<documentid="C059315E6533A5706BF0801B3B03DD2D"ID="10.11646/zootaxa.4273.2.5SLASH11211"ID-CLB-Dataset="32916"ID-DOI="10.11646/zootaxa.4273.2.5"ID-GBIF-Dataset="58cea2d1-846c-4754-999e-bb49270cfb85"ID-ISSN="1175-5326"ID-Zenodo-Dep="801976"ID-ZooBank="1A2E1716-5EAA-4268-B30B-1F5B1BB23BFF"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"checkinTime="1496389609248"checkinUser="plazi"docAuthor="Komai, Tomoyuki"docDate="2017"docId="9E5987D1022BFFCB75F0FC7CFCF0F9BB"docLanguage="en"docName="zootaxa.4273.2.5SLASH11211.pdf"docOrigin="Zootaxa 4273 (2)"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Catapaguroides fragilis Melin 1939"docType="treatment"docVersion="26"lastPageNumber="243"masterDocId="6260FFA9022AFFC37567FFD5FFA7FF97"masterDocTitle="Redescription of the pagurid hermit crab Catapaguroides fragilis (Melin, 1939) and descriptions of two new species from deep-sea off the Ryukyu Islands, Japan (Crustacea: Decapoda: Anomura)"masterLastPageNumber="257"masterPageNumber="235"pageNumber="236"updateTime="1732724065403"updateUser="ExternalLinkService">
<mods:titleid="463D8B2AC071B09F8FD1882C90271A91">Redescription of the pagurid hermit crab Catapaguroides fragilis (Melin, 1939) and descriptions of two new species from deep-sea off the Ryukyu Islands, Japan (Crustacea: Decapoda: Anomura)</mods:title>
<bibRefCitationid="72614B36022BFFC274A5FC7CFDF0FC54"author="Melin"box="[450,599,937,963]"pageId="1"pageNumber="236"refString="Melin, G. (1939) Paguriden und Galatheiden von Prof. Dr. Sixten Bocks Expedition nach den Bonin-Inseln 1914. Kongliga Svenska Uetenskapsakademiens Handlinger, Series 3, 18, 1 - 119."type="journal article"year="1939">Melin, 1939</bibRefCitation>
<bibRefCitationid="72614B36022BFFC27490FBEDFDD1FBD9"author="Melin"box="[503,630,1080,1102]"pageId="1"pageNumber="236"refString="Melin, G. (1939) Paguriden und Galatheiden von Prof. Dr. Sixten Bocks Expedition nach den Bonin-Inseln 1914. Kongliga Svenska Uetenskapsakademiens Handlinger, Series 3, 18, 1 - 119."type="journal article"year="1939">Melin, 1939</bibRefCitation>
<bibRefCitationid="72614B36022BFFC274B7FB83FD3EFBFB"author="Saint"box="[464,665,1110,1132]"pageId="1"pageNumber="236"refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne- Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum National d'Histoire Naturelle, Paris, Series 2, 39, 923 - 954, 1100 - 1119. [dated 1967, published 1968]"type="journal article"year="1968">Saint Laurent 1968</bibRefCitation>
<bibRefCitationid="72614B36022BFFC27621FB83FC74FBFB"author="Miyake"box="[838,979,1110,1132]"pageId="1"pageNumber="236"refString="Miyake, S. (1978) The Crustacean Anomura of Sagami Bay. Hoikusha, Tokyo, 200 pp. (English text) + 161 pp. (Japanese text), 4 pls."type="book"year="1978">Miyake 1978</bibRefCitation>
<bibRefCitationid="72614B36022BFFC27198FB83FEF0FB1C"author="McLaughlin"pageId="1"pageNumber="236"refString="McLaughlin, P. A. & Pittman, C. (2002) Reinstatement and rediagnosis of Catapaguroides setosus and description of a second Hawaiian species of the genus (Decapoda: Anomura: Paguridea: Paguridae). Pacific Science, 56, 41 - 48. https: // doi. org / 10.1353 / psc. 2002.0006"type="journal article"year="2002">McLaughlin & Pittman 2002</bibRefCitation>
<bibRefCitationid="72614B36022BFFC27625FBA0FB95FB1C"author="Okuno"box="[834,1074,1141,1163]"pageId="1"pageNumber="236"refString="Okuno, J. & Arima, H. (2006) Additional records of inshore hermit crabs (Crustacea, Decapoda, Anomura) from Izu-ohshima Island, the northern Izu Islands, Japan. Bulletin of the Biogeographical Society of Japan, 61, 29 - 43. [in Japanese with English abstract]"type="journal article"year="2006">Okuno & Arima 2006</bibRefCitation>
<bibRefCitationid="72614B36022BFFC2747EFB41FDACFB3D"author="Komai"box="[281,523,1172,1194]"pageId="1"pageNumber="236"refString="Komai, T. & Rahayu, D. L. (2013) Species of the pagurid hermit crab genus Catapaguroides A. Milne-Edwards & Bouvier, 1892 (Crustacea: Decapoda: Anomura) from the Bohol Sea, the Philippines, with descriptions of eight new species. Raffles Bulletin of Zoology, 61, 143 - 188."type="journal article"year="2013">Komai & Rahayu 2013</bibRefCitation>
<collectionCodeid="70E1AE02022BFFC2762EFB02FC07FB78"box="[841,928,1239,1263]"country="Canada"httpUri="http://grbio.org/cool/8z6v-f8is"name="Saskatchewan Museum of Natural History"pageId="1"pageNumber="236">SMNH</collectionCode>
<collectingDateid="720AE9EF022BFFC2714EFB2EFB66FA84"box="[1065,1217,1274,1299]"pageId="1"pageNumber="236"value="1914-07-29">29 July 1914</collectingDate>
Ten pairs of biserial phyllobranchiate gills; 2 very small, but lamellate arthrobranchs on maxilliped 3; no pleurobranch on either wall of thoracomere 7 (above base of pereopod 4).
<figureCitationid="8ECB2A42022BFFC27478F9EBFEF9F9C0"box="[287,350,1598,1623]"captionStart="FIGURE 1"captionStartId="2.[151,250,1709,1731]"captionTargetBox="[167,1415,361,1640]"captionTargetId="figure@2.[159,1444,337,1679]"captionTargetPageId="2"captionText="FIGURE 1. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, shield (including carapace lateral lobes) and cephalic appendages, dorsal view; B, left antennal peduncle, ventral view; C, left maxilliped 3, lateral view; D, same, ischium, ventral view; E, left pereopod 4, lateral view; F, thoracic sternite 6, ventral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, outer view."httpUri="https://zenodo.org/record/801978/files/figure.png"pageId="1"pageNumber="236">Fig. 1</figureCitation>
A) approximately as long as wide; anterior margins between rostral lobe and lateral projections concave; anterolateral margins slightly terraced or sloping, posterior margin roundly truncate; dorsal surface almost glabrous, with few tufts of short setae laterally. Rostrum rounded. Lateral projections triangular, produced to level of rostrum, each with small marginal or submarginal spine. Carapace lateral lobes moderately wide, well calcified.
