treatments-xml/data/94/72/87/947287DFFFA9FF867EAA12DBFD44FA80.xml

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<document id="62C61925FFD4453C7B80010534E2B1B1" ID-DOI="10.11646/zootaxa.5026.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5300736" ID-ZooBank="44594360-3408-4C4C-A4E6-1025F171A49F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630039285316" checkinUser="plazi" docAuthor="Michie, Laura A., Barnes, R. S. K., Clark, Paul F., Bennett, Wayne A. &amp; Cragg, Simon M." docDate="2021" docId="947287DFFFA9FF867EAA12DBFD44FA80" docLanguage="en" docName="zootaxa.5026.4.2.pdf" docOrigin="Zootaxa 5026 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Austruca perplexa" docType="treatment" docVersion="4" lastPageNumber="488" masterDocId="684BFFA7FFA1FF8E7E3D124CFF94FFB1" masterDocTitle="Distinguishing ten sympatric species of fiddler crab (Decapoda: Ocypodidae) using a suite of phenotypic characteristics" masterLastPageNumber="506" masterPageNumber="480" pageNumber="488" updateTime="1699139032236" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DA866B9F1E93B7467097C89070841EE8">Distinguishing ten sympatric species of fiddler crab (Decapoda: Ocypodidae) using a suite of phenotypic characteristics</mods:title>
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<mods:namePart id="4D6EB5B32441E62D0E9757EC35CBFF59">Michie, Laura A.</mods:namePart>
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<mods:namePart id="DB9C5A9B996E7085DEC040CC69D5A7EA">Barnes, R. S. K.</mods:namePart>
<mods:affiliation id="EBAAE6DBDD5A4D70B492E5AD4327DB9D">Department of Zoology and Conservation Research Institute, University of Cambridge, CB 2 3 EJ, UK.</mods:affiliation>
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<mods:namePart id="BC24376858266A93CEA6BF4CD3E612E1">Clark, Paul F.</mods:namePart>
<mods:affiliation id="0E9F78A51B8C28B7D4E6D2BED8EF12EA">Department of Life Sciences, The Natural History Museum, Cromwell Rad, London, SW 7 5 BD, UK.</mods:affiliation>
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<mods:namePart id="D1FF6B36A468316E7157B17C6AE5B5F7">Bennett, Wayne A.</mods:namePart>
<mods:affiliation id="E599D6476F740A1258170F4267A03D40">University of West Florida, 11000 University Parkway, Pensacola, US.</mods:affiliation>
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<mods:namePart id="3D0EF772088977E02FF91A19A484FAE9">Cragg, Simon M.</mods:namePart>
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<mods:affiliation id="44E4C183418B1ABC9A198BC1CE5F96ED">simon. cragg @ port. ac. uk; https: // orcid. org / 0000 - 0003 - 1082 - 7653</mods:affiliation>
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<treatment id="947287DFFFA9FF867EAA12DBFD44FA80" ID-DOI="http://doi.org/10.5281/zenodo.5307758" ID-Zenodo-Dep="5307758" LSID="urn:lsid:plazi:treatment:947287DFFFA9FF867EAA12DBFD44FA80" httpUri="http://treatment.plazi.org/id/947287DFFFA9FF867EAA12DBFD44FA80" lastPageNumber="488" pageId="8" pageNumber="488">
<subSubSection id="54C16542FFA9FF867EAA12DBFD54FF00" box="[151,704,151,178]" pageId="8" pageNumber="488" type="nomenclature">
<paragraph id="1C6436C9FFA9FF867EAA12DBFD54FF00" blockId="8.[151,704,151,214]" box="[151,704,151,178]" pageId="8" pageNumber="488">
<heading id="472C81A5FFA9FF867EAA12DBFD54FF00" bold="true" box="[151,704,151,178]" fontSize="11" level="1" pageId="8" pageNumber="488" reason="1">
<taxonomicName id="DBDB4D4AFFA9FF867EAA12DBFD54FF00" ID-CoL="JYNY" authority="(H. Milne Edwards, 1852)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1852" box="[151,704,151,178]" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="488" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="2EAFEADBFFA9FF867EAA12DBFD54FF00" bold="true" box="[151,704,151,178]" pageId="8" pageNumber="488">
<emphasis id="2EAFEADBFFA9FF867EAA12DBFEE1FF03" bold="true" box="[151,373,151,178]" italics="true" pageId="8" pageNumber="488">Austruca perplexa</emphasis>
