306 lines
35 KiB
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306 lines
35 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/jhr.96.101873" ID-Pensoft-Pub="1314-2607-96-241" ID-Pensoft-UUID="A5722C06212C5BA7A2099B77C6A8DF54" ID-ZooBank="15A2B06B92F34E70AC8F6FEABF365E71" ModsDocID="1314-2607-96-241" checkinTime="1684817117864" checkinUser="pensoft" docAuthor="Wood, Thomas J." docDate="2023" docId="B5F8A1DAD91CD7A3C7E6C949E49EC900" docLanguage="en" docName="JourHymenoptRes 96: 241-484" docOrigin="Journal of Hymenoptera Research 96" docPubDate="2023-05-22" docSource="http://dx.doi.org/10.3897/jhr.96.101873" docTitle="Ovandrena limbata subsp. subgen. nov." docType="treatment" docVersion="1" id="A5722C06212C5BA7A2099B77C6A8DF54" lastPageNumber="241" masterDocId="A5722C06212C5BA7A2099B77C6A8DF54" masterDocTitle="The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera, Andrenidae)" masterLastPageNumber="484" masterPageNumber="241" pageNumber="241" updateTime="1684817117864" updateUser="pensoft">
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<mods:titleInfo id="4587FA101CC1128F0911D774F4078004">
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<mods:title id="0B449A82A9573A858DDBD55DD4146A0A">The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera, Andrenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F68F9A74B2A844DD9BE9C74AA7363E2E" type="personal">
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<mods:roleTerm id="678851BA1A97ADA8BB3EDB030ABD3381">Author</mods:roleTerm>
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<mods:namePart id="7BBCD4EBCBB72F7EB481A435BED79F0D">Wood, Thomas J.</mods:namePart>
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<mods:nameIdentifier id="94AF331DEE5ECF5EAF05F5F50BEA945C" type="ORCID">https://orcid.org/0000-0001-5653-224X</mods:nameIdentifier>
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<mods:affiliation id="47BF18E60667A13E802B98C663134BC9">University of Mons, Research Institute for Biosciences, Laboratory of Zoology, Place du Parc 20, 7000, Mons, Belgium</mods:affiliation>
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<mods:nameIdentifier id="BE10EE8929EA16E34E0954929AFB9C03" type="email">thomasjames.wood@umons.ac.be</mods:nameIdentifier>
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<mods:typeOfResource id="17A97681657C6E7170E131F6BD8DD494">text</mods:typeOfResource>
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<mods:title id="9797855C1AE38047700EF4B5E0DA3744">Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part id="7D22B1B071870AC3B644196865AB5F08">
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<mods:date id="D71078DF54B84C6CB2DF638DABB87EDA">2023</mods:date>
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<mods:detail id="CCC06D88BD7B9BCE5427B908B2CE0BF1" type="pubDate">
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<mods:number id="1B7135D49698E420C86FAE4FD33023E1">2023-05-22</mods:number>
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</mods:detail>
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<mods:detail id="C539F830B1A775EAC068BA500C63C7E8" type="volume">
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<mods:number id="842F0BFC915E3257C368C8A8F0D5E512">96</mods:number>
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</mods:detail>
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<mods:start id="7AB7CD5BA659E81604E9E0379CF0DC11">241</mods:start>
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<mods:end id="212FD54D1C16EBC711D12E5B4A4765C4">484</mods:end>
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<mods:url id="BC57733C87E8E35A18DEBC01A9259A88">http://dx.doi.org/10.3897/jhr.96.101873</mods:url>
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</mods:location>
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<mods:classification id="8240573003EA554958E59BE26F92D560">journal article</mods:classification>
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<mods:identifier id="F8A5B53332011404031AFB06F7EB7938" type="DOI">http://dx.doi.org/10.3897/jhr.96.101873</mods:identifier>
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<mods:identifier id="391137C4301675E11864FB2D231DECAE" type="Pensoft-Pub">1314-2607-96-241</mods:identifier>
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</mods:mods>
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<treatment id="B5F8A1DAD91CD7A3C7E6C949E49EC900" LSID="urn:lsid:zoobank.org:act:A122C278-C33D-4790-A34A-FC270AE6A962" httpUri="http://treatment.plazi.org/id/B5F8A1DAD91CD7A3C7E6C949E49EC900" lastPageNumber="241" pageId="0" pageNumber="241">
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<subSubSection id="ABCE327FE036F69A192CD2C99AD6DBE3" pageId="0" pageNumber="241" type="nomenclature">
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<paragraph id="314A5B52896E37578F3F374C28279CAF" pageId="0" pageNumber="241">
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Subgenus
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<taxonomicName id="A269CFEE9C4FA243449FEF13AC34662E" LSID="https://zoobank.org/A122C278-C33D-4790-A34A-FC270AE6A962" family="Andrenidae" genus="Ovandrena" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ovandrena subsp. subgen. nov." order="Hymenoptera" pageId="0" pageNumber="241" rank="subSpecies" species="limbata" subSpecies="subgen. nov.">Ovandrena subgen. nov.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="A1FDA648F10285AB1D3640254E1BD8BB" pageId="0" pageNumber="241" type="type species">
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<paragraph id="4EBBED08D250E6189B8B1EB11044DD57" pageId="0" pageNumber="241">Type species.</paragraph>
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<paragraph id="AF4131E1D476138CB86975692919D0A0" pageId="0" pageNumber="241">
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<taxonomicName id="1EDF539950D6F8B7E02FBBB92176BFA3" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena oviventris" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="oviventris">
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<emphasis id="B39AB1AFE8E54146D55587F1855E7BA6" italics="true" pageId="0" pageNumber="241">Andrena oviventris</emphasis>
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</taxonomicName>
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<normalizedToken id="103F2826F4D4E4225CA8B535A9B5D935" originalValue="Pérez">Perez</normalizedToken>
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, 1895.