<figureCitationid="8ECB2A42022BFFC274EAF927FE6CF89C"box="[397,459,1778,1803]"captionStart="FIGURE 1"captionStartId="2.[151,250,1709,1731]"captionTargetBox="[167,1415,361,1640]"captionTargetId="figure@2.[159,1444,337,1679]"captionTargetPageId="2"captionText="FIGURE 1. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, shield (including carapace lateral lobes) and cephalic appendages, dorsal view; B, left antennal peduncle, ventral view; C, left maxilliped 3, lateral view; D, same, ischium, ventral view; E, left pereopod 4, lateral view; F, thoracic sternite 6, ventral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, outer view."httpUri="https://zenodo.org/record/801978/files/figure.png"pageId="1"pageNumber="236">Fig. 1</figureCitation>
A) long, slender, 0.8–0.9 times as long as shield, subcylindrical, with basal part slightly inflated; dorsal surface mesially with row of 4 tufts of stiff setae; cornea small, rounded, very slightly dilated, corneal width approximately 0.2 of peduncular length. Ocular acicle triangular, terminating in small marginal spine, bearing few short setae subdistally, separated to each other basally by basal width of 1 acicle. Interocular lobe with slightly convex anterior surface.
<figureCitationid="8ECB2A42022BFFC274AEF873FDABF828"box="[457,524,1958,1983]"captionStart="FIGURE 1"captionStartId="2.[151,250,1709,1731]"captionTargetBox="[167,1415,361,1640]"captionTargetId="figure@2.[159,1444,337,1679]"captionTargetPageId="2"captionText="FIGURE 1. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, shield (including carapace lateral lobes) and cephalic appendages, dorsal view; B, left antennal peduncle, ventral view; C, left maxilliped 3, lateral view; D, same, ischium, ventral view; E, left pereopod 4, lateral view; F, thoracic sternite 6, ventral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, outer view."httpUri="https://zenodo.org/record/801978/files/figure.png"pageId="1"pageNumber="236">Fig. 1</figureCitation>
A), when fully extended, overreaching distal corneal margin by 0.8 length of ultimate article. Basal article with spinule on lateral face of statocyst lobe. Ultimate article 0.6–0.7 length of shield, slightly broadened distally, with 2 long plumose setae at dorsolateral distal portion; dorsal surface with row of some widely spaced setae. Dorsal flagellum longer than ultimate peduncular segment; distal portion more than 3 times as long as proximal aesthetasc-bearing portion, consisting of 6 or 7 articles; ventral flagellum consisting of 6 articles.
<figureCitationid="8ECB2A420228FFC174CCF892FE4CF8C8"box="[427,491,1863,1888]"captionStart="FIGURE 1"captionStartId="2.[151,250,1709,1731]"captionTargetBox="[167,1415,361,1640]"captionTargetId="figure@2.[159,1444,337,1679]"captionTargetPageId="2"captionText="FIGURE 1. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, shield (including carapace lateral lobes) and cephalic appendages, dorsal view; B, left antennal peduncle, ventral view; C, left maxilliped 3, lateral view; D, same, ischium, ventral view; E, left pereopod 4, lateral view; F, thoracic sternite 6, ventral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, outer view."httpUri="https://zenodo.org/record/801978/files/figure.png"pageId="2"pageNumber="237">Fig. 1</figureCitation>
A, B) not reaching distal corneal margin. Articles 5 and 4 with few setae. Article 3 with 1 prominent spine at ventromesial distal angle, extending as far as dorsolateral distal angle of article 2. Article 2 with dorsolateral distal angle produced into simple spine far falling short of midlength of article 4, occasionally with 1 subterminal spinule on mesial margin; dorsomesial distal angle with small spine. Article 1 unarmed. Antennal acicle moderately slender, gently arcuate, overreaching corneal base, occasionally reaching distal corneal margin, terminating in small spine, with row of short to long stiff setae mesially. Antennal flagellum exceeding 6 times of shield length, reaching to tip of extended right cheliped in males; articles each principally with 2 or 3 setae on distal margin, those in proximal half of flagellum each with setae of 2 or more length of 1 article.
A–D) somewhat elongate; propodalcarpal articulation with slight clockwise rotation. Chela about 2.5 times as long as wide, widest at base of dactylus. Dactylus articulating moderately obliquely with palm, about 0.6 length of palm, gently curved ventrally; dorsal surface convex, unarmed, dorsomesial margin not delimited, armed with 1 small spine proximally; cutting edge with 3 blunt, triangular calcareous teeth, terminating in calcareous claw; scattered tufts of short to long setae on surfaces. Palm 1.8 times as long as wide, subequal in length to carpus, with scattered tufts of short to long setae on surfaces (ventral setae longest); dorsal surface gently convex transversely, with dorsomesial row of 8 or 9 small spines and 1 dorsoproximal spine or tubercle, otherwise unarmed; dorsolateral and dorsomesial margins not delimited; ventral surface slightly convex. Fixed finger with 2 blunt calcareous teeth on cutting edge, terminating in calcareous claw. Carpus subcylindrical, about 1.2 length of merus, slightly broadened distally, about 2 times longer than distal width; surfaces with scattered tufts of long setae; dorsal surface with row of 7–9 small spines or tubercles on midline (often in double row) and 1 small mesial spine proximal to midlength; ventrolateral distal angle and distomesial angle each with tiny spine. Merus with transverse rows of stiff setae on dorsal surface; dorsodistal margin with small spine mesially, partially obscured by stiff setae; lateral surface with several tufts of setae adjacent to ventrolateral margin, ventrolateral margin with 2 or 3 small spines distally; mesial surface mostly glabrous, ventromesial margin with 2 small spines at distal angle and 1 subdistal spine; ventral surface with scattered tufts of long setae. Ischium unarmed, but bearing long bristle-like setae on dorsal margin and mesial face.
