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<mods:title id="033368E33739796951A49B3CDCE5364D">Annotated checklist of the land snail fauna from southern Cambodia (Mollusca, Gastropoda)</mods:title>
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<mods:namePart id="346AEC86A600F0195E398F7DE30F8A39">Sutcharit, Chirasak</mods:namePart>
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<mods:namePart id="B2854335047A70CB2C27FC68E4C5D232">Thach, Phanara</mods:namePart>
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<mods:namePart id="93CFFBE8FA60C31E094E16EE281AFA71">Chhuoy, Samol</mods:namePart>
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<mods:namePart id="D95CA4D3A20BDD01DA9A33E803392C94">Ngor, Peng Bun</mods:namePart>
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<mods:namePart id="9465553943F4C04CF03029FC5AF1BBF3">Jeratthitikul, Ekgachai</mods:namePart>
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<mods:namePart id="C1508DC0F7D5337F4CA9C0C01412BA94">Siriwut, Warut</mods:namePart>
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<mods:namePart id="FFC679FC7213A8B2F75893324C329088">Srisonchai, Ruttapon</mods:namePart>
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<mods:namePart id="287037B3139B8DB808E9E4A5CF714909">Ng, Ting Hui</mods:namePart>
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<mods:namePart id="6F2CBDEAB0CAA884D3F9428DE769E87D">Pholyotha, Arthit</mods:namePart>
<mods:affiliation id="C7928F1D35A40EEA053E35C8D219FF2C">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="13A570F10BBA8C918C33D65A8B1A2747">Jirapatrasilp, Parin</mods:namePart>
<mods:affiliation id="3441D44827B704B79786224565E11B60">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<treatment id="A8BA2B42822F538E889B28DC9E7AA101" ID-GBIF-Taxon="165073725" LSID="urn:lsid:plazi:treatment:A8BA2B42822F538E889B28DC9E7AA101" httpUri="http://treatment.plazi.org/id/A8BA2B42822F538E889B28DC9E7AA101" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Cyclophoridae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda">
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<paragraph id="F01B940C7CFDFCECEC2125AEA7CF5C51" pageId="0" pageNumber="1">
<taxonomicName id="F8D2DAA0D26BE632788E46DDF7EB35EF" ID-CoL="7RSTR" LSID="A8BA2B42-822F-538E-889B-28DC9E7AA101" authority="(Morlet, 1885)" baseAuthorityName="Morlet" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Lagocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagocheilus klobukowskii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="klobukowskii">Lagocheilus klobukowskii (Morlet, 1885)</taxonomicName>
<figureCitation id="2C8C394A07D9665DBFC6F8F44901E591" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Living snails of A Lagocheilus klobukowskii (Morlet, 1885) B Lagocheilus landesi (Morlet, 1885) C Pupina crosseana Morlet, 1883 D Valiguna siamensis (Martens, 1867) E, F Valiguna sp. E notum or dorsal view and F hyponotum or ventral view G Succinea tenuis Morelet, 1865 and H Hypselostoma cambodjense Benthem Jutting, 1962. All not to scale." figureDoi="10.3897/zookeys.948.51671.figure3" httpUri="https://binary.pensoft.net/fig/430960" pageId="0" pageNumber="1">Figs 3A</figureCitation>
<figureCitation id="2B24B06D81D6D2DA49430BCA1B8E1CA7" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Lagocheilus klobukowskii (Morlet, 1885) A yellowish morph and B purplish-black morph C Pupina crosseana Morlet, 1883 D Succinea tenuis Morelet, 1865 and E, F Hypselostoma benetuitum Vermeulen et al., 2019 E from locality no. 11 (type locality) and F from locality no. 17." figureDoi="10.3897/zookeys.948.51671.figure7" httpUri="https://binary.pensoft.net/fig/430964" pageId="0" pageNumber="1">, 7A, B</figureCitation>
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<taxonomicName id="5506EAB0DBE03A966D1B594C1B5E9072" authorityName="Morlet" authorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclophorus klobukowskii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="klobukowskii">Cyclophorus klobukowskii</taxonomicName>
Morlet, 1885[1884]: 391, 392, pl. 12, fig. 1. Type locality: Near the Kamchay rapids, around the 
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cave (Kampot-Hatien road); commonly found on mountains, in forests, up to Compong-Som, and on the banks of 
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. 
<bibRefCitation id="186B72E7CDF3F5D8F9EE8A86052AB15F" author="Fischer, PH" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="40 - 64" refId="B26" refString="Fischer, PH, 1973a. Les mollusques testaces du Cambodge. Premier partie: Introduction et Gasteropodes Prosobranches. Journal de Conchyliologie 90: 40 - 64" title="Les mollusques testaces du Cambodge. Premier partie: Introduction et Gasteropodes Prosobranches." volume="90" year="1973 a">Fischer 1973a</bibRefCitation>
: 46, 47.
