treatments-xml/data/DF/18/F9/DF18F971FFD40B3CFF4BFD076D5FBED4.xml

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<mods:title id="E26553D49A41DB14445A9A3EF057AB90">Overview of the lady beetle tribe Diomini (Coleoptera: Coccinellidae) and description of a new phytophagous, silk-spinning genus from Costa Rica that induces food bodies on leaves of Piper (Piperaceae)</mods:title>
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<mods:namePart id="57F183905491175DD68C261F15E46098">Vandenberg, Natalia J.</mods:namePart>
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<paragraph id="570E4867FFD40B3DFF4BFD076DA7BF3F" blockId="22.[151,499,766,826]" box="[151,499,766,792]" pageId="22" pageNumber="277">
<heading id="0C46FF0BFFD40B3DFF4BFD076DA7BF3F" bold="true" box="[151,499,766,792]" fontSize="11" level="1" pageId="22" pageNumber="277" reason="1">
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<taxonomicName id="90B133E4FFD40B3DFF4BFD076DDBBF3F" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[151,399,766,792]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="277" phylum="Arthropoda" rank="species" species="nanita" status="sp. nov.">
<emphasis id="65C59475FFD40B3DFF4BFD076DDBBF3F" bold="true" box="[151,399,766,792]" italics="true" pageId="22" pageNumber="277">Moiradiomus nanita</emphasis>
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<taxonomicNameLabel id="7EF6290EFFD40B3DFE4AFD076DA7BF3F" box="[406,499,766,792]" pageId="22" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="1FAB1BECFFD40B3DFF4BFCD86DFFBF1D" box="[151,427,801,826]" pageId="22" pageNumber="277" type="description">
<paragraph id="570E4867FFD40B3DFF4BFCD86DFFBF1D" blockId="22.[151,499,766,826]" box="[151,427,801,826]" pageId="22" pageNumber="277">
(
<figureCitation id="CF8A54E2FFD40B3DFF43FCD86D72BF1D" box="[159,294,801,826]" captionStart="FIGURES 17–20" captionStartId="14.[151,264,1965,1988]" captionTargetBox="[151,1436,260,1911]" captionTargetId="figure@14.[151,1436,260,1911]" captionTargetPageId="14" captionText="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.2623573" httpUri="https://zenodo.org/record/2623573/files/figure.png" pageId="22" pageNumber="277">Figs. 19, 20</figureCitation>
, 
<figureCitation id="CF8A54E2FFD40B3DFEEEFCDB6D05BF1D" box="[306,337,802,826]" captionStart="FIGURES 25–28" captionStartId="15.[151,264,1975,1998]" captionTargetBox="[151,1436,1466,1918]" captionTargetId="figure@15.[151,1436,1461,1924]" captionTargetPageId="15" captionText="FIGURES 25–28. Diagrammatic ventral view of Moiradiomus prosterna (setae, color patterns, and surface punctation not indicated): 25, M. clotho; 26, M. lachesis; 27, M. atopos; 28, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623578" httpUri="https://zenodo.org/record/2623578/files/figure.png" pageId="22" pageNumber="277">28</figureCitation>
, 
<figureCitation id="CF8A54E2FFD40B3DFE81FCDB6D2FBF1D" box="[349,379,802,826]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="22" pageNumber="277">32</figureCitation>
, 
<figureCitation id="CF8A54E2FFD40B3DFE5AFCDB6DF7BF1D" box="[390,419,802,826]" captionStart="FIGURES 40–42" captionStartId="20.[151,264,1957,1980]" captionTargetBox="[151,1436,193,1936]" captionTargetId="figure@20.[151,1436,193,1936]" captionTargetPageId="20" captionText="FIGURES 40–42. Tents of Moiradiomus spp.: 40, M. lachesis on Piper lanosibracteum; 41, M. nanita on P. reticulatum; 42, M. atropos on P. friedrichsthalii." figureDoi="http://doi.org/10.5281/zenodo.2623584" httpUri="https://zenodo.org/record/2623584/files/figure.png" pageId="22" pageNumber="277">41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1FAB1BECFFD40B3DFF4BFC906CA8B812" pageId="22" pageNumber="277" type="diagnosis">
