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<document id="9D52DB1B70D7A186CE4696F55DBFF66F" ID-CLB-Dataset="22249" ID-DOI="10.5852/ejt.2013.48" ID-GBIF-Dataset="02866e62-ab61-4cf4-9d98-0248e251084e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3822836" ID-ZooBank="E7080722-E348-448D-96E5-D537F4865BB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589219740188" checkinUser="carolina" docAuthor="Thuy, Ben" docDate="2013" docId="99789763658085ADD35D2786FCF39B9A" docLanguage="en" docName="ejt-2013-48.pdf.imf" docOrigin="European Journal of Taxonomy 48" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophiojagtus Thuy 2013, gen. nov." docType="treatment" docUuid="0B5FB6EE-42F6-43D5-BC67-41AE35B82EE3" docUuidSource="ZooBank" docVersion="10" lastPageNumber="215" masterDocId="6541EF1B6555857BD1302529FFF89F78" masterDocTitle="Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)" masterLastPageNumber="242" masterPageNumber="1" pageNumber="214" updateTime="1698839159859" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C64AAF4EE7150F29D5E8E9EE8837D79E">Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)</mods:title>
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<mods:roleTerm id="C3A5723716209F008DD796D3CBF3734A">Author</mods:roleTerm>
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<mods:namePart id="550EA6CF8468ED4F83AAC12FB70C84F6">Thuy, Ben</mods:namePart>
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<mods:title id="623C7305D79C8782E14C5CA21C708184">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="29A4C18EBD8F290BD4F3996351D3B54B">
<mods:date id="D24B6DC84B507F7DA06758302B61D7CB">2013</mods:date>
<mods:detail id="FD9D0277CED47AF390E2C9A0B37D68D3" type="pubDate">
<mods:number id="23A1C7E09A22EE6B84619296EB835546">2013-07-02</mods:number>
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<mods:detail id="7F4231AC784C40B14118547FA91D1D0F" type="issue">
<mods:number id="44E02513D0FA9900BF66B144D89141A3">48</mods:number>
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<mods:classification id="9000C04FA87B150BD79A2E24963E825C">journal article</mods:classification>
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<mods:identifier id="E92F94CDD0B3636F9F1826CCCC20E0C5" type="DOI">10.5852/ejt.2013.48</mods:identifier>
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<treatment id="99789763658085ADD35D2786FCF39B9A" ID-DOI="http://doi.org/10.5281/zenodo.3844198" ID-GBIF-Taxon="164044812" ID-Zenodo-Dep="3844198" LSID="urn:lsid:zoobank.org:act:0B5FB6EE-42F6-43D5-BC67-41AE35B82EE3" httpUri="http://treatment.plazi.org/id/99789763658085ADD35D2786FCF39B9A" lastPageId="214" lastPageNumber="215" pageId="213" pageNumber="214">
<subSubSection id="59CB75FE658085AED35D2786FC3E9DB1" box="[621,966,686,713]" pageId="213" pageNumber="214" type="nomenclature">
<paragraph id="116E2675658085AED35D2786FC3E9DB1" blockId="213.[364,1224,686,749]" box="[621,966,686,713]" pageId="213" pageNumber="214">
<heading id="4A269119658085AED35D2786FC3E9DB1" box="[621,966,686,713]" centered="true" fontSize="11" level="2" pageId="213" pageNumber="214" reason="2">
Genus
<taxonomicName id="D6D15DF6658085AED3F02787FCAC9DB1" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[704,852,686,713]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="213" pageNumber="214" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA67658085AED3F02787FCAC9DB1" bold="true" box="[704,852,686,713]" italics="true" pageId="213" pageNumber="214">Ophiojagtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658085AED26B2786FC3E9DB1" box="[859,966,687,713]" pageId="213" pageNumber="214" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658085AED05C27FBFB309D95" box="[364,1224,722,749]" pageId="213" pageNumber="214" type="description">
<paragraph id="116E2675658085AED05C27FBFB309D95" blockId="213.[364,1224,686,749]" box="[364,1224,722,749]" pageId="213" pageNumber="214">
<uri id="65402A77658085AED05C27FBFB309D95" box="[364,1224,722,749]" pageId="213" pageNumber="214">
urn:lsid:zoobank.org:act:
