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<document id="7783ECC4AD50BF88739A4B46A3EBABE5" ID-CLB-Dataset="27799" ID-DOI="10.11646/zootaxa.4531.1.1" ID-GBIF-Dataset="5002a2e0-25ad-4052-b7a8-04b44b755507" ID-ISSN="1175-5326" ID-Zenodo-Dep="2614310" ID-ZooBank="2BA3FB6A-EB9F-4EC6-B2D8-5D638C43E7C0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553786401115" checkinUser="plazi" docAuthor="Gómez-Zurita, Jesús" docDate="2018" docId="8670B06C7766FF8FB2F0FF15FEB6FAF2" docLanguage="en" docName="zootaxa.4531.1.1.pdf" docOrigin="Zootaxa 4531 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calligrapha femorata Jacoby J. Gomez-Zurita 1891" docType="treatment" docVersion="8" lastPageNumber="17" masterDocId="7A49C8147768FF9FB267FF82FFFAFF92" masterDocTitle="Systematic revision of Central American Calligrapha Chevrolat of the subgenus Erythrographa subgen. nov. (Coleoptera: Chrysomelidae, Chrysomelinae)" masterLastPageNumber="58" masterPageNumber="1" pageNumber="15" updateTime="1698714807028" updateUser="ExternalLinkService">
<mods:mods id="598225519427D55BC1964A2D6FA53108" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A0634D877B0B1B48B8A952A7ED365B0E">Systematic revision of Central American Calligrapha Chevrolat of the subgenus Erythrographa subgen. nov. (Coleoptera: Chrysomelidae, Chrysomelinae)</mods:title>
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<mods:namePart id="D942000481F724B1C7EAA946025E3AFE">Gómez-Zurita, Jesús</mods:namePart>
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<mods:title id="AE56147005EBA8A07742AFDE12BF61D5">Zootaxa</mods:title>
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<mods:date id="428F3EAB2188DB206FD0B90CEEF7A64A">2018</mods:date>
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<treatment id="8670B06C7766FF8FB2F0FF15FEB6FAF2" ID-DOI="http://doi.org/10.5281/zenodo.5957614" ID-GBIF-Taxon="154520700" ID-Zenodo-Dep="5957614" LSID="urn:lsid:plazi:treatment:8670B06C7766FF8FB2F0FF15FEB6FAF2" httpUri="http://treatment.plazi.org/id/8670B06C7766FF8FB2F0FF15FEB6FAF2" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
<subSubSection id="46C352F17766FF91B2F0FF15FDB9FF20" box="[151,579,151,178]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="0E66017A7766FF91B2F0FF15FDB9FF20" blockId="14.[151,579,151,211]" box="[151,579,151,178]" pageId="14" pageNumber="15">
<heading id="552EB6167766FF91B2F0FF15FDB9FF20" bold="true" box="[151,579,151,178]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="C9D97AF97766FF91B2F0FF15FDB9FF20" authority="Jacoby, 1891" authorityName="Jacoby J. Gomez-Zurita" authorityYear="1891" box="[151,579,151,178]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687766FF91B2F0FF15FDB9FF20" bold="true" box="[151,579,151,178]" pageId="14" pageNumber="15">
<emphasis id="3CADDD687766FF91B2F0FF15FE66FF23" bold="true" box="[151,412,151,177]" italics="true" pageId="14" pageNumber="15">Calligrapha femorata</emphasis>
Jacoby, 1891
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="46C352F17766FF91B2F0FF38FC11FE81" pageId="14" pageNumber="15" type="description">
<paragraph id="0E66017A7766FF91B2F0FF38FEEDFF41" blockId="14.[151,579,151,211]" box="[151,279,186,211]" pageId="14" pageNumber="15">
(
<figureCitation id="96E21DFF7766FF91B2F8FF38FF0EFF41" box="[159,244,186,211]" captionStart="FIGURE 4" captionStartId="15.[151,250,1123,1145]" captionTargetBox="[158,1431,203,1102]" captionTargetId="figure@15.[150,1436,193,1126]" captionTargetPageId="15" captionText="FIGURE 4. Dorsal and lateral views of the holotypes of Calligrapha femorata Jacoby (a) and C. synthesys sp. nov. (b)." httpUri="https://zenodo.org/record/2614330/files/figure.png" pageId="14" pageNumber="15">Figs 4a</figureCitation>
,
<figureCitation id="96E21DFF7766FF91B367FF38FEF4FF41" box="[256,270,186,211]" captionStart="FIGURE 5" captionStartId="17.[151,250,1182,1204]" captionTargetBox="[150,1436,262,1160]" captionTargetId="figure@17.[150,1436,262,1160]" captionTargetPageId="17" captionText="FIGURE 5. Distribution of Calligrapha femorata Jacoby and C. synthesys sp. nov." httpUri="https://zenodo.org/record/2614332/files/figure.png" pageId="14" pageNumber="15">5</figureCitation>
)
</paragraph>
<paragraph id="0E66017A7766FF91B2F0FF7FFC11FE81" blockId="14.[151,1003,253,398]" box="[151,1003,253,275]" pageId="14" pageNumber="15">
<taxonomicName id="C9D97AF97766FF91B2F0FF7FFDF8FE81" authority="Jacoby, 1891" authorityName="Jacoby J. Gomez-Zurita" authorityYear="1891" box="[151,514,253,275]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687766FF91B2F0FF7FFE8FFE81" box="[151,373,253,275]" italics="true" pageId="14" pageNumber="15">Calligrapha femorata</emphasis>
Jacoby, 1891
</taxonomicName>
. Biol. Centr.-Amer., vol. vi, pt. 1, suppl., p. 247.
