25</figureCitation>
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) longer than total length of head and pronotum combined, antennomeres III to X acutely serrate; XI fusiform, twice as long as X.
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</paragraph>
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<paragraph id="6B1936DCA749925CFF4679131FE61416" blockId="14.[151,760,1730,2073]" lastBlockId="14.[828,1437,190,1749]" pageId="14" pageNumber="489">
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Pronotum smooth at apex; postcoxal projections short; prosternal process long, completely separating postcoxal projections. Elytra with rows of deep, large, asetigerous punctures in basal 5/6; apical margins not indented. Mesoventrite with evenly and densely arranged setigerous punctures. Metaventral anterior process tapered and as wide as mesoventral posterior process. Metendosternite (
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<figureCitation id="F39D2A59A749925CFCB77EAF1F9815D6" box="[836,930,222,245]" captionStart="Figs 107–118" captionStartId="13.[151,188,1469,1488]" captionTargetBox="[151,1432,204,1437]" captionTargetId="figure-13@13.[151,1436,204,1437]" captionTargetPageId="13" captionText="Figs 107–118. Metendosternite, terminal parts and genitalia of Cladiscus liaoi sp. nov. (107–110, 113–115, male; 111, 112, 116–118, female). 107 – metendosternite; 108, 111 – pygidium; 109, 112 – ventrite VI; 110 – spicular fork; 113–115 – male genitalia in lateral (113), ventral (114), and dorsal (115) views; 116–118 – apices of ovipositor in lateral (116), ventral (117), and dorsal (118) views." figureDoi="http://doi.org/10.5281/zenodo.4489584" httpUri="https://zenodo.org/record/4489584/files/figure.png" pageId="14" pageNumber="489">Fig. 107</figureCitation>
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) with short furcal laminae; process extending anteriorly, tapered; furcal arms almost straight, extending anterolaterally.
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</paragraph>
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<paragraph id="6B1936DCA749925CFCA97F4F193E1456" blockId="14.[828,1437,190,1749]" pageId="14" pageNumber="489">
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Protibiae (
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<figureCitation id="F39D2A59A749925CFC3B7F4F182F1476" box="[968,1045,318,341]" captionStart="Figs 27–39" captionStartId="4.[151,188,2003,2022]" captionTargetBox="[170,1398,189,1963]" captionTargetId="figure-13@4.[170,1418,189,1973]" captionTargetPageId="4" captionText="Figs 27–39. Male protibiae (27–33) and prothorax in ventral view (34–39). 27, 34 – Cladiscus obeliscus Lewis, 1892; 28, 35 – C. weyersi Kraatz, 1899; 29, 36 – C. sauteri Schenkling, 1912; 30, 37 – C. terminalis Schenkling, 1912; 31, 38 – C. pallidicornis Corporaal & van der Wiel, 1949; 32, 39 – C. hachijoensis sp. nov.; 33 – C. liaoi sp. nov. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4489556" httpUri="https://zenodo.org/record/4489556/files/figure.png" pageId="14" pageNumber="489">Fig. 33</figureCitation>
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) gently widening from base to apex., weakly curved. Tibial spur formula 0–0–0.