<uuid id="65771CA0658085AED3BF27FBFB309D95" box="[655,1224,722,749]" pageId="213" pageNumber="214">0B5FB6EE-42F6-43D5-BC67-41AE35B82EE3</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658085AED18D263CFCD89C26" pageId="213" pageNumber="214" type="type_taxon">
<paragraph id="116E2675658085AED18D263CFEA09C56" blockId="213.[189,344,788,815]" box="[189,344,788,815]" pageId="213" pageNumber="214">
<heading id="4A269119658085AED18D263CFEA09C56" bold="true" box="[189,344,788,815]" fontSize="11" level="3" pageId="213" pageNumber="214" reason="3">
<emphasis id="23A5FA67658085AED18D263CFEA09C56" bold="true" box="[189,344,788,815]" pageId="213" pageNumber="214">
<typeStatus id="CE6A98D7658085AED18D263CFF029C57" box="[189,250,789,815]" pageId="213" pageNumber="214">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658085AED18D266DFCD89C26" blockId="213.[189,800,836,862]" box="[189,800,836,862]" pageId="213" pageNumber="214">
<taxonomicName id="D6D15DF6658085AED18D266DFE5E9C26" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[189,422,836,862]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="213" pageNumber="214" phylum="Echinodermata" rank="species" species="acklesi" status="sp. nov.">
<emphasis id="23A5FA67658085AED18D266DFE5E9C26" box="[189,422,836,862]" italics="true" pageId="213" pageNumber="214">Ophiojagtus acklesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658085AED09E266DFDF19C26" box="[430,521,836,862]" pageId="213" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
by present designation.
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658085AED18D26AFFE359C8B" pageId="213" pageNumber="214" type="discussion">
<paragraph id="116E2675658085AED18D26AFFE209CD8" blockId="213.[189,472,902,928]" box="[189,472,902,928]" pageId="213" pageNumber="214">
<heading id="4A269119658085AED18D26AFFE209CD8" bold="true" box="[189,472,902,928]" fontSize="11" level="3" pageId="213" pageNumber="214" reason="3">
<emphasis id="23A5FA67658085AED18D26AFFE209CD8" bold="true" box="[189,472,902,928]" pageId="213" pageNumber="214">Other species included</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658085AED18D269CFE359C8B" blockId="213.[189,1398,949,1011]" pageId="213" pageNumber="214">
<taxonomicName id="D6D15DF6658085AED18D269CFD759CB7" authority="Hess, 1966" authorityName="von Martens" authorityYear="1867" box="[189,653,949,976]" class="Ophiuroidea" family="Hemieuryalidae" genus="Hemieuryale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="213" pageNumber="214" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658085AED18D269CFEAC9CB7" box="[189,340,949,975]" italics="true" pageId="213" pageNumber="214">Hemieuryale</emphasis>
?
<emphasis id="23A5FA67658085AED05E269FFE0C9CB7" box="[366,500,950,975]" italics="true" pageId="213" pageNumber="214">argoviensis</emphasis>
<bibRefCitation id="75405B84658085AED332269FFD759CB7" author="Hess H." box="[514,653,949,976]" pageId="213" pageNumber="214" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess, 1966</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="D6D15DF6658085AED3AF269CFC679CB7" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[671,927,949,975]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="213" pageNumber="214" phylum="Echinodermata" rank="species" species="irimurai" status="sp. nov.">
<emphasis id="23A5FA67658085AED3AF269CFC679CB7" box="[671,927,949,975]" italics="true" pageId="213" pageNumber="214">Ophiojagtus irimurai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658085AED29B269FFBF49CA8" box="[939,1036,950,976]" pageId="213" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="D6D15DF6658085AED560269CFE309C8B" authority="Kutscher &amp; Jagt, 2000" authorityName="Matsumoto" authorityYear="1915" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiosmilax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="213" pageNumber="214" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658085AED560269CFB1E9CB7" box="[1104,1254,949,975]" italics="true" pageId="213" pageNumber="214">Ophiosmilax</emphasis>
?
