treatments-xml/data/93/28/87/932887D3FFD42C42FF3A2E425DAFF845.xml

<document id="4D87ABE94142C6021728FC25F12077D3" ID-CLB-Dataset="31650" ID-DOI="10.11646/zootaxa.4341.4.5" ID-GBIF-Dataset="6c4545c4-e521-4b2d-99d1-6a29fd12ab80" ID-ISSN="1175-5326" ID-Zenodo-Dep="1040596" ID-ZooBank="0DA889AB-FD1A-403F-932D-BD0298A78251" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509608394247" checkinUser="plazi" docAuthor="Liang, Wei-Ren, Maruyama, Munetoshi &amp; Li, Hou-Feng" docDate="2017" docId="932887D3FFD42C42FF3A2E425DAFF845" docLanguage="en" docName="zootaxa.4341.4.5.pdf" docOrigin="Zootaxa 4341 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zyras (Diaulaconia) artemis Liang &amp; Maruyama &amp; Li 2017, sp. n." docType="treatment" docVersion="8" lastPageNumber="533" masterDocId="6F11FFABFFD52C47FFAD28285E74FFB6" masterDocTitle="Zyras (Diaulaconia) artemis sp. n., a new termitophilous species from Taiwan (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="538" masterPageNumber="528" pageNumber="529" updateTime="1698470682602" updateUser="ExternalLinkService">
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<mods:title id="D2567EB7011701592695839774FDCA84">Zyras (Diaulaconia) artemis sp. n., a new termitophilous species from Taiwan (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="A66BB3B9CF240A58509E9D60FA6DCBE2">Liang, Wei-Ren</mods:namePart>
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<mods:namePart id="D97720275A5A25A92ECF5C635FE43C66">Maruyama, Munetoshi</mods:namePart>
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<treatment id="932887D3FFD42C42FF3A2E425DAFF845" ID-DOI="http://doi.org/10.5281/zenodo.6028469" ID-GBIF-Taxon="137227793" ID-Zenodo-Dep="6028469" LSID="urn:lsid:plazi:treatment:932887D3FFD42C42FF3A2E425DAFF845" httpUri="http://treatment.plazi.org/id/932887D3FFD42C42FF3A2E425DAFF845" lastPageId="5" lastPageNumber="533" pageId="1" pageNumber="529">
<subSubSection id="539B654EFFD42C46FF3A2E425F7FF910" pageId="1" pageNumber="529" type="nomenclature">
<paragraph id="1B3E36C5FFD42C46FF3A2E425D1FF933" blockId="1.[151,875,1642,1702]" box="[151,875,1642,1669]" pageId="1" pageNumber="529">
<heading id="407681A9FFD42C46FF3A2E425D1FF933" bold="true" box="[151,875,1642,1669]" fontSize="11" level="1" pageId="1" pageNumber="529" reason="1">
<emphasis id="29F5EAD7FFD42C46FF3A2E425D1FF933" bold="true" box="[151,875,1642,1669]" pageId="1" pageNumber="529">
<taxonomicName id="DC814D46FFD42C46FF3A2E425D6FF933" ID-CoL="5DCQT" authority="Liang, Maruyama &amp; Li" authorityName="Liang &amp; Maruyama &amp; Li" authorityYear="2017" box="[151,795,1642,1669]" class="Insecta" family="Staphylinidae" genus="Zyras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="529" phylum="Arthropoda" rank="species" species="artemis" status="sp. nov." subGenus="Diaulaconia">
<emphasis id="29F5EAD7FFD42C46FF3A2E425EA8F932" bold="true" box="[151,220,1642,1668]" italics="true" pageId="1" pageNumber="529">Zyras</emphasis>
(
<emphasis id="29F5EAD7FFD42C46FF412E425FF6F932" bold="true" box="[236,386,1642,1668]" italics="true" pageId="1" pageNumber="529">Diaulaconia</emphasis>
) 
<emphasis id="29F5EAD7FFD42C46FE3E2E425F98F932" bold="true" box="[403,492,1642,1668]" italics="true" pageId="1" pageNumber="529">artemis</emphasis>
Liang, Maruyama &amp; Li
</taxonomicName>
, 
<taxonomicNameLabel id="32C657ACFFD42C46FC842E435D1FF933" box="[809,875,1643,1669]" pageId="1" pageNumber="529" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="1B3E36C5FFD42C46FF3A2EA55F7FF910" blockId="1.[151,875,1642,1702]" box="[151,267,1677,1702]" pageId="1" pageNumber="529">
(
<figureCitation id="83BA2A40FFD42C46FF322EA55F76F910" box="[159,258,1677,1702]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="3.[151,250,2009,2031]" captionStartId-1="4.[151,250,1511,1533]" captionStartId-2="5.[151,250,1666,1688]" captionStartId-3="6.[151,250,1502,1524]" captionStartId-4="7.[151,250,1790,1812]" captionStartId-5="8.[151,250,1936,1958]" captionTargetBox-1="[227,1359,187,1467]" captionTargetBox-2="[151,1436,185,1642]" captionTargetBox-3="[165,1419,208,1471]" captionTargetBox-4="[171,1424,516,1745]" captionTargetBox-5="[182,1415,745,1902]" captionTargetId-1="figure@4.[151,1436,161,1489]" captionTargetId-2="figure@5.[151,1436,185,1642]" captionTargetId-3="figure@6.[151,1436,193,1478]" captionTargetId-4="figure@7.[151,1436,481,1766]" captionTargetId-5="figure@8.[151,1436,736,1914]" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 1. Habitus of Zyras (Diaulaconia) artemis sp. n., male, dorsal vieW." captionText-1="FIGURE 2. Zyras (Diaulaconia) artemis sp. n. A, forebody, dorsal vieW; B, rigHt antenna, lateral vieW; C, ditto, dorsal vieW; D, frons of tHe female WitHout granulations; E, frons of tHe male WitH granulations." captionText-2="FIGURE 3. Zyras (Diaulaconia) artemis sp. n. A, male 4 th tergite WitH a pair of projeCtions on posterior margin, large individual, dorsal vieW; B, ditto, projeCtion bilobed apiCally, lateral vieW; C, ditto, small individual, dorsal vieW; D, ditto, projeCtion WitH no bilobed apiCally, lateral vieW; E, male 7 th tergite WitH apiCal median tuberCle, dorsal vieW; F, ditto, lateral vieW; G, female 7 th tergite WitH no small median tuberCle, dorsal vieW." captionText-3="FIGURE 4. Zyras (Diaulaconia) artemis sp. n., male. A, eigHt abdominal tergite, dorsal vieW; B, 8 th abdominal sternite, ventral vieW; C, median lobe of aedeagus, ventral vieW; D, ditto, tHe pHotograpH; E, ditto, lateral vieW; F, ditto, tHe pHotograpH." captionText-4="FIGURE 5. Zyras (Diaulaconia) artemis sp. n., female. A, eigHt abdominal tergite, dorsal vieW; B 8 th abdominal sternite, ventral vieW; C, spermatHeCa; D, tHe pHotograpH of tHe apiCal part and spermatHeCal gland; E, tHe pHotograpH of tHe inner Wall of apiCal 1 / 3 of tHe basal part; F, tHe base of tHe basal part and tHe spermatHeCal duCt." captionText-5="FIGURE 6. Variations of tHe spermatHeCa of Zyras (Diaulaconia) artemis sp. n. LoCality of eaCH speCimens: A, Gaupo (Taoyuan Co.); B, Hsilin trail (Hualien Co.); C, DaHasHan (Pingtung Co.); D, E, Dakeng (TaiCHung City); F, Xiaping (Nantou Co.); G, H, LuansHan (Taitung Co.); I, SHeding (Pingtung Co.); J, K, PengHu Islands (PengHu Co.)." httpUri-1="https://zenodo.org/record/1040598/files/figure.png" httpUri-2="https://zenodo.org/record/1040600/files/figure.png" httpUri-3="https://zenodo.org/record/1040602/files/figure.png" httpUri-4="https://zenodo.org/record/1040604/files/figure.png" httpUri-5="https://zenodo.org/record/1040606/files/figure.png" pageId="1" pageNumber="529">Figs 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="539B654EFFD42C45FF3A2EFD5A83FCEA" lastPageId="2" lastPageNumber="530" pageId="1" pageNumber="529" type="materials_examined">