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</paragraph>
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</subSubSection>
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<subSubSection id="BD59E9E5A858D4EA461A0DE52D13BCE4" pageId="0" pageNumber="241" type="diagnosis">
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<paragraph id="5D95615A7DED1CD78E9058BF013886F7" pageId="0" pageNumber="241">Diagnosis.</paragraph>
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<paragraph id="38A364132778F425FDC18DC574464D14" pageId="0" pageNumber="241">
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Through the combination of slightly upturned fore margin of the clypeus, broad fovea occupying at least
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<normalizedToken id="0D064D3E9D4F67FA2A5973805341CD66" originalValue="½">1/2</normalizedToken>
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the space between the lateral ocellus and the inner margin of the compound eye, weak but distinct humeral angle, unmodified posterior face of the hind femur (without teeth, carinae, or spines), simple hind tibial spur (not broadened basally or medially), dark integument, black male clypeus, and essential absence of defining features it falls very close to members of the
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<taxonomicName id="F008B519E3B3AF1AB98340C504F8F603" family="Andrenidae" genus="Blandandrena" kingdom="Animalia" lsidName="Blandandrena relata" order="Hymenoptera" pageId="0" pageNumber="241" rank="species" species="relata">
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<emphasis id="7BB006BEFEDC9DBE12EBB030309AB32A" italics="true" pageId="0" pageNumber="241">Blandandrena relata</emphasis>
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</taxonomicName>
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-group and to
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<taxonomicName id="6DB68F2A98DCF1FC394D4C60884C9D4C" family="Andrenidae" genus="Blandandrena" kingdom="Animalia" lsidName="Blandandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="3C145572055085763CEEB586AAE36B4B" italics="true" pageId="0" pageNumber="241">Blandandrena</emphasis>
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</taxonomicName>
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that were formerly lumped together under the subgenus
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<taxonomicName id="09CF016CD558FA4517E8B69A9B1753D1" family="Andrenidae" genus="Blandandrena" kingdom="Animalia" lsidName="Blandandrena (Poliandrena)" order="Hymenoptera" pageId="0" pageNumber="241" rank="subGenus" subGenus="Poliandrena">Blandandrena Poliandrena</taxonomicName>
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(see above).
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</paragraph>
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<paragraph id="E6B711E96B210A467C1AD6ADFDFA4CE2" pageId="0" pageNumber="241">
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In the female sex,
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<taxonomicName id="60201B87D1E649BAEC15C2C807CEB3DF" family="Andrenidae" genus="Ovandrena" kingdom="Animalia" lsidName="Ovandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="30AB518BE5A5FCF9916751BB5749FBBE" italics="true" pageId="0" pageNumber="241">Ovandrena</emphasis>
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</taxonomicName>
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species differ from these groups by only a single major character: the structure of the propodeal triangle (Fig.
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<figureCitation id="94EA3D780B3B97E47F8AD1DC9C7CCE28" captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Andrena (Ovandrena) subgen. nov. characters. Andrena (Ovandrena) marsae Schmiedeknecht, 1900, female A propodeal triangle; Andrena (Ovandrena) oviventris Perez, 1895 B female propodeal triangle C female scutal hairs, profile view F male genital capsule; Andrena (Ovandrena) farinosa Perez, 1895 D female scutal hairs, profile view E male genital capsule." figureDoi="10.3897/jhr.96.101873.figure45" httpUri="https://binary.pensoft.net/fig/853437" pageId="0" pageNumber="241">45A, B</figureCitation>
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).