<figureCitationid="8ECB2A420229FFC074ABFC01FDABFC7B"box="[460,524,980,1005]"captionStart="FIGURE 3"captionStartId="6.[151,250,1936,1958]"captionTargetBox="[198,1380,209,1925]"captionTargetId="figure@6.[170,1414,193,1932]"captionTargetPageId="6"captionText="FIGURE 3. Catapaguroides fragilis (Melin, 1939), A – C, female (sl 1.18 mm), CBM-ZC 9018; C – E, male (sl 1.54 mm), CBM-ZC 13594. A, right cheliped, mesial view; B, same, lateral view, setae omitted; C, same, chela and carpus, dorsal view; D, left cheliped, mesial view; E, same, lateral view, setae omitted; F, same, chela and carpus, dorsal view, setae omitted."httpUri="https://zenodo.org/record/801982/files/figure.png"pageId="3"pageNumber="238">Fig. 3</figureCitation>
A–C) not particularly elongate; setation less developed. Chela about 2.4 times as long as wide, ovate in general outlines. Dactylus 0.8 length of palm, gently curved ventrally; dorsal surface unarmed, dorsomesial margin not delimited, with 1 small spine proximally; cutting edge with 3 blunt, triangular calcareous teeth, terminating in tiny calcareous claw. Palm 1.4 times as long as wide, subequal in length to carpus; dorsal surface gently convex, with dorsomesial row of 5 or 6 small spines and 1 small but conspicuous dorsoproximal spine; dorsolateral and dorsomesial margins not delimited; ventral surface convex. Fixed finger with 2 blunt calcareous teeth on cutting edge, terminating in calcareous claw. Carpus slightly longer than merus, slightly broadened distally, about 1.7 times longer than distal width; dorsal surface with row of 4 small spines on midline and 1 mesial spine at about midlength; ventrolateral distal distomesial angles each with small spine. Merus with 3 spines on ventrolateral margin distally; ventromesial margin somewhat expanded, with 2 spines at distal angle and 1 subdistal spine; ventral surface concave, with scattered tufts of long setae. Ischium as in male cheliped.
<figureCitationid="8ECB2A420229FFC07415FA8AFE11FAEF"box="[370,438,1375,1400]"captionStart="FIGURE 3"captionStartId="6.[151,250,1936,1958]"captionTargetBox="[198,1380,209,1925]"captionTargetId="figure@6.[170,1414,193,1932]"captionTargetPageId="6"captionText="FIGURE 3. Catapaguroides fragilis (Melin, 1939), A – C, female (sl 1.18 mm), CBM-ZC 9018; C – E, male (sl 1.54 mm), CBM-ZC 13594. A, right cheliped, mesial view; B, same, lateral view, setae omitted; C, same, chela and carpus, dorsal view; D, left cheliped, mesial view; E, same, lateral view, setae omitted; F, same, chela and carpus, dorsal view, setae omitted."httpUri="https://zenodo.org/record/801982/files/figure.png"pageId="3"pageNumber="238">Fig. 3</figureCitation>
D–F) moderately slender, reaching midlength of right chela in males; propodal-carpal articulation without noticeable rotation. Chela not arched, 3.2 times longer than wide. Dactylus 0.9–1.0 times as long as palm, unarmed, with numerous tufts of short to long setae on surfaces; dorsomesial margin not delimited, unarmed; cutting edge with row of minute corneous teeth, terminating in small corneous claw. Palm 0.8 length of carpus, with scattered tufts of long setae on surfaces; dorsal surface slightly convex, with 2 or 3 small dorsomesial spines (second spine sometimes larger than others), otherwise spineless. Fixed finger with row of minute corneous teeth at least on distal half of cutting edge, terminating in small corneous claw. Carpus slightly widened distally, about 2.6 times longer than distal width, with small spine at dorsomesial distal angle; dorsal surface with 2–5 small spines on midline, and with numerous tufts of long stiff setae; lateral, mesial and ventral surfaces also with scattered tufts of long setae, ventrolateral distal angle with tiny spine; ventral surface slightly convex. Merus with some tufts of short to long stiff setae on dorsal surface, dorsodistal margin unarmed; ventrolateral margin generally convex, with 2 small spines distally; ventromesial margin also convex, with 2 small spines subdistally; ventral surface with scattered long setae. Ischium with tufts of long bristle-like setae on dorsal surface and distomesial margin.
<figureCitationid="8ECB2A420229FFC077E8F882FD71F8E7"box="[655,726,1879,1904]"captionStart="FIGURE 4"captionStartId="7.[151,250,1873,1895]"captionTargetBox="[204,1377,231,1824]"captionTargetId="figure@7.[149,1434,193,1840]"captionTargetPageId="7"captionText="FIGURE 4. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, right pereopod 2, lateral view; B, left pereopod 3, lateral view; C, D, dactyli of right pereopod 2 and left pereopod 3, mesial view."httpUri="https://zenodo.org/record/801984/files/figure.png"pageId="3"pageNumber="238">Fig. 4</figureCitation>
A, B) long and slender, right second reaching extended right cheliped in males. Dactyli 1.1–1.2 (second) or 1.4–1.5 (third) times as long as propodi, about 13–14 times as long as wide, straight in dorsal view, weakly curved ventrally in lateral view, terminating in long, slender corneous claws; dorsal margins each with row of moderately long bristle-like setae; mesial faces each with row of stiff setae on midline and row of 2–4 (second) or 4–7 slender spiniform setae adjacent to ventral margin (
<figureCitationid="8ECB2A42022EFFC771D8FF42FAA5FF27"box="[1215,1282,151,176]"captionStart="FIGURE 4"captionStartId="7.[151,250,1873,1895]"captionTargetBox="[204,1377,231,1824]"captionTargetId="figure@7.[149,1434,193,1840]"captionTargetPageId="7"captionText="FIGURE 4. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, right pereopod 2, lateral view; B, left pereopod 3, lateral view; C, D, dactyli of right pereopod 2 and left pereopod 3, mesial view."httpUri="https://zenodo.org/record/801984/files/figure.png"pageId="4"pageNumber="239">Fig. 4</figureCitation>
C, D); ventral margin unarmed. Propodi each with long, slender spiniform setae flanked by 2–4 shorter spiniform or bristle-like setae (male second) or 1 small spiniform setae (female second and third in both sexes) on ventrodistal margin mesially; dorsal margins each with tufts of moderately long stiff setae; ventral surface with sparse long setae. Carpi usually with tiny dorsodistal spine; sparse tufts moderately long setae on dorsal margins. Meri each with 1 tiny distal spine on ventrolateral margin (second) or unarmed (third); dorsal margins each with row of tufts of moderately long setae; ventral margins with tufts of long setae. Ischia each with sparse setae on dorsal and ventral margins. Second pereopods with conspicuous notch on ventral margin at articulation between merus and ischium. Female with unpaired left gonopore.
<figureCitationid="8ECB2A42022EFFC77402FE09FE0CFE63"box="[357,427,476,501]"captionStart="FIGURE 1"captionStartId="2.[151,250,1709,1731]"captionTargetBox="[167,1415,361,1640]"captionTargetId="figure@2.[159,1444,337,1679]"captionTargetPageId="2"captionText="FIGURE 1. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, shield (including carapace lateral lobes) and cephalic appendages, dorsal view; B, left antennal peduncle, ventral view; C, left maxilliped 3, lateral view; D, same, ischium, ventral view; E, left pereopod 4, lateral view; F, thoracic sternite 6, ventral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, outer view."httpUri="https://zenodo.org/record/801978/files/figure.png"pageId="4"pageNumber="239">Fig. 1</figureCitation>
E) weakly semichelate; dactylus terminating in large corneous claw; propodal rasp consisting of single row of several corneous scales. Pereopod 5 semichelate.
