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<document id="B06DD7717D77D6A12EB144EC0187B666" ID-DOI="10.5281/zenodo.4619638" ID-ISSN="2345-7600" ID-RefBankID="FF8BFFD82661FFFFFFA5FF89FFB0FFE1" ID-Zenodo-Dep="4619638" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616096590396" checkinUser="carolina" docAuthor="Daugeron, Christophe &amp; Grootaert, Patrick" docDate="2005" docId="7E7087CC0145FF89FCE5F94C3EDE5B90" docLanguage="en" docName="RafflesBZool.53.2.211-220.pdf" docOrigin="Raffles Bulletin of Zoology 53 (2)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Empis (Coptophlebia) neesoonensis Daugeron &amp; Grootaert 2005, new species" docType="treatment" docVersion="3" lastPageNumber="213" masterDocId="8249FFB40145FF8BFF97FF993F0D5848" masterDocTitle="Empidine Dance Flies From Singapore And Southern Malaysia (Diptera: Empididae: Empidinae)" masterLastPageNumber="220" masterPageNumber="211" pageNumber="211" updateTime="1698891084545" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="DACD5E9C76DCF092102C5D5F7967186C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F3651E8CA5E1C0BF464ACF39F2456B68">
<mods:title id="D325E4220ED93DB1BAEB0B5130D3CED1">Empidine Dance Flies From Singapore And Southern Malaysia (Diptera: Empididae: Empidinae)</mods:title>
</mods:titleInfo>
<mods:name id="D3A8E701298FDFF3A34FC0FEC624F764" type="personal">
<mods:role id="0710B9A13C2CF3671BA117624E9AB8C1">
<mods:roleTerm id="CE4D8A49DA822D96A5325390743E072A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BD241371930F605AE14D7042B9832BDC">Daugeron, Christophe</mods:namePart>
</mods:name>
<mods:name id="9A7B2853E465F87275C75DE5F69B02AD" type="personal">
<mods:role id="D2B1E0AE8EB797C8C3C7DC38C0F27BE4">
<mods:roleTerm id="E10E04F136C48389FC5520D1A9704980">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0C9A8261185ACE5429B1C82B5B8D935A">Grootaert, Patrick</mods:namePart>
</mods:name>
<mods:typeOfResource id="53E05CEC38241F1DF797E1282C6AC9D9">text</mods:typeOfResource>
<mods:relatedItem id="8A61CABB1D8EB58E5FCE3D1C966C3327" type="host">
<mods:titleInfo id="79D3E3B69EE2D4FCE3BD90CB73109B18">
<mods:title id="50619584B2F20F42A6DF5CFF81EEA23D">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="C4865ACA8CED01109153078CF13984CB">
<mods:date id="1E448AA48036F18A9CE107F42913D649">2005</mods:date>
<mods:detail id="C4C3F3A44B566F3692258CFF10979B42" type="pubDate">
<mods:number id="06A6DBD1726E74A6C6CE8AEF60715E5E">2005-12-31</mods:number>
</mods:detail>
<mods:detail id="929D6D6842ADD754E113B4325C545A9D" type="volume">
<mods:number id="7CF50E69F1C0C3963FCD78B679BD98A8">53</mods:number>
</mods:detail>
<mods:detail id="BE481F260D39D0B569DDCA3C0FC2E961" type="issue">
<mods:number id="CE7822631760C877D2D00E4E28E6FC29">2</mods:number>
</mods:detail>
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<mods:start id="3B96A772A0E6EDA803B2C3C827C47F70">211</mods:start>
<mods:end id="125BFC84C796307F42B97D41C1888F5A">220</mods:end>
</mods:extent>
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<mods:classification id="264C913E9DF0ACAFCCBBEA70655F6557">journal article</mods:classification>
<mods:identifier id="0912AF4EF5F3EA79C80B16775AAEBC14" type="DOI">10.5281/zenodo.4619638</mods:identifier>
<mods:identifier id="0AD5DBCF1522E3E415A77965CDC2F8CE" type="ISSN">2345-7600</mods:identifier>
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<mods:identifier id="378B51101D722BDF2913698EF76A0099" type="Zenodo-Dep">4619638</mods:identifier>