<emphasis id="23A5FA67658085AED5CF269CFA8E9CB7" box="[1279,1398,949,975]" italics="true" pageId="213" pageNumber="214">alternatus</emphasis>
<bibRefCitation id="75405B84658085AED18D26F0FE309C8B" author="Kutscher M. &amp; Jagt J. W. M." box="[189,456,985,1011]" pageId="213" pageNumber="214" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt, 2000</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658085AED18D2132FA8F9A43" pageId="213" pageNumber="214" type="diagnosis">
<paragraph id="116E2675658085AED18D2132FECE9B4D" blockId="213.[189,310,1051,1077]" box="[189,310,1051,1077]" pageId="213" pageNumber="214">
<heading id="4A269119658085AED18D2132FECE9B4D" bold="true" box="[189,310,1051,1077]" fontSize="11" level="3" pageId="213" pageNumber="214" reason="3">
<emphasis id="23A5FA67658085AED18D2132FECE9B4D" bold="true" box="[189,310,1051,1077]" pageId="213" pageNumber="214">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658085AED18D2163FA8F9A43" blockId="213.[189,1399,1098,1340]" pageId="213" pageNumber="214">Ophiacanthid with stout, thick, strongly curved LAPs generally with a large height/width ratio; outer surface devoid of ornament elements; no spurs on outer proximal and inner distal edges; ventral portion of LAPs long to extremely long, strongly protruding ventralwards and often with widened ventral tip; proximal edge commonly with proximally pointing protrudion; large, ear-shaped spine articulations composed of thick, continuous volute, freestanding on bulging distal portion of LAP and not bordered proximally by a ridge-like structure; broad, well-defined ridge on the inner side of the LAPs, generally with dorso-proximally pointing dorsal part; tentacle notch large but poorly defined and generally shallow.</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658085AED18D204DFEC49A8C" pageId="213" pageNumber="214" type="etymology">
<paragraph id="116E2675658085AED18D204DFEBC9A06" blockId="213.[189,324,1380,1406]" box="[189,324,1380,1406]" pageId="213" pageNumber="214">
<heading id="4A269119658085AED18D204DFEBC9A06" bold="true" box="[189,324,1380,1406]" fontSize="11" level="3" pageId="213" pageNumber="214" reason="3">
<emphasis id="23A5FA67658085AED18D204DFEBC9A06" bold="true" box="[189,324,1380,1406]" pageId="213" pageNumber="214">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658085AED18D20BAFEC49A8C" blockId="213.[189,1399,1427,1524]" pageId="213" pageNumber="214">Genus named in honour of my friend and colleague John W.M. Jagt for his generous support in the course of the present project and for his outstanding contributions to ophiuroid palaeontology; gender masculine.</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658085ADD18D2334FCF39B9A" lastPageId="214" lastPageNumber="215" pageId="213" pageNumber="214" type="discussion">
<paragraph id="116E2675658085AED18D2334FED7994F" blockId="213.[189,303,1565,1591]" box="[189,303,1565,1591]" pageId="213" pageNumber="214">
<heading id="4A269119658085AED18D2334FED7994F" bold="true" box="[189,303,1565,1591]" fontSize="11" level="3" pageId="213" pageNumber="214" reason="3">
<emphasis id="23A5FA67658085AED18D2334FED7994F" bold="true" box="[189,303,1565,1591]" pageId="213" pageNumber="214">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658085AED18D2365FD6B9861" blockId="213.[189,1399,1612,1818]" pageId="213" pageNumber="214">
The Mesozoic ophiuroid fossil record includes a highly distinctive
<typeStatus id="CE6A98D7658085AED2882365FC13991E" box="[952,1003,1612,1638]" pageId="213" pageNumber="214">type</typeStatus>
of dissociated LAPs characterised by a thick and massive general aspect, very long, conspicuous ventral portion, large, ear-shaped spine articulations freestanding on the bulging distal portion of the LAP and composed of a thick, continuous volute, and the absence of outer surface ornament and spurs. This combination of characters is not found in any other ophiuroid LAP
<typeStatus id="CE6A98D7658085AED36423F5FD72998E" box="[596,650,1756,1782]" pageId="213" pageNumber="214">type</typeStatus>
. A new genus,
<taxonomicName id="D6D15DF6658085AED20B23F2FC34998D" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[827,972,1755,1781]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="213" pageNumber="214" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA67658085AED20B23F2FC34998D" box="[827,972,1755,1781]" italics="true" pageId="213" pageNumber="214">Ophiojagtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658085AED2E423F5FBB9998E" box="[980,1089,1756,1782]" pageId="213" pageNumber="214" rank="genus">gen. nov.</taxonomicNameLabel>
, is thus introduced here to accommodate the LAP
<typeStatus id="CE6A98D7658085AED0E02229FDFB9862" box="[464,515,1792,1818]" pageId="213" pageNumber="214">type</typeStatus>
in question.