<paragraph id="1B3E36C5FFD42C45FF3A2EFD5A83FCEA" blockId="1.[151,1437,1749,2026]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="530" pageId="1" pageNumber="529">
<emphasis id="29F5EAD7FFD42C46FF3A2EFD5C66F958" bold="true" box="[151,530,1749,1774]" pageId="1" pageNumber="529">Type materials (163 specimens).</emphasis>
Holotype: male, Taiwan: Pingtung Co.: Sheding: 21.9646˚N; 120.8270˚E, 
<quantity id="DC799B20FFD42C46FAF82EFD5BECF958" box="[1365,1432,1749,1774]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="529" unit="m" value="165.0">165m</quantity>
, 
<date id="6F3F1005FFD42C46FF3A2ED15F6DF8A4" box="[151,281,1785,1810]" pageId="1" pageNumber="529" value="2015-11-18">18.XI.2015</date>
, by high-intensity discharge lamp (HID light trap), W.-R. Liang &amp; Y.-C. Lan, deposited in NMNS. Paratypes: Hualien Co.: Danongdafu: 23.6185˚N; 121.4009˚E, 
<quantity id="DC799B20FFD42C46FCF52F355DEEF880" box="[856,922,1821,1846]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="529" unit="m" value="160.0">160m</quantity>
, 
<date id="6F3F1005FFD42C46FC082F355A6AF880" box="[933,1054,1821,1846]" pageId="1" pageNumber="529" value="2016-10-20">20.X.2016</date>
, by ultraviolet light trap (UV light trap), W.-R. Liang &amp; Y.-Z. Huang, 1♂ (dissected) (NMNS); Hsilin trail: 23.8275˚N, 121.4250˚E, 
<quantity id="DC799B20FFD42C46FAF82F6A5BECF8EC" box="[1365,1432,1858,1882]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="529" unit="m" value="270.0">270m</quantity>
, 
<date id="6F3F1005FFD42C46FF3A2F4D5F7BF8C8" box="[151,271,1893,1918]" pageId="1" pageNumber="529" value="2016-10-19">19.X.2016</date>
, by UV light trap, W.-R. Liang &amp; Y.-Z. Huang, 1♀ (dissected) (NCHU). Nantou Co.: Xiaping: 23.7759˚N, 120.6720˚E, 
<quantity id="DC799B20FFD42C46FE172FA15F88F814" box="[442,508,1929,1954]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="529" unit="m" value="150.0">150m</quantity>
, 
<date id="6F3F1005FFD42C46FDA02FA15CA5F814" box="[525,721,1929,1954]" pageId="1" pageNumber="529" value="2015-04-25" valueMax="2015-05-30" valueMin="2015-04-25">25.IV–30.V.2015</date>
, by flight interception trap (FIT), W.-R. Liang, 1♂, 1♀ (all dissected) (KUM); 
<date id="6F3F1005FFD42C46FEDF2F855C38F870" box="[370,588,1965,1990]" pageId="1" pageNumber="529" value="2015-06-23" valueMax="2015-07-15" valueMin="2015-06-23">23.VI–15.VII.2015</date>
, by FIT, 3♂♂ (NMNS); 
<date id="6F3F1005FFD42C46FCCB2F855DA0F870" box="[870,980,1965,1990]" pageId="1" pageNumber="529" value="2016-01-28">28.I.2016</date>
, near the foraging site of 
<taxonomicName id="DC814D46FFD42C46FB5B2F865BE8F870" baseAuthorityName="Shiraki" baseAuthorityYear="1909" box="[1270,1436,1966,1990]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="1" pageNumber="529" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="29F5EAD7FFD42C46FB5B2F865BE8F870" box="[1270,1436,1966,1990]" italics="true" pageId="1" pageNumber="529">O. formosanus</emphasis>
</taxonomicName>
in the rotten wood, 1♂, 1♀ (NCHU); 
<date id="6F3F1005FFD42C46FDDE2FF95D7CF85C" box="[627,776,2001,2026]" pageId="1" pageNumber="529" value="2016-08-28">28.VIII.2016</date>
, 1♂ (NMNS); 23.7760˚N, 120.6723˚E, 
<quantity id="DC799B20FFD42C46FB432FFA5B45F85C" box="[1262,1329,2002,2026]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="529" unit="m" value="147.0">147m</quantity>
, 
<date id="6F3F1005FFD42C45FAE92FF95F6AFF06" lastPageId="2" lastPageNumber="530" pageId="1" pageNumber="529" value="2015-07-15" valueMax="2015-08-02" valueMin="2015-07-15">15.VII– 2.VIII.2015</date>
, by UV light trap, 4♂♂, 5♀♀ (NCHU); 23.7748˚N, 120.6722˚E, 
<quantity id="DC799B20FFD72C45FBBF28B05A21FF06" box="[1042,1109,152,176]" metricMagnitude="2" metricUnit="m" metricValue="1.49" pageId="2" pageNumber="530" unit="m" value="149.0">149m</quantity>
, 
<date id="6F3F1005FFD72C45FBCF28BF5B35FF06" box="[1122,1345,151,176]" pageId="2" pageNumber="530" value="2015-07-15" valueMax="2015-08-02" valueMin="2015-07-15">15.VII–2.VIII.2015</date>
, by UV light trap, 10♂♂, 11♀♀ (KUM); Shueili, 23.7661˚N, 120.8560˚E, 
<quantity id="DC799B20FFD72C45FC2B28945DBDFF62" box="[902,969,188,212]" metricMagnitude="2" metricUnit="m" metricValue="4.04" pageId="2" pageNumber="530" unit="m" value="404.0">404m</quantity>
, 
<date id="6F3F1005FFD72C45FC7928945AECFF63" box="[980,1176,188,213]" pageId="2" pageNumber="530" value="2016-04-29" valueMax="2016-05-16" valueMin="2016-04-29">29.IV–16.V.2016</date>
, by Malaise trap, H.-T. Yeh, 1♂, 3♀♀ (NTU); 
<date id="6F3F1005FFD72C45FE6F28F75C0FFF4E" box="[450,635,223,248]" pageId="2" pageNumber="530" value="2016-05-16" valueMax="2016-06-01" valueMin="2016-05-16">16.V–1.VI.2016</date>
, 1♂, 2♀♀ (KUM); 
<date id="6F3F1005FFD72C45FCD228F75A36FF4E" box="[895,1090,223,248]" pageId="2" pageNumber="530" value="2016-06-01" valueMax="2016-06-15" valueMin="2016-06-01">1.VI–15.VI.2016</date>
, 2♀♀ (NTU); Qingshuigou: 23.7615˚N, 120.7927˚E, 
<quantity id="DC799B20FFD72C45FE1C292C5F80FEAA" box="[433,500,260,284]" metricMagnitude="2" metricUnit="m" metricValue="4.34" pageId="2" pageNumber="530" unit="m" value="434.0">434m</quantity>
, 
<date id="6F3F1005FFD72C45FDAD292C5CB0FEAB" box="[512,708,260,285]" pageId="2" pageNumber="530" value="2016-04-29" valueMax="2016-05-16" valueMin="2016-04-29">29.IV–16.V.2016</date>
, by Malaise trap, H.-T. Yeh, 5♀♀ (KUM); 
<date id="6F3F1005FFD72C45FB1E292C5B18FEAB" box="[1203,1388,260,285]" pageId="2" pageNumber="530" value="2016-05-16" valueMax="2016-06-01" valueMin="2016-05-16">16.V–1.VI.2016</date>
, 1♂ (KUM); 
<date id="6F3F1005FFD72C45FF57290F5FE3FEF6" box="[250,407,295,320]" pageId="2" pageNumber="530" value="2016-06-01" valueMax="2016-06-15" valueMin="2016-06-01">1–15.VI.2016</date>
, 2♂♂, 1♀ (KUM); 
