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<mods:title id="B5D8848A14937936375B4EDE98EBDA72">Nomenclatural notes and typifications in Campanula versicolor (Campanulaceae) and related names</mods:title>
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<mods:namePart id="33D9FB789FEAFFFDBEE5E0301CC1B4C5">Lakušić, Dmitar</mods:namePart>
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<mods:namePart id="8BE7CA376D8C5F2EC968F0FB8BDC0330">Pietro, Romeo Di</mods:namePart>
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<taxonomicName id="FACDD5E0F674FFAE898CFD126B767EAA" ID-CoL="QC6C" authority="Andrews (1804" authorityName="Andrews. B. Color" authorityYear="1804" box="[136,576,660,687]" class="Magnoliopsida" family="Campanulaceae" genus="Campanula" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="265" phylum="Tracheophyta" rank="species" species="versicolor">
<emphasis id="0FB97271F674FFAE898CFD1268BE7EAA" bold="true" box="[136,392,660,687]" italics="true" pageId="1" pageNumber="265">Campanula versicolor</emphasis>
<bibRefCitation id="595CD392F674FFAE888BFD106B767EAA" author="Andrews, H. C." box="[399,576,662,687]" pageId="1" pageNumber="265" refId="ref4865" refString="Andrews, H. C. (1804) The botanist's repository, comprising colour'd engravings of new and rare plants, Vol. 6. T. Bensley, London, 396 pp. https: // doi. org / 10.5962 / bhl. title. 51972" type="book" year="1804">Andrews (1804</bibRefCitation>
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: t. 396)
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<emphasis id="0FB97271F674FFAE898CFD3A69DB7ED4" bold="true" box="[136,237,700,721]" pageId="1" pageNumber="265">
<typeStatus id="E27610C1F674FFAE898CFD3A69DB7ED4" box="[136,237,700,721]" pageId="1" pageNumber="265" type="lectotype">Lectotype</typeStatus>
</emphasis>
(designated here):—ILLUSTRATION. plate CCCXCVI [tab. 396] in Andrews, 
<emphasis id="0FB97271F674FFAE8AEBFD3A6D687ED4" box="[1007,1118,700,721]" italics="true" pageId="1" pageNumber="265">Bot. Repos.</emphasis>
6: t. 396 (1804)(available at http:// plantillustrations.org/illustration.php?id_illustration=111483&amp;mobile=0&amp;code_category_taxon=1)(
<figureCitation id="A5F6B2E6F674FFAE8D7BFD666DFA7EF0" box="[1151,1228,736,757]" captionStart="FIGURE 1" captionStartId="1.[136,229,1875,1896]" captionTargetBox="[151,1436,927,1850]" captionTargetId="figure-230@1.[151,1436,927,1850]" captionTargetPageId="1" captionText="FIGURE 1. A. Lectotype of Campanula versicolor Andrews. B. Color illustration of C. versicolor Sibthorp." figureDoi="http://doi.org/10.5281/zenodo.13696516" httpUri="https://zenodo.org/record/13696516/files/figure.png" pageId="1" pageNumber="265">Fig. 1A</figureCitation>
).
</paragraph>
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<emphasis id="0FB97271F674FFAE898CFC8269EE7F1C" bold="true" box="[136,216,772,793]" pageId="1" pageNumber="265">
<typeStatus id="E27610C1F674FFAE898CFC8269EE7F1C" box="[136,216,772,793]" pageId="1" pageNumber="265" type="epitype">Epitype</typeStatus>
</emphasis>
(designated here):— 
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. Parnas [Mt Parnassus], stene, krečnjak [limestone cliffs], 
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, 
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, 
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a.s.l., 
<date id="497388A3F674FFAE89F2FCAE68A67F38" box="[246,400,808,829]" pageId="1" pageNumber="265" value="2014-08-21">21 August 2014</date>
, 
<emphasis id="0FB97271F674FFAE889EFCAE6A1F7F38" box="[410,809,808,829]" italics="true" pageId="1" pageNumber="265">D. Lakušić, N. Kuzmanović &amp; I. Janković</emphasis>
40045 (BEOU!)(
<figureCitation id="A5F6B2E6F674FFAE8AD6FCAE6D3A7F38" box="[978,1036,808,829]" captionStart="FIGURE 2" captionStartId="2.[136,229,2029,2050]" captionTargetBox="[155,1432,190,2004]" captionTargetId="figure-15@2.[155,1432,190,2004]" captionTargetPageId="2" captionText="FIGURE 2. Epitype of Campanula versicolor Andrews (BEOU 40045!)." figureDoi="http://doi.org/10.5281/zenodo.13696520" httpUri="https://zenodo.org/record/13696520/files/figure.png" pageId="1" pageNumber="265">Fig. 2</figureCitation>
).