<paragraph id="570E4867FFD40B3DFF4BFC906CA8B812" blockId="22.[151,1437,873,2017]" pageId="22" pageNumber="277">
<emphasis id="65C59475FFD40B3DFF4BFC906D45BFA5" bold="true" box="[151,273,873,898]" pageId="22" pageNumber="277">Diagnosis.</emphasis>
This species is readily distinguished from its congeners by the male genitalia (
<figureCitation id="CF8A54E2FFD40B3DFB47FC9068BABFA5" box="[1179,1262,873,898]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="22" pageNumber="277">Fig. 32</figureCitation>
) with its short, broad paramere (length only 2× width), basal lobe in ventral view strongly tapered toward apex with rounded apical knob, and penis tube with slight angulation and irregular swelling near distal ½. In addition, the adult form (
<figureCitation id="CF8A54E2FFD40B3DFF43FC2C6D7EBFC9" box="[159,298,981,1006]" captionStart="FIGURES 17–20" captionStartId="14.[151,264,1965,1988]" captionTargetBox="[151,1436,260,1911]" captionTargetId="figure@14.[151,1436,260,1911]" captionTargetPageId="14" captionText="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.2623573" httpUri="https://zenodo.org/record/2623573/files/figure.png" pageId="22" pageNumber="277">Figs. 19, 20</figureCitation>
) is smaller and relatively shorter than in the other species, and the prosternal carinae (
<figureCitation id="CF8A54E2FFD40B3DFAC8FC2C693CBFC9" box="[1300,1384,981,1006]" captionStart="FIGURES 25–28" captionStartId="15.[151,264,1975,1998]" captionTargetBox="[151,1436,1466,1918]" captionTargetId="figure@15.[151,1436,1461,1924]" captionTargetPageId="15" captionText="FIGURES 25–28. Diagrammatic ventral view of Moiradiomus prosterna (setae, color patterns, and surface punctation not indicated): 25, M. clotho; 26, M. lachesis; 27, M. atopos; 28, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623578" httpUri="https://zenodo.org/record/2623578/files/figure.png" pageId="22" pageNumber="277">Fig. 28</figureCitation>
) are nearly parallel, framing a subtrapezoidal depression. It is also distinguished by its larval host plant: 
<taxonomicName id="90B133E4FFD40B3DFB23FC0369C8B835" box="[1279,1436,1017,1042]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="22" pageNumber="277" phylum="Tracheophyta" rank="species" species="reticulatum">
<emphasis id="65C59475FFD40B3DFB23FC0369C8B835" box="[1279,1436,1017,1042]" italics="true" pageId="22" pageNumber="277">P. reticulatum</emphasis>
</taxonomicName>
L., 1753.
</paragraph>
</subSubSection>
<subSubSection id="1FAB1BECFFD40B3CFF1BFBB86879BDF4" lastPageId="23" lastPageNumber="278" pageId="22" pageNumber="277" type="description">
<paragraph id="570E4867FFD40B3DFF1BFBB86912BBBE" blockId="22.[151,1437,873,2017]" pageId="22" pageNumber="277">
<emphasis id="65C59475FFD40B3DFF1BFBB86EEEB87D" bold="true" box="[199,698,1089,1114]" pageId="22" pageNumber="277">
Description of 
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(male) (
<figureCitation id="CF8A54E2FFD40B3DFD8FFBB86EF8B87D" box="[595,684,1089,1114]" captionStart="FIGURES 17–20" captionStartId="14.[151,264,1965,1988]" captionTargetBox="[151,1436,260,1911]" captionTargetId="figure@14.[151,1436,260,1911]" captionTargetPageId="14" captionText="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.2623573" httpUri="https://zenodo.org/record/2623573/files/figure.png" pageId="22" pageNumber="277">Fig. 19</figureCitation>
).