<date id="6F3F1005FFD72C45FDDB290F5D36FEF6" box="[630,834,295,320]" pageId="2" pageNumber="530" value="2016-06-30" valueMax="2016-07-01" valueMin="2016-06-30">30.VI–1.VII.2016</date>
, 1♂ (NTU); 
<date id="6F3F1005FFD72C45FC7C290F5AF6FEF6" box="[977,1154,295,320]" pageId="2" pageNumber="530" value="2016-08-02" valueMax="2016-08-17" valueMin="2016-08-02">2–17.VIII.2016</date>
, 1♀ (NTU). Penghu Co.: Penghu Islands [Peng Hwu Isls.], 
<date id="6F3F1005FFD72C45FD8929645CB6FED2" box="[548,706,332,356]" pageId="2" pageNumber="530" value="1948-06-04" valueMax="1948-06-13" valueMin="1948-06-04">4–13.VI.1948</date>
, C.-W. Chen &amp; L-T. Chen, 8♂♂, 7♀♀ (2♂♂, 2♀♀ dissected) (TARI). Pingtung Co.: same data as holotype, 7♂♂, 6♀♀ (4♂♂, 4♀♀ dissected) (KUM, NCHU, NMNS); Sheding: 21.9613˚N; 120.8236˚E, 
<quantity id="DC799B20FFD72C45FD8D29BC5C16FE1A" box="[544,610,404,428]" metricMagnitude="2" metricUnit="m" metricValue="1.93" pageId="2" pageNumber="530" unit="m" value="193.0">193m</quantity>
, 
<date id="6F3F1005FFD72C45FDDD29BC5D71FE1B" box="[624,773,404,429]" pageId="2" pageNumber="530" value="2015-08-24">24.VIII.2015</date>
, by sifting from the foraging site of 
<taxonomicName id="DC814D46FFD72C45FB0629BD5B26FE1A" baseAuthorityName="Shiraki" baseAuthorityYear="1909" box="[1195,1362,404,428]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="29F5EAD7FFD72C45FB0629BD5B26FE1A" box="[1195,1362,404,428]" italics="true" pageId="2" pageNumber="530">O. formosanus</emphasis>
</taxonomicName>
in the leaf litter, Naomichi Tsuji &amp; W.-R. Liang, 9♂♂, 8♀♀ (1♂, 3♀♀ dissected) (KUM, NCHU, NMNS); Dahanshan: 22.4073˚N; 120.6644˚E, 
<quantity id="DC799B20FFD72C45FE1B29F45F8DFE43" box="[438,505,476,501]" metricMagnitude="2" metricUnit="m" metricValue="4.63" pageId="2" pageNumber="530" unit="m" value="463.0">463m</quantity>
, 
<date id="6F3F1005FFD72C45FDA529F45CFEFE43" box="[520,650,476,501]" pageId="2" pageNumber="530" value="2015-11-21">21.XI.2015</date>
, by UV light trap, W.-R. Liang, 1♀ (dissected) (NCHU); Formosa: Kuraru [Kueitzuchiao], 
<date id="6F3F1005FFD72C45FE0B29D75C11FDAE" box="[422,613,511,536]" pageId="2" pageNumber="530" value="1932-08-21" valueMax="1932-08-25" valueMin="1932-08-21">21–25.VIII.1932</date>
, Y. Miwa, 1♀ (TARI). Taichung City: Dakeng: 24.1758˚N; 120.7776˚E, 
<quantity id="DC799B20FFD72C45FF3A2A0C5EADFD8A" box="[151,217,548,572]" metricMagnitude="2" metricUnit="m" metricValue="3.78" pageId="2" pageNumber="530" unit="m" value="378.0">378m</quantity>
, 
<date id="6F3F1005FFD72C45FF482A0C5F0EFD8B" box="[229,378,548,573]" pageId="2" pageNumber="530" value="2016-08-19">19.VIII.2016</date>
, by UV light trap, W.-R. Liang, 1♂, 3♀♀ (all dissected) (NCHU); 
<date id="6F3F1005FFD72C45FBCB2A0C5A8FFD8B" box="[1126,1275,548,573]" pageId="2" pageNumber="530" value="2016-08-27">27.VIII.2016</date>
, 3♂♂, 1♀ (all dissected) (NCHU); Wanfeng Hill, 
<date id="6F3F1005FFD72C45FD8C2A605C03FDD6" box="[545,631,584,608]" pageId="2" pageNumber="530" value="1984-10">X.1984</date>
, K.-S. Lin &amp; K.-C. Chou, by Malaise trap, 9♂♂, 9♀♀ (TARI); Hsinshe, 
<date id="6F3F1005FFD72C45FF3A2A445F44FD33" box="[151,304,620,645]" pageId="2" pageNumber="530" value="1955-12-05" valueMax="1955-12-08" valueMin="1955-12-05">5–8.XII.1955</date>
, 1♂, 2♀♀ (TARI). Taitung Co.: Basian Cave: 23.3995˚N, 121.4788˚E, 
<quantity id="DC799B20FFD72C45FBD92A445ADDFD32" box="[1140,1193,620,644]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="530" unit="m" value="43.0">43m</quantity>
, 
<date id="6F3F1005FFD72C45FB152A445B44FD33" box="[1208,1328,620,645]" pageId="2" pageNumber="530" value="2016-10-21">21.X.2016</date>
, near the foraging site of 
<taxonomicName id="DC814D46FFD72C45FEF82AB85F8BFD1E" baseAuthorityName="Shiraki" baseAuthorityYear="1909" box="[341,511,656,680]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="29F5EAD7FFD72C45FEF82AB85F8BFD1E" box="[341,511,656,680]" italics="true" pageId="2" pageNumber="530">O. formosanus</emphasis>
</taxonomicName>
in the rotten wood, W.-R. Liang &amp; Y.-Z. Huang, 3♂♂ (1 dissected) (NCHU); Taitung [Formosa: Taito], 
<date id="6F3F1005FFD72C45FE782A9C5CEDFD7A" box="[469,665,692,717]" pageId="2" pageNumber="530" value="1919-02-25" valueMax="1919-03-27" valueMin="1919-02-25">25.II–27.III.1919</date>
, S. Inamura, J. Sonan, and M. Yoshino, 1♂ (dissected), 
<taxonomicName id="DC814D46FFD72C45FAF12A9D5F39FD46" authorityName=", Fauvel" authorityYear="1905" class="Insecta" family="Staphylinidae" genus="Zyras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="compressicornis">
<emphasis id="29F5EAD7FFD72C45FAF12A9D5F39FD46" italics="true" pageId="2" pageNumber="530">Zyras compressicornis</emphasis>
</taxonomicName>
(det. T. Shiraki), 
<taxonomicName id="DC814D46FFD72C45FD942AF15CA9FD46" authorityName="Liang &amp; Maruyama &amp; Li" authorityYear="2017" box="[569,733,729,752]" class="Insecta" family="Staphylinidae" genus="Zyras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="29F5EAD7FFD72C45FD942AF15CA9FD46" box="[569,733,729,752]" italics="true" pageId="2" pageNumber="530">Zyras artemis</emphasis>
</taxonomicName>
(det. W.-R. Liang, 2017) (TARI); Luanshan: 22.9139˚N, 121.1839˚E, 
<quantity id="DC799B20FFD72C45FE832AD45F04FCA3" box="[302,368,764,789]" metricMagnitude="2" metricUnit="m" metricValue="6.05" pageId="2" pageNumber="530" unit="m" value="605.0">605m</quantity>
, 
<date id="6F3F1005FFD72C45FE2C2AD45C70FCA3" box="[385,516,764,789]" pageId="2" pageNumber="530" value="2015-11-14">14.XI.2015</date>
, by UV light trap, W.-R. Liang, 8♂♂, 10♀♀ (2♂♂, 2♀♀ dissected) (KUM, NCHU, NMNS). Taoyuan Co.: Gaupo: 24.7634˚N, 121.3486˚E, 
<quantity id="DC799B20FFD72C45FC2C2B375DB7FC8E" box="[897,963,799,824]" metricMagnitude="2" metricUnit="m" metricValue="5.14" pageId="2" pageNumber="530" unit="m" value="514.0">514m</quantity>
, 
<date id="6F3F1005FFD72C45FC7C2B375A24FC8E" box="[977,1104,799,824]" pageId="2" pageNumber="530" value="2016-04-12">12.IV.2016</date>
, near the foraging site of 
<taxonomicName id="DC814D46FFD72C45FA2C2B085F6EFCEA" baseAuthorityName="Shiraki" baseAuthorityYear="1909" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="29F5EAD7FFD72C45FA2C2B085F6EFCEA" italics="true" pageId="2" pageNumber="530">O. formosanus</emphasis>
</taxonomicName>
in the rotten wood, W.-R. Liang &amp; Y.-Z. Huang, 3♂♂, 1♀ (1♂, 1♀ dissected) (NCHU).