</paragraph>
</subSubSection>
<subSubSection id="46C352F17766FF91B2F0FE9EFE55FDC1" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="0E66017A7766FF91B2F0FE9EFD41FEA0" blockId="14.[151,1003,253,398]" box="[151,699,284,306]" pageId="14" pageNumber="15">
<taxonomicName id="C9D97AF97766FF91B2F0FE9EFE1FFEA0" authority=": Weise, 1916" authorityName="Weise" authorityYear="1916" box="[151,485,284,306]" class="Insecta" family="Chrysomelidae" genus="Polyspila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687766FF91B2F0FE9EFEA2FEA0" box="[151,344,284,306]" italics="true" pageId="14" pageNumber="15">Polyspila femorata</emphasis>
:
<bibRefCitation id="6A487C8B7766FF91B302FE9EFE1FFEA0" author="Weise, J." box="[357,485,284,306]" pageId="14" pageNumber="15" refId="ref58324" refString="Weise, J. (1916) Coleopterorum Catalogus. Pars 68. Chrysomelidae: 12. Chrysomelinae. W. Junk, Berlin, 255 pp." type="book" year="1916">Weise, 1916</bibRefCitation>
</taxonomicName>
. Coleopt. Cat., p. 39.
</paragraph>
<paragraph id="0E66017A7766FF91B2F0FEB8FC5EFEC2" blockId="14.[151,1003,253,398]" box="[151,932,314,336]" pageId="14" pageNumber="15">
<taxonomicName id="C9D97AF97766FF91B2F0FEB8FDBFFEC2" authority=": Blackwelder, 1946" authorityName="Blackwelder" authorityYear="1946" box="[151,581,314,336]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687766FF91B2F0FEB8FE86FEC2" box="[151,380,314,336]" italics="true" pageId="14" pageNumber="15">Calligrapha femorata:</emphasis>
<bibRefCitation id="6A487C8B7766FF91B3E4FEB8FDBFFEC2" author="Blackwelder, R. E." box="[387,581,314,336]" pageId="14" pageNumber="15" pagination="551 - 763" refId="ref56857" refString="Blackwelder, R. E. (1946) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and Suth America. Part 4. United States National Museum Bulletin, 185, 551 - 763." type="journal article" year="1946">Blackwelder, 1946</bibRefCitation>
</taxonomicName>
.
<collectingCountry id="76CE41EA7766FF91B029FEB8FD81FEC2" box="[590,635,314,336]" name="United States of America" pageId="14" pageNumber="15">U.S.</collectingCountry>
Natl. Mus. Bull. 185, p. 674.
</paragraph>
<paragraph id="0E66017A7766FF91B2F0FEDBFCA9FEFD" blockId="14.[151,1003,253,398]" box="[151,851,345,367]" pageId="14" pageNumber="15">
<taxonomicName id="C9D97AF97766FF91B2F0FEDBFDF4FEFD" authority=": Wilcox, 1975" authorityName="Wilcox" authorityYear="1975" box="[151,526,345,367]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687766FF91B2F0FEDBFE8EFEFD" box="[151,372,345,367]" italics="true" pageId="14" pageNumber="15">Calligrapha femorata</emphasis>
:
<bibRefCitation id="6A487C8B7766FF91B3E5FEDBFDF4FEFD" author="Wilcox, J. A." box="[386,526,345,367]" pageId="14" pageNumber="15" refId="ref58390" refString="Wilcox, J. A. (1975) Checklist of the beetles of Canada, United States, Mexico, Central America and the West Indies. Vol. 1. Pt. 7. The leaf beetles. Biological Research Institute of America, New York, 166 pp. [red version]" type="book" year="1975">Wilcox, 1975</bibRefCitation>
</taxonomicName>
. Checklist Chryomelidae, p. 66.