(H. Milne Edwards, 1852)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="54C16542FFA9FF867EAA12F7FEBEFF67" box="[151,298,187,214]" pageId="8" pageNumber="488" type="description">
<paragraph id="1C6436C9FFA9FF867EAA12F7FEBEFF67" blockId="8.[151,704,151,214]" box="[151,298,187,214]" pageId="8" pageNumber="488">
(
<figureCitation id="84E02A4CFFA9FF867EA212F7FEB6FF67" box="[159,290,187,214]" captionStart="FIGURE 5" captionStartId="9.[151,250,1885,1909]" captionTargetBox="[196,1391,181,1859]" captionTargetId="figure-20@9.[196,1391,181,1861]" captionTargetPageId="9" captionText="FIGURE 5. Austruca perplexa H. Milne Edwards, 1852: ad Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of right G1; e, mesial view; f, lateral view; CLSM images of apical part of G1; g, mesial view; h, lateral view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300746" httpUri="https://zenodo.org/record/5300746/files/figure.png" pageId="8" pageNumber="488">Figure 5aj</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFA9FF867EAA134FFE45FEF0" pageId="8" pageNumber="488" type="materials_examined">
<paragraph id="1C6436C9FFA9FF867EAA134FFE45FEF0" blockId="8.[151,1437,259,1329]" pageId="8" pageNumber="488">
<emphasis id="2EAFEADBFFA9FF867EAA134FFE14FEAC" bold="true" box="[151,384,259,285]" pageId="8" pageNumber="488">Material examined.</emphasis>
<materialsCitation id="ACB33C94FFA9FF867FBB134FFE5AFEF0" collectingDate="2014-08-26" collectorName="L. Michie" country="Indonesia" latitude="-5.497222" location="Kaledupa" longLatPrecision="21" longitude="123.755554" pageId="8" pageNumber="488" specimenCount="40" specimenCount-female="20" specimenCount-male="20">
<specimenCount id="0ADDFD40FFA9FF867FBB134FFE29FEAF" box="[390,445,259,286]" pageId="8" pageNumber="488" type="female">20 ♀</specimenCount>
,
<specimenCount id="0ADDFD40FFA9FF867FF6134FFD90FEAF" box="[459,516,259,286]" pageId="8" pageNumber="488" type="male">20 ♂</specimenCount>
,
<location id="19046012FFA9FF867C32134FFDE9FEAC" LSID="urn:lsid:plazi:treatment:947287DFFFA9FF867EAA12DBFD44FA80:19046012FFA9FF867C32134FFDE9FEAC" box="[527,637,259,285]" country="Indonesia" latitude="-5.497222" longLatPrecision="21" longitude="123.755554" name="Kaledupa" pageId="8" pageNumber="488">Kaledupa</location>
,
<collectingCountry id="64CC7659FFA9FF867CB5134FFD6CFEAC" box="[648,760,259,285]" name="Indonesia" pageId="8" pageNumber="488">Indonesia</collectingCountry>
,
<geoCoordinate id="79EF500EFFA9FF867D3F134FFC13FEAF" box="[770,903,259,286]" degrees="5" direction="south" minutes="29" orientation="latitude" pageId="8" pageNumber="488" precision="15" seconds="50" value="-5.497222">5º 29 50S</geoCoordinate>
,
<geoCoordinate id="79EF500EFFA9FF867DAF134FFBA1FEAC" box="[914,1077,259,286]" degrees="123" direction="east" minutes="45" orientation="longitude" pageId="8" pageNumber="488" precision="15" seconds="20" value="123.755554">123º 45 20E</geoCoordinate>
, mud flat in front of mangroves, coll.
<collectorName id="B12E531FFFA9FF867EF21364FED4FEF0" box="[207,320,295,322]" pageId="8" pageNumber="488">L. Michie</collectorName>
,
<date id="68651009FFA9FF867F71136BFE5AFEF0" box="[332,462,295,321]" pageId="8" pageNumber="488" value="2014-08-26">
<collectingDate id="7821E9E1FFA9FF867F71136BFE5AFEF0" box="[332,462,295,321]" pageId="8" pageNumber="488" value="2014-08-26">26.08.2014</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFA9FF867EFA1307FB6CFB84" pageId="8" pageNumber="488" type="diagnosis">
<paragraph id="1C6436C9FFA9FF867EFA1307FC69FE1C" blockId="8.[151,1437,259,1329]" pageId="8" pageNumber="488">
<emphasis id="2EAFEADBFFA9FF867EFA1307FEAEFED4" bold="true" box="[199,314,331,357]" pageId="8" pageNumber="488">Diagnosis</emphasis>
. Carapace: slightly broader than long; smooth; front broad; anterolateral angles moderately acute and slightly produced; anterolateral as well as dorsolateral margins short or nearly absent; anterolateral borders slightly convergent. Major cheliped: long dactyl and pollex; pollex with central tooth.