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<taxonomicName id="CE66542AC05DDD7FDDC63659D63DA0E9" family="Andrenidae" genus="Ovandrena" kingdom="Animalia" lsidName="Ovandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="12C0E7714AE73E833E32F10DC3858D4A" italics="true" pageId="0" pageNumber="241">Ovandrena</emphasis>
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</taxonomicName>
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species have scutal hairs that are not as short and clearly squamous as in members of the
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<taxonomicName id="AD9AA2A333A86EB5D2D3C5D5A7C8C2AA" authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Aenandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aenandrena" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="genus">
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<emphasis id="7AAB3DF37C45208F800222219F902551" italics="true" pageId="0" pageNumber="241">Aenandrena</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="AECF153631B05BE583D1C7F32DBA8259" family="Andrenidae" genus="Limbandrena" kingdom="Animalia" lsidName="Limbandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="D8A0E0A73040E41167F69C3125BC0EE0" italics="true" pageId="0" pageNumber="241">Limbandrena</emphasis>
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</taxonomicName>
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, or
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<taxonomicName id="B1B0F304A47F46F13CEDB81C9C66D2D6" authorityName="Hedicke" authorityYear="1933" class="Insecta" family="Andrenidae" genus="Lepidandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lepidandrena" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="genus">
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<emphasis id="387FB8C1A0C1AB8B56D0E85FAADC2304" italics="true" pageId="0" pageNumber="241">Lepidandrena</emphasis>
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</taxonomicName>
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Hedicke, 1933, but they are partially squamous in some species (Fig.
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<figureCitation id="8BAB898DE7DBD8B3CB73F6251A9FA294" captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Andrena (Ovandrena) subgen. nov. characters. Andrena (Ovandrena) marsae Schmiedeknecht, 1900, female A propodeal triangle; Andrena (Ovandrena) oviventris Perez, 1895 B female propodeal triangle C female scutal hairs, profile view F male genital capsule; Andrena (Ovandrena) farinosa Perez, 1895 D female scutal hairs, profile view E male genital capsule." figureDoi="10.3897/jhr.96.101873.figure45" httpUri="https://binary.pensoft.net/fig/853437" pageId="0" pageNumber="241">45C, D</figureCitation>
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), whereas the scutal hairs in the
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<taxonomicName id="297C9AFD4FBF9ED78544E88119778ADE" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena relata" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="relata">
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<emphasis id="B46C62F1B970DB74E5C93ACF063B6E9A" italics="true" pageId="0" pageNumber="241">Andrena relata</emphasis>
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</taxonomicName>
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-group and
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<taxonomicName id="82DDE2CEF31288D58A6579DD8115679E" family="Andrenidae" genus="Blandandrena" kingdom="Animalia" lsidName="Blandandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="E1E044C0BB3F475DB7BE28D62FFD2454" italics="true" pageId="0" pageNumber="241">Blandandrena</emphasis>
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</taxonomicName>
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are typically simple, though in some species such as
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<taxonomicName id="5855CC199512E8A835ADA2DEED4E9B22" class="Insecta" family="Plantaginaceae" kingdom="Animalia" lsidName="A. corax" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="corax">
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<emphasis id="0922F3697B7954B194A2F96E775680D9" italics="true" pageId="0" pageNumber="241">A. corax</emphasis>
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</taxonomicName>
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the hairs can be semi-squamous. The structure of the propodeal triangle is therefore the only character that allows consistent separation. In
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<taxonomicName id="A22CB73A4D195610C5CF0101A1B715DC" family="Andrenidae" genus="Ovandrena" kingdom="Animalia" lsidName="Ovandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="6DF84C4DAAA5166B8FEACF86D7966C17" italics="true" pageId="0" pageNumber="241">Ovandrena</emphasis>
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</taxonomicName>
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, the propodeal triangle is a broad and well-defined triangle (lateral margins extending almost to the lateral edges of the metanotum) that is defined by raised carinae; the internal surface is covered in raised irregular carinae of a similar width, therefore appearing homogenous and consistent (Fig.