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</paragraph>
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<paragraph id="6B1936DCA749925CFCA97F0F19041736" blockId="14.[828,1437,190,1749]" pageId="14" pageNumber="489">
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Apical margin of abdominal ventrite V emarginated broadly. Apical margin of pygidium (
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<figureCitation id="F39D2A59A749925CFB257FEF190E1496" box="[1238,1332,414,437]" captionStart="Figs 107–118" captionStartId="13.[151,188,1469,1488]" captionTargetBox="[151,1432,204,1437]" captionTargetId="figure-13@13.[151,1436,204,1437]" captionTargetPageId="13" captionText="Figs 107–118. Metendosternite, terminal parts and genitalia of Cladiscus liaoi sp. nov. (107–110, 113–115, male; 111, 112, 116–118, female). 107 – metendosternite; 108, 111 – pygidium; 109, 112 – ventrite VI; 110 – spicular fork; 113–115 – male genitalia in lateral (113), ventral (114), and dorsal (115) views; 116–118 – apices of ovipositor in lateral (116), ventral (117), and dorsal (118) views." figureDoi="http://doi.org/10.5281/zenodo.4489584" httpUri="https://zenodo.org/record/4489584/files/figure.png" pageId="14" pageNumber="489">Fig. 108</figureCitation>
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) rounded posteriorly; struts short. Apical margin of ventrite VI (
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<figureCitation id="F39D2A59A749925CFA827FCF1F5F14D6" captionStart="Figs 107–118" captionStartId="13.[151,188,1469,1488]" captionTargetBox="[151,1432,204,1437]" captionTargetId="figure-13@13.[151,1436,204,1437]" captionTargetPageId="13" captionText="Figs 107–118. Metendosternite, terminal parts and genitalia of Cladiscus liaoi sp. nov. (107–110, 113–115, male; 111, 112, 116–118, female). 107 – metendosternite; 108, 111 – pygidium; 109, 112 – ventrite VI; 110 – spicular fork; 113–115 – male genitalia in lateral (113), ventral (114), and dorsal (115) views; 116–118 – apices of ovipositor in lateral (116), ventral (117), and dorsal (118) views." figureDoi="http://doi.org/10.5281/zenodo.4489584" httpUri="https://zenodo.org/record/4489584/files/figure.png" pageId="14" pageNumber="489">Fig. 109</figureCitation>
|
|||
|
|
) weakly emarginated. Spicular fork (
|
|||
|
|
<figureCitation id="F39D2A59A749925CFB0C7FAF196614D6" box="[1279,1372,478,501]" captionStart="Figs 107–118" captionStartId="13.[151,188,1469,1488]" captionTargetBox="[151,1432,204,1437]" captionTargetId="figure-13@13.[151,1436,204,1437]" captionTargetPageId="13" captionText="Figs 107–118. Metendosternite, terminal parts and genitalia of Cladiscus liaoi sp. nov. (107–110, 113–115, male; 111, 112, 116–118, female). 107 – metendosternite; 108, 111 – pygidium; 109, 112 – ventrite VI; 110 – spicular fork; 113–115 – male genitalia in lateral (113), ventral (114), and dorsal (115) views; 116–118 – apices of ovipositor in lateral (116), ventral (117), and dorsal (118) views." figureDoi="http://doi.org/10.5281/zenodo.4489584" httpUri="https://zenodo.org/record/4489584/files/figure.png" pageId="14" pageNumber="489">Fig. 110</figureCitation>
|
|||
|
|
) with intraspicular plate, which is 1/3 of spicular fork.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6B1936DCA749925CFCA97C6F1FD71796" blockId="14.[828,1437,190,1749]" pageId="14" pageNumber="489">
|
|||
|
|
CM (
|
|||
|
|