</paragraph>
<paragraph id="0E66017A7766FF91B2F0FEFAFC31FE1C" blockId="14.[151,1003,253,398]" box="[151,971,376,398]" pageId="14" pageNumber="15">
<taxonomicName id="C9D97AF97766FF91B2F0FEFAFD6CFE1C" authority=": Benitez-Garcia et al., 2017" authorityName="Benitez-Garcia" authorityYear="2017" box="[151,662,376,398]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687766FF91B2F0FEFAFE8EFE1C" box="[151,372,376,398]" italics="true" pageId="14" pageNumber="15">Calligrapha femorata</emphasis>
:
<bibRefCitation id="6A487C8B7766FF91B3E5FEFAFD6CFE1C" author="Benitez-Garcia, B. &amp; Lopez-Perez, S. &amp; Zaragoza-Caballero, S." box="[386,662,376,398]" pageId="14" pageNumber="15" pagination="335 - 348" refId="ref56802" refString="Benitez-Garcia, B., Lopez-Perez, S. &amp; Zaragoza-Caballero, S. (2017) Sinopsis de los generos mexicanos de Chrysomelinae (Coleoptera: Chrysomelidae). Revista Mexicana de Biodiversidad, 88, 335 - 348. https: // doi. org / 10.1016 / j. rmb. 2017.03.026" type="journal article" year="2017">
Benítez-García
<emphasis id="3CADDD687766FF91B046FEFBFDB4FE1C" box="[545,590,376,398]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
., 2017
</bibRefCitation>
</taxonomicName>
. Rev. Mex. Biodiv. 88, p. 339.
</paragraph>
<paragraph id="0E66017A7766FF91B2F0FE37FE55FDC1" blockId="14.[151,1437,437,2029]" pageId="14" pageNumber="15">
This species was described based on a single specimen, as explicitly mentioned in the original description, published in the
<date id="7A6727BA7766FF91B30EFE54FDECFE7D" box="[361,534,470,495]" pageId="14" pageNumber="15" value="1891-02">February 1891</date>
supplement of the
<emphasis id="3CADDD687766FF91B16BFE54FBA4FE7D" box="[780,1118,470,495]" italics="true" pageId="14" pageNumber="15">Biologia Centrali-Americana</emphasis>
(Jacoby
<date id="7A6727BA7766FF91B6B4FE55FAACFE7D" box="[1235,1366,471,495]" pageId="14" pageNumber="15" value="1888" valueMax="1892">1888–1892</date>
). The information provided in the description fits in every detail the only specimen originally labelled as
<taxonomicName id="C9D97AF97766FF91B774FE75FF06FDA3" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687766FF91B774FE75FF06FDA3" italics="true" pageId="14" pageNumber="15">Calligrapha femorata</emphasis>
</taxonomicName>
in the beetle collection of the Natural History Museum (London), and this is recognized as Jacoby's
<typeStatus id="D162BFD87766FF91B2F0FDB8FF03FDC1" box="[151,249,570,595]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
for this species.
</paragraph>
</subSubSection>
<subSubSection id="46C352F17766FF90B2A0FDDEFC1EF84E" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="0E66017A7766FF91B2A0FDDEFCDFFD25" blockId="14.[151,1437,437,2029]" pageId="14" pageNumber="15">
<materialsCitation id="BEB10B277766FF91B2A0FDDEFCDFFD25" ID-GBIF-Occurrence="2114053526" collectionCode="NHM" collectorName="H. H. Smith" location="Guerrero" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Guerrero" typeStatus="holotype">
<typeStatus id="D162BFD87766FF91B2A0FDDEFEC8FDE7" box="[199,306,604,629]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
, by original designation: Xautipa,
<collectingRegion id="CC1DCF987766FF91B0D2FDDEFCE7FDE6" box="[693,797,604,628]" country="Mexico" name="Guerrero" pageId="14" pageNumber="15">Guerrero</collectingRegion>
,
<collectorName id="A32C64AC7766FF91B14FFDDFFC53FDE7" box="[808,937,604,629]" pageId="14" pageNumber="15">H.H. Smith</collectorName>
/
<emphasis id="3CADDD687766FF91B1D9FDDEFBE9FDE7" box="[958,1043,604,629]" italics="true" pageId="14" pageNumber="15">Calligr.</emphasis>
<taxonomicName id="C9D97AF97766FF91B67DFDDEFB4BFDE6" authority="Jac." authorityName="Jacoby J. Gomez-Zurita" authorityYear="1891" box="[1050,1201,604,628]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687766FF91B67DFDDEFB85FDE6" box="[1050,1151,604,628]" italics="true" pageId="14" pageNumber="15">femorata</emphasis>
Jac.
</taxonomicName>
[blue] / Sp. figured / Godman-Salvin Coll., Biol. Centr.-Amer. (
<collectionCode id="68C899BF7766FF91B01FFDFCFD45FD04" box="[632,703,638,662]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="14" pageNumber="15" type="Herbarium">NHM</collectionCode>
). Only two basal antennomeres of right antennae remaining, and right meso-onychium and left metatarsus are also missing.
