188 lines
17 KiB
XML
188 lines
17 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.669.12608" ID-GBIF-Dataset="dbfccdc4-e5a8-42d2-a706-cfa0062ebe65" ID-PMC="PMC5523371" ID-Pensoft-Pub="1313-2970-669-29" ID-PubMed="28769651" ID-ZBK="7BC01A4ED22C44B38D2DA0DF2CD53AA4" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-669-29" ModsDocOrigin="ZooKeys 669" ModsDocTitle="Taxonomic revision of the endemic Bornean genera Anexodus Pascoe and Pantilema Aurivillius (Coleoptera, Cerambycidae, Lamiinae)" checkinTime="1492772871763" checkinUser="pensoft" docAuthor="Gabris, Radim, Trnka, Filip, Wahab, Rodzay Abdul & Kundrata, Robin" docDate="2017" docId="CD57FE8BC8FCC74CE0F84A5AEAD21984" docLanguage="en" docName="ZooKeys 669: 29-51" docOrigin="ZooKeys 669" docSource="http://dx.doi.org/10.3897/zookeys.669.12608" docTitle="Anexodus tufi Gabris, Trnka, Wahab & Kundrata, 2017, sp. n." docType="treatment" docUuid="85AFEBB7-502B-4303-8C86-B9DD45AD8F1F" docUuidSource="ZooBank" docVersion="5" lastPageNumber="33" masterDocId="BD7AE331C048FFACFF95FFF3FFF6FFFF" masterDocTitle="Taxonomic revision of the endemic Bornean genera Anexodus Pascoe and Pantilema Aurivillius (Coleoptera, Cerambycidae, Lamiinae)" masterLastPageNumber="51" masterPageNumber="29" pageNumber="31" updateTime="1668164414139" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the endemic Bornean genera Anexodus Pascoe and Pantilema Aurivillius (Coleoptera, Cerambycidae, Lamiinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gabris, Radim</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Trnka, Filip</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wahab, Rodzay Abdul</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kundrata, Robin</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>669</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>29</mods:start>
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<mods:end>51</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.669.12608</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.669.12608</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-669-29</mods:identifier>
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<mods:identifier type="ZBK">7BC01A4ED22C44B38D2DA0DF2CD53AA4</mods:identifier>
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<mods:identifier type="ZooBank">7BC01A4ED22C44B38D2DA0DF2CD53AA4</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="128687983" LSID="urn:lsid:zoobank.org:act:85AFEBB7-502B-4303-8C86-B9DD45AD8F1F" httpUri="http://treatment.plazi.org/id/CD57FE8BC8FCC74CE0F84A5AEAD21984" lastPageId="4" lastPageNumber="33" pageId="2" pageNumber="31">
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<subSubSection pageId="2" pageNumber="31" type="nomenclature">
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<paragraph pageId="2" pageNumber="31">
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<taxonomicName LSID="http://zoobank.org/85AFEBB7-502B-4303-8C86-B9DD45AD8F1F" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus tufi" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="tufi">Anexodus tufi</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="31">sp. n.</taxonomicNameLabel>
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Figs 7-8, 15-16, 20, 26-27, 31, 33, 40
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="32" type="type material">
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<paragraph pageId="3" pageNumber="32">
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<pageBreakToken pageId="3" pageNumber="32" start="start">Type</pageBreakToken>
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material.
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</paragraph>
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<paragraph pageId="3" pageNumber="32">
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Holotype, male, "BRUNEI, Ulu Temburong NP / Kuala Belalong FSC /
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.5465555">4°32'47.6"N</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="115.1575">115°09'27"E</geoCoordinate>
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/ I. H. Tuf leg. II.2013" (UBDC); paratype, female, same data as holotype (UPOL); 2 paratypes, females, "BRUNEI, Ulu Temburong NP / Kuala Belalong FSC /
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.5465555">4°32'47.6"N</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="115.1575">115°09'27"E</geoCoordinate>
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/ Z.
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<normalizedToken originalValue="Mačát">Macat</normalizedToken>
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leg. I.2014" (BMNH); paratype, female, "BRUNEI, Ulu Temburong NP / Kuala Belalong FSC /
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.5465555">4°32'47.6"N</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="115.1575">115°09'27"E</geoCoordinate>
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/ O.