</mods:mods>
<treatment id="7E7087CC0145FF89FCE5F94C3EDE5B90" LSID="urn:lsid:plazi:treatment:7E7087CC0145FF89FCE5F94C3EDE5B90" httpUri="http://treatment.plazi.org/id/7E7087CC0145FF89FCE5F94C3EDE5B90" lastPageId="2" lastPageNumber="213" pageId="0" pageNumber="211">
<subSubSection id="BEC365510145FF8BFCE5F94C3A9E5EA5" box="[882,1427,1749,1773]" pageId="0" pageNumber="211" type="nomenclature">
<paragraph id="F66636DA0145FF8BFCE5F94C3A9E5EA5" blockId="0.[882,1427,1749,1805]" box="[882,1427,1749,1773]" pageId="0" pageNumber="211">
<heading id="AD2E81B60145FF8BFCE5F94C3A9E5EA5" bold="true" box="[882,1427,1749,1773]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="211" reason="2">
<emphasis id="C4ADEAC80145FF8BFCE5F94C3A9E5EA5" bold="true" box="[882,1427,1749,1773]" pageId="0" pageNumber="211">
<taxonomicName id="31D94D590145FF8BFCE5F94C3A0D5EA5" ID-CoL="8BZ6V" authority="Daugeron &amp; Grootaert, 2005" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[882,1280,1749,1773]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="211" phylum="Arthropoda" rank="species" species="neesoonensis" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="C4ADEAC80145FF8BFCE5F94C3CB55EA5" bold="true" box="[882,952,1749,1773]" italics="true" pageId="0" pageNumber="211">Empis</emphasis>
(
<emphasis id="C4ADEAC80145FF8BFC5EF94C3B505EA5" bold="true" box="[969,1117,1749,1773]" italics="true" pageId="0" pageNumber="211">Coptophlebia</emphasis>
)
<emphasis id="C4ADEAC80145FF8BFBFAF94C3A0D5EA5" bold="true" box="[1133,1280,1749,1773]" italics="true" pageId="0" pageNumber="211">neesoonensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="DF9E57B30145FF8BFA99F94C3A9E5EA5" box="[1294,1427,1749,1773]" pageId="0" pageNumber="211" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BEC365510145FF8BFBABF96F3BC55F45" box="[1084,1224,1782,1805]" pageId="0" pageNumber="211" type="description">
<paragraph id="F66636DA0145FF8BFBABF96F3BC55F45" blockId="0.[882,1427,1749,1805]" box="[1084,1224,1782,1805]" pageId="0" pageNumber="211">
(
<figureCitation id="6EE22A5F0145FF8BFBD3F96F3BAB5F45" box="[1092,1190,1782,1805]" captionStart="Fig" captionStartId="1.[113,149,2063,2084]" captionTargetBox="[117,1448,682,1971]" captionTargetId="figure-9@1.[102,1460,631,1988]" captionTargetPageId="1" captionText="Fig. 1. Male fore leg in frontal view: A, tibia and first tarsomere of E. (C.) neesoonensis; B, tibia and tarsus of E. (C.) producta; C, tibia and first tarsomere of E. (C.) tortuosa; D, tibia and first tarsomere of E. (C.) verruca." figureDoi="http://doi.org/10.5281/zenodo.4619640" httpUri="https://zenodo.org/record/4619640/files/figure.png" pageId="0" pageNumber="211">Figs. 1A</figureCitation>
,
<figureCitation id="6EE22A5F0145FF8BFB25F96F3BCD5F45" box="[1202,1216,1782,1805]" captionStart="Fig" captionStartId="2.[136,172,2093,2114]" captionTargetBox="[144,1464,1104,2005]" captionTargetId="figure-25@2.[113,1493,1063,2049]" captionTargetPageId="2" captionText="Fig. 2. Male hypopygium of E. (C.) neesoonensis, lateral view. Scale = 4.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4619642" httpUri="https://zenodo.org/record/4619642/files/figure.png" pageId="0" pageNumber="211">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BEC365510145FF8BFCD4F8AC3CAE5079" pageId="0" pageNumber="211" type="materials_examined">