</materialsCitation>
</paragraph>
<paragraph id="0E66017A7766FF91B2A0FD3DFE07FCED" blockId="14.[151,1437,437,2029]" pageId="14" pageNumber="15">
Habitus (
<figureCitation id="96E21DFF7766FF91B357FD3DFE7FFD4A" box="[304,389,703,728]" captionStart="FIGURE 4" captionStartId="15.[151,250,1123,1145]" captionTargetBox="[158,1431,203,1102]" captionTargetId="figure@15.[150,1436,193,1126]" captionTargetPageId="15" captionText="FIGURE 4. Dorsal and lateral views of the holotypes of Calligrapha femorata Jacoby (a) and C. synthesys sp. nov. (b)." httpUri="https://zenodo.org/record/2614330/files/figure.png" pageId="14" pageNumber="15">Fig. 4a</figureCitation>
): Length:
<quantity id="C921AC9F7766FF91B39BFD42FD9EFD45" box="[508,612,704,728]" metricMagnitude="-3" metricUnit="m" metricValue="8.38" pageId="14" pageNumber="15" unit="mm" value="8.38">8.38 mm</quantity>
, width:
<quantity id="C921AC9F7766FF91B0A4FD3DFCD7FD45" box="[707,813,703,728]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="14" pageNumber="15" unit="mm" value="5.4">5.40 mm</quantity>
. Body ovoid, moderately convex. Head and pronotum dark metallic green; scutellum, dark markings of elytra, epipleura, and apex of mandibes very dark brown, almost black with slight bronzy reflection; ventral surfaces and large frontal and posterior femoral spots (occupying basal 2/3 of femora) black with bluish-greenish metallic reflections; apical margin of clypeus, labrum, mandibles, antennae, mouth segments, ground of femora, tibiae, tarsi and fine margins of abdominal ventrites reddish orange; pale areas of elytra dark yellow.
</paragraph>
<paragraph id="0E66017A7766FF90B2A0FC05FC1EF84E" blockId="14.[151,1437,437,2029]" lastBlockId="15.[151,1437,1187,2012]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Head large, deeply inserted in pronotum, rather strongly densely punctured on frons, around frontal and clypeal suture, above antennal calli and around eyes; supraocular furrow as wide impressed region above upper margin of eye. Clypeus punctured apically and at sides. Labrum long (W/L = 1.98), smooth, shiny, with sides regularly round and weakly emarginate anteriorly. Mandibles strong, long, protruding twice length of labrum, weakly concave at sides basally, with few large, deep punctures at sides, umpunctured and shiny frontally. Last palpomere of maxillary palpi strongly dilated apically, apex cut almost straight and sides regularly curved towards narrowed base; previous palpomere as broad at apex as base of last palpomere, markedly narrowing externally towards base; first maxillary palpomere club-shaped. Antennae slender, weakly clavate, reaching behind humeri; scape strongly dilated from very narrow base; pedicel small, as short club, half as long as scape; third antennomere slender, slightly clavate, slightly longer than scape; fourth antennomere slender, 1.5x as long as second; antennomeres 4–6 progressively shorter and more rugose, with sixth antennomere slightly longer than second; antennomeres 7–11 slightly more incrassate but slender, with surface finely rugose and densely pubescent; seventh antennomere as long as fifth, eighth and ninth subequal, slightly longer than fourth, and tenth and eleventh progressively longer, with eleventh as long as third; eighth antennomere about 1.5x as long as wide at apex. Pronotum subtrapezoidal, transverse (W/L = 1.94), narrower than base of elytra; sides nearly straight at basal 2/3, weakly converging and gently curved at apical 1/3 towards moderately protruding anterior angles, finely margined; anterior border feebly concave bewteen angles, narrowly margined; basal border of pronotum bisinuate, unmargined; surface smooth, shiny, with fine microgranulation and sparse, moderately strong punctures on disc, unevenly distributed, stronger and sometimes confluent at sides and angles, elongate and partially aligned with basal border at sides of pronotum. Hypomera weakly convex on disc, smooth and shiny, unpunctured, with some transverse wrinkles basally; hypomeral suture deep and continuous, curved at basal angle and weakly and gradually diverging to base of anterior angles. Prosternum transversely convex at middle, slightly depressed at sides, finely margined anteriorly, with strong, rugose punctures at sides and before procoxae; prosternal process narrow between coxae, convex, slightly spatulate apically, punctured at sides, smooth medially. Mesepimera and