</paragraph>
</subSubSection>
<subSubSection id="539B654EFFD72C43FF6A2B4F5AF1F933" lastPageId="4" lastPageNumber="532" pageId="2" pageNumber="530" type="description">
<paragraph id="1B3E36C5FFD72C45FF6A2B4F5B1CFA1A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="530">
<emphasis id="29F5EAD7FFD72C45FF6A2B4F5F2DFC36" bold="true" box="[199,345,871,896]" pageId="2" pageNumber="530">Description.</emphasis>
Body (
<figureCitation id="83BA2A40FFD72C45FE062B4F5F9EFC36" box="[427,490,871,896]" captionStart="FIGURE 1" captionStartId="3.[151,250,2009,2031]" captionText="FIGURE 1. Habitus of Zyras (Diaulaconia) artemis sp. n., male, dorsal vieW." pageId="2" pageNumber="530">Fig. 1</figureCitation>
) slender. Head dark blackish-brown. Antennae, pronotum, elytra, and legs orange to reddish-brown. Abdomen blackish-brown with posterior margins of 3rd to 7th segments dark reddish-brown. Head capsule widest behind eyes, surface finely wrinkled, moderately punctate with short setae. Eyes large and laterally prominent, length of eyes 2.08–2.17 times as long as head width. Antennae (
<figureCitation id="83BA2A40FFD72C45FB9E2BFE5AC1FC59" box="[1075,1205,982,1007]" captionStart="FIGURE 2" captionStartId="4.[151,250,1511,1533]" captionTargetBox="[227,1359,187,1467]" captionTargetId="figure@4.[151,1436,161,1489]" captionTargetPageId="4" captionText="FIGURE 2. Zyras (Diaulaconia) artemis sp. n. A, forebody, dorsal vieW; B, rigHt antenna, lateral vieW; C, ditto, dorsal vieW; D, frons of tHe female WitHout granulations; E, frons of tHe male WitH granulations." httpUri="https://zenodo.org/record/1040598/files/figure.png" pageId="2" pageNumber="530">Fig. 2B, C</figureCitation>
) slightly exceeding combined length of head and pronotum; 1 
<taxonomicName id="DC814D46FFD72C45FDC72BD25D78FBA3" box="[618,780,1018,1045]" pageId="2" pageNumber="530" rank="subSpecies" subSpecies="antennomere">st antennomere</taxonomicName>
longest, longer than 2nd and 3rd combined; 2nd antennomere shorter than 3rd and narrowest, 3rd antennomere longer and slightly narrower than 4th; 4th to 10th antennomeres similar in shape and length; 11th pointed and approximately as long as 3rd; relative lengths of antennomeres: 26:10:15:10:10:10:10:10:11:11:18. Pronotum (
<figureCitation id="83BA2A40FFD72C45FD152C475D6FFB31" box="[696,795,1135,1160]" captionStart="FIGURE 2" captionStartId="4.[151,250,1511,1533]" captionTargetBox="[227,1359,187,1467]" captionTargetId="figure@4.[151,1436,161,1489]" captionTargetPageId="4" captionText="FIGURE 2. Zyras (Diaulaconia) artemis sp. n. A, forebody, dorsal vieW; B, rigHt antenna, lateral vieW; C, ditto, dorsal vieW; D, frons of tHe female WitHout granulations; E, frons of tHe male WitH granulations." httpUri="https://zenodo.org/record/1040598/files/figure.png" pageId="2" pageNumber="530">Fig. 2A</figureCitation>
) subtrapeziform with anterior and posterior margin smoothly convex; 1.2 times as wide as long, widest just behind anterior margin, narrowed posteriorly; surface moderately punctate with short and light colour setae, with 4 black macrosetae on pronotal disc and with 40-50 black macrosetae along the whole margin of disc. Scutellum slightly wrinkled and hairy. Elytra (
<figureCitation id="83BA2A40FFD72C45FB722CF35B4CFB45" box="[1247,1336,1243,1268]" captionStart="FIGURE 2" captionStartId="4.[151,250,1511,1533]" captionTargetBox="[227,1359,187,1467]" captionTargetId="figure@4.[151,1436,161,1489]" captionTargetPageId="4" captionText="FIGURE 2. Zyras (Diaulaconia) artemis sp. n. A, forebody, dorsal vieW; B, rigHt antenna, lateral vieW; C, ditto, dorsal vieW; D, frons of tHe female WitHout granulations; E, frons of tHe male WitH granulations." httpUri="https://zenodo.org/record/1040598/files/figure.png" pageId="2" pageNumber="530">Fig. 2A</figureCitation>
) slightly widened apically, surface densely punctate with short and light coloured setae. Elytron with about 20 black macrosetae on main surface and 15 black macrosetae on the lateral margin. Legs hairy, hind tibia approximately as long as elytra; approximate relative lengths of tarsomeres: 14:14:12: 
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fore tarsi; 24:17:13:13: 
<quantity id="DC799B20FFD72C45FB4A2D6F5B53FAD6" box="[1255,1319,1351,1376]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" pageId="2" pageNumber="530" unit="in" value="27.0">27 in</quantity>
mid tarsi; 30:19:16:15: 
<quantity id="DC799B20FFD72C45FE8C2D455F2BFA30" box="[289,351,1389,1414]" metricMagnitude="-1" metricUnit="m" metricValue="7.112" pageId="2" pageNumber="530" unit="in" value="28.0">28 in</quantity>
hind tarsi. Abdomen subparallel-sided, nearly as wide as elytra; 3rd to 7th tergites glabrous except for 1–2 rows of setae along each posterior area; 9th tergite with 3 macrosetae; 10th tergite with 6–8 macrosetae.
</paragraph>
<paragraph id="1B3E36C5FFD72C45FF6A2D935AC0F94A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="530">
<emphasis id="29F5EAD7FFD72C45FF6A2D935F7FFA62" bold="true" box="[199,267,1467,1492]" pageId="2" pageNumber="530">Male.</emphasis>
Frons of head with minute granulations (
<figureCitation id="83BA2A40FFD72C45FD582D935D24FA62" box="[757,848,1467,1492]" captionStart="FIGURE 2" captionStartId="4.[151,250,1511,1533]" captionTargetBox="[227,1359,187,1467]" captionTargetId="figure@4.[151,1436,161,1489]" captionTargetPageId="4" captionText="FIGURE 2. Zyras (Diaulaconia) artemis sp. n. A, forebody, dorsal vieW; B, rigHt antenna, lateral vieW; C, ditto, dorsal vieW; D, frons of tHe female WitHout granulations; E, frons of tHe male WitH granulations." httpUri="https://zenodo.org/record/1040598/files/figure.png" pageId="2" pageNumber="530">Fig. 2E</figureCitation>
). 4th tergite with a pair of projections on posterior margin (
<figureCitation id="83BA2A40FFD72C45FF582DF75F1BFA4E" box="[245,367,1503,1528]" captionStart="FIGURE 3" captionStartId="5.[151,250,1666,1688]" captionTargetBox="[151,1436,185,1642]" captionTargetId="figure@5.[151,1436,185,1642]" captionTargetPageId="5" captionText="FIGURE 3. Zyras (Diaulaconia) artemis sp. n. A, male 4 th tergite WitH a pair of projeCtions on posterior margin, large individual, dorsal vieW; B, ditto, projeCtion bilobed apiCally, lateral vieW; C, ditto, small individual, dorsal vieW; D, ditto, projeCtion WitH no bilobed apiCally, lateral vieW; E, male 7 th tergite WitH apiCal median tuberCle, dorsal vieW; F, ditto, lateral vieW; G, female 7 th tergite WitH no small median tuberCle, dorsal vieW." httpUri="https://zenodo.org/record/1040600/files/figure.png" pageId="2" pageNumber="530">Fig. 3A, C</figureCitation>
); these projections apically bilobed in lateral view in large individuals (
<figureCitation id="83BA2A40FFD72C45FB3C2DF75A9CFA4E" box="[1169,1256,1503,1528]" captionStart="FIGURE 3" captionStartId="5.[151,250,1666,1688]" captionTargetBox="[151,1436,185,1642]" captionTargetId="figure@5.[151,1436,185,1642]" captionTargetPageId="5" captionText="FIGURE 3. Zyras (Diaulaconia) artemis sp. n. A, male 4 th tergite WitH a pair of projeCtions on posterior margin, large individual, dorsal vieW; B, ditto, projeCtion bilobed apiCally, lateral vieW; C, ditto, small individual, dorsal vieW; D, ditto, projeCtion WitH no bilobed apiCally, lateral vieW; E, male 7 th tergite WitH apiCal median tuberCle, dorsal vieW; F, ditto, lateral vieW; G, female 7 th tergite WitH no small median tuberCle, dorsal vieW." httpUri="https://zenodo.org/record/1040600/files/figure.png" pageId="2" pageNumber="530">Fig. 3B</figureCitation>
), while in small individuals can be much reduced, not bilobed apically (