</emphasis>
Length 
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, width 
<quantity id="9049E582FFD40B3DFC06FBBB686CB87E" box="[986,1080,1090,1114]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="277" unit="mm" value="0.9">0.9 mm</quantity>
. Form shortened, oval weakly convex. Metathoracic wing present. Dorsal surfaces feebly shining, distinctly punctate, pubescent with fimbriate borders. Head, pronotum, appendages including mouthparts, straw yellow, with meso- metacoxae slightly darkened; base of pronotum in front of scutellar shield with light brown smudge; scutellar shield light brown; elytron light brown with apex narrowly yellow brown, epipleuron yellow brown, nearly transparent; pubescence shiny, off white. Venter predominantly light brown; prosternum pale yellow; mesepimeron, mesepisternum paler than surrounding sclerites except posterior margin of mesepimeron narrowly darkened; abdomen light brown, lighter, yellower toward apex; tarsal claw, mandibular apex dark reddish amber. Dorsal punctation moderately coarse; punctures separated by approximately 1× diameter, on elytron equal in width to eye facet, finer on head, pronotum. Dorsal pubescence semi erect, moderately dense, evenly distributed, lacking distinct setal pattern; individual setae equal to about 2/3 to 1× length of scutellar shield, arcuate; elytral setae mostly directed posteriorly except directed outwardly at sides of body, posterolaterally near suture. Head large, 0.80× width of pronotum; eye large, finely facetted, with interfacetal setae, weakly notched near antennal insertion by small rounded ocular canthus; inner orbits arcuate, strongly diverging toward vertex, weakly diverging toward clypeus, with minimum separation at slightly below midlength; interocular distance 1.2× width of eye in frontal view. Antenna composed of 10 antennomeres; antennomere 3 elongate, about 1.8× length of antennomere 4; last 4 antennomeres forming gradual club expanding apically from antennomere 7–9; antennomere 10 subrhomboidal, slightly narrower, slightly longer than 9. Maxillary palp with terminal palpomere short, strongly expanded distally; with oblique apex. Pronotum transverse, width 2.0× length, weakly, evenly convex. Elytron in dorsal view with lateral margin weakly arcuate; elytral apices dehiscent. Ventral surfaces pubescent, distinctly punctate, except glabrous, impunctate in posterior 4/5ths of metacoxal plate; punctation coarser, denser on abdomen. Prosternum (
<figureCitation id="CF8A54E2FFD40B3DFB52F8E868B4BB0D" box="[1166,1248,1809,1834]" captionStart="FIGURES 25–28" captionStartId="15.[151,264,1975,1998]" captionTargetBox="[151,1436,1466,1918]" captionTargetId="figure@15.[151,1436,1461,1924]" captionTargetPageId="15" captionText="FIGURES 25–28. Diagrammatic ventral view of Moiradiomus prosterna (setae, color patterns, and surface punctation not indicated): 25, M. clotho; 26, M. lachesis; 27, M. atopos; 28, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623578" httpUri="https://zenodo.org/record/2623578/files/figure.png" pageId="22" pageNumber="277">Fig. 28</figureCitation>
) with carinae of intercoxal process extending to apex, subparallel, framing subtrapezoidal depression. Suture between abdominal ventrites 1 and 2 distinct, linear; 5 
<superScript id="A0C4E52FFFD40B3DFDCDF8A16E49BB41" attach="left" box="[529,541,1880,1894]" fontSize="6" pageId="22" pageNumber="277">th</superScript>
ventrite with apex truncate; 6 
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ventrite with apex shallowly, roundly emarginate in median ½. Tarsal claw with broad scythe-like inner tooth extending from base to apical 1/3rd or beyond.
</paragraph>
<paragraph id="570E4867FFD40B3CFF1BF85D6F2ABCDF" blockId="22.[151,1437,873,2017]" lastBlockId="23.[151,1437,151,755]" lastPageId="23" lastPageNumber="278" pageId="22" pageNumber="277">
<emphasis id="65C59475FFD40B3DFF1BF85D6DB3BB9A" bold="true" box="[199,487,1956,1981]" pageId="22" pageNumber="277">
Male genitalia (
<figureCitation id="CF8A54E2FFD40B3DFE5FF85D6D8EBB9A" box="[387,474,1956,1981]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="22" pageNumber="277">Fig. 32</figureCitation>
).