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<figureCitation id="8213BEFC725ABEACA9E7296060C7CEF7" captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Andrena (Ovandrena) subgen. nov. characters. Andrena (Ovandrena) marsae Schmiedeknecht, 1900, female A propodeal triangle; Andrena (Ovandrena) oviventris Perez, 1895 B female propodeal triangle C female scutal hairs, profile view F male genital capsule; Andrena (Ovandrena) farinosa Perez, 1895 D female scutal hairs, profile view E male genital capsule." figureDoi="10.3897/jhr.96.101873.figure45" httpUri="https://binary.pensoft.net/fig/853437" pageId="0" pageNumber="241">45A, B</figureCitation>
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). In the
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<taxonomicName id="D1BBB95BD930F15D5FCA08530B9AD29C" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena relata" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="relata">
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<emphasis id="F3B0B9A79F47D7C675EE843283656EC6" italics="true" pageId="0" pageNumber="241">Andrena relata</emphasis>
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</taxonomicName>
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-group and
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<taxonomicName id="0763BC58D0427A2521E05D2D69B7CF17" family="Andrenidae" genus="Blandandrena" kingdom="Animalia" lsidName="Blandandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="C91662DDC6B80D0D39F0163BA05A32D5" italics="true" pageId="0" pageNumber="241">Blandandrena</emphasis>
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</taxonomicName>
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, the propodeal triangle varies from smooth and undefined to weakly defined with at most irregular rugosity on its internal surface, but never approaches the condition in
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<taxonomicName id="32B8FF416FD2F1A8D35AD9F03A721CDA" family="Andrenidae" genus="Ovandrena" kingdom="Animalia" lsidName="Ovandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="75602E141DC8B54E52BA79DC22CBAE1E" italics="true" pageId="0" pageNumber="241">Ovandrena</emphasis>
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</taxonomicName>
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. The combination of these two characters therefore always allows recognition. They can also be partially recognised by the extremely dense and fine punctation of the terga, punctures separated by <0.5 puncture diameters; in comparison groups, the punctures are never this fine or this dense.
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</paragraph>
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<caption id="734C06DEDAF4FC0CE46DF6C0299FBC19" doi="10.3897/jhr.96.101873.figure45" httpUri="https://binary.pensoft.net/fig/853437" pageId="0" pageNumber="241" start="Figure 45" startId="F45">
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<paragraph id="A45AF48B8A97ED1553F163F0525A6A36" pageId="0" pageNumber="241">
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<emphasis id="0AB5EE7D6C11AF97A8B76E2F9554312F" bold="true" pageId="0" pageNumber="241">Figure 45.</emphasis>
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<taxonomicName id="0818529597A15AC3DEA985EB3F7FC939" authority="" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Ovandrena)" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="subGenus" subGenus="Ovandrena">Andrena (Ovandrena)</taxonomicName>
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subgen. nov. characters.
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<taxonomicName id="918ECDFAE9B5CBEB5B9EFE6B46918E07" authorityName="Schmiedeknecht" authorityYear="1900" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Ovandrena) marsae" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="marsae" subGenus="Ovandrena">Andrena (Ovandrena) marsae</taxonomicName>
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Schmiedeknecht, 1900, female
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<emphasis id="0FFA0B4F06E6584175A951BA98E7FA89" bold="true" pageId="0" pageNumber="241">A</emphasis>
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propodeal triangle;
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<taxonomicName id="F5BB6FFEAB4DEAF556FDD95888AA9A70" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Ovandrena) oviventris" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="oviventris" subGenus="Ovandrena">Andrena (Ovandrena) oviventris</taxonomicName>
|
||
|
|
<normalizedToken id="01D22DD3D727C78FD1316F1C9F44D554" originalValue="Pérez">Perez</normalizedToken>
|
||
|
|
, 1895
|
||
|
|
<emphasis id="61BCDE55E4F6F85338C0BFA1D57EAC13" bold="true" pageId="0" pageNumber="241">B</emphasis>
|
||
|
|
female propodeal triangle
|
||
|
|
<emphasis id="3CB795FC1C71738A69C2678EAA4B07C7" bold="true" pageId="0" pageNumber="241">C</emphasis>
|
||
|
|
female scutal hairs, profile view
|
||
|
|
<emphasis id="03024C5BD62237215AA553F22C067EF0" bold="true" pageId="0" pageNumber="241">F</emphasis>
|
||
|
|
male genital capsule;
|
||
|
|
<taxonomicName id="C157C022C9D75204C2445D5428445994" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Ovandrena) farinosa" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="farinosa" subGenus="Ovandrena">Andrena (Ovandrena) farinosa</taxonomicName>
|
||
|
|
<normalizedToken id="E4DE873C12741E859BF82A651A8CDDB3" originalValue="Pérez">Perez</normalizedToken>
|
||
|
|
, 1895
|
||
|
|
<emphasis id="CD211F59A59924AFF28F6B4FFFFB5521" bold="true" pageId="0" pageNumber="241">D</emphasis>
|
||
|
|
female scutal hairs, profile view
|
||
|
|
<emphasis id="1D93A65B43E755384AF4EABD432AC270" bold="true" pageId="0" pageNumber="241">E</emphasis>
|
||
|
|
male genital capsule.