<figureCitation id="83BA2A40FFD72C45FD532E2E5D2CF9A9" box="[766,856,1542,1567]" captionStart="FIGURE 3" captionStartId="5.[151,250,1666,1688]" captionTargetBox="[151,1436,185,1642]" captionTargetId="figure@5.[151,1436,185,1642]" captionTargetPageId="5" captionText="FIGURE 3. Zyras (Diaulaconia) artemis sp. n. A, male 4 th tergite WitH a pair of projeCtions on posterior margin, large individual, dorsal vieW; B, ditto, projeCtion bilobed apiCally, lateral vieW; C, ditto, small individual, dorsal vieW; D, ditto, projeCtion WitH no bilobed apiCally, lateral vieW; E, male 7 th tergite WitH apiCal median tuberCle, dorsal vieW; F, ditto, lateral vieW; G, female 7 th tergite WitH no small median tuberCle, dorsal vieW." httpUri="https://zenodo.org/record/1040600/files/figure.png" pageId="2" pageNumber="530">Fig. 3D</figureCitation>
). 7th tergite with apical median tubercle (
<figureCitation id="83BA2A40FFD72C45FAB12E2E5BFBF9A9" box="[1308,1423,1542,1567]" captionStart="FIGURE 3" captionStartId="5.[151,250,1666,1688]" captionTargetBox="[151,1436,185,1642]" captionTargetId="figure@5.[151,1436,185,1642]" captionTargetPageId="5" captionText="FIGURE 3. Zyras (Diaulaconia) artemis sp. n. A, male 4 th tergite WitH a pair of projeCtions on posterior margin, large individual, dorsal vieW; B, ditto, projeCtion bilobed apiCally, lateral vieW; C, ditto, small individual, dorsal vieW; D, ditto, projeCtion WitH no bilobed apiCally, lateral vieW; E, male 7 th tergite WitH apiCal median tuberCle, dorsal vieW; F, ditto, lateral vieW; G, female 7 th tergite WitH no small median tuberCle, dorsal vieW." httpUri="https://zenodo.org/record/1040600/files/figure.png" pageId="2" pageNumber="530">Fig. 3E, F</figureCitation>
). 8th tergite (
<figureCitation id="83BA2A40FFD72C45FEB32E055F0AF9F0" box="[286,382,1581,1606]" captionStart="FIGURE 4" captionStartId="6.[151,250,1502,1524]" captionTargetBox="[165,1419,208,1471]" captionTargetId="figure@6.[151,1436,193,1478]" captionTargetPageId="6" captionText="FIGURE 4. Zyras (Diaulaconia) artemis sp. n., male. A, eigHt abdominal tergite, dorsal vieW; B, 8 th abdominal sternite, ventral vieW; C, median lobe of aedeagus, ventral vieW; D, ditto, tHe pHotograpH; E, ditto, lateral vieW; F, ditto, tHe pHotograpH." httpUri="https://zenodo.org/record/1040602/files/figure.png" pageId="2" pageNumber="530">Fig. 4A</figureCitation>
) with posterior margin emarginate and forming pair of projections, with about 20–30 macrosetae; posterior to posterolateral margin fringed with thick and light colour sensory setae. 8th sternite (
<figureCitation id="83BA2A40FFD72C45FAC32E7C5ECDF926" captionStart="FIGURE 4" captionStartId="6.[151,250,1502,1524]" captionTargetBox="[165,1419,208,1471]" captionTargetId="figure@6.[151,1436,193,1478]" captionTargetPageId="6" captionText="FIGURE 4. Zyras (Diaulaconia) artemis sp. n., male. A, eigHt abdominal tergite, dorsal vieW; B, 8 th abdominal sternite, ventral vieW; C, median lobe of aedeagus, ventral vieW; D, ditto, tHe pHotograpH; E, ditto, lateral vieW; F, ditto, tHe pHotograpH." httpUri="https://zenodo.org/record/1040602/files/figure.png" pageId="2" pageNumber="530">Fig. 4B</figureCitation>
) with posterior margin slightly emarginate or rounded, with about 50 macrosetae; thick and light colour sensory setae fringed densely on posterior to posterolateral margin. Aedeagus (
<figureCitation id="83BA2A40FFD72C45FC1E2EB45A59F902" box="[947,1069,1692,1717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1502,1524]" captionTargetBox="[165,1419,208,1471]" captionTargetId="figure@6.[151,1436,193,1478]" captionTargetPageId="6" captionText="FIGURE 4. Zyras (Diaulaconia) artemis sp. n., male. A, eigHt abdominal tergite, dorsal vieW; B, 8 th abdominal sternite, ventral vieW; C, median lobe of aedeagus, ventral vieW; D, ditto, tHe pHotograpH; E, ditto, lateral vieW; F, ditto, tHe pHotograpH." httpUri="https://zenodo.org/record/1040602/files/figure.png" pageId="2" pageNumber="530">Fig. 4C, D</figureCitation>
) elongate pear-shaped in ventral view, pointed at apex; apical lobe with two lateral projections of internal sac, these projections are slender and curved ventrally (
<figureCitation id="83BA2A40FFD72C45FEF02ECC5FA5F94A" box="[349,465,1764,1789]" captionStart="FIGURE 4" captionStartId="6.[151,250,1502,1524]" captionTargetBox="[165,1419,208,1471]" captionTargetId="figure@6.[151,1436,193,1478]" captionTargetPageId="6" captionText="FIGURE 4. Zyras (Diaulaconia) artemis sp. n., male. A, eigHt abdominal tergite, dorsal vieW; B, 8 th abdominal sternite, ventral vieW; C, median lobe of aedeagus, ventral vieW; D, ditto, tHe pHotograpH; E, ditto, lateral vieW; F, ditto, tHe pHotograpH." httpUri="https://zenodo.org/record/1040602/files/figure.png" pageId="2" pageNumber="530">Fig. 4E, F</figureCitation>
); apical sclerite and copulatory piece of inner sac as in 
<figureCitation id="83BA2A40FFD72C45FB902ECC5AC4F94A" box="[1085,1200,1764,1789]" captionStart="FIGURE 4" captionStartId="6.[151,250,1502,1524]" captionTargetBox="[165,1419,208,1471]" captionTargetId="figure@6.[151,1436,193,1478]" captionTargetPageId="6" captionText="FIGURE 4. Zyras (Diaulaconia) artemis sp. n., male. A, eigHt abdominal tergite, dorsal vieW; B, 8 th abdominal sternite, ventral vieW; C, median lobe of aedeagus, ventral vieW; D, ditto, tHe pHotograpH; E, ditto, lateral vieW; F, ditto, tHe pHotograpH." httpUri="https://zenodo.org/record/1040602/files/figure.png" pageId="2" pageNumber="530">Fig. 4C–F</figureCitation>
.
</paragraph>
<paragraph id="1B3E36C5FFD72C44FF6A2F225D52FF4E" blockId="2.[151,1437,151,2013]" lastBlockId="3.[151,1436,151,248]" lastPageId="3" lastPageNumber="531" pageId="2" pageNumber="530">
<emphasis id="29F5EAD7FFD72C45FF6A2F225F51F895" bold="true" box="[199,293,1802,1827]" pageId="2" pageNumber="530">Female.</emphasis>
Frons of head smooth, lacking granulation (
<figureCitation id="83BA2A40FFD72C45FC8A2F225DF1F895" box="[807,901,1802,1827]" captionStart="FIGURE 2" captionStartId="4.[151,250,1511,1533]" captionTargetBox="[227,1359,187,1467]" captionTargetId="figure@4.[151,1436,161,1489]" captionTargetPageId="4" captionText="FIGURE 2. Zyras (Diaulaconia) artemis sp. n. A, forebody, dorsal vieW; B, rigHt antenna, lateral vieW; C, ditto, dorsal vieW; D, frons of tHe female WitHout granulations; E, frons of tHe male WitH granulations." httpUri="https://zenodo.org/record/1040598/files/figure.png" pageId="2" pageNumber="530">Fig. 2D</figureCitation>
). 3rd to 7th tergite simple, lacking projection or small median tubercle on posterior margin (
<figureCitation id="83BA2A40FFD72C45FDD22F195CADF8FC" box="[639,729,1841,1866]" captionStart="FIGURE 3" captionStartId="5.[151,250,1666,1688]" captionTargetBox="[151,1436,185,1642]" captionTargetId="figure@5.[151,1436,185,1642]" captionTargetPageId="5" captionText="FIGURE 3. Zyras (Diaulaconia) artemis sp. n. A, male 4 th tergite WitH a pair of projeCtions on posterior margin, large individual, dorsal vieW; B, ditto, projeCtion bilobed apiCally, lateral vieW; C, ditto, small individual, dorsal vieW; D, ditto, projeCtion WitH no bilobed apiCally, lateral vieW; E, male 7 th tergite WitH apiCal median tuberCle, dorsal vieW; F, ditto, lateral vieW; G, female 7 th tergite WitH no small median tuberCle, dorsal vieW." httpUri="https://zenodo.org/record/1040600/files/figure.png" pageId="2" pageNumber="530">Fig. 3G</figureCitation>
). 8th tergite (
<figureCitation id="83BA2A40FFD72C45FCCF2F195DCDF8FC" box="[866,953,1841,1866]" captionStart="FIGURE 5" captionStartId="7.[151,250,1790,1812]" captionTargetBox="[171,1424,516,1745]" captionTargetId="figure@7.[151,1436,481,1766]" captionTargetPageId="7" captionText="FIGURE 5. Zyras (Diaulaconia) artemis sp. n., female. A, eigHt abdominal tergite, dorsal vieW; B 8 th abdominal sternite, ventral vieW; C, spermatHeCa; D, tHe pHotograpH of tHe apiCal part and spermatHeCal gland; E, tHe pHotograpH of tHe inner Wall of apiCal 1 / 3 of tHe basal part; F, tHe base of tHe basal part and tHe spermatHeCal duCt." httpUri="https://zenodo.org/record/1040604/files/figure.png" pageId="2" pageNumber="530">Fig. 5A</figureCitation>