<paragraph id="F66636DA0145FF8BFCD4F8AC3C845FCC" blockId="0.[835,1474,1845,1924]" pageId="0" pageNumber="211">
<emphasis id="C4ADEAC80145FF8BFCD4F8AC3B2E5F02" bold="true" box="[835,1059,1845,1866]" italics="true" pageId="0" pageNumber="211">Material examined. –</emphasis>
<materialsCitation id="46B13C870145FF8BFBBAF8AC3C845FCC" collectingDate="2002-12-04" collectionCode="ZRC" collectorName="P. Grootaert" country="Singapore" location="Singapore" pageId="0" pageNumber="211" specimenCount="1" typeStatus="holotype">
<typeStatus id="296288780145FF8BFBBAF8AC3B845F02" box="[1069,1161,1845,1866]" pageId="0" pageNumber="211" type="holotype">Holotype</typeStatus>
<emphasis id="C4ADEAC80145FF8BFB05F8AF3B975F03" box="[1170,1178,1846,1867]" italics="true" pageId="0" pageNumber="211">-</emphasis>
male,
<collectingCountry id="8ECE764A0145FF8BFB73F8AC3A465F02" box="[1252,1355,1845,1866]" name="Singapore" pageId="0" pageNumber="211">Singapore</collectingCountry>
, Nee Soon, primary rain forest, sample 22054, coll.
<collectorName id="5B2C530C0145FF8BFB5AF8CB3A4A5F2F" box="[1229,1351,1874,1895]" pageId="0" pageNumber="211">P. Grootaert</collectorName>
,
<date id="8267101A0145FF8BFAC5F8CB3ACF5F2F" box="[1362,1474,1874,1895]" pageId="0" pageNumber="211" value="2002-12-04">
<collectingDate id="9223E9F20145FF8BFAC5F8CB3ACF5F2F" box="[1362,1474,1874,1895]" pageId="0" pageNumber="211" value="2002-12-04">4 Dec.2002</collectingDate>
</date>
(
<collectionCode id="90C8AE1F0145FF8BFCDDF8F63C735FCC" box="[842,894,1903,1924]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="0" pageNumber="211">ZRC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F66636DA0145FF8BFCD4F8303BFD5FF6" blockId="0.[835,1264,1961,1982]" box="[835,1264,1961,1982]" pageId="0" pageNumber="211">
<materialsCitation id="46B13C870145FF8BFCD4F8303BE15FF6" box="[835,1260,1961,1982]" collectionCode="ZRC" pageId="0" pageNumber="211" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="296288780145FF8BFCD4F8303C955FF6" box="[835,920,1961,1982]" pageId="0" pageNumber="211" type="paratype">Paratype</typeStatus>
<emphasis id="C4ADEAC80145FF8BFC08F8303CA65FF6" box="[927,939,1961,1982]" italics="true" pageId="0" pageNumber="211">–</emphasis>
<specimenCount id="E0DFFD530145FF8BFC25F8303B0A5FF6" box="[946,1031,1961,1982]" pageId="0" pageNumber="211" type="female">1 female</specimenCount>
with same data (
<collectionCode id="90C8AE1F0145FF8BFB26F8303BEB5FF6" box="[1201,1254,1961,1982]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="0" pageNumber="211">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F66636DA0145FF8BFCD4F87B3CAE5079" blockId="0.[835,1474,2018,2097]" pageId="0" pageNumber="211">
Other material
<emphasis id="C4ADEAC80145FF8BFC46F87B3CD05FBF" box="[977,989,2018,2039]" italics="true" pageId="0" pageNumber="211">–</emphasis>
<materialsCitation id="46B13C870145FF8BFC75F87B3CAD5079" collectingDate="2003-11-22" collectorName="P. Grootaert" country="Singapore" location="Singapore" pageId="0" pageNumber="211" specimenCount="1" specimenCount-male="1">
<specimenCount id="E0DFFD530145FF8BFC75F87B3B2F5FBF" box="[994,1058,2018,2039]" pageId="0" pageNumber="211" type="male">1 male</specimenCount>
in bad condition (head missing),
<collectingCountry id="8ECE764A0145FF8BFACCF87B3AB35FBF" box="[1371,1470,2018,2039]" name="Singapore" pageId="0" pageNumber="211">Singapore</collectingCountry>
, NUS campus, sample 23082, coll.