</paragraph>
<paragraph id="116E2675658085ADD18D2268FE879D7D" blockId="213.[189,1399,1857,2027]" lastBlockId="214.[189,1399,267,518]" lastPageId="214" lastPageNumber="215" pageId="213" pageNumber="214">
Previous records which can be assigned to this new genus include
<taxonomicName id="D6D15DF6658085AED28E2268FBAD9823" authorityName="von Martens" authorityYear="1867" box="[958,1109,1857,1883]" class="Ophiuroidea" family="Hemieuryalidae" genus="Hemieuryale" kingdom="Animalia" order="Ophiurida" pageId="213" pageNumber="214" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658085AED28E2268FBAD9823" box="[958,1109,1857,1883]" italics="true" pageId="213" pageNumber="214">Hemieuryale</emphasis>
</taxonomicName>
?
<taxonomicName id="D6D15DF6658085AED559226BFA8F9823" authority="Hess, 1966" authorityName="Hess" authorityYear="1966" box="[1129,1399,1857,1884]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="213" pageNumber="214" phylum="Echinodermata" rank="species" species="argoviensis">
<emphasis id="23A5FA67658085AED559226BFB179823" box="[1129,1263,1858,1883]" italics="true" pageId="213" pageNumber="214">argoviensis</emphasis>
<bibRefCitation id="75405B84658085AED5C5226BFA8F9823" author="Hess H." box="[1269,1399,1857,1884]" pageId="213" pageNumber="214" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess, 1966</bibRefCitation>
</taxonomicName>
from the Oxfordian of
<collectingCountry id="69C666E5658085AED0FA224CFDAE9807" box="[458,598,1893,1919]" name="Switzerland" pageId="213" pageNumber="214">Switzerland</collectingCountry>
and
<collectingCountry id="69C666E5658085AED3A0224FFD1898F8" box="[656,736,1894,1920]" name="France" pageId="213" pageNumber="214">France</collectingCountry>
(
<bibRefCitation id="75405B84658085AED3C0224CFC8898F8" author="Hess H." box="[752,880,1893,1920]" pageId="213" pageNumber="214" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess 1966</bibRefCitation>
),
<taxonomicName id="D6D15DF6658085AED2B6224CFBE49807" authorityName="Matsumoto" authorityYear="1915" box="[902,1052,1893,1919]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiosmilax" kingdom="Animalia" order="Ophiurida" pageId="213" pageNumber="214" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658085AED2B6224CFBE49807" box="[902,1052,1893,1919]" italics="true" pageId="213" pageNumber="214">Ophiosmilax</emphasis>
</taxonomicName>
?