) similar to male, slightly shorter and wider than in male. 8th sternite (
<figureCitation id="83BA2A40FFD72C45FE1E2F705C7DF8C7" box="[435,521,1880,1905]" captionStart="FIGURE 5" captionStartId="7.[151,250,1790,1812]" captionTargetBox="[171,1424,516,1745]" captionTargetId="figure@7.[151,1436,481,1766]" captionTargetPageId="7" captionText="FIGURE 5. Zyras (Diaulaconia) artemis sp. n., female. A, eigHt abdominal tergite, dorsal vieW; B 8 th abdominal sternite, ventral vieW; C, spermatHeCa; D, tHe pHotograpH of tHe apiCal part and spermatHeCal gland; E, tHe pHotograpH of tHe inner Wall of apiCal 1 / 3 of tHe basal part; F, tHe base of tHe basal part and tHe spermatHeCal duCt." httpUri="https://zenodo.org/record/1040604/files/figure.png" pageId="2" pageNumber="530">Fig. 5B</figureCitation>
) with posterior margin rounded, with about 50 macrosetae; thick and light colour sensory setae fringed sparsely on posterior and posterolateral margin. Spermatheca (
<figureCitation id="83BA2A40FFD72C45FBC62F545A95F823" box="[1131,1249,1916,1941]" captionStart="FIGURE 5" captionStartId="7.[151,250,1790,1812]" captionTargetBox="[171,1424,516,1745]" captionTargetId="figure@7.[151,1436,481,1766]" captionTargetPageId="7" captionText="FIGURE 5. Zyras (Diaulaconia) artemis sp. n., female. A, eigHt abdominal tergite, dorsal vieW; B 8 th abdominal sternite, ventral vieW; C, spermatHeCa; D, tHe pHotograpH of tHe apiCal part and spermatHeCal gland; E, tHe pHotograpH of tHe inner Wall of apiCal 1 / 3 of tHe basal part; F, tHe base of tHe basal part and tHe spermatHeCal duCt." httpUri="https://zenodo.org/record/1040604/files/figure.png" pageId="2" pageNumber="530">Fig. 5C–F</figureCitation>
, 
<figureCitation id="83BA2A40FFD72C45FB5D2F545A8BF823" box="[1264,1279,1916,1941]" captionStart="FIGURE 6" captionStartId="8.[151,250,1936,1958]" captionTargetBox="[182,1415,745,1902]" captionTargetId="figure@8.[151,1436,736,1914]" captionTargetPageId="8" captionText="FIGURE 6. Variations of tHe spermatHeCa of Zyras (Diaulaconia) artemis sp. n. LoCality of eaCH speCimens: A, Gaupo (Taoyuan Co.); B, Hsilin trail (Hualien Co.); C, DaHasHan (Pingtung Co.); D, E, Dakeng (TaiCHung City); F, Xiaping (Nantou Co.); G, H, LuansHan (Taitung Co.); I, SHeding (Pingtung Co.); J, K, PengHu Islands (PengHu Co.)." httpUri="https://zenodo.org/record/1040606/files/figure.png" pageId="2" pageNumber="530">6</figureCitation>
) anfractuous, basal part highly variable in shape, at least 10 times as long as apical part, rounded and slightly emarginate at base (
<figureCitation id="83BA2A40FFD72C45FF322FEC5F63F86B" box="[159,279,1988,2013]" captionStart="FIGURE 5" captionStartId="7.[151,250,1790,1812]" captionTargetBox="[171,1424,516,1745]" captionTargetId="figure@7.[151,1436,481,1766]" captionTargetPageId="7" captionText="FIGURE 5. Zyras (Diaulaconia) artemis sp. n., female. A, eigHt abdominal tergite, dorsal vieW; B 8 th abdominal sternite, ventral vieW; C, spermatHeCa; D, tHe pHotograpH of tHe apiCal part and spermatHeCal gland; E, tHe pHotograpH of tHe inner Wall of apiCal 1 / 3 of tHe basal part; F, tHe base of tHe basal part and tHe spermatHeCal duCt." httpUri="https://zenodo.org/record/1040604/files/figure.png" pageId="2" pageNumber="530">Fig. 5C, F</figureCitation>
), widest at base and suddenly constricted at basal 1/3, inner wall smooth; approximate median 1/3 of basal part thinnest, anfractuous and varied in shape (
<figureCitation id="83BA2A40FFD62C44FD5A28BF5D49FF06" box="[759,829,151,176]" captionStart="FIGURE 6" captionStartId="8.[151,250,1936,1958]" captionTargetBox="[182,1415,745,1902]" captionTargetId="figure@8.[151,1436,736,1914]" captionTargetPageId="8" captionText="FIGURE 6. Variations of tHe spermatHeCa of Zyras (Diaulaconia) artemis sp. n. LoCality of eaCH speCimens: A, Gaupo (Taoyuan Co.); B, Hsilin trail (Hualien Co.); C, DaHasHan (Pingtung Co.); D, E, Dakeng (TaiCHung City); F, Xiaping (Nantou Co.); G, H, LuansHan (Taitung Co.); I, SHeding (Pingtung Co.); J, K, PengHu Islands (PengHu Co.)." httpUri="https://zenodo.org/record/1040606/files/figure.png" pageId="3" pageNumber="531">Fig. 6</figureCitation>
), inner wall smooth; apical 1/3 of basal part slightly curved, the inner wall unsmooth (
<figureCitation id="83BA2A40FFD62C44FDB728945CE1FF62" box="[538,661,188,213]" captionStart="FIGURE 5" captionStartId="7.[151,250,1790,1812]" captionTargetBox="[171,1424,516,1745]" captionTargetId="figure@7.[151,1436,481,1766]" captionTargetPageId="7" captionText="FIGURE 5. Zyras (Diaulaconia) artemis sp. n., female. A, eigHt abdominal tergite, dorsal vieW; B 8 th abdominal sternite, ventral vieW; C, spermatHeCa; D, tHe pHotograpH of tHe apiCal part and spermatHeCal gland; E, tHe pHotograpH of tHe inner Wall of apiCal 1 / 3 of tHe basal part; F, tHe base of tHe basal part and tHe spermatHeCal duCt." httpUri="https://zenodo.org/record/1040604/files/figure.png" pageId="3" pageNumber="531">Fig. 5C, E</figureCitation>
); connection of basal part and apical part bended about 90 degrees; apical part short, its inner wall curved apicad (
<figureCitation id="83BA2A40FFD62C44FD3328F75D6DFF4E" box="[670,793,223,248]" captionStart="FIGURE 5" captionStartId="7.[151,250,1790,1812]" captionTargetBox="[171,1424,516,1745]" captionTargetId="figure@7.[151,1436,481,1766]" captionTargetPageId="7" captionText="FIGURE 5. Zyras (Diaulaconia) artemis sp. n., female. A, eigHt abdominal tergite, dorsal vieW; B 8 th abdominal sternite, ventral vieW; C, spermatHeCa; D, tHe pHotograpH of tHe apiCal part and spermatHeCal gland; E, tHe pHotograpH of tHe inner Wall of apiCal 1 / 3 of tHe basal part; F, tHe base of tHe basal part and tHe spermatHeCal duCt." httpUri="https://zenodo.org/record/1040604/files/figure.png" pageId="3" pageNumber="531">Fig. 5C, D</figureCitation>
).
</paragraph>
<caption id="4FFE664DFFD62C44FF3A2FF15DC4F859" box="[151,944,2009,2031]" pageId="3" pageNumber="531">
<paragraph id="1B3E36C5FFD62C44FF3A2FF15DC4F859" blockId="3.[151,944,2009,2031]" box="[151,944,2009,2031]" pageId="3" pageNumber="531">
<emphasis id="29F5EAD7FFD62C44FF3A2FF15F67F858" bold="true" box="[151,275,2009,2031]" pageId="3" pageNumber="531">FIGURE 1.</emphasis>
Habitus of 
<taxonomicName id="DC814D46FFD62C44FE242FF25CDFF859" authority="Liang &amp; Maruyama &amp; Li, 2017" authorityName="Liang &amp; Maruyama &amp; Li" authorityYear="2017" box="[393,683,2009,2031]" class="Insecta" family="Staphylinidae" genus="Zyras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="531" phylum="Arthropoda" rank="species" species="artemis" status="sp. nov." subGenus="Diaulaconia">
<emphasis id="29F5EAD7FFD62C44FE242FF25FB7F859" box="[393,451,2010,2031]" italics="true" pageId="3" pageNumber="531">Zyras</emphasis>
(
<emphasis id="29F5EAD7FFD62C44FE7F2FF15C25F859" box="[466,593,2009,2031]" italics="true" pageId="3" pageNumber="531">Diaulaconia</emphasis>
) 
<emphasis id="29F5EAD7FFD62C44FDF22FF25CDFF859" box="[607,683,2010,2031]" italics="true" pageId="3" pageNumber="531">artemis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32C657ACFFD62C44FD1C2FF25C9EF859" box="[689,746,2010,2031]" pageId="3" pageNumber="531" rank="species">
<emphasis id="29F5EAD7FFD62C44FD1C2FF25C9EF859" bold="true" box="[689,746,2010,2031]" pageId="3" pageNumber="531">sp. n.</emphasis>
</taxonomicNameLabel>
, male, dorsal vieW.