|
||
|
|
</paragraph>
|
||
|
|
</caption>
|
||
|
|
<paragraph id="F99FB436F9BB679BE215AE846DB835C0" pageId="0" pageNumber="241">
|
||
|
|
Males can be recognised through their combination of dark clypeus with upturned fore margin, distinctive propodeal triangle as in the female sex, pronotum with weak or strong humeral angle, A3 exceeding A4 but shorter than A4+5, and genital capsule which is compact with pronounced and rounded gonocoxal teeth (Fig.
|
||
|
|
<figureCitation id="CD987F051B91C67BB3C6E9230AEFAE13" captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Andrena (Ovandrena) subgen. nov. characters. Andrena (Ovandrena) marsae Schmiedeknecht, 1900, female A propodeal triangle; Andrena (Ovandrena) oviventris Perez, 1895 B female propodeal triangle C female scutal hairs, profile view F male genital capsule; Andrena (Ovandrena) farinosa Perez, 1895 D female scutal hairs, profile view E male genital capsule." figureDoi="10.3897/jhr.96.101873.figure45" httpUri="https://binary.pensoft.net/fig/853437" pageId="0" pageNumber="241">45E, F</figureCitation>
|
||
|
|
). Members of the
|
||
|
|
<taxonomicName id="11388DF24B68CD4213AAFBC5737B4F34" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena relata" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="relata">
|
||
|
|
<emphasis id="7CD1BBAEC22FF7D20659A0D273AC2394" italics="true" pageId="0" pageNumber="241">Andrena relata</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
-group often have a yellow-marked clypeus, but for species showing a dark clypeus (e.g.
|
||
|
|
<taxonomicName id="8E9FAFC9C701C86E76C74D2C10FDBE6C" class="Insecta" family="Plantaginaceae" kingdom="Animalia" lsidName="A. corax" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="corax">
|
||
|
|
<emphasis id="50753EC0D74070E9F470B7B523D51A66" italics="true" pageId="0" pageNumber="241">A. corax</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
), the clearly defined triangular propodeal triangle allows separation (smooth to at most weakly defined in the
|
||
|
|
<taxonomicName id="AC679FB8B043822E7E78F5DE2903C539" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena relata" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="relata">
|
||
|
|
<emphasis id="C03C51D72C2413DD1E4E30ABB6AC4B9E" italics="true" pageId="0" pageNumber="241">Andrena relata</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
-group). Separation can be made from
|
||
|
|
<taxonomicName id="73FC748A3304EBBC06A9EF39F7D6F234" family="Andrenidae" genus="Blandandrena" kingdom="Animalia" lsidName="Blandandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
|
||
|
|
<emphasis id="95461B4B3A0AD775240FFD4F3A60A7F1" italics="true" pageId="0" pageNumber="241">Blandandrena</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
by the genital capsule and the propodeal triangle, as detailed in the diagnosis section for that subgenus.
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="F4BB4ABABD1E76445E1DB2F65C104059" pageId="0" pageNumber="241" type="description">
|
||
|
|
<paragraph id="137BB8ED07157ED4995FF732D05F35D8" pageId="0" pageNumber="241">Description.</paragraph>
|
||
|
|
<paragraph id="B3AD65A68AC2FAD35E3EA357FA31EDC8" pageId="0" pageNumber="241">
|
||
|
|
Small to moderately sized bees (7-11 mm) typically with dark integument, one species with red tergal markings; male clypeus always dark. Head broad, 1.3-1.4 times broader than long, compound eyes with inner margins weakly converging apically. Gena slightly exceeding width of compound eye; ocelloccipital distance short, slightly less than to slightly more than diameter of lateral ocellus. Facial fovea moderately broad, occupying
|
||
|
|
<normalizedToken id="E135F2C4FAD2E582BE8510ED90E607EE" originalValue="½">1/2</normalizedToken>
|
||
|
|
distance between lateral ocellus and compound eye. Female scutum, scutellum, and metanotum covered with shortish light brown to whitish semi-squamous hairs (Fig.
|
||
|
|
<figureCitation id="70901C1059F84A115D3477DC41DD1578" captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Andrena (Ovandrena) subgen. nov. characters. Andrena (Ovandrena) marsae Schmiedeknecht, 1900, female A propodeal triangle; Andrena (Ovandrena) oviventris Perez, 1895 B female propodeal triangle C female scutal hairs, profile view F male genital capsule; Andrena (Ovandrena) farinosa Perez, 1895 D female scutal hairs, profile view E male genital capsule." figureDoi="10.3897/jhr.96.101873.figure45" httpUri="https://binary.pensoft.net/fig/853437" pageId="0" pageNumber="241">45C, D</figureCitation>
|
||
|
|
). Pronotum laterally with weak to strong humeral angle. Dorsolateral surface of propodeum with regular and dense network of raised reticulation; propodeal triangle wide, lateral margins extending almost to lateral edges of metanotum, laterally defined by straight raised carinae, thus strongly triangular (Fig.