mesanepisterna finely microreticulate, punctured at angles. Metanepisterna markedly narrow posteriorly, with strong and elongated scattered punctures. Metaventrite convex, shiny and leathery, with fine scattered punctation on disc, impressed and slightly rugose at sides, punctured at anterior angles. Scutellum longer than wide at base (W/L = 0.84), lancetshaped, glossy, unpunctured. Elytra long and parallel, with humeri marked, widest at middle, and regularly curved at apical third; surface with very fine uneven punctation on pale areas, and with strong punctures around and within dark markings; premarginal row of punctures confused in subhumeral area, regular and gradually approaching margin of elytron toward sutural angle; scutellar row long, with some ten spaced punctures reaching basal 1/3 of elytron. Dark markings on elytra expand clearly beyond associated puncture features: (i) sutural stripe broad, completely surrounding scutellum at base by twice its width, occupying interval between scutellar row and sutural row of punctures, reaching sutural angle; (ii) subsutural stripe entirely confluent with sutural stripe from base, at level with apical 1/3 of scutellum, to apex of elytra, expanded laterally as preapical lobe shortly before sutural angle; (iii) arcuate band entire, externally concave, mostly confluent internally with subsutural stripe except briefly at divergent ends; (iv) humeral spot large, long, nearly re
<typeStatus id="D162BFD87767FF90B09CF9F5FCCAF91D" box="[763,816,1655,1679]" pageId="15" pageNumber="16">type</typeStatus>
); one spot round, large, shortly confluent laterally with broad dark margin of elytron slightly behind midlateral spot; one spot round, slightly smaller (broken into three small spots on left elytron of
<typeStatus id="D162BFD87767FF90B395F93BFDDCF943" box="[498,550,1721,1745]" pageId="15" pageNumber="16">type</typeStatus>
), on apical concavity of arcuate band; and one large, irregular elongate spot reaching from near apex of arcuate band to dark lateral margin of elytron at level with base of spot of apical declivity. Epipleura wide at base, microsculptured, glossy, unpunctured. Femora slender, very finely microreticulate, with fine and scattered punctures. Tibiae straight and slender, shorter than femora, gradually widening apically and weakly impressed externally at apical half by apically expanded furrow; smooth, with scattered punctation basally, finely rugose and pubescent apically, with dense golden setae expanding apically on tibiae internally. Tarsi slender, shorter than corresponding tibiae. First abdominal ventrite convex, as long as metaventrite medially, shiny, with sparse fine punctation and short fine hairs; remaining ventrites finely microreticulate, with denser, stronger punctation, feebly impressed at sides.
</paragraph>
</subSubSection>
<caption id="5AA651F27767FF90B2F0FBE1FAACFBEB" box="[151,1366,1123,1145]" httpUri="https://zenodo.org/record/2614330/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1123,1145]" targetBox="[158,1431,203,1102]" targetPageId="15">
<paragraph id="0E66017A7767FF90B2F0FBE1FAACFBEB" blockId="15.[151,1366,1123,1145]" box="[151,1366,1123,1145]" pageId="15" pageNumber="16">
<emphasis id="3CADDD687767FF90B2F0FBE1FEE8FBEA" bold="true" box="[151,274,1123,1145]" pageId="15" pageNumber="16">FIGURE 4.</emphasis>
Dorsal and lateral views of the holotypes of
<taxonomicName id="C9D97AF97767FF90B0BEFBE1FBF9FBEB" authority="Jacoby" authorityName="Jacoby J. Gomez-Zurita" authorityYear="1891" box="[729,1027,1123,1145]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687767FF90B0BEFBE1FC4DFBEB" box="[729,951,1123,1145]" italics="true" pageId="15" pageNumber="16">Calligrapha femorata</emphasis>
Jacoby
</taxonomicName>
(a) and
<taxonomicName id="C9D97AF97767FF90B630FBE1FB2EFBEB" authority="Gómez-Zurita, 2018" authorityName="Gómez-Zurita" authorityYear="2018" box="[1111,1236,1123,1145]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="synthesys" status="sp. nov.">
<emphasis id="3CADDD687767FF90B630FBE1FB2EFBEB" box="[1111,1236,1123,1145]" italics="true" pageId="15" pageNumber="16">C. synthesys</emphasis>
</taxonomicName>
<emphasis id="3CADDD687767FF90B6BCFBE1FAD1FBEA" bold="true" box="[1243,1323,1123,1144]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="279E60137767FF90B6BCFBE1FAD1FBEA" box="[1243,1323,1123,1144]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(b).