<collectorName id="5B2C530C0145FF8BFB16F8663BF9505C" box="[1153,1268,2047,2068]" pageId="0" pageNumber="211">P. Grootaert</collectorName>
,
<date id="8267101A0145FF8BFB6CF8663A79505C" box="[1275,1396,2047,2068]" pageId="0" pageNumber="211" value="2003-11-22">
<collectingDate id="9223E9F20145FF8BFB6CF8663A79505C" box="[1275,1396,2047,2068]" pageId="0" pageNumber="211" value="2003-11-22">22 Nov.2003</collectingDate>
</date>
(KBIN- IRScNB)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BEC365510144FF8AFFE6FF583FF258B0" pageId="1" pageNumber="212" type="etymology">
<paragraph id="F66636DA0144FF8AFFE6FF583FF258B0" blockId="1.[113,752,193,248]" pageId="1" pageNumber="212">
<emphasis id="C4ADEAC80144FF8AFFE6FF583E0E5891" bold="true" box="[113,259,193,217]" italics="true" pageId="1" pageNumber="212">Etymology. –</emphasis>
The name of the species is derived from the type-locality.
</paragraph>
</subSubSection>
<subSubSection id="BEC365510144FF8AFFE6FEB83EC159D0" pageId="1" pageNumber="212" type="diagnosis">
<paragraph id="F66636DA0144FF8AFFE6FEB83EC159D0" blockId="1.[113,752,289,408]" pageId="1" pageNumber="212">
<emphasis id="C4ADEAC80144FF8AFFE6FEB83FF65971" bold="true" box="[113,251,289,313]" italics="true" pageId="1" pageNumber="212">Diagnosis. –</emphasis>
Small blackish species with all bristles black, wing clear with median veins almost not visible, fore tibia deformed with about 6 spine-like anterodorsal bristles in basal half not longer than tibia depth.
</paragraph>
</subSubSection>
<subSubSection id="BEC365510144FF89FFE6FE583FF55AD0" lastPageId="2" lastPageNumber="213" pageId="1" pageNumber="212" type="description">
<paragraph id="F66636DA0144FF8AFFE6FE583FB65A70" blockId="1.[113,753,449,568]" pageId="1" pageNumber="212">
<emphasis id="C4ADEAC80144FF8AFFE6FE583EAF5990" box="[113,418,449,473]" italics="true" pageId="1" pageNumber="212">
<emphasis id="C4ADEAC80144FF8AFFE6FE583E6C5991" bold="true" box="[113,353,449,473]" italics="true" pageId="1" pageNumber="212">Description. – Male –</emphasis>
Head
</emphasis>
. Occiput, antenna dark brown, second and third flagellomeres aristiform. Face blackish. Palpus apparently brown. Labrum length about twice head height.