|
||
|
|
<figureCitation id="6EFE0212B65D4E71A51AA995009C5FF8" captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Andrena (Ovandrena) subgen. nov. characters. Andrena (Ovandrena) marsae Schmiedeknecht, 1900, female A propodeal triangle; Andrena (Ovandrena) oviventris Perez, 1895 B female propodeal triangle C female scutal hairs, profile view F male genital capsule; Andrena (Ovandrena) farinosa Perez, 1895 D female scutal hairs, profile view E male genital capsule." figureDoi="10.3897/jhr.96.101873.figure45" httpUri="https://binary.pensoft.net/fig/853437" pageId="0" pageNumber="241">45A, B</figureCitation>
|
||
|
|
). Internal surface of propodeum covered by raised irregular carinae of a similar width, therefore homogenous and consistent. Forewing with nervulus interstitial to weakly antefurcal. Hind tibial spurs simple, not broadened basally or medially. Terga densely punctate, in females punctures extremely dense and fine, separated by <0.5 puncture diameters, in males punctures slightly sparser and coarser, separated by 0.5-1 puncture diameters. Male genital capsule compact, consistent across species, with gonocoxae strongly produced into apical rounded teeth (Fig.
|
||
|
|
<figureCitation id="74593776BA8237F61B6BA3DB2196DDAA" captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Andrena (Ovandrena) subgen. nov. characters. Andrena (Ovandrena) marsae Schmiedeknecht, 1900, female A propodeal triangle; Andrena (Ovandrena) oviventris Perez, 1895 B female propodeal triangle C female scutal hairs, profile view F male genital capsule; Andrena (Ovandrena) farinosa Perez, 1895 D female scutal hairs, profile view E male genital capsule." figureDoi="10.3897/jhr.96.101873.figure45" httpUri="https://binary.pensoft.net/fig/853437" pageId="0" pageNumber="241">45E, F</figureCitation>
|
||
|
|
). Gonostyli apically spatulate, with a more or less strongly raised inner margin. Penis valves basally narrow or moderately broad, in both cases becoming narrow medially.
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="41BC85BB3903DD6C69446857E7CAEF90" pageId="0" pageNumber="241" type="etymology">
|
||
|
|
<paragraph id="7B74FC42150E5E65D44938A9B08064D1" pageId="0" pageNumber="241">Etymology.</paragraph>
|
||
|
|
<paragraph id="95175FB9A1FBE87B6EF1B57E37DC01F3" pageId="0" pageNumber="241">
|
||
|
|
The name is taken from the type species for the genus,
|
||
|
|
<taxonomicName id="30701A6914B99276C41F68AC16CEE8AC" class="Insecta" family="Plantaginaceae" kingdom="Animalia" lsidName="A. oviventris" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="oviventris">
|
||
|
|
<emphasis id="32C586EBE9CBFF2347D9C5DB1BB14AB8" italics="true" pageId="0" pageNumber="241">A. oviventris</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
. It derives from the Latin
|
||
|
|
<emphasis id="A7E1EC0A1EC1477014B33AD377606168" italics="true" pageId="0" pageNumber="241">ovum</emphasis>
|
||
|
|
meaning egg, in reference to the egg-shaped metasoma. The gender is feminine.