</paragraph>
</caption>
<subSubSection id="46C352F17778FF8FB2A0FF15FDFDFE87" pageId="16" pageNumber="17" type="distribution">
<paragraph id="0E66017A7778FF8FB2A0FF15FDFDFE87" blockId="16.[151,1436,151,310]" pageId="16" pageNumber="17">
<emphasis id="3CADDD687778FF8FB2A0FF15FE9AFF22" bold="true" box="[199,352,151,176]" pageId="16" pageNumber="17">Distribution.</emphasis>
This species is rare in collections and so far it is only known from specimens collected in
<collectingCountry id="76CE41EA7778FF8FB2F0FF3BFF15FF40" box="[151,239,185,210]" name="Mexico" pageId="16" pageNumber="17">Mexico</collectingCountry>
, in sthe outhwestern slopes of the Sierra Madre Occidental and Transvolcanic belt (
<figureCitation id="96E21DFF7778FF8FB6D8FF3BFAFDFF40" box="[1215,1287,185,210]" captionStart="FIGURE 5" captionStartId="17.[151,250,1182,1204]" captionTargetBox="[150,1436,262,1160]" captionTargetId="figure@17.[150,1436,262,1160]" captionTargetPageId="17" captionText="FIGURE 5. Distribution of Calligrapha femorata Jacoby and C. synthesys sp. nov." httpUri="https://zenodo.org/record/2614332/files/figure.png" pageId="16" pageNumber="17">Fig. 5</figureCitation>
). Until more distribution data becomes available for this species, it can be considered a typical element of the Mexican Transition zone (
<bibRefCitation id="6A487C8B7778FF8FB334FF7EFE01FE87" author="Morrone, J. J." box="[339,507,252,277]" pageId="16" pageNumber="17" pagination="467 - 494" refId="ref57956" refString="Morrone, J. J. (2006) Biogeographic areas and transition zones of Latin America and the Caribbean islands based on panbiogeographic and cladistic analyses of the entomofauna. Annual Review of Entomology, 51, 467 - 494. https: // doi. org / 10.1146 / annurev. ento. 50.071803.130447" type="journal article" year="2006">Morrone 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="46C352F17778FF8FB2A0FE9FFE51FD54" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="0E66017A7778FF8FB2A0FE9FFD84FEA4" blockId="16.[151,1436,151,310]" box="[199,638,285,310]" pageId="16" pageNumber="17">
<emphasis id="3CADDD687778FF8FB2A0FE9FFE53FEA4" bold="true" box="[199,425,285,310]" pageId="16" pageNumber="17">Material examined</emphasis>
(eight specimens).
</paragraph>
<paragraph id="0E66017A7778FF8FB2F0FEE2FEF7FEEA" blockId="16.[151,1437,352,1376]" box="[151,269,352,376]" pageId="16" pageNumber="17">
<collectingCountry id="76CE41EA7778FF8FB2F0FEE2FEF3FEEA" box="[151,265,352,376]" name="Mexico" pageId="16" pageNumber="17">MEXICO</collectingCountry>
.
</paragraph>
<paragraph id="0E66017A7778FF8FB2A0FE03FE51FD54" blockId="16.[151,1437,352,1376]" pageId="16" pageNumber="17">
<emphasis id="3CADDD687778FF8FB2A0FE03FEE3FE08" bold="true" box="[199,281,385,410]" pageId="16" pageNumber="17">FSCA:</emphasis>
(1) one specimen:
<collectingCountry id="76CE41EA7778FF8FB398FE03FDA2FE08" box="[511,600,385,410]" name="Mexico" pageId="16" pageNumber="17">Mexico</collectingCountry>
,
<collectingRegion id="CC1DCF987778FF8FB000FE00FD37FE0B" box="[615,717,386,409]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="16" pageNumber="17">Veracruz</collectingRegion>
, Fortín de las
<collectingCountry id="76CE41EA7778FF8FB11CFE03FC3EFE08" box="[891,964,385,410]" name="Indonesia" pageId="16" pageNumber="17">Flores</collectingCountry>
, at Microwave Station,
<date id="7A6727BA7778FF8FB6B9FE03FAA2FE08" box="[1246,1368,385,410]" pageId="16" pageNumber="17" value="1992-06-30">30.vi.1992</date>
, B.K. Dozier.
<emphasis id="3CADDD687778FF8FB293FE20FEC4FE29" bold="true" box="[244,318,418,443]" pageId="16" pageNumber="17">MCZ:</emphasis>
(1) one specimen: Juquila,
<collectingCountry id="76CE41EA7778FF8FB0ECFE20FD19FE29" box="[651,739,418,443]" name="Mexico" pageId="16" pageNumber="17">Mexico</collectingCountry>
, Sallé Coll., 1st Jacoby Coll.,
<taxonomicName id="C9D97AF97778FF8FB628FE20FECCFE4F" authority="Jac. J. Gomez-Zurita" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687778FF8FB628FE20FAB2FE29" box="[1103,1352,418,443]" italics="true" pageId="16" pageNumber="17">Calligrapha femorata</emphasis>
Jac. J. Gómez-Zurita
</taxonomicName>
det. 2010; (2) one specimen: Juquila, Mohr, Jacoby 2nd Coll.,
<taxonomicName id="C9D97AF97778FF8FB19DFE46FAC0FE4E" authority="Jac. J." authorityName="Jac. J." box="[1018,1338,452,477]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687778FF8FB19DFE46FB15FE4E" box="[1018,1263,452,477]" italics="true" pageId="16" pageNumber="17">Calligrapha femorata</emphasis>
Jac. J.
</taxonomicName>
Gómez- Zurita det. 2010.