</emphasis>
Basal lobe in ventral view (
<figureCitation id="CF8A54E2FFD40B3DFCF7F85D6FD8BB9A" box="[811,908,1956,1981]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="22" pageNumber="277">Fig. 32b</figureCitation>
) strongly, unevenly tapered toward apex, with small knob at distal most point, slightly shorter than paramere; in lateral view (
<figureCitation id="CF8A54E2FFD40B3DFBCCF831683BBBC7" box="[1040,1135,1992,2017]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="22" pageNumber="277">Fig. 32a</figureCitation>
) nearly flat in basal 2/3rds sinuous in apical third with upturned apex; paramere short and broad, strongly expanded toward obliquely rounded apex, about 3.0× as long as wide, slightly longer than longest setae. Penis (
<figureCitation id="CF8A54E2FFD50B3CFBD7FF456839BCF3" box="[1035,1133,188,213]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="23" pageNumber="278">Fig. 32c</figureCitation>
) lacking apical flagellum, slender with irregular swelling at apical ½; capsule wedge shaped.
</paragraph>
<paragraph id="570E4867FFD50B3CFF1BFEFD6875BD40" blockId="23.[151,1437,151,755]" pageId="23" pageNumber="278">
<emphasis id="65C59475FFD50B3CFF1BFEFD6DC1BD3A" bold="true" box="[199,405,260,285]" pageId="23" pageNumber="278">
Female (
<figureCitation id="CF8A54E2FFD50B3CFEECFEFD6DDCBD3A" box="[304,392,260,285]" captionStart="FIGURES 17–20" captionStartId="14.[151,264,1965,1988]" captionTargetBox="[151,1436,260,1911]" captionTargetId="figure@14.[151,1436,260,1911]" captionTargetPageId="14" captionText="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.2623573" httpUri="https://zenodo.org/record/2623573/files/figure.png" pageId="23" pageNumber="278">Fig. 20</figureCitation>
).
</emphasis>
Similar to male except pronotum light brown; prosternum, mesepimeron, mesepisternum medium brown. 5 
<superScript id="A0C4E52FFFD50B3CFEBEFEDE6D3ABD12" attach="left" box="[354,366,295,309]" fontSize="6" pageId="23" pageNumber="278">th</superScript>
abdominal ventrite with posterior margin truncate; 6 
<superScript id="A0C4E52FFFD50B3CFC13FEDE6F8FBD12" attach="left" box="[975,987,295,309]" fontSize="6" pageId="23" pageNumber="278">th</superScript>
ventrite with posterior margin arcuate. Tarsal claw with short triangular tooth near base, not extending beyond apical ½.
</paragraph>
<paragraph id="570E4867FFD50B3CFF1BFE8B6D9EBD88" blockId="23.[151,1437,151,755]" pageId="23" pageNumber="278">
<emphasis id="65C59475FFD50B3CFF1BFE8B6D68BDAC" bold="true" box="[199,316,370,395]" pageId="23" pageNumber="278">Variation.</emphasis>
Length 
<quantity id="9049E582FFD50B3CFE7EFE8B6E64BDAD" box="[418,560,370,395]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="23" pageNumber="278" unit="mm" value="1.3" valueMax="1.5" valueMin="1.1">1.1–1.5 mm</quantity>
. Individuals vary slightly in the extent and degree of dark pigmentation on dorsal and ventral surfaces.