<figureCitation id="F39D2A59A749925CFC617C6F181F1716" box="[914,1061,542,565]" captionStart="Figs 107–118" captionStartId="13.[151,188,1469,1488]" captionTargetBox="[151,1432,204,1437]" captionTargetId="figure-13@13.[151,1436,204,1437]" captionTargetPageId="13" captionText="Figs 107–118. Metendosternite, terminal parts and genitalia of Cladiscus liaoi sp. nov. (107–110, 113–115, male; 111, 112, 116–118, female). 107 – metendosternite; 108, 111 – pygidium; 109, 112 – ventrite VI; 110 – spicular fork; 113–115 – male genitalia in lateral (113), ventral (114), and dorsal (115) views; 116–118 – apices of ovipositor in lateral (116), ventral (117), and dorsal (118) views." figureDoi="http://doi.org/10.5281/zenodo.4489584" httpUri="https://zenodo.org/record/4489584/files/figure.png" pageId="14" pageNumber="489">Figs 113–115</figureCitation>
|
|||
|
|
) bearing fine spines. Tegmen shorter than phallus; phallobase incised dorsally; phallobasic struts reaching phallobasic apodeme, which is 3/4 as long as tegmen. Phallus flattened, divaricated in apical 1/3, rounded at apex.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6B1936DCA749925CFCA97CCF1FB71656" blockId="14.[828,1437,190,1749]" pageId="14" pageNumber="489">
|
|||
|
|
<emphasis id="59D2EACEA749925CFCA97CCF1F8917F6" bold="true" box="[858,947,702,725]" italics="true" pageId="14" pageNumber="489">Female.</emphasis>
|
|||
|
|
Similar to male (
|
|||
|
|
<figureCitation id="F39D2A59A749925CFB837CCF188817F6" box="[1136,1202,702,725]" captionStart="Figs 1–13" captionStartId="2.[151,188,2001,2020]" captionTargetBox="[174,1418,197,1920]" captionTargetId="figure-13@2.[166,1421,189,1930]" captionTargetPageId="2" captionText="Figs 1–13. Habitus and head of Cladiscus spp. (1–5, 6, 8, 10, 12, male; 7, 9, 11, 13, female). 1 – Cladiscus obeliscus Lewis, 1892; 2 – C. weyersi Kraatz, 1899; 3 – C. sauteri Schenkling, 1912; 4 – C. terminalis Schenkling, 1912; 5 – C. pallidicornis Corporaal & van der Wiel, 1949; 6, 7, 10, 11 – C. hachijoensis sp. nov. (6, 10, holotype; 7, 11, paratype); 8, 9, 12, 13 – C. liaoi sp. nov. (8, 12, holotype; 9, 13, paratype). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4489552" httpUri="https://zenodo.org/record/4489552/files/figure.png" pageId="14" pageNumber="489">Fig. 9</figureCitation>
|
|||
|
|
), but EyD/EyW 2.30; antennae (
|
|||
|
|
<figureCitation id="F39D2A59A749925CFC5A7CAF1FC217D6" box="[937,1016,734,757]" captionStart="Figs 14–26" captionStartId="3.[150,187,900,919]" captionTargetBox="[151,1434,200,863]" captionTargetId="figure-472@3.[151,1437,200,871]" captionText="Figs 14–26. Antennae (14, 16, 17, 19, 21, 23, 25 – male; 15, 18, 20, 22, 24, 26 – female). 14, 15 – Cladiscus obeliscus Lewis, 1892; 16 – C. weyersi Kraatz, 1899; 17, 18 – C. sauteri Schenkling, 1912; 19, 20 – C. terminalis Schenkling, 1912; 21, 22 – C. pallidicornis Corporaal & van der Wiel, 1949; 23, 24 – C. hachijoensis sp. nov.; 25, 26 – C. liaoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4551445" httpUri="https://zenodo.org/record/4551445/files/figure.png" pageId="14" pageNumber="489">Fig. 26</figureCitation>
|
|||
|
|
) weakly serrate; elytra pointed posteriorly at apex; central portion of metaventrite with pair of bundles of elongate setae-like trichobothria; apical margin of abdominal ventrite V truncated broadly and deeply in middle.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6B1936DCA749925CFCA97D0F188C16F6" blockId="14.[828,1437,190,1749]" pageId="14" pageNumber="489">