Male with sexual tube of medium length (3–4 times of coxal length) extending from coxa of right pereopod 5 (
<figureCitationid="8ECB2A42022EFFC775F8FD5AFF47FD3F"box="[159,224,655,680]"captionStart="FIGURE 1"captionStartId="2.[151,250,1709,1731]"captionTargetBox="[167,1415,361,1640]"captionTargetId="figure@2.[159,1444,337,1679]"captionTargetPageId="2"captionText="FIGURE 1. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, shield (including carapace lateral lobes) and cephalic appendages, dorsal view; B, left antennal peduncle, ventral view; C, left maxilliped 3, lateral view; D, same, ischium, ventral view; E, left pereopod 4, lateral view; F, thoracic sternite 6, ventral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, outer view."httpUri="https://zenodo.org/record/801978/files/figure.png"pageId="4"pageNumber="239">Fig. 1</figureCitation>
G), directed from right to left across ventral body surface and far overreaching lateral margin of left coxa, terminating in rounded tip; left coxa with very short, papilla-like sexual tube. Median lobe on thoracic sternite 8 (
<figureCitationid="8ECB2A42022EFFC775F8FD02FF79FD67"box="[159,222,727,752]"captionStart="FIGURE 1"captionStartId="2.[151,250,1709,1731]"captionTargetBox="[167,1415,361,1640]"captionTargetId="figure@2.[159,1444,337,1679]"captionTargetPageId="2"captionText="FIGURE 1. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, shield (including carapace lateral lobes) and cephalic appendages, dorsal view; B, left antennal peduncle, ventral view; C, left maxilliped 3, lateral view; D, same, ischium, ventral view; E, left pereopod 4, lateral view; F, thoracic sternite 6, ventral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, outer view."httpUri="https://zenodo.org/record/801978/files/figure.png"pageId="4"pageNumber="239">Fig. 1</figureCitation>
G) partially obscured by numerous, often curled, long setae.
<figureCitationid="8ECB2A42022EFFC77443FD29FEC1FC83"box="[292,358,764,789]"captionStart="FIGURE 1"captionStartId="2.[151,250,1709,1731]"captionTargetBox="[167,1415,361,1640]"captionTargetId="figure@2.[159,1444,337,1679]"captionTargetPageId="2"captionText="FIGURE 1. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, shield (including carapace lateral lobes) and cephalic appendages, dorsal view; B, left antennal peduncle, ventral view; C, left maxilliped 3, lateral view; D, same, ischium, ventral view; E, left pereopod 4, lateral view; F, thoracic sternite 6, ventral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, outer view."httpUri="https://zenodo.org/record/801978/files/figure.png"pageId="4"pageNumber="239">Fig. 1</figureCitation>
H) slightly longer than wide, narrowed posteriorly; no trace of lateral indentations; posterior lobes strongly asymmetrical, separated by deep median cleft; left terminal margin with 2 or 3 minute spines and laterally directed, acute outer angle; right terminal margin strongly oblique, with 1–3 or 2 minute spines and acute or subacute outer angle.
<emphasisid="2484EAD5022EFFC775A0FC59FE14FC32"bold="true"box="[199,435,908,933]"pageId="4"pageNumber="239">Colouration in life.</emphasis>
Carapace reddish. Ocular peduncles with red and white longitudinal stripes on light orange background. Antennular peduncle translucent yellowish or orangish. Antennal peduncle translucent. Chelipeds generally whitish or translucent, with scattered small red spots on chelae, carpi and meri. Ambulatory legs generally translucent, each with row of small red spots along dorsal and ventral margins of dactylus to merus.
, six specimens, four males and two females, were available for examination. They are generally very similar except for sexual differences. Like vast majority of the other species in
<taxonomicNameid="D1F04D44022EFFC775D3FBB1FEC0FBEB"authorityName="A. Milne-Edwards & Bouvier"authorityYear="1892"box="[180,359,1124,1148]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="4"pageNumber="239"phylum="Arthropoda"rank="genus">
, the propodus of the pereopod 2 of males is provided with a set of spiniform setae, consisting of one long spiniform setae flanked by two to four shorter spiniform or bristle-like setae, on the ventrodistal margin. In females, there is only one small spiniform seta on the ventrodistal margin of the pereopod 2 propodus. The right cheliped is much more elongate in males than in females; spines on the palm and carpus are more numerous and smaller in males than in females (for example, dorsomesial spines on the palm are seven to nine in males, five or six in females).
<materialsCitationid="A6983C9A022EFFC775A0FA72FDE1F8DA"ID-GBIF-Occurrence="1563183613"collectorName="de Saint Laurent & De Saint Laurent & Although de Saint Laurent"country="French Polynesia"county="Although de Saint Laurent"elevation="40"location="Mururoan"municipality="Only"pageId="4"pageNumber="239"specimenCount="40"stateProvince="Kumamoto Prefecture"typeStatus="holotype">
was originally described on the basis of a single male specimen by
<bibRefCitationid="72614B36022EFFC7703EFA72FF46FA73"author="Melin"pageId="4"pageNumber="239"refString="Melin, G. (1939) Paguriden und Galatheiden von Prof. Dr. Sixten Bocks Expedition nach den Bonin-Inseln 1914. Kongliga Svenska Uetenskapsakademiens Handlinger, Series 3, 18, 1 - 119."type="journal article"year="1939">Melin (1939)</bibRefCitation>
collected from the Ogasawara (= Bonin) Islands [as
<taxonomicNameid="D1F04D44022EFFC7774DFA25FD79F99F"authorityName="A. Milne-Edwards & Bouvier"authorityYear="1892"box="[554,734,1520,1544]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="4"pageNumber="239"phylum="Arthropoda"rank="genus">
<bibRefCitationid="72614B36022EFFC771C0FA3AFAE2F99F"box="[1191,1349,1519,1544]"pageId="4"pageNumber="239"refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne- Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum National d'Histoire Naturelle, Paris, Series 2, 39, 923 - 954, 1100 - 1119. [dated 1967, published 1968]"type="journal article">Saint Laurent</bibRefCitation>
</collectorName>
(1968) transferred the Melin’s taxon to
<taxonomicNameid="D1F04D44022EFFC77498F9C1FD14F9BB"authorityName="A. Milne-Edwards & Bouvier"authorityYear="1892"box="[511,691,1556,1580]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="4"pageNumber="239"phylum="Arthropoda"rank="genus">
<bibRefCitationid="72614B36022EFFC771B7F9C1FA3BF9BA"author="Edmondson"box="[1232,1436,1556,1581]"pageId="4"pageNumber="239"refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20, 185 - 243."type="journal article"year="1951">
<bibRefCitationid="72614B36022EFFC77749F9E2FD6FF9C7"box="[558,712,1591,1616]"pageId="4"pageNumber="239"refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne- Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum National d'Histoire Naturelle, Paris, Series 2, 39, 923 - 954, 1100 - 1119. [dated 1967, published 1968]"type="journal article">Saint Laurent</bibRefCitation>
<locationid="132F601C022EFFC771DDF970FA8DF92B"LSID="urn:lsid:plazi:treatment:9E5987D1022BFFCB75F0FC7CFCF0F9BB:132F601C022EFFC771DDF970FA8DF92B"box="[1210,1322,1701,1724]"country="French Polynesia"county="Although de Saint Laurent"municipality="Only"name="Mururoan"pageId="4"pageNumber="239"stateProvince="Kumamoto Prefecture">Mururoan</location>
<bibRefCitationid="72614B36022EFFC774FDF912FD90F977"box="[410,567,1735,1760]"pageId="4"pageNumber="239"refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne- Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum National d'Histoire Naturelle, Paris, Series 2, 39, 923 - 954, 1100 - 1119. [dated 1967, published 1968]"type="journal article">Saint Laurent</bibRefCitation>
</collectorName>
(1968).