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<normalizedToken originalValue="Machač">Machac</normalizedToken>
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leg. II.2015" (UPOL).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="32" type="differential dagnosis">
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<paragraph pageId="3" pageNumber="32">Differential dagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="32">
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This species is similar to
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<taxonomicName lsidName="A. aquilus" pageId="3" pageNumber="32" rank="species" species="aquilus">A. aquilus</taxonomicName>
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in having yellowish stripes on the dorsal body surface and labrum with the whole surface punctured (Fig. 20), but it differs in shape of the scape (apex simple in
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<taxonomicName lsidName="A. tufi" pageId="3" pageNumber="32" rank="species" species="tufi">A. tufi</taxonomicName>
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sp. n., apex with distinct lateral hook-shaped projection in
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<taxonomicName lsidName="A. aquilus" pageId="3" pageNumber="32" rank="species" species="aquilus">A. aquilus</taxonomicName>
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; Figs 21-22, 26-27), length of the parameres (relatively longer in
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<taxonomicName lsidName="A. aquilus" pageId="3" pageNumber="32" rank="species" species="aquilus">A. aquilus</taxonomicName>
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; Figs 28, 31), and shape of the spermatheca (simply elongated in
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<taxonomicName lsidName="A. tufi" pageId="3" pageNumber="32" rank="species" species="tufi">A. tufi</taxonomicName>
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sp. n., widened at second half in
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<taxonomicName lsidName="A. aquilus" pageId="3" pageNumber="32" rank="species" species="aquilus">A. aquilus</taxonomicName>
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; Figs 32-33).
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</paragraph>
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<caption pageId="3" pageNumber="32">
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<paragraph pageId="3" pageNumber="32">
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Figures 17-27.
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="genus">Anexodus</taxonomicName>
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spp.: 17-20 Head, frontal view: 17
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus aquilus" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="aquilus">Anexodus aquilus</taxonomicName>
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Pascoe, holotype male 18
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus sarawakensis" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="sarawakensis">Anexodus sarawakensis</taxonomicName>
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Sudre, holotype male 19
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus syptakovae" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="syptakovae">Anexodus syptakovae</taxonomicName>
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sp. n., holotype male 20
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus tufi" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="tufi">Anexodus tufi</taxonomicName>
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sp. n., holotype male 21-27 Antenna: 21
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus aquilus" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="aquilus">Anexodus aquilus</taxonomicName>
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Pascoe, male 22
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus aquilus" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="aquilus">Anexodus aquilus</taxonomicName>
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Pascoe, female 23
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus sarawakensis" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="sarawakensis">Anexodus sarawakensis</taxonomicName>
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Sudre, male 24
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus sarawakensis" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="sarawakensis">Anexodus sarawakensis</taxonomicName>
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Sudre, female 25
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus syptakovae" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="syptakovae">Anexodus syptakovae</taxonomicName>
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sp. n., male 26
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus tufi" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="tufi">Anexodus tufi</taxonomicName>
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sp. n., male 27
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus tufi" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="tufi">Anexodus tufi</taxonomicName>
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sp. n., female. Not to scale.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="33" pageId="3" pageNumber="32" type="description">
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<paragraph pageId="3" pageNumber="32">Description.</paragraph>
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<paragraph pageId="3" pageNumber="32">Holotype (male). BL 7.4 mm, BW 2.3 mm. Body brown with yellowish stripes ranging from vertex through sides of pronotum to basal part of elytra, densely clothed with very short brown pubescence. Head about as wide as anterior margin of pronotum. Labrum transverse, its surface with moderately sparse punctures (Fig. 20). Eyes moderately emarginate at antennal articulations (Figs 15-16). Antennae 0.9 times as long as body; scape gradually slightly widened towards apex, thickest at apical part, apex simple (Figs 26-27); the relative ratio of antennomere lengths 3.2: 1.5: 1.0: 1.2: 1.0: 0.9: 0.5: 0.5: 0.5: 0.5: 1.0.</paragraph>