|
|||
|
|
Pygidium (
|
|||
|
|
<figureCitation id="F39D2A59A749925CFC277D0F180B16B6" box="[980,1073,894,917]" captionStart="Figs 107–118" captionStartId="13.[151,188,1469,1488]" captionTargetBox="[151,1432,204,1437]" captionTargetId="figure-13@13.[151,1436,204,1437]" captionTargetPageId="13" captionText="Figs 107–118. Metendosternite, terminal parts and genitalia of Cladiscus liaoi sp. nov. (107–110, 113–115, male; 111, 112, 116–118, female). 107 – metendosternite; 108, 111 – pygidium; 109, 112 – ventrite VI; 110 – spicular fork; 113–115 – male genitalia in lateral (113), ventral (114), and dorsal (115) views; 116–118 – apices of ovipositor in lateral (116), ventral (117), and dorsal (118) views." figureDoi="http://doi.org/10.5281/zenodo.4489584" httpUri="https://zenodo.org/record/4489584/files/figure.png" pageId="14" pageNumber="489">Fig. 111</figureCitation>
|
|||
|
|
) rounded posteriorly, with struts short. Ventrite VI (
|
|||
|
|
<figureCitation id="F39D2A59A749925CFBE17DEF184B1696" box="[1042,1137,926,949]" captionStart="Figs 107–118" captionStartId="13.[151,188,1469,1488]" captionTargetBox="[151,1432,204,1437]" captionTargetId="figure-13@13.[151,1436,204,1437]" captionTargetPageId="13" captionText="Figs 107–118. Metendosternite, terminal parts and genitalia of Cladiscus liaoi sp. nov. (107–110, 113–115, male; 111, 112, 116–118, female). 107 – metendosternite; 108, 111 – pygidium; 109, 112 – ventrite VI; 110 – spicular fork; 113–115 – male genitalia in lateral (113), ventral (114), and dorsal (115) views; 116–118 – apices of ovipositor in lateral (116), ventral (117), and dorsal (118) views." figureDoi="http://doi.org/10.5281/zenodo.4489584" httpUri="https://zenodo.org/record/4489584/files/figure.png" pageId="14" pageNumber="489">Fig. 112</figureCitation>
|
|||
|
|
) rounded posteriorly, with apodeme 2/3 as long as ventrite VI.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6B1936DCA749925CFCA97DAF196F1116" blockId="14.[828,1437,190,1749]" pageId="14" pageNumber="489">
|
|||
|
|
Spermatheca membranous; Sc lacking. CPr (
|
|||
|
|
<figureCitation id="F39D2A59A749925CFA997DAF1FA31136" captionStart="Figs 107–118" captionStartId="13.[151,188,1469,1488]" captionTargetBox="[151,1432,204,1437]" captionTargetId="figure-13@13.[151,1436,204,1437]" captionTargetPageId="13" captionText="Figs 107–118. Metendosternite, terminal parts and genitalia of Cladiscus liaoi sp. nov. (107–110, 113–115, male; 111, 112, 116–118, female). 107 – metendosternite; 108, 111 – pygidium; 109, 112 – ventrite VI; 110 – spicular fork; 113–115 – male genitalia in lateral (113), ventral (114), and dorsal (115) views; 116–118 – apices of ovipositor in lateral (116), ventral (117), and dorsal (118) views." figureDoi="http://doi.org/10.5281/zenodo.4489584" httpUri="https://zenodo.org/record/4489584/files/figure.png" pageId="14" pageNumber="489">Figs 116–118</figureCitation>
|
|||
|
|
) distally trifurcate; central process large; dorsolateral and ventrolateral processes relatively short.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6B1936DCA749925CFCCF7A4F197E10B6" blockId="14.[828,1437,190,1749]" pageId="14" pageNumber="489">
|
|||
|
|
<emphasis id="59D2EACEA749925CFCCF7A4F18A51176" bold="true" box="[828,1183,1086,1109]" pageId="14" pageNumber="489">
|
|||
|
|
Measurements and ratios.