</paragraph>
<paragraph id="76CCECA3ED61C131A6926F77A7B6B9CB" pageId="0" pageNumber="1">
<taxonomicName id="3C16E4E4D4780E2F5979AAF38C8BDA90" baseAuthorityName="Morlet" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Lagocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagocheilus klobukowskii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="klobukowskii">Lagocheilus klobukowskii</taxonomicName>
: 
<bibRefCitation id="24D4362C019378C0F3B5F71CE72818CA" DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
: 19, 20, figs 9b, c, 18c.
</paragraph>
</subSubSection>
<subSubSection id="209CED525198244ECC1BAED0AEF20B44" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="C058D509549D1E43AD48F5501A0693B3" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="E9850AED149A03845AF44B1323EC14F5" pageId="0" pageNumber="1">
Locality no. 9: CUMZ-CM044 (7 shells), CUMZ-CM045 (12 specimens in ethanol). Locality no. 10: CUMZ-CM068 (3 specimens in ethanol). Locality no. 11: CUMZ-CM079 (2 shells). Locality no. 13: CUMZ-CM128 (3 shells), CM129 (53 specimens in ethanol; Fig. 
<figureCitation id="A926408207AF9BE9BAA6396C9B086E71" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Living snails of A Lagocheilus klobukowskii (Morlet, 1885) B Lagocheilus landesi (Morlet, 1885) C Pupina crosseana Morlet, 1883 D Valiguna siamensis (Martens, 1867) E, F Valiguna sp. E notum or dorsal view and F hyponotum or ventral view G Succinea tenuis Morelet, 1865 and H Hypselostoma cambodjense Benthem Jutting, 1962. All not to scale." figureDoi="10.3897/zookeys.948.51671.figure3" httpUri="https://binary.pensoft.net/fig/430960" pageId="0" pageNumber="1">3A</figureCitation>
). Locality no. 17: CUMZ-CM137 (18 shells; Fig. 
<figureCitation id="1B9627C9B14B73334A40875C5C5A652A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Lagocheilus klobukowskii (Morlet, 1885) A yellowish morph and B purplish-black morph C Pupina crosseana Morlet, 1883 D Succinea tenuis Morelet, 1865 and E, F Hypselostoma benetuitum Vermeulen et al., 2019 E from locality no. 11 (type locality) and F from locality no. 17." figureDoi="10.3897/zookeys.948.51671.figure7" httpUri="https://binary.pensoft.net/fig/430964" pageId="0" pageNumber="1">7A, B</figureCitation>
). The snails were found to live on the ground among leaf litter and decaying wood, on tree trunks and limestone wall.
</paragraph>
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<paragraph id="F2C56C9E76176A5E3EA4567E2E0D3853" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="CA18502695032116D7838DE0381495E5" pageId="0" pageNumber="1">
Cambodia and Laos (
<bibRefCitation id="BB00F8EF38B76B808CCAC6059EECF730" DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
).
</paragraph>
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<subSubSection id="EB593F3E882CB6AA54BB5D2B71C0E6D4" pageId="0" pageNumber="1" type="remarks">
<paragraph id="5045987CE587AB3B270AA06E0EC7D907" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="589452FCD207D71A8CC7BFF152A781B3" pageId="0" pageNumber="1">
This species was described from 
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grotte de 
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(route de Kampot 
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Hatien) 
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. We collected topotypic specimens that tend to have a variable shell colour from yellowish (Fig. 
<figureCitation id="19610899C4147CF68BFA486EA69EB708" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Lagocheilus klobukowskii (Morlet, 1885) A yellowish morph and B purplish-black morph C Pupina crosseana Morlet, 1883 D Succinea tenuis Morelet, 1865 and E, F Hypselostoma benetuitum Vermeulen et al., 2019 E from locality no. 11 (type locality) and F from locality no. 17." figureDoi="10.3897/zookeys.948.51671.figure7" httpUri="https://binary.pensoft.net/fig/430964" pageId="0" pageNumber="1">7A</figureCitation>
, see fig. 9b in 
<bibRefCitation id="2E7D289751B212A5769F6E23E2CECE0B" DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
for the syntype) to purplish-black (Fig. 
<figureCitation id="6E536A4749FB6A73B6E03A517674C5E8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Lagocheilus klobukowskii (Morlet, 1885) A yellowish morph and B purplish-black morph C Pupina crosseana Morlet, 1883 D Succinea tenuis Morelet, 1865 and E, F Hypselostoma benetuitum Vermeulen et al., 2019 E from locality no. 11 (type locality) and F from locality no. 17." figureDoi="10.3897/zookeys.948.51671.figure7" httpUri="https://binary.pensoft.net/fig/430964" pageId="0" pageNumber="1">7B</figureCitation>
). This limestone associated species has a wide distribution from southern Cambodia to eastern Laos (
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). The snails are commonly found in montane forest, living on decaying wood, on tree trunks and exposed limestone.