<emphasis id="3CADDD687778FF8FB33AFE67FE51FE6C" bold="true" box="[349,427,485,510]" pageId="16" pageNumber="17">NHM:</emphasis>
(1) one specimen: Juquila,
<collectingCountry id="76CE41EA7778FF8FB08FFE67FCBAFE6C" box="[744,832,485,510]" name="Mexico" pageId="16" pageNumber="17">Mexico</collectingCountry>
, Sallé Coll., 697, Godman-Salvin Coll., Biol. Centr.- Amer.
<emphasis id="3CADDD687778FF8FB28EFD84FEC9FD8D" bold="true" box="[233,307,518,543]" pageId="16" pageNumber="17">NMB:</emphasis>
(1) one specimen: Ventanas,
<collectingRegion id="CC1DCF987778FF8FB0FDFD85FD05FD8D" box="[666,767,519,543]" country="Mexico" name="Durango" pageId="16" pageNumber="17">Durango</collectingRegion>
, Höge,
<taxonomicName id="C9D97AF97778FF8FB107FD84FAE0FD8D" authority="Jac. J. Bechyne" box="[864,1306,518,543]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="suffriani">
<emphasis id="3CADDD687778FF8FB107FD84FBAFFD8D" box="[864,1109,518,543]" italics="true" pageId="16" pageNumber="17">Calligrapha suffriani</emphasis>
Jac. J. Bechyné
</taxonomicName>
det. 1954,
<emphasis id="3CADDD687778FF8FB2F0FDA5FEBFFDD2" box="[151,325,551,576]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="C9D97AF97778FF8FB2F0FDA5FEDAFDD2" authorityName="Chevrolat" authorityYear="1836" box="[151,288,551,576]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Calligrapha</taxonomicName>
nr.
</emphasis>
<taxonomicName id="C9D97AF97778FF8FB32BFDAAFD59FDD2" authority="Jac. J. Gomez-Zurita" box="[332,675,551,576]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687778FF8FB32BFDAAFE4BFDD2" box="[332,433,552,576]" italics="true" pageId="16" pageNumber="17">femorata</emphasis>
Jac. J. Gómez-Zurita
</taxonomicName>
det. 2010.
<emphasis id="3CADDD687778FF8FB144FDA5FC7FFDD2" bold="true" box="[803,901,551,576]" pageId="16" pageNumber="17">NMNH:</emphasis>
(1) one specimen:
<collectingCountry id="76CE41EA7778FF8FB638FDA5FB4DFDD2" box="[1119,1207,551,576]" name="Mexico" pageId="16" pageNumber="17">Mexico</collectingCountry>
,
<collectingRegion id="CC1DCF987778FF8FB6A4FDAAFAD0FDAD" box="[1219,1322,552,575]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="16" pageNumber="17">Veracruz</collectingRegion>
, Fortín de las
<collectingCountry id="76CE41EA7778FF8FB2A6FDCBFEF0FDF0" box="[193,266,585,610]" name="Indonesia" pageId="16" pageNumber="17">Flores</collectingCountry>
, 2900’, iii.7.1966, in bromeliads, George E. Ball, D.R. Whitehead collectors,
<taxonomicName id="C9D97AF97778FF8FB6C5FDCBFE55FD11" authority="Jacoby J. Gomez-Zurita" authorityName="Jacoby J. Gomez-Zurita" authorityYear="1891" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687778FF8FB6C5FDCBFA66FDF3" box="[1186,1436,585,610]" italics="true" pageId="16" pageNumber="17">Calligrapha femorata</emphasis>
Jacoby J. Gómez-Zurita
</taxonomicName>
det. 2011.
<emphasis id="3CADDD687778FF8FB05BFDE8FD4DFD11" bold="true" box="[572,695,618,643]" pageId="16" pageNumber="17">TAMUIC:</emphasis>
(1) two specimens: [X0533905, X0534193],
<collectingCountry id="76CE41EA7778FF8FB6BFFDE8FACAFD11" box="[1240,1328,618,643]" name="Mexico" pageId="16" pageNumber="17">Mexico</collectingCountry>
,
<collectingRegion id="CC1DCF987778FF8FB726FDE9FA62FD11" box="[1345,1432,619,643]" country="Mexico" name="Oaxaca" pageId="16" pageNumber="17">Oaxaca</collectingRegion>
, Hwy131, 15 mi N San Gabriel Mixtepec,
<date id="7A6727BA7778FF8FB014FD0EFD0EFD37" box="[627,756,652,677]" pageId="16" pageNumber="17" value="1987-07-11">11.vii.1987</date>
, Kovarik &amp; Schaffner coll.,
<taxonomicName id="C9D97AF97778FF8FB651FD0EFECCFD54" authority="Jacoby J. Gomez-Zurita" authorityName="Jacoby J. Gomez-Zurita" authorityYear="1891" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="3CADDD687778FF8FB651FD0EFAD6FD36" box="[1078,1324,652,677]" italics="true" pageId="16" pageNumber="17">Calligrapha femorata</emphasis>
Jacoby J. Gómez-Zurita
</taxonomicName>
det. 2011.