|
|||
|
|
<emphasis id="59D2EACEA749925CFB947A4F18A51176" bold="true" box="[1127,1183,1086,1109]" italics="true" pageId="14" pageNumber="489">Male</emphasis>
|
|||
|
|
</emphasis>
|
|||
|
|
(n Y 1): BL:
|
|||
|
|
<quantity id="AC5E9B39A749925CFADA7A4F19A21176" box="[1321,1432,1086,1109]" metricMagnitude="-2" metricUnit="m" metricValue="1.018" pageId="14" pageNumber="489" unit="mm" value="10.18">10.18 mm</quantity>
|
|||
|
|
; PL:
|
|||
|
|
<quantity id="AC5E9B39A749925CFC987A2F1FEA1156" box="[875,976,1118,1141]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="14" pageNumber="489" unit="mm" value="2.18">2.18 mm</quantity>
|
|||
|
|
; PWA:
|
|||
|
|
<quantity id="AC5E9B39A749925CFBD57A2F18B11156" box="[1062,1163,1118,1141]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="14" pageNumber="489" unit="mm" value="1.28">1.28 mm</quantity>
|
|||
|
|
; PWM:
|
|||
|
|
<quantity id="AC5E9B39A749925CFB1A7A2F19741156" box="[1257,1358,1118,1141]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="489" unit="mm" value="0.48">0.48 mm</quantity>
|
|||
|
|
; PWB:
|
|||
|
|
<quantity id="AC5E9B39A749925CFCCF7A0F1F9A11B6" box="[828,928,1150,1173]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="14" pageNumber="489" unit="mm" value="1.03">1.03 mm</quantity>
|
|||
|
|
; EL: 8.00 mm; EW:
|
|||
|
|
<quantity id="AC5E9B39A749925CFB777A0F18D211B6" box="[1156,1256,1150,1173]" metricMagnitude="-3" metricUnit="m" metricValue="1.83" pageId="14" pageNumber="489" unit="mm" value="1.83">1.83 mm</quantity>
|
|||
|
|
; EyD:
|
|||
|
|
<quantity id="AC5E9B39A749925CFAC77A0F19A211B6" box="[1332,1432,1150,1173]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="489" unit="mm" value="0.35">0.35 mm</quantity>
|
|||
|
|
; EyW:
|
|||
|
|
<quantity id="AC5E9B39A749925CFC8D7AEF1FE41196" box="[894,990,1182,1205]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="14" pageNumber="489" unit="mm" value="0.63">0.63 mm</quantity>
|
|||
|
|
; EL/PL: 3.68; EW/PWA: 1.43; EW/PWM: 3.84; EW/PWB: 1.78; PWA/PWB: 1.24; PWA/PWM: 2.68; PWB/PWM: 2.16; EyD/EyW: 0.56.
|
|||
|
|
<emphasis id="59D2EACEA749925CFB327AAF192E11D6" bold="true" box="[1217,1300,1246,1269]" italics="true" pageId="14" pageNumber="489">Female</emphasis>
|
|||
|
|
(n Y 1): BL:
|
|||
|
|
<quantity id="AC5E9B39A749925CFCCF7A8F1FA71036" box="[828,925,1278,1301]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="14" pageNumber="489" unit="mm" value="13.4">13.4 mm</quantity>
|
|||
|
|
; PL:
|
|||
|
|
<quantity id="AC5E9B39A749925CFC217A8F18091036" box="[978,1075,1278,1301]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="14" pageNumber="489" unit="mm" value="2.18">2.18 mm</quantity>
|
|||
|
|
; PWA:
|
|||
|
|
<quantity id="AC5E9B39A749925CFB727A8F18D81036" box="[1153,1250,1278,1301]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="14" pageNumber="489" unit="mm" value="1.28">1.28 mm</quantity>
|
|||
|
|
; PWM:
|
|||
|
|
<quantity id="AC5E9B39A749925CFAC47A8F19A21036" box="[1335,1432,1278,1301]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="14" pageNumber="489" unit="mm" value="0.98">0.98 mm</quantity>
|
|||
|
|
; PWB:
|
|||
|
|