<bibRefCitationid="72614B36022EFFC777C6F912FCEAF977"author="Miyake"box="[673,845,1735,1760]"pageId="4"pageNumber="239"refString="Miyake, S. (1978) The Crustacean Anomura of Sagami Bay. Hoikusha, Tokyo, 200 pp. (English text) + 161 pp. (Japanese text), 4 pls."type="book"year="1978">
<locationid="132F601C022EFFC777C6F912FD51F977"LSID="urn:lsid:plazi:treatment:9E5987D1022BFFCB75F0FC7CFCF0F9BB:132F601C022EFFC777C6F912FD51F977"box="[673,758,1735,1760]"country="French Polynesia"county="Although de Saint Laurent"municipality="Only"name="Miyake"pageId="4"pageNumber="239"stateProvince="Kumamoto Prefecture">Miyake</location>
<locationid="132F601C022EFFC774E3F938FE4FF893"LSID="urn:lsid:plazi:treatment:9E5987D1022BFFCB75F0FC7CFCF0F9BB:132F601C022EFFC774E3F938FE4FF893"box="[388,488,1773,1796]"country="French Polynesia"county="Although de Saint Laurent"municipality="Only"name="Japanese"pageId="4"pageNumber="239"stateProvince="Kumamoto Prefecture">Japanese</location>
main islands (
<locationid="132F601C022EFFC777F2F939FC85F892"LSID="urn:lsid:plazi:treatment:9E5987D1022BFFCB75F0FC7CFCF0F9BB:132F601C022EFFC777F2F939FC85F892"box="[661,802,1772,1797]"country="French Polynesia"county="Although de Saint Laurent"municipality="Only"name="Sagami Bay"pageId="4"pageNumber="239"stateProvince="Kumamoto Prefecture">Sagami Bay</location>
<bibRefCitationid="72614B36022EFFC77196F939FEE6F8BF"author="McLaughlin"pageId="4"pageNumber="239"refString="McLaughlin, P. A. & Pittman, C. (2002) Reinstatement and rediagnosis of Catapaguroides setosus and description of a second Hawaiian species of the genus (Decapoda: Anomura: Paguridea: Paguridae). Pacific Science, 56, 41 - 48. https: // doi. org / 10.1353 / psc. 2002.0006"type="journal article"year="2002">
<locationid="132F601C022EFFC77196F939FADBF892"LSID="urn:lsid:plazi:treatment:9E5987D1022BFFCB75F0FC7CFCF0F9BB:132F601C022EFFC77196F939FADBF892"box="[1265,1404,1772,1797]"country="French Polynesia"county="Although de Saint Laurent"municipality="Only"name="McLaughlin"pageId="4"pageNumber="239"stateProvince="Kumamoto Prefecture">McLaughlin</location>
&
<locationid="132F601C022EFFC775F0F8DAFF49F8BF"LSID="urn:lsid:plazi:treatment:9E5987D1022BFFCB75F0FC7CFCF0F9BB:132F601C022EFFC775F0F8DAFF49F8BF"box="[151,238,1807,1832]"country="French Polynesia"county="Although de Saint Laurent"municipality="Only"name="Pittman"pageId="4"pageNumber="239"stateProvince="Kumamoto Prefecture">Pittman</location>
(2002)
</bibRefCitation>
clarified that
<locationid="132F601C022EFFC77487F8DAFDDFF8BF"LSID="urn:lsid:plazi:treatment:9E5987D1022BFFCB75F0FC7CFCF0F9BB:132F601C022EFFC77487F8DAFDDFF8BF"box="[480,632,1807,1832]"country="French Polynesia"county="Although de Saint Laurent"municipality="Only"name="Edmondson's"pageId="4"pageNumber="239"stateProvince="Kumamoto Prefecture">Edmondson’s</location>
(1951)
<taxonomicNameid="D1F04D44022EFFC777B4F8C5FC78F8BF"authorityName="McLaughlin & Pittman"authorityYear="2002"box="[723,991,1808,1832]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="4"pageNumber="239"phylum="Arthropoda"rank="species"species="setosus">
to show morphological differences in this appendage in discriminating
<taxonomicNameid="D1F04D44022EFFC7713BF88DFB75F8E7"authorityName="McLaughlin & Pittman"authorityYear="2002"box="[1116,1234,1880,1904]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="4"pageNumber="239"phylum="Arthropoda"rank="species"species="setosus">
<bibRefCitationid="72614B36022EFFC775F0F8A9FE12F802"author="Komai"box="[151,437,1916,1941]"pageId="4"pageNumber="239"refString="Komai, T. & Takeda, M. (2006) A review of the pagurid hermit crab (Decapoda: Anomura: Paguroidea) fauna of the Sagami Sea, Central Japan. Memoirs of the National Science Museum, Tokyo, 41, 71 - 144."type="journal article"year="2006">
<bibRefCitationid="72614B36022EFFC7717EF8A9FB63F803"author="Miyake"box="[1049,1220,1916,1941]"pageId="4"pageNumber="239"refString="Miyake, S. (1978) The Crustacean Anomura of Sagami Bay. Hoikusha, Tokyo, 200 pp. (English text) + 161 pp. (Japanese text), 4 pls."type="book"year="1978">
<taxonomicNameid="D1F04D44022EFFC774A5F874FCF0F82E"authority="de Saint Laurent, 1968"authorityName="de Saint Laurent"authorityYear="1968"box="[450,855,1952,1977]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="4"pageNumber="239"phylum="Arthropoda"rank="species"species="japonicus">
<bibRefCitationid="72614B36022EFFC77710F875FCF0F82E"author="Saint"box="[631,855,1952,1977]"pageId="4"pageNumber="239"refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne- Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum National d'Histoire Naturelle, Paris, Series 2, 39, 923 - 954, 1100 - 1119. [dated 1967, published 1968]"type="journal article"year="1968">Saint Laurent, 1968</bibRefCitation>
<bibRefCitationid="72614B36022EFFC77158F811FAD6F84A"author="Miyake"box="[1087,1393,1988,2013]"pageId="4"pageNumber="239"refString="Miyake, S. (1978) The Crustacean Anomura of Sagami Bay. Hoikusha, Tokyo, 200 pp. (English text) + 161 pp. (Japanese text), 4 pls."type="book"year="1978">
<taxonomicNameid="D1F04D44022FFFC67424FF4DFE76FF38"authorityName="de Saint Laurent"authorityYear="1968"box="[323,465,152,175]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="5"pageNumber="240"phylum="Arthropoda"rank="species"species="japonicus">
, particularly in the non-inflated corneas and the lack of scattered small spines on the dorsal surface of the right palm.
