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<document id="573B134420C41D3438E09F18C5D06805" ID-DOI="10.5281/zenodo.7225407" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1702531225240" checkinUser="valdenar" docAuthor="Grzelak, Katarzyna &amp; Sørensen, Martin V." docDate="2022" docId="7E3687A4FFA2D256FC8A218CFE54FB58" docLanguage="en" docName="ZoolStud.55.32.1-21.pdf" docOrigin="Zoological studies (Zool. Stud.) 55 (32)" docSource="http://dx.doi.org/10.5281/zenodo.7225407" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Echinoderes serratulus Grzelak &amp; Sørensen 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="17" masterDocId="820FFFDCFFA8D244FFFC2531FFEBFA2E" masterDocTitle="Echinoderes galadrielae Grzelak &amp; amp; Sørensen 2022, sp. nov." masterLastPageNumber="32" masterPageNumber="32" pageNumber="11" updateTime="1707938672798" updateUser="ExternalLinkService">
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<mods:title id="62B28CEB0B2A3BEDCE123417D8C3D202">Echinoderes galadrielae Grzelak &amp; amp; Sørensen 2022, sp. nov.</mods:title>
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<mods:namePart id="B32EE2332F3CF370A207A063F667B0C8">Sørensen, Martin V.</mods:namePart>
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<treatment id="7E3687A4FFA2D256FC8A218CFE54FB58" LSID="urn:lsid:plazi:treatment:7E3687A4FFA2D256FC8A218CFE54FB58" httpUri="http://treatment.plazi.org/id/7E3687A4FFA2D256FC8A218CFE54FB58" lastPageId="18" lastPageNumber="17" pageId="10" pageNumber="11">
<subSubSection id="BE856539FFA2D24EFC8A218CFA8BFF02" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="F62036B2FFA2D24EFC8A218CFA8BFF02" blockId="10.[855,1447,1213,1324]" pageId="10" pageNumber="11">
<heading id="AD6881DEFFA2D24EFC8A218CFA62FEF8" bold="true" box="[886,1417,1213,1238]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="11" reason="2">
<emphasis id="C4EBEAA0FFA2D24EFC8A218CFA62FEF8" bold="true" box="[886,1417,1213,1238]" pageId="10" pageNumber="11">
<taxonomicName id="319F4D31FFA2D24EFC8A218CFB75FEF8" ID-CoL="7Z5ST" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[886,1182,1213,1238]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="10" pageNumber="11" phylum="Kinorhyncha" rank="species" species="serratulus" status="sp. nov.">
<emphasis id="C4EBEAA0FFA2D24EFC8A218CFB75FEF8" bold="true" box="[886,1182,1213,1238]" italics="true" pageId="10" pageNumber="11">Echinoderes serratulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFD857DBFFA2D24EFB59218CFAE3FEF8" box="[1189,1288,1213,1238]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
Yamasaki
</emphasis>
</heading>
<uri id="820E3AB0FFA2D24EFCAB21EDFB3EFF21" pageId="10" pageNumber="11">
urn:lsid:zoobank.org:act:
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</uri>
[New Japanese name: Nokogiri togekawa]
</paragraph>
</subSubSection>
<subSubSection id="BE856539FFA2D254FC732061FA8FF898" lastPageId="16" lastPageNumber="17" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="F62036B2FFA2D24EFC732061FBFEFC87" blockId="10.[850,1452,1360,1961]" pageId="10" pageNumber="11">
<emphasis id="C4EBEAA0FFA2D24EFC732061FB98FF47" box="[911,1139,1360,1385]" italics="true" pageId="10" pageNumber="11">Material examined</emphasis>
: All specimens collected
<date id="82211072FFA2D24EFCAE2041FBBEFFA7" box="[850,1109,1392,1417]" pageId="10" pageNumber="11" value="2014-11-25">25 November 2014</date>
at
<quantity id="31679B57FFA2D24EFB7B2041FB21FFA7" box="[1159,1226,1392,1417]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="m" value="10.0">10 m</quantity>
depth near Hon Mieu Island, Nha Trang,
<collectingCountry id="8E887622FFA2D24EFB6520A1FAEAFF87" box="[1177,1281,1424,1449]" name="Vietnam" pageId="10" pageNumber="11">Vietnam</collectingCountry>
(
<geoCoordinate id="93AB5075FFA2D24EFAE920A1FA4CFF87" box="[1301,1447,1424,1449]" degrees="12" direction="north" minutes="11.60" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="12.193334">12°11.60'N</geoCoordinate>
,
<geoCoordinate id="93AB5075FFA2D24EFCAE2081FC17FFE7" box="[850,1020,1456,1481]" degrees="109" direction="east" minutes="13.96" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="109.232666">109°13.96'E</geoCoordinate>
).