</paragraph>
<paragraph id="570E4867FFD50B3CFF1BFE436879BDF4" blockId="23.[151,1437,151,755]" box="[199,1069,442,467]" pageId="23" pageNumber="278">
<emphasis id="65C59475FFD50B3CFF1BFE436DCEBDF4" bold="true" box="[199,410,442,467]" pageId="23" pageNumber="278">Trophic relations.</emphasis>
Larvae develop on food bodies of 
<taxonomicName id="90B133E4FFD50B3CFCFDFE456FE8BDF4" box="[801,956,442,467]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="23" pageNumber="278" phylum="Tracheophyta" rank="species" species="reticulatum">
<emphasis id="65C59475FFD50B3CFCFDFE456FE8BDF4" box="[801,956,442,467]" italics="true" pageId="23" pageNumber="278">P. reticulatum</emphasis>
</taxonomicName>
(
<figureCitation id="CF8A54E2FFD50B3CFC10FE43684ABDF4" box="[972,1054,442,467]" captionStart="FIGURES 40–42" captionStartId="20.[151,264,1957,1980]" captionTargetBox="[151,1436,193,1936]" captionTargetId="figure@20.[151,1436,193,1936]" captionTargetPageId="20" captionText="FIGURES 40–42. Tents of Moiradiomus spp.: 40, M. lachesis on Piper lanosibracteum; 41, M. nanita on P. reticulatum; 42, M. atropos on P. friedrichsthalii." figureDoi="http://doi.org/10.5281/zenodo.2623584" httpUri="https://zenodo.org/record/2623584/files/figure.png" pageId="23" pageNumber="278">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1FAB1BECFFD50B3CFF1BFE276D16BE44" pageId="23" pageNumber="278" type="etymology">
<paragraph id="570E4867FFD50B3CFF1BFE276D16BE44" blockId="23.[151,1437,151,755]" pageId="23" pageNumber="278">
<emphasis id="65C59475FFD50B3CFF1BFE276D1ABDD0" bold="true" box="[199,334,478,503]" pageId="23" pageNumber="278">Etymology.</emphasis>
Nanita, proper noun in apposition, from Nan (a version of Ann) + ita (diminutive). A reference to an old nursery rhyme or cantrip used to encourage a coccinellid to fly from your fingertip: 
<emphasis id="65C59475FFD50B3CFB42FDFB6D16BE44" italics="true" pageId="23" pageNumber="278">Ladybird, ladybird, fly away home. Your house is on fire. Your children all roam, except little Nan, who sits in a pan, weaving gold laces as fast as she can.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="1FAB1BECFFD50B3CFF1BFD976D5FBED4" pageId="23" pageNumber="278" type="materials_examined">
<paragraph id="570E4867FFD50B3CFF1BFD976D5FBED4" blockId="23.[151,1437,151,755]" pageId="23" pageNumber="278">
<emphasis id="65C59475FFD50B3CFF1BFD976D2EBEA0" bold="true" box="[199,378,622,647]" pageId="23" pageNumber="278">Type material.</emphasis>
<materialsCitation id="E7D9423AFFD50B3CFE5BFD976D5FBED4" ID-GBIF-Occurrence="2234228121" collectingDate="2013-02" collectingDateMax="2017-01" collectingDateMin="2013-02" collectionCode="USNM, MZUCR" collectorName="P. Hanson" country="Costa Rica" elevation="100" location="Selva" pageId="23" pageNumber="278" specimenCount="18" specimenCount-female="5" specimenCount-male="4" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="880AF6C5FFD50B3CFE5BFD976DBBBEA0" box="[391,495,622,647]" pageId="23" pageNumber="278" type="holotype">Holotype</typeStatus>
(male) with labels: “ 
<collectingCountry id="2FA608F7FFD50B3CFD33FD966FF4BEA0" box="[751,928,623,647]" name="Costa Rica" pageId="23" pageNumber="278">COSTA RICA</collectingCountry>
: 
<collectingRegion id="95758685FFD50B3CFC73FD976858BEA0" box="[943,1036,622,647]" country="Costa Rica" name="Heredia" pageId="23" pageNumber="278">Heredia</collectingRegion>
: OTS-La 
<location id="526E1EBCFFD50B3CFB59FD976892BEA0" LSID="urn:lsid:plazi:treatment:DF18F971FFD40B3CFF4BFD076D5FBED4:526E1EBCFFD50B3CFB59FD976892BEA0" box="[1157,1222,622,647]" country="Costa Rica" name="Selva" pageId="23" pageNumber="278" stateProvince="Heredia">Selva</location>
, 
<quantity id="9049E582FFD50B3CFB0BFD966943BEA0" box="[1239,1303,623,647]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="278" unit="m" value="100.0">
<elevation id="DC9CAF54FFD50B3CFB0BFD966943BEA0" box="[1239,1303,623,647]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="278" unit="m" value="100.0">100m</elevation>
</quantity>
<date id="230F6EA7FFD50B3CFAFFFD96692CBEA0" box="[1315,1400,623,647]" pageId="23" pageNumber="278" value="2013-02">
<collectingDate id="334B974FFFD50B3CFAFFFD96692CBEA0" box="[1315,1400,623,647]" pageId="23" pageNumber="278" value="2013-02">II.2013</collectingDate>
</date>
, 
<collectorName id="FA442DB1FFD50B3CFA54FD896CB8BE8D" pageId="23" pageNumber="278">P. Hanson</collectorName>
/ ex. 