<bibRefCitationid="72614B36022FFFC6777FFF69FC8BFF42"author="Okuno"box="[536,812,188,213]"pageId="5"pageNumber="240"refString="Okuno, J. & Arima, H. (2006) Additional records of inshore hermit crabs (Crustacea, Decapoda, Anomura) from Izu-ohshima Island, the northern Izu Islands, Japan. Bulletin of the Biogeographical Society of Japan, 61, 29 - 43. [in Japanese with English abstract]"type="journal article"year="2006">
, but neither provided detailed morphological description.
<bibRefCitationid="72614B36022FFFC676E5FF0AFB07FF6F"author="Komai"box="[898,1184,223,248]"pageId="5"pageNumber="240"refString="Komai, T. & Rahayu, D. L. (2013) Species of the pagurid hermit crab genus Catapaguroides A. Milne-Edwards & Bouvier, 1892 (Crustacea: Decapoda: Anomura) from the Bohol Sea, the Philippines, with descriptions of eight new species. Raffles Bulletin of Zoology, 61, 143 - 188."type="journal article"year="2013">
<taxonomicNameid="D1F04D44022FFFC670E5FED1FD9CFED7"authority="Komai & Rahayu, 2013"authorityName="Komai & Rahayu"authorityYear="2013"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="5"pageNumber="240"phylum="Arthropoda"rank="species"species="crassimanus">
<bibRefCitationid="72614B36022FFFC6744DFEF2FD9CFED7"author="Komai"box="[298,571,295,320]"pageId="5"pageNumber="240"refString="Komai, T. & Rahayu, D. L. (2013) Species of the pagurid hermit crab genus Catapaguroides A. Milne-Edwards & Bouvier, 1892 (Crustacea: Decapoda: Anomura) from the Bohol Sea, the Philippines, with descriptions of eight new species. Raffles Bulletin of Zoology, 61, 143 - 188."type="journal article"year="2013">Komai & Rahayu, 2013</bibRefCitation>
<taxonomicNameid="D1F04D44022FFFC676D4FEFDFBC1FED7"authorityName="A. Milne-Edwards & Bouvier"authorityYear="1892"box="[947,1126,296,320]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="5"pageNumber="240"phylum="Arthropoda"rank="genus">
<bibRefCitationid="72614B36022FFFC6747BFE99FD84FEF3"author="Komai"box="[284,547,332,357]"pageId="5"pageNumber="240"refString="Komai, T. & Rahayu, D. L. (2013) Species of the pagurid hermit crab genus Catapaguroides A. Milne-Edwards & Bouvier, 1892 (Crustacea: Decapoda: Anomura) from the Bohol Sea, the Philippines, with descriptions of eight new species. Raffles Bulletin of Zoology, 61, 143 - 188."type="journal article"year="2013">
, male (sl 1.54 mm), CBM-ZC 13594. A, right pereopod 2, lateral view; B, left pereopod 3, lateral view; C, D, dactyli of right pereopod 2 and left pereopod 3, mesial view.
<bibRefCitationid="72614B36022DFFC47702F878FC19F851"author="McLaughlin"box="[613,958,1965,1990]"pageId="7"pageNumber="242"refString="McLaughlin, P. A. & Pittman, C. (2002) Reinstatement and rediagnosis of Catapaguroides setosus and description of a second Hawaiian species of the genus (Decapoda: Anomura: Paguridea: Paguridae). Pacific Science, 56, 41 - 48. https: // doi. org / 10.1353 / psc. 2002.0006"type="journal article"year="2002">McLaughlin & Pittman (2002)</bibRefCitation>
<materialsCitationid="A6983C9A0222FFCB7043FF69FD4DFC82"ID-GBIF-Occurrence="1563183619"collectionCode="SMNH"collectorName="de Saint Laurent & E. D. Akerman & McLaughlin & Pittman & However & The & As & Melin's & Melin & Japanese & De Saint Laurent"country="Sweden"location="Stockholm"pageId="8"pageNumber="243"specimenCount="5"stateProvince="Stockholm"typeStatus="holotype">
<collectionCodeid="70E1AE020222FFCB75F9FF35FF56FF6F"box="[158,241,224,248]"country="Canada"httpUri="http://grbio.org/cool/8z6v-f8is"name="Saskatchewan Museum of Natural History"pageId="8"pageNumber="243">SMNH</collectionCode>
Type-291), corresponding to the “mutilated” specimen examined by
<bibRefCitationid="72614B360222FFCB7144FF0AFB1CFF60"box="[1059,1211,223,248]"pageId="8"pageNumber="243"refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne- Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum National d'Histoire Naturelle, Paris, Series 2, 39, 923 - 954, 1100 - 1119. [dated 1967, published 1968]"type="journal article">Saint Laurent</bibRefCitation>
</collectorName>
(1968). During this study, I examined a color image of the body and left cheliped of the
<collectorNameid="BB0553110222FFCB71B2FED0FAD9FE8A"box="[1237,1406,260,285]"pageId="8"pageNumber="243">E. D. Akerman</collectorName>
of
<collectionCodeid="70E1AE020222FFCB75F0FEFDFF4BFED7"box="[151,236,296,320]"country="Canada"httpUri="http://grbio.org/cool/8z6v-f8is"name="Saskatchewan Museum of Natural History"pageId="8"pageNumber="243">SMNH</collectionCode>
. I confirmed that the additional specimens examined in this study agree well with the
in the present condition, including the small, non-inflated corneas, the presence of multiple mid-dorsal spines on the left cheliped carpus and the relatively short right sexual tube without an elongate, anteriorly directed distal portion.