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<paragraph pageId="3" pageNumber="32">Prothorax as long as wide, laterally with one distinct tubercle; pronotal disc with a pair of moderately distinct tubercles near middle and one median at second half and one indistinct median at anterior half; pronotal tubercles punctate. Prosternum in front of coxae 0.9 times shorter than diameter of coxal cavity. Scutellum transverse, two times as wide as long. Elytra elongate, 1.8 times as long as wide at widest part, 1.9 times as long as pronotum, widest near middle; each elytron with three rows of distinct, longitudinally elongate tubercles (Figs 7-8, 15-16), inner row forming a distinct ridge basally; sparsely covered with large deep punctures arranged in slightly irregular rows. Legs long, slender; relative lengths of metatarsomeres 1.0: 0.6: 1.0: 2.1.</paragraph>
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<paragraph lastPageId="4" lastPageNumber="33" pageId="3" pageNumber="32">
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Male genitalia with tegmen elongate, widest before middle, basally with short strut; parameres elongate, 3.3 times longer than wide, apically with long setae (Fig. 31). Pe
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<pageBreakToken pageId="4" pageNumber="33" start="start">nis</pageBreakToken>
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subparallel-sided, apically truncate; dorsal struts diverged from about 1/3 of penis length. Internal sac long, with paired small medial sclerites and distinct flagellar sclerites.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="33" type="intraspecific variability">
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<paragraph pageId="4" pageNumber="33">Intraspecific variability.</paragraph>
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<paragraph pageId="4" pageNumber="33">The male paratype is larger (body length 9.4 mm, body width 2.8 mm).</paragraph>
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<paragraph pageId="4" pageNumber="33">
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Female. Most characters same as for males. BL 8.5-10.2 mm, BW 2.7-3.4. Antennae shorter, 0.7 times as long as body, with relatively longer scape and pedicel (length ratio of antennomeres
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<normalizedToken originalValue="I–III">I-III</normalizedToken>
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: 3.4-3.6: 1.7-1.8: 1.0). Elytra 2.2-2.3 times as long pronotum. Fifth ventrite with apex truncate. Spermatheca sclerotized, slender, elongate, curved, gradually tapered toward apex; sclerotized part of spermathecal duct simple (Fig. 33).
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</paragraph>
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<caption pageId="4" pageNumber="33">
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<paragraph pageId="4" pageNumber="33">
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Figures 28-34.
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">Anexodus</taxonomicName>
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species: 28-31 Aedeagus, ventral view: 28
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus aquilus" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="aquilus">Anexodus aquilus</taxonomicName>
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Pascoe, holotype 29
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus sarawakensis" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="sarawakensis">Anexodus sarawakensis</taxonomicName>
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Sudre, holotype 30
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus syptakovae" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="syptakovae">Anexodus syptakovae</taxonomicName>
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sp. n., holotype 31
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus tufi" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="tufi">Anexodus tufi</taxonomicName>
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sp. n., holotype 32-34 Spermatheca: 32
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus aquilus" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="aquilus">Anexodus aquilus</taxonomicName>
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Pascoe 33
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus tufi" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="tufi">Anexodus tufi</taxonomicName>
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sp. n. 34
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<taxonomicName class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus sarawakensis" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="sarawakensis">Anexodus sarawakensis</taxonomicName>
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Sudre. Not to scale.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="33" type="distribution">
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<paragraph pageId="4" pageNumber="33">Distribution.</paragraph>
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<paragraph pageId="4" pageNumber="33">
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Brunei (Fig. 43). All the specimens in the type series are from the vicinity of the Kuala Belalong Field Studies Centre (KBFSC) in the Ulu Temburong National Park (Fig. 41) which has been described in detail by
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<bibRefCitation author="Sevcik, J" journalOrPublisher="ZooKeys" pageId="5" pageNumber="34" pagination="57 - 77" title="Diversity of Manota Williston (Diptera, Mycetophilidae) in Ulu Temburong National Park, Brunei." url="https://doi.org/10.3897/zookeys.428.7912" volume="428" year="2014">
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<normalizedToken originalValue="Ševčík">Sevcik</normalizedToken>
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et al. (2014)
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</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="33" type="etymology">
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<paragraph pageId="4" pageNumber="33">Etymology.</paragraph>
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<paragraph pageId="4" pageNumber="33">This species is named after Mr. I. H. Tuf (UPOL, Czech Republic; Fig. 42), who collected a part of the type series.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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