<typeStatus id="29248810FFA2D24EFBE92081FB7BFFE7" box="[1045,1168,1456,1481]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
: adult female (ZIHU-5053), mounted in Fluoromount G
<superScript id="01EA9BFAFFA2D24EFB1420FFFB1FFFF3" attach="left" box="[1256,1268,1486,1501]" fontSize="6" pageId="10" pageNumber="11">®</superScript>
.
<typeStatus id="29248810FFA2D24EFAFE20E1FA8EFFC7" box="[1282,1381,1488,1513]" pageId="10" pageNumber="11" type="allotype">Allotype</typeStatus>
: adult male (ZIHU-5054), mounted in Fluoromount G
<superScript id="01EA9BFAFFA2D24EFA6420DFFA4FFFD3" attach="right" box="[1432,1444,1518,1533]" fontSize="6" pageId="10" pageNumber="11">®</superScript>
.
<typeStatus id="29248810FFA2D24EFCAE2321FC3BFC07" box="[850,976,1552,1577]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="E099FD3BFFA2D24EFC212321FBA5FC07" box="[989,1102,1552,1577]" count="1" pageId="10" pageNumber="11" type="adult">one adult</specimenCount>
female and
<specimenCount id="E099FD3BFFA2D24EFB172321FAB2FC07" box="[1259,1369,1552,1577]" count="2" pageId="10" pageNumber="11" type="adult">two adult</specimenCount>
males (ZIHU-5055-5057), mounted in Fluoromount G
<superScript id="01EA9BFAFFA2D24EFA64231FFA4FFC13" attach="left" box="[1432,1444,1582,1597]" fontSize="6" pageId="10" pageNumber="11">®</superScript>
. Additional material:
<specimenCount id="E099FD3BFFA2D24EFBC22361FB18FC47" box="[1086,1267,1616,1641]" count="4" pageId="10" pageNumber="11" type="generic">four specimens</specimenCount>
for
<collectionCode id="908EAE77FFA2D24EFAD82361FAB5FC44" box="[1316,1374,1616,1642]" pageId="10" pageNumber="11">SEM</collectionCode>
(
<specimenCount id="E099FD3BFFA2D24EFA902361FC65FCA7" count="3" pageId="10" pageNumber="11" type="adult">three adult</specimenCount>
females and
<specimenCount id="E099FD3BFFA2D24EFBC32341FB58FCA7" box="[1087,1203,1648,1673]" count="1" pageId="10" pageNumber="11" type="adult">one adult</specimenCount>
male), mounted on aluminum stubs.
</paragraph>
<paragraph id="F62036B2FFA2D24EFC732381FA9FFD27" blockId="10.[850,1452,1360,1961]" pageId="10" pageNumber="11">
<emphasis id="C4EBEAA0FFA2D24EFC732381FBFBFCE7" box="[911,1040,1712,1737]" italics="true" pageId="10" pageNumber="11">Etymology</emphasis>
: The species name
<taxonomicName id="319F4D31FFA2D24EFAEF2381FA66FCE7" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1299,1421,1712,1737]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="10" pageNumber="11" phylum="Kinorhyncha" rank="species" species="serratulus">
<emphasis id="C4EBEAA0FFA2D24EFAEF2381FA66FCE7" box="[1299,1421,1712,1737]" italics="true" pageId="10" pageNumber="11">serratulus</emphasis>
</taxonomicName>
is from Latin, meaning serration, referring to the species conspicuous primary pectinate fringe.