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<emphasis id="0FB97271F674FFAE898CFCCA684F7F64" bold="true" box="[136,377,844,865]" pageId="1" pageNumber="265">Protologue information</emphasis>
:—“It was first raised from seeds brought to this country from 
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, by the late and much regretted Professor Sibthorp”.
</paragraph>
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<paragraph id="3D72AE63F674FFAE898CF8D56DAA7B6D" blockId="1.[136,1180,1875,1896]" box="[136,1180,1875,1896]" pageId="1" pageNumber="265">
<emphasis id="0FB97271F674FFAE898CF8D569C87B6D" bold="true" box="[136,254,1875,1896]" pageId="1" pageNumber="265">FIGURE 1.</emphasis>
A. Lectotype of 
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<emphasis id="0FB97271F674FFAE889BF8D56B407B6D" box="[415,630,1875,1896]" italics="true" pageId="1" pageNumber="265">Campanula versicolor</emphasis>
Andrews. B. Color
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illustration of 
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<emphasis id="0FB97271F674FFAE8ABBF8D56D0A7B6D" box="[959,1084,1875,1896]" italics="true" pageId="1" pageNumber="265">C. versicolor</emphasis>
Sibthorp.
</taxonomicName>
</paragraph>
</caption>
<caption id="69B2FEEBF677FFAD898CF86B6A7C7407" ID-DOI="http://doi.org/10.5281/zenodo.13696520" ID-Zenodo-Dep="13696520" box="[136,842,2029,2050]" httpUri="https://zenodo.org/record/13696520/files/figure.png" pageId="2" pageNumber="266" startId="2.[136,229,2029,2050]" targetBox="[155,1432,190,2004]" targetPageId="2" targetType="figure">
<paragraph id="3D72AE63F677FFAD898CF86B6A7C7407" blockId="2.[136,842,2029,2050]" box="[136,842,2029,2050]" pageId="2" pageNumber="266">
<emphasis id="0FB97271F677FFAD898CF86B69C87407" bold="true" box="[136,254,2029,2050]" pageId="2" pageNumber="266">FIGURE 2.</emphasis>
Epitype of 
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<emphasis id="0FB97271F677FFAD886AF86B6B737407" box="[366,581,2029,2050]" italics="true" pageId="2" pageNumber="266">Campanula versicolor</emphasis>
Andrews
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(BEOU 40045!).
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<paragraph id="3D72AE63F676FFAC898CFF266A977E6C" blockId="3.[136,1452,160,797]" pageId="3" pageNumber="267">
<emphasis id="0FB97271F676FFAC898CFF2669FF7CBC" bold="true" box="[136,201,160,185]" pageId="3" pageNumber="267">Note:</emphasis>
—The color illustration is published in the protologue together with the description, being part of original material. According to 
<bibRefCitation id="595CD392F676FFAC888DFF426BAB7CD8" author="Stafleu, F. A. &amp; Cowan, R. S." box="[393,669,196,221]" pageId="3" pageNumber="267" refId="ref6327" refString="Stafleu, F. A. &amp; Cowan, R. S. (1976) Taxonomic literature, Vol. 1. Ed. 2. Bohn, Scheltema &amp; Holkema, Utrecht, 1136 pp. Available from: http: // www. biodiversitylibrary. org / item / 103414" type="book" year="1976">Stafleu &amp; Cowan (1976)</bibRefCitation>
, the drawings in the Botanist‘s repository are made by Andrews based on the living specimens, and no herbarium material is known to exist for any of his drawings. The plant was raised from the seeds brought from 
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by Prof. Sibthorp (
<bibRefCitation id="595CD392F676FFAC8BF5FE8A6AAD7D20" author="Andrews, H. C." box="[753,923,268,293]" pageId="3" pageNumber="267" refId="ref4865" refString="Andrews, H. C. (1804) The botanist's repository, comprising colour'd engravings of new and rare plants, Vol. 6. T. Bensley, London, 396 pp. https: // doi. org / 10.5962 / bhl. title. 51972" type="book" year="1804">Andrews 1804</bibRefCitation>
). The description was made based on the living specimen as well. However, this species was discovered by John Sibthorp during his first field trip to the Mediterranean regions— 