</paragraph>
<paragraph id="F66636DA0144FF89FCBBFF583E625930" blockId="1.[812,1452,193,568]" lastBlockId="2.[136,776,193,376]" lastPageId="2" lastPageNumber="213" pageId="1" pageNumber="212">
<emphasis id="C4ADEAC80144FF8AFCBBFF583C715890" box="[812,892,193,216]" italics="true" pageId="1" pageNumber="212">Thorax</emphasis>
brown, scutum subshiny. Acrostichals missing; dorsocentrals uniserial ending with 1 strong, long bristle in prescutellar depression.
<emphasis id="C4ADEAC80144FF8AFBA7FE983B6E5950" box="[1072,1123,257,280]" italics="true" pageId="1" pageNumber="212">Legs</emphasis>
(
<figureCitation id="6EE22A5F0144FF8AFBE7FE983BCF5950" box="[1136,1218,257,280]" captionStart="Fig" captionStartId="1.[113,149,2063,2084]" captionTargetBox="[117,1448,682,1971]" captionTargetId="figure-9@1.[102,1460,631,1988]" captionTargetPageId="1" captionText="Fig. 1. Male fore leg in frontal view: A, tibia and first tarsomere of E. (C.) neesoonensis; B, tibia and tarsus of E. (C.) producta; C, tibia and first tarsomere of E. (C.) tortuosa; D, tibia and first tarsomere of E. (C.) verruca." figureDoi="http://doi.org/10.5281/zenodo.4619640" httpUri="https://zenodo.org/record/4619640/files/figure.png" pageId="1" pageNumber="212">Fig. 1A</figureCitation>
) brown. Left mid and hind legs missing. All femora with rather short bristles. Fore tibia deformed, somewhat S-shaped with about 6 spine-like anterodorsal bristles in basal half; first fore tarsomere with 1 strong, long anterolateral bristle at base. Mid tibia with 3 strong, rather long dorsal bristles (at basal quarter, about at middle, at tip); first mid tarsomere with 1 strong, long anterodorsal bristle at apical tip. Hind tibia swollen apically with strong, long dorsal bristles, several distinct ventral bristles at apical half; hind tarsus swollen, first four tarsomeres with long dorsal bristles.
<emphasis id="C4ADEAC80147FF89FE1DFF583ECF5890" box="[394,450,193,216]" italics="true" pageId="2" pageNumber="213">Wing</emphasis>
(length =
<quantity id="31219B3F0147FF89FDBEFF583D775890" box="[553,634,193,216]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="213" unit="mm" value="2.2">2.2 mm</quantity>
) clear. Discal cell and median veins almost not visible. A1 abbreviated. Halter brown.
<emphasis id="C4ADEAC80147FF89FEB7FE983E885950" box="[288,389,257,280]" italics="true" pageId="2" pageNumber="213">Abdomen</emphasis>
brown, sternites with pair of distinct bristles posteriorly.
<emphasis id="C4ADEAC80147FF89FEEDFEB83D015970" box="[378,524,289,312]" italics="true" pageId="2" pageNumber="213">Hypopygium</emphasis>
(
<figureCitation id="6EE22A5F0147FF89FDB6FEB83D605970" box="[545,621,289,312]" captionStart="Fig" captionStartId="2.[136,172,2093,2114]" captionTargetBox="[144,1464,1104,2005]" captionTargetId="figure-25@2.[113,1493,1063,2049]" captionTargetPageId="2" captionText="Fig. 2. Male hypopygium of E. (C.) neesoonensis, lateral view. Scale = 4.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4619642" httpUri="https://zenodo.org/record/4619642/files/figure.png" pageId="2" pageNumber="213">Fig. 2</figureCitation>
). Epandrium triangular with strong, long bristles at tip. Hypandrium pointed. Phallus thin.