</paragraph>
<paragraph id="F62036B2FFA2D24FFC732221FEF7FBF8" blockId="10.[850,1452,1360,1961]" lastBlockId="11.[136,738,253,1974]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="C4EBEAA0FFA2D24EFC732221FBE1FD07" box="[911,1034,1808,1833]" italics="true" pageId="10" pageNumber="11">Diagnosis</emphasis>
:
<taxonomicName id="319F4D31FFA2D24EFBE12221FB53FD07" authorityName="Claparede" authorityYear="1863" box="[1053,1208,1808,1833]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="10" pageNumber="11" phylum="Kinorhyncha" rank="genus">
<emphasis id="C4EBEAA0FFA2D24EFBE12221FB53FD07" box="[1053,1208,1808,1833]" italics="true" pageId="10" pageNumber="11">Echinoderes</emphasis>
</taxonomicName>
with short acicular spine middorsally on segment 4 and sublaterally on segments 6 and 7; lateroventral tubules on segments 5 and 8, midlateral tubules on segment 9, and laterodorsal tubules on segment 10; large sieve plates on segment 9; primary pectinate fringes ventrally on segments 1-9 conspicuous, with longer fringe tips; tips of ventromedial primary pectinate fringes on segments 2-5 obliquely orientated, pointing towards midventral line; lateral terminal spines short and plump, length 12-15% of trunk length.
</paragraph>
<paragraph id="F62036B2FFA3D24FFF3924ECFD3FF858" blockId="11.[136,738,253,1974]" pageId="11" pageNumber="12">
<emphasis id="C4EBEAA0FFA3D24FFF3924ECFEA1FBD8" box="[197,330,477,502]" italics="true" pageId="11" pageNumber="12">Description</emphasis>
: Adult with head, neck and eleven trunk segments (
<figureCitation id="6EA42A37FFA3D24FFEA824CCFE0AF838" box="[340,481,509,534]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFE0C24CCFDFEF838" box="[496,533,509,534]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9A</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFDDE24CCFDB3F838" box="[546,600,509,534]" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">10A</figureCitation>
). See table 3 for measurements, and table 4 for positions of cuticular structures (sensory spots, glandular cell outlets, acicular spines, tubules and sieve plates).
</paragraph>
<paragraph id="F62036B2FFA3D24FFF39274CFE38FFD8" blockId="11.[136,738,253,1974]" pageId="11" pageNumber="12">
Head consists of retractable mouth cone and introvert (
<figureCitation id="6EA42A37FFA3D24FFEB927ACFE5EF899" box="[325,437,669,695]" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">Figs. 10</figureCitation>
A-C). Mouth cone with inner oral styles and nine outer oral styles. Exact number and arrangement of inner oral styles not observed. Each outer oral style composed of rectangular basal part and triangular distal part (
<figureCitation id="6EA42A37FFA3D24FFF30260CFED4F978" box="[204,319,829,854]" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">Fig. 10B</figureCitation>
). Basal parts of outer oral styles alternate in size: five large anterior to odd sectors of introvert, and four slightly smaller ones in even sectors (
<figureCitation id="6EA42A37FFA3D24FFECA26ACFE5AF998" box="[310,433,925,950]" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">Figs. 10B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFE4326ACFE34F998" box="[447,479,925,950]" captionStart="Fig" captionStartId="16.[174,208,1925,1945]" captionTargetBox="[430,1192,972,1900]" captionTargetId="figure-340@16.[430,1192,969,1900]" captionTargetPageId="16" captionText="Fig. 11. Diagram of mouth cone, introvert, and placids in Echinoderes serratulus sp. nov. Grey area and heavy line arcs show mouth cone and placids, respectively. The table lists the scalid arrangement by sector." figureDoi="http://doi.org/10.5281/zenodo.10659231" httpUri="https://zenodo.org/record/10659231/files/figure.png" pageId="11" pageNumber="12">11</figureCitation>