</paragraph>
</subSubSection>
<subSubSection id="46C352F17778FF8FB2A0FD4CFEB6FAF2" pageId="16" pageNumber="17" type="description">
<paragraph id="0E66017A7778FF8FB2A0FD4CFEB6FAF2" blockId="16.[151,1437,352,1376]" pageId="16" pageNumber="17">
<emphasis id="3CADDD687778FF8FB2A0FD4CFEC6FD75" bold="true" box="[199,316,718,743]" pageId="16" pageNumber="17">Variation.</emphasis>
Martin Jacoby knew a single specimen of his new taxon, which displayed the highly conspicuous dark metallic spots on reddish femora. In fact, among the specimens in the collection of this author currently in the Natural History Museum (London), there was another specimen of this species (I found it mixed with representatives of
<taxonomicName id="C9D97AF97778FF8FB30BFCB1FDA7FCD9" authority="Stal" authorityName="Stal" authorityYear="1859" box="[364,605,818,843]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="stillatipennis">
<emphasis id="3CADDD687778FF8FB30BFCB1FDD9FCD9" box="[364,547,818,843]" italics="true" pageId="16" pageNumber="17">C. stillatipennis</emphasis>
Stål
</taxonomicName>
). Interestingly, the specimen shows remarkable differences to the
<typeStatus id="D162BFD87778FF8FB702FCB1FA62FCD9" box="[1381,1432,819,843]" pageId="16" pageNumber="17">type</typeStatus>
, including the lack of darkened spots on femora, but also differences in coloration, heavier and more confluent elytral markings, although humeral markings in particular are basally free from basal margin of elytra. Moreover, this specimen shows hypomeral sutures almost obliterated, and more slender proportions (length:
<quantity id="C921AC9F7778FF8FB684FC15FAB3FC3C" box="[1251,1353,919,943]" metricMagnitude="-3" metricUnit="m" metricValue="8.13" pageId="16" pageNumber="17" unit="mm" value="8.13">8.13 mm</quantity>
, width:
<quantity id="C921AC9F7778FF8FB2F0FC3AFEF9FC42" box="[151,259,952,976]" metricMagnitude="-3" metricUnit="m" metricValue="4.98" pageId="16" pageNumber="17" unit="mm" value="4.98">4.98 mm</quantity>
), but otherwise it matches the
<typeStatus id="D162BFD87778FF8FB017FC3AFD5AFC42" box="[624,672,952,976]" pageId="16" pageNumber="17">type</typeStatus>
in important, diagnostic characters, such as the long labrum, the pronotum parallel-sided and much narrower than the elytra, the number and arrangements of markings on elytra, the subhumeral spot fused with humeral marking, and the midlateral spot expanded to form a wide elytral dark margin. In the Museum of Comparative Zoology (Cambridge MA) there are other specimens from the same source and almost identical to the one in London, but they are closer to the
<typeStatus id="D162BFD87778FF8FB1EBFBBCFC46FBC4" box="[908,956,1086,1110]" pageId="16" pageNumber="17">type</typeStatus>
in having their humeral marking confluent with basal margin of elytron and deeply furrowed hypomeral suture, an important diagnostic character for this entire lineage of
<taxonomicName id="C9D97AF97778FF8FB332FBFDFE24FB0A" authorityName="Chevrolat" authorityYear="1836" box="[341,478,1151,1176]" class="Insecta" family="Chrysomelidae" genus="Calligrapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="3CADDD687778FF8FB332FBFDFE24FB0A" box="[341,478,1151,1176]" italics="true" pageId="16" pageNumber="17">Calligrapha</emphasis>
</taxonomicName>
. One of the specimens at MCZ, however, departs again from the
<typeStatus id="D162BFD87778FF8FB6D8FB02FB15FB0A" box="[1215,1263,1152,1176]" pageId="16" pageNumber="17">type</typeStatus>
in the shape of pronotum, with sinuous sides, still narrower at base than base of elytra and slightly cordiform; elytral markings in this specimen are even heavier than those in the
<typeStatus id="D162BFD87778FF8FB091FB41FCD3FB49" box="[758,809,1219,1243]" pageId="16" pageNumber="17">type</typeStatus>
, leaving very reduced pale areas, and their femora, particularly metafemora are largely darkened basally. Finally, the specimen found in the beetle collection of the Smithsonian Institution still has another peculiarity: the specimen is darkened and it is difficult to recognize coloration, but it looks as if its legs were uniformly dark with metallic shine; otherwise, the specimens conforms well to the
<typeStatus id="D162BFD87778FF8FB373FACAFEB2FAF2" box="[276,328,1352,1376]" pageId="16" pageNumber="17">type</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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