</paragraph>
<paragraph id="1C6436C9FFA9FF867EFA13FBFC77FD8C" blockId="8.[151,1437,259,1329]" pageId="8" pageNumber="488">
<emphasis id="2EAFEADBFFA9FF867EFA13FBFECCFE60" bold="true" box="[199,344,439,465]" pageId="8" pageNumber="488">Colouration</emphasis>
(
<figureCitation id="84E02A4CFFA9FF867F5A13FBFE67FE63" box="[359,499,439,466]" captionStart="FIGURE 5" captionStartId="9.[151,250,1885,1909]" captionTargetBox="[196,1391,181,1859]" captionTargetId="figure-20@9.[196,1391,181,1861]" captionTargetPageId="9" captionText="FIGURE 5. Austruca perplexa H. Milne Edwards, 1852: ad Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of right G1; e, mesial view; f, lateral view; CLSM images of apical part of G1; g, mesial view; h, lateral view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300746" httpUri="https://zenodo.org/record/5300746/files/figure.png" pageId="8" pageNumber="488">Figure 5ad</figureCitation>
). Similar to
<taxonomicName id="DBDB4D4AFFA9FF867CBC13FBFD6AFE60" box="[641,766,439,465]" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="488" phylum="Arthropoda" rank="species" species="cryptica">
<emphasis id="2EAFEADBFFA9FF867CBC13FBFD6AFE60" box="[641,766,439,465]" italics="true" pageId="8" pageNumber="488">A. cryptica</emphasis>
</taxonomicName>
. Adult male. Carapace: mainly black with white transverse markings varying in size and number, anterior ventral surface of carapace and third maxilliped white. Major cheliped: dactyl and pollex white with manus and carpus yellow or white, merus, ischium, basis and coxa yellow. Minor cheliped &amp; ambulatory legs: yellow, white, grey, occasionally black.
</paragraph>
<paragraph id="1C6436C9FFA9FF867EFA100BFC4FFD18" blockId="8.[151,1437,259,1329]" pageId="8" pageNumber="488">Adult female. Carapace: varies greatly, most often mottled light brown/russet, grey or white, occasionally same as male carapace, anterior ventral surface of carapace and third maxilliped most often white or grey, occasionally black with ischium of third maxilliped white. Ambulatory legs: like males.</paragraph>
<paragraph id="1C6436C9FFA9FF867EFA10FFFBC3FD40" blockId="8.[151,1437,259,1329]" pageId="8" pageNumber="488">Juvenile male. Carapace: usually mottled light brown/russet with pattern and dark colours developing with maturity. Major cheliped: usually entirely white and develops colour with maturation.</paragraph>
<paragraph id="1C6436C9FFA9FF867EFA10B7FE47FC8B" blockId="8.[151,1437,259,1329]" pageId="8" pageNumber="488">
Juvenile female. Same pattern as juvenile males, sometimes difficult to distinguish juveniles of
<taxonomicName id="DBDB4D4AFFA9FF867AC610B7FAE2FCA4" box="[1275,1398,763,789]" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="488" phylum="Arthropoda" rank="species" species="cryptica">
<emphasis id="2EAFEADBFFA9FF867AC610B7FAE2FCA4" box="[1275,1398,763,789]" italics="true" pageId="8" pageNumber="488">A. cryptica</emphasis>
</taxonomicName>
,
<taxonomicName id="DBDB4D4AFFA9FF867BB910B7FE9CFC88" baseAuthorityName="Rathbun" baseAuthorityYear="1924" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="488" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="2EAFEADBFFA9FF867BB910B7FE9CFC88" italics="true" pageId="8" pageNumber="488">A. mjoebergi</emphasis>
</taxonomicName>
and
<taxonomicName id="DBDB4D4AFFA9FF867F03116CFE5FFC88" baseAuthorityName="H. -T. Shih, Naruse &amp; P. K. L. Ng" baseAuthorityYear="2010" box="[318,459,799,826]" class="Malacostraca" family="Ocypodidae" genus="Gelasimus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="488" phylum="Arthropoda" rank="species" species="jocelynae">
<emphasis id="2EAFEADBFFA9FF867F03116CFE5FFC88" box="[318,459,799,826]" italics="true" pageId="8" pageNumber="488">G. jocelynae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1C6436C9FFA9FF867EFA110FFE21FC78" blockId="8.[151,1437,259,1329]" pageId="8" pageNumber="488">
<emphasis id="2EAFEADBFFA9FF867EFA110FFE45FCEC" bold="true" box="[199,465,835,861]" pageId="8" pageNumber="488">Gonopod morphology</emphasis>