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="732A10EBB2A2679067C07860C866E31F" pageId="0" pageNumber="241" type="included species">
|
||
|
|
<paragraph id="1E3DA60CE0B01D4A73AFB0E6A27C8F13" pageId="0" pageNumber="241">Included species.</paragraph>
|
||
|
|
<paragraph id="F35062E324A0EA933687D565893BE6C5" pageId="0" pageNumber="241">
|
||
|
|
<taxonomicName id="74710FC2A6ADDBC44ED75763DE391629" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena farinosa" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="farinosa">
|
||
|
|
<emphasis id="78DC6EB616B96834095FA67B8799CA49" italics="true" pageId="0" pageNumber="241">Andrena farinosa</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
<normalizedToken id="C50DCD8CCEE9EC02D73F98199EF6FF06" originalValue="Pérez">Perez</normalizedToken>
|
||
|
|
, 1895 (Spain and France),
|
||
|
|
<taxonomicName id="1D0AA55F1E58673E9C02E0D478EC2F9D" authorityName="Wood" authorityYear="2020" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena farinosoides" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="farinosoides">
|
||
|
|
<emphasis id="55C3C55B3AA74F83D88D6AB2D1D81234" italics="true" pageId="0" pageNumber="241">Andrena farinosoides</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
Wood, 2020 (Morocco),
|
||
|
|
<taxonomicName id="CC0713C85416A80BC102A74AFFA65240" authorityName="Schmiedeknecht" authorityYear="1900" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena marsae" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="marsae">
|
||
|
|
<emphasis id="89563D5FD03FF9ABDB2A6C83EF60DF63" italics="true" pageId="0" pageNumber="241">Andrena marsae</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
Schmiedeknecht, 1900 (Morocco, Algeria, Tunisia), and
|
||
|
|
<taxonomicName id="BFD8394C5BAD73D026EA7E1D1093491D" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena oviventris" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="oviventris">
|
||
|
|
<emphasis id="D9D62087641E2DABEC5226323781BB8A" italics="true" pageId="0" pageNumber="241">Andrena oviventris</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
(Morocco, Algeria, Portugal, Spain, France). The subgenus is therefore currently restricted to the Western Mediterranean, and the centre of diversity is Morocco. The status of
|
||
|
|
<taxonomicName id="70611623C79E4E83D7556FB4DA626FE6" class="Insecta" family="Plantaginaceae" genus="A." kingdom="Animalia" lsidName="A." order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis id="3AD91764ECE4FEAE26438645B2D9F2F2" italics="true" pageId="0" pageNumber="241">A.</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
(incertae sedis)
|
||
|
|
<emphasis id="122ED9C6A2DB4A2F148D8A9939C0D40A" italics="true" pageId="0" pageNumber="241">inusitata</emphasis>
|
||
|
|
Pisanty, 2022 must be resolved through genetic analysis (see
|
||
|
|
<bibRefCitation id="D01E08B9275D9284004F1B8DD7910895" DOI="https://doi.org/10.11646/zootaxa.5185.1.1" author="Pisanty, G" journalOrPublisher="Zootaxa" pageId="0" pageNumber="241" pagination="1 - 109" refId="B68" refString="Pisanty, G, Scheuchl, E, Martin, T, Cardinal, S, Wood, TJ, 2022a. Twenty-five new species of mining bees (Hymenoptera: Andrenidae: Andrena) from Israel and the Levant. Zootaxa 5185 (1): 1 - 109, DOI: https://doi.org/10.11646/zootaxa.5185.1.1" title="Twenty-five new species of mining bees (Hymenoptera: Andrenidae: Andrena) from Israel and the Levant." url="https://doi.org/10.11646/zootaxa.5185.1.1" volume="5185" year="2022 a">Pisanty et al. 2022a</bibRefCitation>
|
||
|
|
).
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="5A076F1DA77EC628A694A0CB5ECAD04A" pageId="0" pageNumber="241" type="materials_examined">
|
||
|
|
<paragraph id="C7B6F76F82C258E13F6CE3E1DB51881C" pageId="0" pageNumber="241">Material examined.</paragraph>
|
||
|
|
<paragraph id="25A97A0D1FFA00805558744A007362AE" pageId="0" pageNumber="241">
|
||
|
|
<emphasis id="C15AF5EBAB9CAD1D4DCB0E338EA83F33" bold="true" pageId="0" pageNumber="241">
|
||
|
|
<taxonomicName id="154BA74EB7A273BEF298F51A561D7A38" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena oviventris" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="oviventris">
|
||
|
|
<emphasis id="B9C70D4D5F6133875EE4A0BE1E0DF44E" italics="true" pageId="0" pageNumber="241">Andrena oviventris</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
:
|
||
|
|
<materialsCitation id="AF7217B6B8A86D7D3C016068D1C2803F" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Teniet el Had" country="Algeria" latitude="35.8727" location="Algeria" longLatPrecision="6" longitude="2.0007" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
|
||
|
|
<collectingCountry id="DDB9207CC55FC6401C468CA43E748ABF" name="Algeria">Algeria</collectingCountry>
|
||
|
|
:
|
||
|
|
<collectorName id="81D0DECD6F7CFD82AAACFBC43B8508AC">Teniet el Had</collectorName>
|
||
|
|
[
|
||
|
|
<geoCoordinate id="25495C6884AEBA4479DFD7423CBFC0CA" degrees="35.8727" direction="north" orientation="latitude" precision="5" value="35.8727">35.8727°N</geoCoordinate>
|
||
|
|
,
|
||
|
|
<geoCoordinate id="FF3419CB37A8EF7E6576FFF6F83D3FDF" degrees="2.0007" direction="east" orientation="longitude" precision="5" value="2.0007">2.0007°E</geoCoordinate>
|
||
|
|
],
|
||
|
|
<specimenCount id="2F5FFA1B5F63AF05CC8E90C7C63933F2" type="female">1♀</specimenCount>
|
||
|
|
, MNHN (
|
||
|
|
<typeStatus id="F7B0A73422F606B8F942A15500895ABE">lectotype</typeStatus>
|
||
|
|
; Fig.