<taxonomicName id="D6D15DF6658085AED501224CFF0098DB" authority="Kutscher &amp; Jagt, 2000" authorityName="Kutscher &amp; Jagt, 2000 in Jagt" authorityYear="2000" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="213" pageNumber="214" phylum="Echinodermata" rank="species" species="alternatus">
<emphasis id="23A5FA67658085AED501224CFB509807" box="[1073,1192,1893,1919]" italics="true" pageId="213" pageNumber="214">alternatus</emphasis>
<bibRefCitation id="75405B84658085AED59F224CFF0098DB" author="Kutscher M. &amp; Jagt J. W. M." pageId="213" pageNumber="214" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt, 2000</bibRefCitation>
</taxonomicName>
from the Campanian and Maastrichtian of
<collectingCountry id="69C666E5658085AED23B22A0FC8198DB" box="[779,889,1929,1955]" name="Germany" pageId="213" pageNumber="214">Germany</collectingCountry>
,
<collectingCountry id="69C666E5658085AED2B922A0FC0E98DB" box="[905,1014,1929,1955]" name="Denmark" pageId="213" pageNumber="214">Denmark</collectingCountry>
and
<collectingCountry id="69C666E5658085AED50722A0FB6498DB" box="[1079,1180,1929,1955]" name="Belgium" pageId="213" pageNumber="214">Belgium</collectingCountry>
(
<bibRefCitation id="75405B84658085AED59E22A0FF0198BF" author="Kutscher M. &amp; Jagt J. W. M." pageId="213" pageNumber="214" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt 2000</bibRefCitation>
) and records of the latter form from the Turonian of the
<collectingCountry id="69C666E5658085AED2BC2284FBBE98BF" box="[908,1094,1965,1991]" name="Czech Republic" pageId="213" pageNumber="214">Czech Republic</collectingCountry>
and
<collectingCountry id="69C666E5658085AED54E2284FB2D98BF" box="[1150,1237,1965,1991]" name="Tunisia" pageId="213" pageNumber="214">Tunisia</collectingCountry>
(
<bibRefCitation id="75405B84658085AED5D42282FA9398BF" author="Storc R." box="[1252,1387,1963,1991]" pageId="213" pageNumber="214" pagination="395 - 401" refId="ref128214" refString="Storc R. 2004. The Late Cretaceous brittle star Ophiosmilax? alternatus from Europe and northern Africa. Geobios 37 (3): 395 - 401. http: // dx. doi. org / 10.1016 / j. geobios. 2003.01.010" type="journal article" year="2004">Štorc 2004</bibRefCitation>
). Astonishingly, the striking similarities between these two species have not been noted previously. As a consequence, they were assigned to completely different genera belonging to unrelated families (
<bibRefCitation id="75405B84658385ADD41F2422FEC69E33" author="Smith A. B. &amp; Paterson G. L. J. &amp; Lafay B." pageId="214" pageNumber="215" pagination="213 - 243" refId="ref127806" refString="Smith A. B., Paterson G. L. J. &amp; Lafay B. 1995. Ophiuroid phylogeny and higher taxonomy: morphological, molecular and palaeontological perspectives. Zoological Journal of the Linnean Society 114 (2): 213 - 243. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1995. tb 00117 c. x" type="journal article" year="1995">
Smith
<emphasis id="23A5FA67658385ADD18D2418FF009E33" box="[189,248,305,331]" italics="true" pageId="214" pageNumber="215">et al.</emphasis>
1995
</bibRefCitation>
). On account of fundamental differences in spine articulation structure, assignment either to the
<taxonomicName id="D6D15DF6658385ADD1DB247FFDA39E08" authority="(Hemieuryale)" authorityName="Verrill" authorityYear="1899" baseAuthorityName="Hemieuryale" box="[235,603,342,368]" class="Ophiuroidea" family="Hemieuryalidae" kingdom="Animalia" order="Ophiurida" pageId="214" pageNumber="215" phylum="Echinodermata" rank="family">
Hemieuryalidae (
<emphasis id="23A5FA67658385ADD08A247FFDA99E08" box="[442,593,342,368]" italics="true" pageId="214" pageNumber="215">Hemieuryale</emphasis>
)
</taxonomicName>
or the
<taxonomicName id="D6D15DF6658385ADD385247FFC9B9E08" authorityName="Ljungman" authorityYear="1867" box="[693,867,342,368]" class="Ophiuroidea" family="Ophiomyxidae" kingdom="Animalia" order="Ophiurida" pageId="214" pageNumber="215" phylum="Echinodermata" rank="family">Ophiomyxidae</taxonomicName>
(
<taxonomicName id="D6D15DF6658385ADD248247FFB129E09" authority="Matsumoto, 1915" authorityName="Matsumoto" authorityYear="1915" box="[888,1258,342,369]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiosmilax" kingdom="Animalia" order="Ophiurida" pageId="214" pageNumber="215" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658385ADD248247FFBF69E08" box="[888,1038,342,368]" italics="true" pageId="214" pageNumber="215">Ophiosmilax</emphasis>
Matsumoto, 1915
</taxonomicName>
) is tenable. Rather, the presence of large, ear-shaped spine articulations with a sigmoidal fold in combination with a series of small perforations on the inner side rather than a single large one near the ventral part of the ridge is suggestive of strong ophiacanthid affinities, however atypical this LAP
<typeStatus id="CE6A98D7658385ADD54424EFFB5F9E98" box="[1140,1191,454,480]" pageId="214" pageNumber="215">type</typeStatus>
might appear for an ophiacanthid.