<bibRefCitationid="72614B360222FFCB77C9FE41FBAAFE3B"author="McLaughlin"box="[686,1037,404,429]"pageId="8"pageNumber="243"refString="McLaughlin, P. A. & Pittman, C. (2002) Reinstatement and rediagnosis of Catapaguroides setosus and description of a second Hawaiian species of the genus (Decapoda: Anomura: Paguridea: Paguridae). Pacific Science, 56, 41 - 48. https: // doi. org / 10.1353 / psc. 2002.0006"type="journal article"year="2002">
<bibRefCitationid="72614B360222FFCB75F9FE09FDC9FE63"author="McLaughlin"box="[158,622,476,501]"pageId="8"pageNumber="243"refString="McLaughlin, P. A. & Pittman, C. (2002) Reinstatement and rediagnosis of Catapaguroides setosus and description of a second Hawaiian species of the genus (Decapoda: Anomura: Paguridea: Paguridae). Pacific Science, 56, 41 - 48. https: // doi. org / 10.1353 / psc. 2002.0006"type="journal article"year="2002">
McLaughlin & Pittman 2002:
<figureCitationid="8ECB2A420222FFCB749AFE09FDFEFE63"box="[509,601,476,501]"captionStart="FIGURE 1"captionStartId="2.[151,250,1709,1731]"captionTargetBox="[167,1415,361,1640]"captionTargetId="figure@2.[159,1444,337,1679]"captionTargetPageId="2"captionText="FIGURE 1. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, shield (including carapace lateral lobes) and cephalic appendages, dorsal view; B, left antennal peduncle, ventral view; C, left maxilliped 3, lateral view; D, same, ischium, ventral view; E, left pereopod 4, lateral view; F, thoracic sternite 6, ventral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, outer view."httpUri="https://zenodo.org/record/801978/files/figure.png"pageId="8"pageNumber="243">Figure 1</figureCitation>
E
</bibRefCitation>
) shows the dactylus of the left cheliped being about half-length of the palm.
<bibRefCitationid="72614B360222FFCB7402FDF1FE5AFDAB"box="[357,509,548,573]"pageId="8"pageNumber="243"refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne- Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum National d'Histoire Naturelle, Paris, Series 2, 39, 923 - 954, 1100 - 1119. [dated 1967, published 1968]"type="journal article">Saint Laurent</bibRefCitation>
</collectorName>
(1968), a discrepancy is seen in the length of the dactylus of pereopod 2 between
(1939) original description and the present specimens.
<bibRefCitationid="72614B360222FFCB763BFD92FC57FDF7"author="Melin"box="[860,1008,583,608]"pageId="8"pageNumber="243"refString="Melin, G. (1939) Paguriden und Galatheiden von Prof. Dr. Sixten Bocks Expedition nach den Bonin-Inseln 1914. Kongliga Svenska Uetenskapsakademiens Handlinger, Series 3, 18, 1 - 119."type="journal article"year="1939">
far as I aware, there is no other congeneric species having the dactylus of the pereopod 2 being subequal in the length to the propodus, and thus the observation by
<bibRefCitationid="72614B360222FFCB7420FD02FE7AFD67"author="Melin"box="[327,477,727,752]"pageId="8"pageNumber="243"refString="Melin, G. (1939) Paguriden und Galatheiden von Prof. Dr. Sixten Bocks Expedition nach den Bonin-Inseln 1914. Kongliga Svenska Uetenskapsakademiens Handlinger, Series 3, 18, 1 - 119."type="journal article"year="1939">
<bibRefCitationid="72614B360222FFCB77AFFD02FCC6FD78"box="[712,865,727,752]"pageId="8"pageNumber="243"refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne- Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum National d'Histoire Naturelle, Paris, Series 2, 39, 923 - 954, 1100 - 1119. [dated 1967, published 1968]"type="journal article">Saint Laurent</bibRefCitation>
<taxonomicNameid="D1F04D440222FFCB779FFCF5FCCDFCAF"authorityName="McLaughlin & Pittman"authorityYear="2002"box="[760,874,800,824]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="8"pageNumber="243"phylum="Arthropoda"rank="species"species="setosus">
are now assessed more precisely. The structure of the right chela is quite different between the two species. For example, the palm is non-operculiform with dorsomesial margin of the dactylus and dorsolateral margin of the palm being all rounded in
A– C); the dorsomesial margin of the dactylus bears a small proximal spine. In contrast, the right chela in
<taxonomicNameid="D1F04D440222FFCB7076FC59FA25FC33"authorityName="McLaughlin & Pittman"authorityYear="2002"box="[1297,1410,908,932]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="8"pageNumber="243"phylum="Arthropoda"rank="species"species="setosus">
is suboperculiform and entirely arched; the dorsomesial margin of the dactylus has a prominent thin ridge, but spineless; the dorsolateral margin is bordered with a slightly elevated ridge extending to the tip of the fixed finger (
<bibRefCitationid="72614B360222FFCB75F9FC22FDDDFB87"author="McLaughlin"box="[158,634,1015,1040]"pageId="8"pageNumber="243"refString="McLaughlin, P. A. & Pittman, C. (2002) Reinstatement and rediagnosis of Catapaguroides setosus and description of a second Hawaiian species of the genus (Decapoda: Anomura: Paguridea: Paguridae). Pacific Science, 56, 41 - 48. https: // doi. org / 10.1353 / psc. 2002.0006"type="journal article"year="2002">McLaughlin & Pittman 2002: 42, fig. 1A</bibRefCitation>
). The carpus of the right cheliped is armed with a longitudinal row of four to nine spines or tubercles on the dorsal surface in
A, B, D), whereas there are only three very small, blunt tubercles in
<taxonomicNameid="D1F04D440222FFCB74D3FB95FD8FFBCF"authorityName="McLaughlin & Pittman"authorityYear="2002"box="[436,552,1088,1112]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="8"pageNumber="243"phylum="Arthropoda"rank="species"species="setosus">
<bibRefCitationid="72614B360222FFCB7707FBEAFC17FBCF"author="McLaughlin"box="[608,944,1087,1112]"pageId="8"pageNumber="243"refString="McLaughlin, P. A. & Pittman, C. (2002) Reinstatement and rediagnosis of Catapaguroides setosus and description of a second Hawaiian species of the genus (Decapoda: Anomura: Paguridea: Paguridae). Pacific Science, 56, 41 - 48. https: // doi. org / 10.1353 / psc. 2002.0006"type="journal article"year="2002">McLaughlin & Pittman 2002</bibRefCitation>
: 42). As shown by