<quantity id="AC5E9B39A749925CFC707B6F1FDC1016" box="[899,998,1310,1333]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" pageId="14" pageNumber="489" unit="mm" value="1.43">1.43 mm</quantity>
|
|||
|
|
; EL:
|
|||
|
|
<quantity id="AC5E9B39A749925CFBEC7B6F18BB1016" box="[1055,1153,1310,1333]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="14" pageNumber="489" unit="mm" value="10.9">10.9 mm</quantity>
|
|||
|
|
; EW:
|
|||
|
|
<quantity id="AC5E9B39A749925CFB317B6F191E1016" box="[1218,1316,1310,1333]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="14" pageNumber="489" unit="mm" value="2.2">2.20 mm</quantity>
|
|||
|
|
; EyD:
|
|||
|
|
<quantity id="AC5E9B39A749925CFA9D7B6F1F531076" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="14" pageNumber="489" unit="mm" value="1.15">1.15 mm</quantity>
|
|||
|
|
; EyW:
|
|||
|
|
<quantity id="AC5E9B39A749925CFC447B4F18201076" box="[951,1050,1342,1365]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="489" unit="mm" value="0.5">0.50 mm</quantity>
|
|||
|
|
; EL/PL: 4.36; EW/PWA: 1.21; EW/ PWM: 2.26; EW/PWB: 1.54; PWA/PWB: 1.28; PWA/ PWM: 1.87; PWB/PWM: 1.46; EyD/EyW: 2.30.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="23BC6557A749925CFCCF7BEF1FDC1376" pageId="14" pageNumber="489" type="diagnosis">
|
|||
|
|
<paragraph id="6B1936DCA749925CFCCF7BEF1FDC1376" blockId="14.[828,1437,190,1749]" pageId="14" pageNumber="489">
|
|||
|
|
<emphasis id="59D2EACEA749925CFCCF7BEF18041096" bold="true" box="[828,1086,1438,1461]" pageId="14" pageNumber="489">Differential diagnosis.</emphasis>
|
|||
|
|
This new species is similar to
|
|||
|
|
<taxonomicName id="ACA64D5FA749925CFCCF7BCF1FF010F6" authorityName="Schenkling" authorityYear="1912" box="[828,970,1470,1493]" class="Insecta" family="Cleridae" genus="Cladiscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="489" phylum="Arthropoda" rank="species" species="terminalis">
|
|||
|
|
<emphasis id="59D2EACEA749925CFCCF7BCF1FF010F6" box="[828,970,1470,1493]" italics="true" pageId="14" pageNumber="489">C. terminalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, but it is distinguishable by the following characters: EyD/EyW
|
|||
|
|
<quantity id="AC5E9B39A749925CFBDA7BAF184810D6" box="[1065,1138,1502,1525]" metricMagnitude="-2" metricUnit="m" metricValue="1.4224" pageId="14" pageNumber="489" unit="in" value="0.56">0.56 in</quantity>
|
|||
|
|
male,
|
|||
|
|
<quantity id="AC5E9B39A749925CFB4B7BAF193B10D6" box="[1208,1281,1502,1525]" metricMagnitude="-2" metricUnit="m" metricValue="5.842" pageId="14" pageNumber="489" unit="in" value="2.3">2.30 in</quantity>
|
|||
|
|
female; PWA/ PWB 1.24–1.28; female antennae (
|
|||
|
|
<figureCitation id="F39D2A59A749925CFB357B8F19221336" box="[1222,1304,1534,1557]" captionStart="Figs 14–26" captionStartId="3.[150,187,900,919]" captionTargetBox="[151,1434,200,863]" captionTargetId="figure-472@3.[151,1437,200,871]" captionText="Figs 14–26. Antennae (14, 16, 17, 19, 21, 23, 25 – male; 15, 18, 20, 22, 24, 26 – female). 14, 15 – Cladiscus obeliscus Lewis, 1892; 16 – C. weyersi Kraatz, 1899; 17, 18 – C. sauteri Schenkling, 1912; 19, 20 – C. terminalis Schenkling, 1912; 21, 22 – C. pallidicornis Corporaal & van der Wiel, 1949; 23, 24 – C. hachijoensis sp. nov.; 25, 26 – C. liaoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4551445" httpUri="https://zenodo.org/record/4551445/files/figure.png" pageId="14" pageNumber="489">Fig. 26</figureCitation>