<taxonomicName id="90B133E4FFD50B3CFEF7FD6D6E3FBE8C" authority="&quot; (USNM)" baseAuthorityName="USNM" box="[299,619,658,683]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="23" pageNumber="278" phylum="Tracheophyta" rank="species" species="reticulatum">
<emphasis id="65C59475FFD50B3CFEF7FD6D6DA4BE8C" box="[299,496,658,683]" italics="true" pageId="23" pageNumber="278">Piper reticulatum</emphasis>
” (
<collectionCode id="31A0D0A2FFD50B3CFDD7FD6A6E36BE8C" box="[523,610,659,683]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="278" type="Museum">USNM</collectionCode>
)
</taxonomicName>
; 
<specimenCount id="41B783EEFFD50B3CFDAAFD6A6EA2BE8C" box="[630,758,659,683]" pageId="23" pageNumber="278" type="generic" typeStatus="paratypes">9 paratypes</specimenCount>
(
<specimenCount id="41B783EEFFD50B3CFCDAFD6A6F34BE8C" box="[774,864,658,683]" pageId="23" pageNumber="278" type="male">4 males</specimenCount>
, 
<specimenCount id="41B783EEFFD50B3CFCB7FD6B6F89BE8C" box="[875,989,658,683]" pageId="23" pageNumber="278" type="female">5 females</specimenCount>
): 8 with same labels as 
<typeStatus id="880AF6C5FFD50B3CFB38FD6B691EBE8C" box="[1252,1354,658,683]" pageId="23" pageNumber="278" type="holotype">holotype</typeStatus>
; 1 with labels: “ 
<collectingCountry id="2FA608F7FFD50B3CFF2FFD4E6DCBBEE8" box="[243,415,695,719]" name="Costa Rica" pageId="23" pageNumber="278">COSTA RICA</collectingCountry>
: 
<collectingRegion id="95758685FFD50B3CFE70FD416E7FBEE8" box="[428,555,696,719]" country="Costa Rica" name="Puntarenas" pageId="23" pageNumber="278">Puntarenas</collectingRegion>
: Rincon de la Osa, 
<date id="230F6EA7FFD50B3CFCD3FD4E6F0FBEE8" box="[783,859,695,719]" pageId="23" pageNumber="278" value="2017-01">
<collectingDate id="334B974FFFD50B3CFCD3FD4E6F0FBEE8" box="[783,859,695,719]" pageId="23" pageNumber="278" value="2017-01">I.2017</collectingDate>
</date>
, 
<collectorName id="FA442DB1FFD50B3CFCB4FD416F8FBEE8" box="[872,987,696,719]" pageId="23" pageNumber="278">P. Hanson</collectorName>
/ ex. 
<taxonomicName id="90B133E4FFD50B3CFBFCFD4168B3BEE8" box="[1056,1255,694,719]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="23" pageNumber="278" phylum="Tracheophyta" rank="species" species="reticulatum">
<emphasis id="65C59475FFD50B3CFBFCFD4168B3BEE8" box="[1056,1255,694,719]" italics="true" pageId="23" pageNumber="278">Piper reticulatum</emphasis>
</taxonomicName>
” (5, 
<collectionCode id="31A0D0A2FFD50B3CFAF9FD4E692EBEE8" box="[1317,1402,695,719]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="278" type="Museum">USNM</collectionCode>
; 4, 
<collectionCode id="31A0D0A2FFD50B3CFF4BFD226D54BED4" box="[151,256,731,755]" pageId="23" pageNumber="278">MZUCR</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>