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and 
<collectingCountry id="45DAEEF3F676FFAC887EFED268CC7D68" box="[378,506,340,365]" name="Turkey" pageId="3" pageNumber="267">Asia Minor</collectingCountry>
in 1786–1787. Unfortunately, he died soon after the return from the second field trip made in between 1794–1795, before he was able to publish the large part of his work from the Mediterranean field trips, intended to be the part of his 
<emphasis id="0FB97271F676FFAC8B1DFE1A6B997DB0" box="[537,687,412,437]" italics="true" pageId="3" pageNumber="267">Flora graeca</emphasis>
. During this first field trip, Sibthorp was accompanied by the artist Ferdinand Bauer, who made the botanical illustration including the one of 
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<emphasis id="0FB97271F676FFAC8ADDFE466D5B7DDC" box="[985,1133,448,473]" italics="true" pageId="3" pageNumber="267">C. versicolor</emphasis>
</taxonomicName>
(
<figureCitation id="A5F6B2E6F676FFAC8D79FE466DF87DDC" box="[1149,1230,448,473]" captionStart="FIGURE 1" captionStartId="1.[136,229,1875,1896]" captionTargetBox="[151,1436,927,1850]" captionTargetId="figure-230@1.[151,1436,927,1850]" captionTargetPageId="1" captionText="FIGURE 1. A. Lectotype of Campanula versicolor Andrews. B. Color illustration of C. versicolor Sibthorp." figureDoi="http://doi.org/10.5281/zenodo.13696516" httpUri="https://zenodo.org/record/13696516/files/figure.png" pageId="3" pageNumber="267">Fig. 1B</figureCitation>
)(
<bibRefCitation id="595CD392F676FFAC8DDAFE4669F27DF8" author="Stafleu, F. A. &amp; Cowan, R. S." pageId="3" pageNumber="267" refId="ref6380" refString="Stafleu, F. A. &amp; Cowan, R. S. (1985) Taxonomic literature, Vol. 5. Ed. 2. Bohn, Scheltema &amp; Holkema, Utrecht, 1066 pp. https: // doi. org / 10.5962 / bhl. title. 48631" type="book" year="1985">Stafleu &amp; Cowan 1985</bibRefCitation>
). The task of preparing and publishing Sibthorp‘s work was undertaken by J. Smith, who issued the first volume of „Flora Graeca prodromus“ two years after Andrews‘ publication. In that book, a short description of 
<taxonomicName id="FACDD5E0F676FFAC8C1DFD8E6C9A7E24" authorityName="Andrews. B. Color" authorityYear="1804" box="[1305,1452,520,545]" class="Magnoliopsida" family="Campanulaceae" genus="Campanula" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="267" phylum="Tracheophyta" rank="species" species="versicolor">
<emphasis id="0FB97271F676FFAC8C1DFD8E6C9A7E24" box="[1305,1452,520,545]" italics="true" pageId="3" pageNumber="267">C. versicolor</emphasis>
</taxonomicName>
is also provided. However, 
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<emphasis id="0FB97271F676FFAC88BEFDAA6B837E40" box="[442,693,556,581]" italics="true" pageId="3" pageNumber="267">Campanula versicolor</emphasis>
Sibthorp
</taxonomicName>
(in 
<bibRefCitation id="595CD392F676FFAC8A4CFDAA6DB17E40" author="Sibthorp, J. &amp; Smith, J. E." box="[840,1159,556,581]" pageId="3" pageNumber="267" refId="ref6291" refString="Sibthorp, J. &amp; Smith, J. E. (1806) Flora Graeca Prodromus, Vol. 1. Typis Richardi Taylor et socii et al., London, 442 pp." type="book" year="1806">Sibthorp &amp; Smith 1806: 138</bibRefCitation>
) is a later homonym of 
<taxonomicName id="FACDD5E0F676FFAC8C96FDAA68527E6C" authority="Andrews" authorityName="Andrews. B. Color" authorityYear="1804" class="Magnoliopsida" family="Campanulaceae" genus="Campanula" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="267" phylum="Tracheophyta" rank="species" species="versicolor">
<emphasis id="0FB97271F676FFAC8C96FDAA69CC7E6C" italics="true" pageId="3" pageNumber="267">C. versicolor</emphasis>
Andrews
</taxonomicName>
and is illegitimate according to Art. 53 of the ICN.