|
||
|
|
<figureCitation id="3C0F4A4E4193B62CF2A7853027AFBD21" captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Andrena (Ovandrena) oviventris Perez, 1895, female lectotype A label details B profile C face, frontal view D terga, dorsal view." figureDoi="10.3897/jhr.96.101873.figure46" httpUri="https://binary.pensoft.net/fig/853438" pageId="0" pageNumber="241">46</figureCitation>
|
||
|
|
)
|
||
|
|
</materialsCitation>
|
||
|
|
</emphasis>
|
||
|
|
.
|
||
|
|
</paragraph>
|
||
|
|
<caption id="185F6407C80244D67D361C1BB47FA2F2" doi="10.3897/jhr.96.101873.figure46" httpUri="https://binary.pensoft.net/fig/853438" pageId="0" pageNumber="241" start="Figure 46" startId="F46">
|
||
|
|
<paragraph id="DC3E76DC3E283DE90935ADB814E23E51" pageId="0" pageNumber="241">
|
||
|
|
<emphasis id="91DB1BD7892F8FD68B5D3C89AAE01222" bold="true" pageId="0" pageNumber="241">Figure 46.</emphasis>
|
||
|
|
<taxonomicName id="22ED63FFFA6BC57043F03D7E111E4BB3" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Ovandrena) oviventris" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="oviventris" subGenus="Ovandrena">Andrena (Ovandrena) oviventris</taxonomicName>
|
||
|
|
<normalizedToken id="4053668D1B893E85F6E6D7E3A51D422B" originalValue="Pérez">Perez</normalizedToken>
|
||
|
|
, 1895, female
|
||
|
|
<typeStatus id="70E00B20A3B6E6C7C5819C267A077315">lectotype</typeStatus>
|
||
|
|
<emphasis id="2009D6640A1FF47AC6FE2EB6AF9A7DCB" bold="true" pageId="0" pageNumber="241">A</emphasis>
|
||
|
|
label details
|
||
|
|
<emphasis id="956CF5023ADF75E50411DBFF59433955" bold="true" pageId="0" pageNumber="241">B</emphasis>
|
||
|
|
profile
|
||
|
|
<emphasis id="AD3E7BF8CE9E6EF4B68C566F235DB4CF" bold="true" pageId="0" pageNumber="241">C</emphasis>
|
||
|
|
face, frontal view
|
||
|
|
<emphasis id="25D4F5A15EC777D78D4BD07FB1E8E4C5" bold="true" pageId="0" pageNumber="241">D</emphasis>
|
||
|
|
terga, dorsal view.
|
||
|
|
</paragraph>
|
||
|
|
</caption>
|
||
|
|
<paragraph id="A911A0D86C69B5E15C01546F7A2B5FE6" pageId="0" pageNumber="241">
|
||
|
|
<emphasis id="82CB9DEAA924C508A73EC74C213BB17D" bold="true" pageId="0" pageNumber="241">
|
||
|
|
<taxonomicName id="C631F94D6B7407A199086F58FB8F17DA" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena farinosa" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="farinosa">
|
||
|
|
<emphasis id="127ED7FB7D765E2AB0306BB26DAA854A" italics="true" pageId="0" pageNumber="241">Andrena farinosa</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
:
|
||
|
|
<materialsCitation id="2ABFF70D6959FF62A294744504E3DC7C" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Spain" location="Murcie" specimenCount="1" specimenCount-female="1" stateProvince="Murcie" typeStatus="lectotype">
|
||
|
|
<collectingCountry id="8AFAE0AFFB898F394436556392A1D5FB" name="Spain">Spain</collectingCountry>
|
||
|
|
:
|
||
|
|
<collectingRegion id="9862890BA81EBB8D1C1DAF047DBB181C" country="Spain" name="Murcia">Murcie</collectingRegion>
|
||
|
|
[
|
||
|
|
<collectingRegion id="5012762174B8C325452E059728CB07CA" country="Spain" name="Murcia">Murcia</collectingRegion>
|
||
|
|
],
|
||
|
|
<specimenCount id="972212EAAC36852DA425BE2F22757089" type="female">1♀</specimenCount>
|
||
|
|
, MNHN (
|
||
|
|
<typeStatus id="2C0D57EC2D451DDFF8C90FDF9159FD22">lectotype</typeStatus>
|
||
|
|
)
|
||
|
|
</materialsCitation>
|
||
|
|
</emphasis>
|
||
|
|
.
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
</treatment>
|
||
|
|
</document>
|