</paragraph>
<caption id="A2A666520144FF8AFFE6F7963CB15009" ID-DOI="http://doi.org/10.5281/zenodo.4619640" ID-Zenodo-Dep="4619640" httpUri="https://zenodo.org/record/4619640/files/figure.png" pageId="1" pageNumber="212" startId="1.[113,149,2063,2084]" targetBox="[117,1448,682,1971]" targetPageId="1">
<paragraph id="F66636DA0144FF8AFFE6F7963CB15009" blockId="1.[113,1451,2063,2113]" pageId="1" pageNumber="212">
Fig. 1. Male fore leg in frontal view: A, tibia and first tarsomere of
<taxonomicName id="31D94D590144FF8AFC9DF7963CD9506C" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[778,980,2063,2084]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="212" phylum="Arthropoda" rank="species" species="neesoonensis" subGenus="Coptophlebia">
<emphasis id="C4ADEAC80144FF8AFC9DF7963C2D506C" box="[778,800,2063,2084]" italics="true" pageId="1" pageNumber="212">E.</emphasis>
(
<emphasis id="C4ADEAC80144FF8AFCB9F7963C48506C" box="[814,837,2063,2084]" italics="true" pageId="1" pageNumber="212">C.</emphasis>
)
<emphasis id="C4ADEAC80144FF8AFCC4F7963CD9506C" box="[851,980,2063,2084]" italics="true" pageId="1" pageNumber="212">neesoonensis</emphasis>
</taxonomicName>
; B, tibia and tarsus of
<taxonomicName id="31D94D590144FF8AFB23F7963A58506C" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[1204,1365,2063,2084]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="212" phylum="Arthropoda" rank="species" species="producta" subGenus="Coptophlebia">
<emphasis id="C4ADEAC80144FF8AFB23F7963BC7506C" box="[1204,1226,2063,2084]" italics="true" pageId="1" pageNumber="212">E.</emphasis>
(
<emphasis id="C4ADEAC80144FF8AFB4FF7963BE2506C" box="[1240,1263,2063,2084]" italics="true" pageId="1" pageNumber="212">C.</emphasis>
)
<emphasis id="C4ADEAC80144FF8AFB6AF7963A58506C" box="[1277,1365,2063,2084]" italics="true" pageId="1" pageNumber="212">producta</emphasis>
</taxonomicName>
; C, tibia and first tarsomere of
<taxonomicName id="31D94D590144FF8AFEDBF7B53EEB5009" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[332,486,2092,2113]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="212" phylum="Arthropoda" rank="species" species="tortuosa" subGenus="Coptophlebia">
<emphasis id="C4ADEAC80144FF8AFEDBF7B53E6F5009" box="[332,354,2092,2113]" italics="true" pageId="1" pageNumber="212">E.</emphasis>
(
<emphasis id="C4ADEAC80144FF8AFEE7F7B53E8A5009" box="[368,391,2092,2113]" italics="true" pageId="1" pageNumber="212">C.</emphasis>
)
<emphasis id="C4ADEAC80144FF8AFE02F7B53EEB5009" box="[405,486,2092,2113]" italics="true" pageId="1" pageNumber="212">tortuosa</emphasis>
</taxonomicName>
; D, tibia and first tarsomere of
<taxonomicName id="31D94D590144FF8AFCB6F7B53CBB5009" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[801,950,2092,2113]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="212" phylum="Arthropoda" rank="species" species="verruca" subGenus="Coptophlebia">
<emphasis id="C4ADEAC80144FF8AFCB6F7B53C3B5009" box="[801,822,2092,2113]" italics="true" pageId="1" pageNumber="212">E.</emphasis>
(
<emphasis id="C4ADEAC80144FF8AFCD2F7B53C515009" box="[837,860,2092,2113]" italics="true" pageId="1" pageNumber="212">C.</emphasis>
)
<emphasis id="C4ADEAC80144FF8AFCFDF7B53CBB5009" box="[874,950,2092,2113]" italics="true" pageId="1" pageNumber="212">verruca</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="F66636DA0147FF89FF1FFE383FF55AD0" blockId="2.[136,775,417,664]" pageId="2" pageNumber="213">
<emphasis id="C4ADEAC80147FF89FF1FFE383FD159F1" bold="true" box="[136,220,417,441]" italics="true" pageId="2" pageNumber="213">Female</emphasis>
similar to male except for the following characters: palpus almost yellow. Dichoptic with all ommatidia of the same size. Fore femora with distinct dorsal pennation at base; first fore tarsomere with short dorsal pennation. Mid and hind femora with long dorsal and ventral pennation. Mid tibia with distinct dorsal pennation and some distinct ventral pennate bristles at base. Hind tibia with long dorsal and ventral pennation.