(
<figureCitation id="84E02A4CFFA9FF867FDF110FFDFBFCEF" box="[482,623,835,862]" captionStart="FIGURE 5" captionStartId="9.[151,250,1885,1909]" captionTargetBox="[196,1391,181,1859]" captionTargetId="figure-20@9.[196,1391,181,1861]" captionTargetPageId="9" captionText="FIGURE 5. Austruca perplexa H. Milne Edwards, 1852: ad Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of right G1; e, mesial view; f, lateral view; CLSM images of apical part of G1; g, mesial view; h, lateral view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300746" httpUri="https://zenodo.org/record/5300746/files/figure.png" pageId="8" pageNumber="488">Figure 5eh</figureCitation>
). G1 with broad termination, directed in dorsolateral direction, mesial border between protuberance and termination with swelling; distal protuberance moderately long, often reaching beyond base of flanges; distal end relatively long mainly due to largely protruding dorsal lobe; pore of sperm channel large, in midline.
</paragraph>
<paragraph id="1C6436C9FFA9FF867EFA119FFB6CFB84" blockId="8.[151,1437,259,1329]" pageId="8" pageNumber="488">
<emphasis id="2EAFEADBFFA9FF867EFA119FFEC1FC5C" bold="true" box="[199,341,979,1005]" pageId="8" pageNumber="488">Gastric mill</emphasis>
(
<figureCitation id="84E02A4CFFA9FF867F5F119FFE77FC5F" box="[354,483,979,1006]" captionStart="FIGURE 5" captionStartId="9.[151,250,1885,1909]" captionTargetBox="[196,1391,181,1859]" captionTargetId="figure-20@9.[196,1391,181,1861]" captionTargetPageId="9" captionText="FIGURE 5. Austruca perplexa H. Milne Edwards, 1852: ad Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of right G1; e, mesial view; f, lateral view; CLSM images of apical part of G1; g, mesial view; h, lateral view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300746" httpUri="https://zenodo.org/record/5300746/files/figure.png" pageId="8" pageNumber="488">Figure 5ij</figureCitation>
). Urocardiac ossicles moderately complex with 34 transverse ridges of median teeth, decreasing in size distally, first one long and strongly arched; stem region has 2 pairs of cusps, first large and rounded, second small and attached to plate dorsally. Lateral tooth plate with 17 comb-shaped teeth.
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFA9FF867EFA1673FAA3FBCF" pageId="8" pageNumber="488" type="biology_ecology">
<paragraph id="1C6436C9FFA9FF867EFA1673FAA3FBCF" blockId="8.[151,1437,259,1329]" pageId="8" pageNumber="488">
<emphasis id="2EAFEADBFFA9FF867EFA1673FEB3FBE8" bold="true" box="[199,295,1087,1113]" pageId="8" pageNumber="488">Biology.</emphasis>
<taxonomicName id="DBDB4D4AFFA9FF867F131673FE68FBE8" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1852" box="[302,508,1087,1113]" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="488" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="2EAFEADBFFA9FF867F131673FE68FBE8" box="[302,508,1087,1113]" italics="true" pageId="8" pageNumber="488">Austruca perplexa</emphasis>
</taxonomicName>
is a small-size species (largest male CL =
<quantity id="DB239B2CFFA9FF867DE41673FBABFBEB" box="[985,1087,1087,1114]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="8" pageNumber="488" unit="mm" value="10.1">10.1 mm</quantity>
, CB =
<quantity id="DB239B2CFFA9FF867AB01673FB67FBEB" box="[1165,1267,1087,1114]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="8" pageNumber="488" unit="mm" value="15.6">15.6 mm</quantity>
, largest female CL =
<quantity id="DB239B2CFFA9FF867EE5162FFEA4FBCF" box="[216,304,1123,1150]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="8" pageNumber="488" unit="mm" value="9.1">9.1 mm</quantity>
, CB =
<quantity id="DB239B2CFFA9FF867F43162FFE72FBCF" box="[382,486,1123,1150]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="8" pageNumber="488" unit="mm" value="14.8">14.8 mm</quantity>
); one of most abundant species on shore, dominates the mid intertidal zone.