</paragraph>
</caption>
<caption id="4FFE664DFFD12C43FF3A2DCF5D95F9AA" httpUri="https://zenodo.org/record/1040598/files/figure.png" pageId="4" pageNumber="532" targetBox="[227,1359,187,1467]" targetPageId="4">
<paragraph id="1B3E36C5FFD12C43FF3A2DCF5D95F9AA" blockId="4.[151,1436,1511,1564]" pageId="4" pageNumber="532">
<emphasis id="29F5EAD7FFD12C43FF3A2DCF5F67FA4A" bold="true" box="[151,275,1511,1533]" pageId="4" pageNumber="532">FIGURE 2.</emphasis>
<taxonomicName id="DC814D46FFD12C43FEB72DC05C49FA4B" authority="Liang &amp; Maruyama &amp; Li, 2017" authorityName="Liang &amp; Maruyama &amp; Li" authorityYear="2017" box="[282,573,1511,1533]" class="Insecta" family="Staphylinidae" genus="Zyras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="species" species="artemis" status="sp. nov." subGenus="Diaulaconia">
<emphasis id="29F5EAD7FFD12C43FEB72DC05F20FA4B" box="[282,340,1512,1533]" italics="true" pageId="4" pageNumber="532">Zyras</emphasis>
(
<emphasis id="29F5EAD7FFD12C43FECE2DCF5F96FA4B" box="[355,482,1511,1533]" italics="true" pageId="4" pageNumber="532">Diaulaconia</emphasis>
) 
<emphasis id="29F5EAD7FFD12C43FE5C2DC05C49FA4B" box="[497,573,1512,1533]" italics="true" pageId="4" pageNumber="532">artemis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32C657ACFFD12C43FDE92DCF5C09FA4A" box="[580,637,1511,1532]" pageId="4" pageNumber="532" rank="species">
<emphasis id="29F5EAD7FFD12C43FDE92DCF5C09FA4A" bold="true" box="[580,637,1511,1532]" pageId="4" pageNumber="532">sp. n.</emphasis>
</taxonomicNameLabel>
A, forebody, dorsal vieW; B, rigHt antenna, lateral vieW; C, ditto, dorsal vieW; D, frons of tHe female WitHout granulations; E, frons of tHe male WitH granulations.
</paragraph>
</caption>
<paragraph id="1B3E36C5FFD12C43FF6A2E605AF1F933" blockId="4.[151,1437,1608,2032]" pageId="4" pageNumber="532">
<emphasis id="29F5EAD7FFD12C43FF6A2E605F09F9D7" bold="true" box="[199,381,1608,1633]" pageId="4" pageNumber="532">Measurements.</emphasis>
BL ≈ 6.9–8.8; FBL ≈ 1.7–2.1; HL: 0.84–0.93; HW: 1.18–1.35; EL: 0.55–0.63; AL: 2.20–2.59; PL: 1.18–1.30; PW: 1.38–1.57; ELL: 1.43–1.61; ELW: 1.72–2.07; HTL: 1.23–1.49 (mm).
</paragraph>
</subSubSection>
<subSubSection id="539B654EFFD12C43FF6A2EB85D82F942" pageId="4" pageNumber="532" type="diagnosis">
<paragraph id="1B3E36C5FFD12C43FF6A2EB85D82F942" blockId="4.[151,1437,1608,2032]" pageId="4" pageNumber="532">
<emphasis id="29F5EAD7FFD12C43FF6A2EB85F35F91F" bold="true" box="[199,321,1680,1705]" pageId="4" pageNumber="532">Diagnosis.</emphasis>
<taxonomicName id="DC814D46FFD12C43FEEA2EBA5F96F91F" authority="Liang &amp; Maruyama &amp; Li, 2017" authorityName="Liang &amp; Maruyama &amp; Li" authorityYear="2017" box="[327,482,1682,1705]" class="Insecta" family="Staphylinidae" genus="Zyras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="species" species="artemis" status="sp. nov.">
<emphasis id="29F5EAD7FFD12C43FEEA2EBA5F96F91F" box="[327,482,1682,1705]" italics="true" pageId="4" pageNumber="532">Zyras artemis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32C657ACFFD12C43FE442EB85C5CF91F" box="[489,552,1680,1705]" pageId="4" pageNumber="532" rank="species">
<emphasis id="29F5EAD7FFD12C43FE442EB85C5CF91F" bold="true" box="[489,552,1680,1705]" pageId="4" pageNumber="532">sp. n.</emphasis>
</taxonomicNameLabel>
is distinguishable from all other 
<taxonomicName id="DC814D46FFD12C43FC3A2EBA5A08F91F" authority="(Diaulaconia)" authorityName="Stephens" authorityYear="1835" baseAuthorityName="Diaulaconia" box="[919,1148,1680,1705]" class="Insecta" family="Staphylinidae" genus="Zyras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="genus">
<emphasis id="29F5EAD7FFD12C43FC3A2EBA5DA3F91F" box="[919,983,1682,1705]" italics="true" pageId="4" pageNumber="532">Zyras</emphasis>
(
<emphasis id="29F5EAD7FFD12C43FC4A2EB85A07F91F" box="[999,1139,1680,1705]" italics="true" pageId="4" pageNumber="532">Diaulaconia</emphasis>
)
</taxonomicName>
species by its remarkable shape of the spermatheca, which basal part anfractuous diversely and at least 10 times as long as apical part, as well as by the shape of the aedeagus and unique projections of 4th tergite in males.
</paragraph>
</subSubSection>
<subSubSection id="539B654EFFD12C43FF6A2ED75D54F846" pageId="4" pageNumber="532" type="etymology">
<paragraph id="1B3E36C5FFD12C43FF6A2ED75CB4F832" blockId="4.[151,1437,1608,2032]" pageId="4" pageNumber="532">
<emphasis id="29F5EAD7FFD12C43FF6A2ED75F3AF8AE" bold="true" box="[199,334,1791,1816]" pageId="4" pageNumber="532">Etymology.</emphasis>
Artemis is the ancient Greek goddess of hunting, wildlife, and childbirth. She is currently seen as the archetype of feminists (
<bibRefCitation id="7F104B34FFD12C43FE792F0B5C13F88A" author="Bolen" box="[468,615,1827,1852]" pageId="4" pageNumber="532" refString="Bolen, J. S. (2014) Artemis: The indomitable spirit in everywoman. Conari Press, San FranCisCo, xix + 253 pp." type="book" year="2014">Bolen, 2014</bibRefCitation>
). The new species is named after Artemis in reference to its predatory behavior, extraordinary reproductive strategy, and in honor of Taiwan’s great progress in gender equality, being the first Asian country legalizing same-sex marriage.
</paragraph>
<paragraph id="1B3E36C5FFD12C43FF6A2FA75D54F846" blockId="4.[151,1437,1608,2032]" pageId="4" pageNumber="532">
<emphasis id="29F5EAD7FFD12C43FF6A2FA75FFEF81E" bold="true" box="[199,394,1935,1960]" pageId="4" pageNumber="532">Biological notes.</emphasis>
<taxonomicName id="DC814D46FFD12C43FE3F2FB85C5AF811" authority="Liang &amp; Maruyama &amp; Li, 2017" authorityName="Liang &amp; Maruyama &amp; Li" authorityYear="2017" box="[402,558,1936,1959]" class="Insecta" family="Staphylinidae" genus="Zyras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="species" species="artemis" status="sp. nov.">
<emphasis id="29F5EAD7FFD12C43FE3F2FB85C5AF811" box="[402,558,1936,1959]" italics="true" pageId="4" pageNumber="532">Zyras artemis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32C657ACFFD12C43FD982FA75C02F81E" box="[565,630,1935,1960]" pageId="4" pageNumber="532" rank="species">
<emphasis id="29F5EAD7FFD12C43FD982FA75C02F81E" bold="true" box="[565,630,1935,1960]" pageId="4" pageNumber="532">sp. n.</emphasis>
</taxonomicNameLabel>
is found to be the obligatory predator of 
<taxonomicName id="DC814D46FFD12C43FBE62FB85A85F81E" baseAuthorityName="Shiraki" baseAuthorityYear="1909" box="[1099,1265,1936,1960]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="29F5EAD7FFD12C43FBE62FB85A85F81E" box="[1099,1265,1936,1960]" italics="true" pageId="4" pageNumber="532">O. formosanus</emphasis>
</taxonomicName>
in 
<collectingCountry id="63967655FFD12C43FABB2FA75B18F81E" box="[1302,1388,1935,1960]" name="Taiwan" pageId="4" pageNumber="532">Taiwan</collectingCountry>
. We also observed its complete life cycle in the laboratory, and confirm its non-feeding larval development. The detail information will be published in other subsequent studies.