</paragraph>
<paragraph id="116E2675658385ADD18D2706FCFF9CE0" blockId="214.[189,1399,559,921]" pageId="214" pageNumber="215">
Affinities within the
<taxonomicName id="D6D15DF6658385ADD08B2706FD8E9D31" baseAuthorityName="Martynov" baseAuthorityYear="2010" box="[443,630,559,585]" class="Ophiuroidea" family="Ophiacanthidae" kingdom="Animalia" order="Ophiurida" pageId="214" pageNumber="215" phylum="Echinodermata" rank="family">Ophiacanthidae</taxonomicName>
, in contrast, are much more difficult to resolve. In spite of the relatively large, poorly defined tentacle notch in the LAPs of
<taxonomicName id="D6D15DF6658385ADD2AB277DFBD49D16" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[923,1068,596,622]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="214" pageNumber="215" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA67658385ADD2AB277DFBD49D16" box="[923,1068,596,622]" italics="true" pageId="214" pageNumber="215">Ophiojagtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658385ADD505277DFB5C9D16" box="[1077,1188,596,622]" pageId="214" pageNumber="215" rank="genus">gen. nov.</taxonomicNameLabel>
, the tentacle pore in the context of the articulated arm was presumably small, considering the extremely long ventral portion of the LAPs. Thus, it seems probable that
<taxonomicName id="D6D15DF6658385ADD21227B6FC4B9DC1" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[802,947,671,697]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="214" pageNumber="215" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA67658385ADD21227B6FC4B9DC1" box="[802,947,671,697]" italics="true" pageId="214" pageNumber="215">Ophiojagtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658385ADD28D27B6FBD39DC1" box="[957,1067,671,697]" pageId="214" pageNumber="215" rank="genus">gen. nov.</taxonomicNameLabel>
belongs to the small-pored ophiacanthids. Closest similarities seem to be shared with the LAPs of
<taxonomicName id="D6D15DF6658385ADD51827EDFB129DA6" authorityName="Thuy" authorityYear="2011" box="[1064,1258,708,734]" class="Ophiuroidea" family="Ophiacanthidae" genus="Inexpectacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="214" pageNumber="215" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658385ADD51827EDFB129DA6" box="[1064,1258,708,734]" italics="true" pageId="214" pageNumber="215">Inexpectacantha</emphasis>
</taxonomicName>
, mainly on account of the massive, thick general aspect, the position of the spine articulations, and the lack of outer surface ornament and of spurs. However, it is likely that these similarities are superficial, considering the significantly different spine articulation morphologies and ridge shapes. The LAPs of
<taxonomicName id="D6D15DF6658385ADD5D5261DFA8E9C36" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[1253,1398,820,846]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="214" pageNumber="215" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA67658385ADD5D5261DFA8E9C36" box="[1253,1398,820,846]" italics="true" pageId="214" pageNumber="215">Ophiojagtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658385ADD18D2673FED49C0C" box="[189,300,858,884]" pageId="214" pageNumber="215" rank="genus">gen. nov.</taxonomicNameLabel>
appear to be so highly derived that the phylogenetic relationships of the genus cannot be further explored on the basis of LAP morphology.