</paragraph>
<paragraph id="3D72AE63F676FFAC89B9FDF26C597F18" blockId="3.[136,1452,160,797]" pageId="3" pageNumber="267">
In addition, for the purpose of the precise application of the name, the specimen preserved at the herbarium of the University of Belgrade (BEOU 40045) is designated here as 
<typeStatus id="E27610C1F676FFAC8A29FD1E6A497EB4" box="[813,895,664,689]" pageId="3" pageNumber="267" type="epitype">epitype</typeStatus>
of 
<taxonomicName id="FACDD5E0F676FFAC8AA0FD1E6D007EB4" authorityName="Andrews. B. Color" authorityYear="1804" box="[932,1078,664,689]" class="Magnoliopsida" family="Campanulaceae" genus="Campanula" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="267" phylum="Tracheophyta" rank="species" species="versicolor">
<emphasis id="0FB97271F676FFAC8AA0FD1E6D007EB4" box="[932,1078,664,689]" italics="true" pageId="3" pageNumber="267">C. versicolor</emphasis>
</taxonomicName>
. The habitus and capsule, as well as the dimension of 
<taxonomicName id="FACDD5E0F676FFAC886DFD3A68CD7ED0" authorityName="Andrews. B. Color" authorityYear="1804" box="[361,507,700,725]" class="Magnoliopsida" family="Campanulaceae" genus="Campanula" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="267" phylum="Tracheophyta" rank="species" species="versicolor">
<emphasis id="0FB97271F676FFAC886DFD3A68CD7ED0" box="[361,507,700,725]" italics="true" pageId="3" pageNumber="267">C. versicolor</emphasis>
</taxonomicName>
, is not evident in the 
<typeStatus id="E27610C1F676FFAC8BE8FD3A6A607ED0" box="[748,854,700,725]" pageId="3" pageNumber="267" type="lectotype">lectotype</typeStatus>
. Thus, an 
<typeStatus id="E27610C1F676FFAC8AC3FD3A6D2F7ED0" box="[967,1049,700,725]" pageId="3" pageNumber="267" type="epitype">epitype</typeStatus>
representing the modern concept of the species is designated to support the 
<typeStatus id="E27610C1F676FFAC8B39FD666B917EFC" box="[573,679,736,761]" pageId="3" pageNumber="267" type="lectotype">lectotype</typeStatus>
. The material was collected on Mt Parnassus (
<collectingCountry id="45DAEEF3F676FFAC8DA2FD666DCF7EFC" box="[1190,1273,736,761]" name="Greece" pageId="3" pageNumber="267">Greece</collectingCountry>
) which was one of the localities visited by Sibthorp during his expedition on the Balkan Peninsula in 1787 (
<bibRefCitation id="595CD392F676FFAC8D83FC826C3B7F18" author="Bruce, M. R." box="[1159,1293,772,797]" pageId="3" pageNumber="267" pagination="353 - 362" refId="ref5005" refString="Bruce, M. R. (1970) John Sibthorp. Taxon 19 (3): 353 - 362. https: // doi. org / 10.2307 / 1219064" type="journal article" year="1970">Bruce 1970</bibRefCitation>
)(
<figureCitation id="A5F6B2E6F676FFAC8C19FC826C547F18" box="[1309,1378,772,797]" captionStart="FIGURE 2" captionStartId="2.[136,229,2029,2050]" captionTargetBox="[155,1432,190,2004]" captionTargetId="figure-15@2.[155,1432,190,2004]" captionTargetPageId="2" captionText="FIGURE 2. Epitype of Campanula versicolor Andrews (BEOU 40045!)." figureDoi="http://doi.org/10.5281/zenodo.13696520" httpUri="https://zenodo.org/record/13696520/files/figure.png" pageId="3" pageNumber="267">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>