</paragraph>
</subSubSection>
<caption id="4FFE664DFFD02C42FF3A2EAA5D2FF94F" httpUri="https://zenodo.org/record/1040600/files/figure.png" pageId="5" pageNumber="533" targetBox="[151,1436,185,1642]" targetPageId="5">
<paragraph id="1B3E36C5FFD02C42FF3A2EAA5D2FF94F" blockId="5.[151,1436,1663,1785]" pageId="5" pageNumber="533">
<emphasis id="29F5EAD7FFD02C42FF3A2EAA5F65F921" bold="true" box="[151,273,1666,1688]" pageId="5" pageNumber="533">FIGURE 3</emphasis>
. 
<taxonomicName id="DC814D46FFD02C42FE8F2EAB5C39F92E" authority="Liang &amp; Maruyama &amp; Li, 2017" authorityName="Liang &amp; Maruyama &amp; Li" authorityYear="2017" box="[290,589,1666,1688]" class="Insecta" family="Staphylinidae" genus="Zyras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="533" phylum="Arthropoda" rank="species" species="artemis" status="sp. nov." subGenus="Diaulaconia">
<emphasis id="29F5EAD7FFD02C42FE8F2EAB5F28F92E" box="[290,348,1667,1688]" italics="true" pageId="5" pageNumber="533">Zyras</emphasis>
(
<emphasis id="29F5EAD7FFD02C42FEC22EAA5F9AF92E" box="[367,494,1666,1688]" italics="true" pageId="5" pageNumber="533">Diaulaconia</emphasis>
) 
<emphasis id="29F5EAD7FFD02C42FDAC2EAB5C39F92E" box="[513,589,1667,1688]" italics="true" pageId="5" pageNumber="533">artemis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32C657ACFFD02C42FDFA2EAA5CE1F921" box="[599,661,1666,1687]" pageId="5" pageNumber="533" rank="species">
<emphasis id="29F5EAD7FFD02C42FDFA2EAA5CE1F921" bold="true" box="[599,661,1666,1687]" pageId="5" pageNumber="533">sp. n.</emphasis>
</taxonomicNameLabel>
A, male 4th tergite WitH a pair of projeCtions on posterior margin, large individual, dorsal vieW; B, ditto, projeCtion bilobed apiCally, lateral vieW; C, ditto, small individual, dorsal vieW; D, ditto, projeCtion WitH no bilobed apiCally, lateral vieW; E, male 7th tergite WitH apiCal median tuberCle, dorsal vieW; F, ditto, lateral vieW; G, female 7th tergite WitH no small median tuberCle, dorsal vieW.
</paragraph>
</caption>
<subSubSection id="539B654EFFD02C42FF6A2F0D5DAFF845" pageId="5" pageNumber="533" type="distribution">
<paragraph id="1B3E36C5FFD02C42FF6A2F0D5DAFF845" blockId="5.[151,1436,1829,2035]" pageId="5" pageNumber="533">
<emphasis id="29F5EAD7FFD02C42FF6A2F0D5F14F888" bold="true" box="[199,352,1829,1854]" pageId="5" pageNumber="533">Distribution.</emphasis>
<taxonomicName id="DC814D46FFD02C42FEC72F0F5C73F888" authority="Liang &amp; Maruyama &amp; Li, 2017" authorityName="Liang &amp; Maruyama &amp; Li" authorityYear="2017" box="[362,519,1831,1854]" class="Insecta" family="Staphylinidae" genus="Zyras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="533" phylum="Arthropoda" rank="species" species="artemis" status="sp. nov.">
<emphasis id="29F5EAD7FFD02C42FEC72F0F5C73F888" box="[362,519,1831,1854]" italics="true" pageId="5" pageNumber="533">Zyras artemis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32C657ACFFD02C42FDBC2F0D5C27F888" box="[529,595,1829,1854]" pageId="5" pageNumber="533" rank="species">
<emphasis id="29F5EAD7FFD02C42FDBC2F0D5C27F888" bold="true" box="[529,595,1829,1854]" pageId="5" pageNumber="533">sp. n.</emphasis>
</taxonomicNameLabel>
is commonly distributed in low altitude from 
<quantity id="DC799B20FFD02C42FBC62F0E5A96F888" box="[1131,1250,1829,1854]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="6.0" metricValueMin="0.0" pageId="5" pageNumber="533" unit="m" value="300.0" valueMax="600.0" valueMin="0.0">0 to 600m</quantity>
around 
<collectingCountry id="63967655FFD02C42FAEE2F0D5BECF888" box="[1347,1432,1829,1854]" name="Taiwan" pageId="5" pageNumber="533">Taiwan</collectingCountry>
, where it is matching the main habitat of its host 
<taxonomicName id="DC814D46FFD02C42FD642F635D05F8D4" baseAuthorityName="Shiraki" baseAuthorityYear="1909" box="[713,881,1866,1890]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="5" pageNumber="533" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="29F5EAD7FFD02C42FD642F635D05F8D4" box="[713,881,1866,1890]" italics="true" pageId="5" pageNumber="533">O. formosanus</emphasis>
</taxonomicName>
.This distribution pattern can be referred to their obligatory relationship, which 
<taxonomicName id="DC814D46FFD02C42FE5E2F465CEEF830" baseAuthorityName="Shiraki" baseAuthorityYear="1909" box="[499,666,1902,1926]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="5" pageNumber="533" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="29F5EAD7FFD02C42FE5E2F465CEEF830" box="[499,666,1902,1926]" italics="true" pageId="5" pageNumber="533">O. formosanus</emphasis>
</taxonomicName>
as the only or main food resource for 
<taxonomicName id="DC814D46FFD02C42FBF82F475ABCF830" authorityName="Liang &amp; Maruyama &amp; Li" authorityYear="2017" box="[1109,1224,1903,1926]" class="Insecta" family="Staphylinidae" genus="Zyras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="533" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="29F5EAD7FFD02C42FBF82F475ABCF830" box="[1109,1224,1903,1926]" italics="true" pageId="5" pageNumber="533">Z. artemis</emphasis>
</taxonomicName>
. There are several small islands around 
<collectingCountry id="63967655FFD02C42FE242FBA5FAAF81D" box="[393,478,1938,1963]" name="Taiwan" pageId="5" pageNumber="533">Taiwan</collectingCountry>
, e.g., Lanyu Island, Green Island, and 
<collectingRegion id="D945F827FFD02C42FC342FBA5D99F81D" box="[921,1005,1938,1963]" country="Taiwan" name="Penghu" pageId="5" pageNumber="533">Penghu</collectingRegion>
Islands. Up to the present, 
<taxonomicName id="DC814D46FFD02C42FA842FBB5BE8F81C" authorityName="Liang &amp; Maruyama &amp; Li" authorityYear="2017" box="[1321,1436,1939,1962]" class="Insecta" family="Staphylinidae" genus="Zyras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="533" phylum="Arthropoda" rank="species" species="artemis">
<emphasis id="29F5EAD7FFD02C42FA842FBB5BE8F81C" box="[1321,1436,1939,1962]" italics="true" pageId="5" pageNumber="533">Z. artemis</emphasis>
</taxonomicName>
has been only recorded in 
<collectingRegion id="D945F827FFD02C42FE662F9D5C6BF878" box="[459,543,1973,1998]" country="Taiwan" name="Penghu" pageId="5" pageNumber="533">Penghu</collectingRegion>
Islands, where 
<taxonomicName id="DC814D46FFD02C42FD7A2F9E5D0BF878" baseAuthorityName="Shiraki" baseAuthorityYear="1909" box="[727,895,1974,1998]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="5" pageNumber="533" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="29F5EAD7FFD02C42FD7A2F9E5D0BF878" box="[727,895,1974,1998]" italics="true" pageId="5" pageNumber="533">O. formosanus</emphasis>
</taxonomicName>
is commonly found, but so far it has not been found on the other two islands, where 
<taxonomicName id="DC814D46FFD02C42FDE92FF35C9DF844" baseAuthorityName="Shiraki" baseAuthorityYear="1909" box="[580,745,2010,2034]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="5" pageNumber="533" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="29F5EAD7FFD02C42FDE92FF35C9DF844" box="[580,745,2010,2034]" italics="true" pageId="5" pageNumber="533">O. formosanus</emphasis>
</taxonomicName>
has been also absent.
</paragraph>
</subSubSection>
</treatment>
</document>