</paragraph>
<paragraph id="116E2675658385ADD18D26EBFCF39B9A" blockId="214.[189,1399,960,1250]" pageId="214" pageNumber="215">
As already hinted at by
<bibRefCitation id="75405B84658385ADD0DF26E9FD729CA4" author="Storc R." box="[495,650,960,988]" pageId="214" pageNumber="215" pagination="395 - 401" refId="ref128214" refString="Storc R. 2004. The Late Cretaceous brittle star Ophiosmilax? alternatus from Europe and northern Africa. Geobios 37 (3): 395 - 401. http: // dx. doi. org / 10.1016 / j. geobios. 2003.01.010" type="journal article" year="2004">Štorc (2004)</bibRefCitation>
, the LAPs here assigned to
<taxonomicName id="D6D15DF6658385ADD2DF26EBFB789CA4" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[1007,1152,962,988]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="214" pageNumber="215" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA67658385ADD2DF26EBFB789CA4" box="[1007,1152,962,988]" italics="true" pageId="214" pageNumber="215">Ophiojagtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658385ADD5BD26EBFB059CA4" box="[1165,1277,962,988]" pageId="214" pageNumber="215" rank="genus">gen. nov.</taxonomicNameLabel>
remained morphologically conservative over considerable time intervals. Although it seems very unlikely that a single species spanned some 20 Ma, as suggested by
<bibRefCitation id="75405B84658385ADD2282122FC549B5F" author="Storc R." box="[792,940,1035,1063]" pageId="214" pageNumber="215" pagination="395 - 401" refId="ref128214" refString="Storc R. 2004. The Late Cretaceous brittle star Ophiosmilax? alternatus from Europe and northern Africa. Geobios 37 (3): 395 - 401. http: // dx. doi. org / 10.1016 / j. geobios. 2003.01.010" type="journal article" year="2004">Štorc (2004)</bibRefCitation>
, it can be challenging, if not impossible, to differentiate stratigraphically close LAP
<typeStatus id="CE6A98D7658385ADD3FD211BFCF39B34" box="[717,779,1074,1100]" pageId="214" pageNumber="215">types</typeStatus>
of
<taxonomicName id="D6D15DF6658385ADD20B211BFC349B34" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[827,972,1074,1100]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="214" pageNumber="215" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA67658385ADD20B211BFC349B34" box="[827,972,1074,1100]" italics="true" pageId="214" pageNumber="215">Ophiojagtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658385ADD2E7211BFBB19B34" box="[983,1097,1074,1100]" pageId="214" pageNumber="215" rank="genus">gen. nov.</taxonomicNameLabel>
, in particular when only few LAPs are available. In the present study, a total of four LAP
<typeStatus id="CE6A98D7658385ADD2E72171FBED9B0A" box="[983,1045,1112,1138]" pageId="214" pageNumber="215">types</typeStatus>
of
<taxonomicName id="D6D15DF6658385ADD571217EFB2A9B09" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[1089,1234,1111,1137]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="214" pageNumber="215" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA67658385ADD571217EFB2A9B09" box="[1089,1234,1111,1137]" italics="true" pageId="214" pageNumber="215">Ophiojagtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658385ADD5EC2171FAB19B0A" box="[1244,1353,1112,1138]" pageId="214" pageNumber="215" rank="genus">gen. nov.</taxonomicNameLabel>
are characterised and identified at the species level. A few additional occurrences of the genus, including those recorded by
<bibRefCitation id="75405B84658385ADD0A12189FDDB9BC4" author="Storc R." box="[401,547,1184,1212]" pageId="214" pageNumber="215" pagination="395 - 401" refId="ref128214" refString="Storc R. 2004. The Late Cretaceous brittle star Ophiosmilax? alternatus from Europe and northern Africa. Geobios 37 (3): 395 - 401. http: // dx. doi. org / 10.1016 / j. geobios. 2003.01.010" type="journal article" year="2004">Štorc (2004)</bibRefCitation>
and
<bibRefCitation id="75405B84658385ADD3692189FCC39BC4" author="Storc R. &amp; Zitt J." box="[601,827,1184,1212]" pageId="214" pageNumber="215" pagination="123 - 140" refId="ref128259" refString="Storc R. &amp; Zitt J. 2008. Late Turonian ophiuroids (Echinodermata) from the Bohemian Cretaceous Basin, Czech Republic. Bulletin of Geosciences 83 (2): 123 - 140. http: // dx. doi. org / 10.3140 / bull. geosci. 2008.02.123" type="journal article" year="2008">Štorc &amp; Žítt (2008)</bibRefCitation>
, could not be assigned with certainty and are thus treated here as indeterminate records of the genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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