</paragraph>
</subSubSection>
<subSubSection id="BEC365510147FF89FF1FFD583EDE5B90" pageId="2" pageNumber="213" type="discussion">
<paragraph id="F66636DA0147FF89FF1FFD583EDE5B90" blockId="2.[136,775,704,984]" pageId="2" pageNumber="213">
<emphasis id="C4ADEAC80147FF89FF1FFD583E095A91" bold="true" box="[136,260,704,729]" pageId="2" pageNumber="213">
<emphasis id="C4ADEAC80147FF89FF1FFD583FE65A91" bold="true" box="[136,235,705,729]" italics="true" pageId="2" pageNumber="213">Remarks</emphasis>
.
<emphasis id="C4ADEAC80147FF89FF60FD583E095A91" box="[247,260,705,729]" italics="true" pageId="2" pageNumber="213">–</emphasis>
</emphasis>
<taxonomicName id="31D94D590147FF89FE9DFD583EED5A90" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[266,480,705,728]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="213" phylum="Arthropoda" rank="species" species="neesoonensis">Empis neesoonensis</taxonomicName>
belongs to the
<taxonomicName id="31D94D590147FF89FD16FD583C0A5A90" authorityName="Meijere" authorityYear="1907" box="[641,775,705,728]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="213" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="C4ADEAC80147FF89FD16FD583C0A5A90" box="[641,775,705,728]" italics="true" pageId="2" pageNumber="213">E. jacobsoni</emphasis>
</taxonomicName>
clade as defined recently (Daugeron &amp; Grootaert, 2005). Within this clade
<taxonomicName id="31D94D590147FF89FED7FC983EE45B50" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[320,489,769,792]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="213" phylum="Arthropoda" rank="species" species="neesoonensis">
<emphasis id="C4ADEAC80147FF89FED7FC983E5D5B50" box="[320,336,769,792]" italics="true" pageId="2" pageNumber="213">E</emphasis>
.
<emphasis id="C4ADEAC80147FF89FECBFC983EE45B50" box="[348,489,769,792]" italics="true" pageId="2" pageNumber="213">neesoonensis</emphasis>
</taxonomicName>
seems to be closely related to
<taxonomicName id="31D94D590147FF89FF34FCB83EC25B70" authority="(Malaysia)" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" baseAuthorityName="Malaysia" box="[163,463,801,824]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="213" phylum="Arthropoda" rank="species" species="cameronensis">
<emphasis id="C4ADEAC80147FF89FF34FCB83E585B70" box="[163,341,801,824]" italics="true" pageId="2" pageNumber="213">E. cameronensis</emphasis>
(
<collectingCountry id="8ECE764A0147FF89FEF5FCB83EC55B70" box="[354,456,801,824]" name="Malaysia" pageId="2" pageNumber="213">Malaysia</collectingCountry>
)
</taxonomicName>
and differs from this species by the following characters: smaller species; anterodorsal spine-like bristles of fore tibia shorter than tibia depth; first mid tarsomere not whitish; median veins almost not visible; epandrium rather pointed at tip, not rounded, with ventral bristles less numerous basally.
</paragraph>
</subSubSection>
</treatment>
</document>
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