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFA9FF867EFA16CBFDA5FB74" pageId="8" pageNumber="488" type="distribution">
<paragraph id="1C6436C9FFA9FF867EFA16CBFDA5FB74" blockId="8.[151,1437,259,1329]" pageId="8" pageNumber="488">
<emphasis id="2EAFEADBFFA9FF867EFA16CBFECDFB10" bold="true" box="[199,345,1159,1185]" pageId="8" pageNumber="488">Distribution</emphasis>
. From Nicobar Is. Eastward, Eastern Indian Ocean,
<collectingCountry id="64CC7659FFA9FF867DA516CBFC68FB10" box="[920,1020,1159,1185]" name="Thailand" pageId="8" pageNumber="488">Thailand</collectingCountry>
to
<collectingCountry id="64CC7659FFA9FF867A2016CBFBF7FB10" box="[1053,1123,1159,1185]" name="China" pageId="8" pageNumber="488">China</collectingCountry>
,
<collectingCountry id="64CC7659FFA9FF867A5116CBFB55FB10" box="[1132,1217,1159,1185]" name="Taiwan" pageId="8" pageNumber="488">Taiwan</collectingCountry>
,
<collectingCountry id="64CC7659FFA9FF867AF616C4FA9AFB13" box="[1227,1294,1160,1186]" name="Japan" pageId="8" pageNumber="488">Japan</collectingCountry>
,
<collectingCountry id="64CC7659FFA9FF867B2A16CBFA0CFB10" box="[1303,1432,1159,1185]" name="Philippines" pageId="8" pageNumber="488">Philippines</collectingCountry>
,
<collectingCountry id="64CC7659FFA9FF867EAA16E7FE93FB74" box="[151,263,1195,1221]" name="Indonesia" pageId="8" pageNumber="488">Indonesia</collectingCountry>
,
<collectingCountry id="64CC7659FFA9FF867F2D16E7FEEEFB74" box="[272,378,1195,1221]" name="Australia" pageId="8" pageNumber="488">Australia</collectingCountry>
, Pacific Islands.
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFA9FF867EFA1683FD44FA80" pageId="8" pageNumber="488" type="discussion">
<paragraph id="1C6436C9FFA9FF867EFA1683FD44FA80" blockId="8.[151,1437,259,1329]" pageId="8" pageNumber="488">
<emphasis id="2EAFEADBFFA9FF867EFA1683FEAFFB58" bold="true" box="[199,315,1231,1257]" pageId="8" pageNumber="488">Remarks.</emphasis>
The specimens examined concur with the description of
<bibRefCitation id="784A4B38FFA9FF867D981683FB0EFB58" author="Naderloo, R. &amp; Turkay, M. &amp; Chen, H. - L." box="[933,1178,1231,1257]" pageId="8" pageNumber="488" pagination="1 - 38" refId="ref14169" refString="Naderloo, R., Turkay, M. &amp; Chen, H. - L. (2010) Taxonomic revision of the wide-front fiddler crabs of the Uca lactea group (Crustacea: Decapoda: Brachyura: Ocypodidae) in the Indo-West Pacific. Zootaxa, 2500 (1), 1 - 38. https: // doi. org / 10.11646 / zootaxa. 2500.1.1" type="journal article" year="2010">
Naderloo
<emphasis id="2EAFEADBFFA9FF867A2F169CFBDCFB58" box="[1042,1096,1231,1257]" italics="true" pageId="8" pageNumber="488">et al.</emphasis>
(2010)
</bibRefCitation>
, specifically the largely protruding dorsal lobe of the G1 and the moderately complex morphology of the gastric mill, which distinguish
<taxonomicName id="DBDB4D4AFFA9FF867BB916BFFF6DFA80" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1852" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="488" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="2EAFEADBFFA9FF867BB916BFFF6DFA80" italics="true" pageId="8" pageNumber="488">A. perplexa</emphasis>
</taxonomicName>
from the other Ambeua
<taxonomicName id="DBDB4D4AFFA9FF867C36175BFD44FA80" box="[523,720,1303,1329]" class="Malacostraca" family="Ocypodidae" genus="Austruca" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="488" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2EAFEADBFFA9FF867C36175BFDE4FA80" box="[523,624,1303,1329]" italics="true" pageId="8" pageNumber="488">Austruca</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>