|
|||
|
|
) as long as male (
|
|||
|
|
<figureCitation id="F39D2A59A749925CFC73786F1FEB1316" box="[896,977,1566,1589]" captionStart="Figs 14–26" captionStartId="3.[150,187,900,919]" captionTargetBox="[151,1434,200,863]" captionTargetId="figure-472@3.[151,1437,200,871]" captionText="Figs 14–26. Antennae (14, 16, 17, 19, 21, 23, 25 – male; 15, 18, 20, 22, 24, 26 – female). 14, 15 – Cladiscus obeliscus Lewis, 1892; 16 – C. weyersi Kraatz, 1899; 17, 18 – C. sauteri Schenkling, 1912; 19, 20 – C. terminalis Schenkling, 1912; 21, 22 – C. pallidicornis Corporaal & van der Wiel, 1949; 23, 24 – C. hachijoensis sp. nov.; 25, 26 – C. liaoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4551445" httpUri="https://zenodo.org/record/4551445/files/figure.png" pageId="14" pageNumber="489">Fig. 25</figureCitation>
|
|||
|
|
); whole elytra yellowish brown, in female pointed at apex.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="23BC6557A749925CFCCF782F18591396" pageId="14" pageNumber="489" type="etymology">
|
|||
|
|
<paragraph id="6B1936DCA749925CFCCF782F18591396" blockId="14.[828,1437,190,1749]" pageId="14" pageNumber="489">
|
|||
|
|
<emphasis id="59D2EACEA749925CFCCF782F1F861356" bold="true" box="[828,956,1630,1653]" pageId="14" pageNumber="489">Etymology.</emphasis>
|
|||
|
|
The species name is dedicated to Mr. Wen -Chuan Liao, the collector of the
|
|||
|
|
<typeStatus id="B41D887EA749925CFB7E780F18D613B6" box="[1165,1260,1662,1685]" pageId="14" pageNumber="489" type="holotype">holotype</typeStatus>
|
|||
|
|
; noun in genitive case standing in apposition.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="23BC6557A749925CFCCF78CF18A113F6" box="[828,1179,1726,1749]" pageId="14" pageNumber="489" type="distribution">
|
|||
|
|
<paragraph id="6B1936DCA749925CFCCF78CF18A113F6" blockId="14.[828,1437,190,1749]" box="[828,1179,1726,1749]" pageId="14" pageNumber="489">
|
|||
|
|
<emphasis id="59D2EACEA749925CFCCF78CF1FFD13F6" bold="true" box="[828,967,1726,1749]" pageId="14" pageNumber="489">Distribution</emphasis>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="F39D2A59A749925CFC2578CF180913F6" box="[982,1075,1726,1749]" captionStart="Fig 126" captionStartId="16.[151,179,1278,1297]" captionTargetBox="[252,1335,189,1259]" captionTargetId="figure-329@16.[252,1335,189,1259]" captionTargetPageId="16" captionText="Fig 126. Distribution map of the species of Cladiscus from Japan and Taiwan." figureDoi="http://doi.org/10.5281/zenodo.4489596" httpUri="https://zenodo.org/record/4489596/files/figure.png" pageId="14" pageNumber="489">Fig. 126</figureCitation>
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).
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<collectingCountry id="13B1764CA749925CFBB678CF18AD13F6" box="[1093,1175,1726,1749]" name="Taiwan" pageId="14" pageNumber="489">Taiwan</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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