<bibRefCitationid="72614B360222FFCB71F5FBEAFECBFBEB"author="McLaughlin"pageId="8"pageNumber="243"refString="McLaughlin, P. A. & Pittman, C. (2002) Reinstatement and rediagnosis of Catapaguroides setosus and description of a second Hawaiian species of the genus (Decapoda: Anomura: Paguridea: Paguridae). Pacific Science, 56, 41 - 48. https: // doi. org / 10.1353 / psc. 2002.0006"type="journal article"year="2002">McLaughlin & Pittman (2002: 42, fig. 1C)</bibRefCitation>
, the left chela is entirely arched in
<taxonomicNameid="D1F04D440222FFCB779DFBB1FCCBFBEB"authorityName="McLaughlin & Pittman"authorityYear="2002"box="[762,876,1124,1148]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="8"pageNumber="243"phylum="Arthropoda"rank="species"species="setosus">
<figureCitationid="8ECB2A420222FFCB7759FB79FDD8FB53"box="[574,639,1196,1221]"captionStart="FIGURE 1"captionStartId="2.[151,250,1709,1731]"captionTargetBox="[167,1415,361,1640]"captionTargetId="figure@2.[159,1444,337,1679]"captionTargetPageId="2"captionText="FIGURE 1. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, shield (including carapace lateral lobes) and cephalic appendages, dorsal view; B, left antennal peduncle, ventral view; C, left maxilliped 3, lateral view; D, same, ischium, ventral view; E, left pereopod 4, lateral view; F, thoracic sternite 6, ventral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, telson, outer view."httpUri="https://zenodo.org/record/801978/files/figure.png"pageId="8"pageNumber="243">Fig. 1</figureCitation>
A), rather than extending beyond them in
<taxonomicNameid="D1F04D440222FFCB710CFB79FB78FB53"authorityName="McLaughlin & Pittman"authorityYear="2002"box="[1131,1247,1196,1220]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="8"pageNumber="243"phylum="Arthropoda"rank="species"species="setosus">
<bibRefCitationid="72614B360222FFCB7197FB79FE18FB7F"author="McLaughlin"pageId="8"pageNumber="243"refString="McLaughlin, P. A. & Pittman, C. (2002) Reinstatement and rediagnosis of Catapaguroides setosus and description of a second Hawaiian species of the genus (Decapoda: Anomura: Paguridea: Paguridae). Pacific Science, 56, 41 - 48. https: // doi. org / 10.1353 / psc. 2002.0006"type="journal article"year="2002">McLaughlin & Pittman 2002: 42, fig. 1A</bibRefCitation>
). The antennular peduncle overreaches the distal corneal margin by the approximately 0.8 length of the ultimate segment in
<taxonomicNameid="D1F04D440222FFCB76ACFB21FB9AFA9B"authorityName="McLaughlin & Pittman"authorityYear="2002"box="[971,1085,1268,1292]"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="8"pageNumber="243"phylum="Arthropoda"rank="species"species="setosus">
<bibRefCitationid="72614B360222FFCB712BFB21FEBEFAA7"author="McLaughlin"pageId="8"pageNumber="243"refString="McLaughlin, P. A. & Pittman, C. (2002) Reinstatement and rediagnosis of Catapaguroides setosus and description of a second Hawaiian species of the genus (Decapoda: Anomura: Paguridea: Paguridae). Pacific Science, 56, 41 - 48. https: // doi. org / 10.1353 / psc. 2002.0006"type="journal article"year="2002">McLaughlin & Pittman 2002: 42, fig. 1A</bibRefCitation>
). The dactyli of the pereopods 3 bear four to six spiniform setae on the mesial face adjacent to the ventral margin in
<figureCitationid="8ECB2A420222FFCB74BAFAE9FD86FAC3"box="[477,545,1340,1365]"captionStart="FIGURE 4"captionStartId="7.[151,250,1873,1895]"captionTargetBox="[204,1377,231,1824]"captionTargetId="figure@7.[149,1434,193,1840]"captionTargetPageId="7"captionText="FIGURE 4. Catapaguroides fragilis (Melin, 1939), male (sl 1.54 mm), CBM-ZC 13594. A, right pereopod 2, lateral view; B, left pereopod 3, lateral view; C, D, dactyli of right pereopod 2 and left pereopod 3, mesial view."httpUri="https://zenodo.org/record/801984/files/figure.png"pageId="8"pageNumber="243">Fig. 4</figureCitation>
C, D), whereas apparently unarmed in
<taxonomicNameid="D1F04D440222FFCB76A8FAE9FF67FAEF"authority="McLaughlin & Pittman 2002: 42"authorityName="McLaughlin & Pittman"authorityPageNumber="42"authorityYear="2002"class="Malacostraca"family="Paguridae"genus="Catapaguroides"kingdom="Animalia"order="Decapoda"pageId="8"pageNumber="243"phylum="Arthropoda"rank="species"species="setosus">
<bibRefCitationid="72614B360222FFCB7128FAE9FF10FAEF"author="McLaughlin"pageId="8"pageNumber="243"refString="McLaughlin, P. A. & Pittman, C. (2002) Reinstatement and rediagnosis of Catapaguroides setosus and description of a second Hawaiian species of the genus (Decapoda: Anomura: Paguridea: Paguridae). Pacific Science, 56, 41 - 48. https: // doi. org / 10.1353 / psc. 2002.0006"type="journal article"year="2002">McLaughlin & Pittman 2002: 42</bibRefCitation>
The right cheliped of the Mururoa female specimen, depicted by de
<bibRefCitationid="72614B360222FFCB76AEFA51FB08FA0A"author="Saint"box="[969,1199,1412,1437]"pageId="8"pageNumber="243"refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne- Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum National d'Histoire Naturelle, Paris, Series 2, 39, 923 - 954, 1100 - 1119. [dated 1967, published 1968]"type="journal article"year="1968">
Saint
<bibRefCitationid="72614B360222FFCB716CFA50FB08FA0A"author="Laurent"box="[1035,1199,1412,1437]"pageId="8"pageNumber="243"refString="Saint Laurent, M. de (1968) Revision des genres Catapaguroides et Cestopagurus et description de quatre genres nouveaux. I. Catapaguroides A. Milne- Edwards et Bouvier et Decaphyllus nov. gen. (Crustaces Decapodes Paguridae). Bulletin du Museum National d'Histoire Naturelle, Paris, Series 2, 39, 923 - 954, 1100 - 1119. [dated 1967, published 1968]"type="journal article"year="1968">Laurent (1968</bibRefCitation>
</bibRefCitation>
: fig. 26), agrees well with those of the present Japanese female specimens of
in the general shape and armature. De Saint Laurent specifically noted the number of spines on the chela and carpus of the left cheliped (one spine on the palm and two spines on the carpus), which could be within range of intraspecific variation. I concur de Saint Laurent’s (1968) identification of the Mururoa specimen with