</paragraph>
<caption id="72B5F498F4B0080F8DED5202ADD13928" doi="10.3897/zookeys.948.51671.figure7" httpUri="https://binary.pensoft.net/fig/430964" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
<paragraph id="9B4EBE2B95019B5B530DE06CEAA82AB2" pageId="0" pageNumber="1">
<emphasis id="F16A8759E57571B7B313A7A381CA9FC4" bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
<emphasis id="0B3F80578227C20CDB8CD42800015065" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
<taxonomicName id="FBCA2D1F373DDD8293C7592089140BB8" baseAuthorityName="Morlet" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Lagocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagocheilus klobukowskii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="klobukowskii">
<emphasis id="95EAF1ECB1D680674CE680FFEBFA9F0A" italics="true" pageId="0" pageNumber="1">Lagocheilus klobukowskii</emphasis>
</taxonomicName>
(Morlet, 1885) 
<emphasis id="5E6C536455F32AAF89A7704B18A56B4C" bold="true" pageId="0" pageNumber="1">A</emphasis>
yellowish morph and 
<emphasis id="50E1FC25823932B285DF0F3E550C3993" bold="true" pageId="0" pageNumber="1">B</emphasis>
purplish-black morph 
<emphasis id="3C9884CEF015197B3EFAC69B06B7929D" bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName id="429B04B2F315198E7D5DFE935058C1A3" authorityName="Morlet" authorityYear="1883" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupina crosseana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="crosseana">
<emphasis id="B0BD164BDA821FFB4FA1567A8C7F2F45" italics="true" pageId="0" pageNumber="1">Pupina crosseana</emphasis>
</taxonomicName>
Morlet, 1883 
<emphasis id="E6C7031A74329BC46BA22B9331F542F3" bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName id="FDF106D4CD41D0B77AD2AEFD98F303D4" authorityName="Morelet" authorityYear="1865" class="Gastropoda" family="Succineidae" genus="Succinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Succinea tenuis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tenuis">
<emphasis id="EA56EAEC43A8C60DB797423389C78CAE" italics="true" pageId="0" pageNumber="1">Succinea tenuis</emphasis>
</taxonomicName>
Morelet, 1865 and 
<emphasis id="A2123374EE16D8CF1412589A1925DA11" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
<taxonomicName id="A50C161FD900B1B811B9F949A626174F" class="Gastropoda" family="Gastrocoptidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma benetuitum" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="benetuitum">
<emphasis id="DEC63173280D700844EC50454FE31089" italics="true" pageId="0" pageNumber="1">Hypselostoma benetuitum</emphasis>
</taxonomicName>
Vermeulen et al., 2019 
<emphasis id="D7D752B468E9AB65688901527BF1E9BE" bold="true" pageId="0" pageNumber="1">E</emphasis>
from locality no. 11 (type locality) and 
<emphasis id="67ABABE12451BB6266CA0B53475C2E17" bold="true" pageId="0" pageNumber="1">F</emphasis>
from locality no. 17.
</paragraph>
</caption>
<paragraph id="33395D93FAD7650106381EE985FA9F8D" pageId="0" pageNumber="1">
<taxonomicName id="9A68B62EFA2287904B81C2648ED5BDA6" baseAuthorityName="Morlet" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Lagocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagocheilus klobukowskii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="klobukowskii">
<emphasis id="C8659D95511D30299252BFB36B953BA5" italics="true" pageId="0" pageNumber="1">Lagocheilus klobukowskii</emphasis>
</taxonomicName>
was originally placed in the genus 
<taxonomicName id="CAA3CE356139B067F48BA386379F7C7D" class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclophorus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="636E199F9E6FEF433C10FD6AF27A8EF9" italics="true" pageId="0" pageNumber="1">Cyclophorus</emphasis>
</taxonomicName>
and later was transferred to the genus 
<taxonomicName id="5B612A90ACB0A077C8450CE74ADC92FE" authorityName="W.T.Blanford" authorityYear="1864" class="Gastropoda" family="Cyclophoridae" genus="Lagocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagocheilus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="B8CC2ECBEB93C58E4AABD37145F61533" italics="true" pageId="0" pageNumber="1">Lagocheilus</emphasis>
</taxonomicName>
(see 
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). The distinguishing characters from the genus 
<taxonomicName id="6C457692231BDA20F3F92C9EC0088970" class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclophorus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="BBC580D51023F2F7FE4AA4019F1FA7C8" italics="true" pageId="0" pageNumber="1">Cyclophorus</emphasis>
</taxonomicName>
are a conic shell, an aperture thickened (not expanded), and a thick calcareous, multispiral and plate-like operculum, whereas 
<taxonomicName id="32DCCEE63A905688F0CBE169B7975607" class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclophorus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="1226024C6FEF906FFC28F2AFCE5F975F" italics="true" pageId="0" pageNumber="1">Cyclophorus</emphasis>
</taxonomicName>
has a turbinate shell, a thick and expanded lip, and a corneous multispiral operculum.
</paragraph>
</subSubSection>
</treatment>
</document>