). Introvert composed of seven rings of spinoscalids and one ring of trichoscalids (
<figureCitation id="6EA42A37FFA3D24FFED726ECFE4DF9D9" box="[299,422,989,1015]" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">Figs. 10C</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFE4F26ECFE39F9D8" box="[435,466,989,1014]" captionStart="Fig" captionStartId="16.[174,208,1925,1945]" captionTargetBox="[430,1192,972,1900]" captionTargetId="figure-340@16.[430,1192,969,1900]" captionTargetPageId="16" captionText="Fig. 11. Diagram of mouth cone, introvert, and placids in Echinoderes serratulus sp. nov. Grey area and heavy line arcs show mouth cone and placids, respectively. The table lists the scalid arrangement by sector." figureDoi="http://doi.org/10.5281/zenodo.10659231" httpUri="https://zenodo.org/record/10659231/files/figure.png" pageId="11" pageNumber="12">11</figureCitation>
). Ring 01 includes ten primary spinoscalids, each with basal sheath and smooth long end-piece (
<figureCitation id="6EA42A37FFA3D24FFE3F212CFDD1FE19" box="[451,570,1053,1079]" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">Fig. 10C</figureCitation>
). Each basal sheath with three overlapping fringes. Proximal fringe extends into three flat projections, like a trident, covering next fringe. Middle fringe with two lateral projections overlapping end-piece. Distal fringe with five threads projecting between two projections of middle fringe. End-piece of primary spinoscalids is longest unit. Rings 02 and 04 with 10 spinoscalids; rings 03 and 05 with 20 spinoscalids. Spinoscalids in rings 02-05 similar in length. Rings 06 and 07 not examined with detail, but ring 06 with at least seven, and ring 07 with at least ten relatively short spinoscalids (
<figureCitation id="6EA42A37FFA3D24FFD8020ACFD3EFF98" box="[636,725,1437,1462]" captionStart="Fig" captionStartId="16.[174,208,1925,1945]" captionTargetBox="[430,1192,972,1900]" captionTargetId="figure-340@16.[430,1192,969,1900]" captionTargetPageId="16" captionText="Fig. 11. Diagram of mouth cone, introvert, and placids in Echinoderes serratulus sp. nov. Grey area and heavy line arcs show mouth cone and placids, respectively. The table lists the scalid arrangement by sector." figureDoi="http://doi.org/10.5281/zenodo.10659231" httpUri="https://zenodo.org/record/10659231/files/figure.png" pageId="11" pageNumber="12">Fig. 11</figureCitation>
). Six trichoscalids attached with trichoscalid plate in sectors 2, 4, 5, 7, 8, and 10.
</paragraph>
<paragraph id="F62036B2FFA3D24FFF3920CCFF38FCB8" blockId="11.[136,738,253,1974]" pageId="11" pageNumber="12">
Neck with 16 placids (
<figureCitation id="6EA42A37FFA3D24FFE2020CCFD9BFC38" box="[476,624,1533,1558]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFD8320CCFD4EFC38" box="[639,677,1533,1558]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9A</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFD4F20CCFD38FC38" box="[691,723,1533,1558]" captionStart="Fig" captionStartId="16.[174,208,1925,1945]" captionTargetBox="[430,1192,972,1900]" captionTargetId="figure-340@16.[430,1192,969,1900]" captionTargetPageId="16" captionText="Fig. 11. Diagram of mouth cone, introvert, and placids in Echinoderes serratulus sp. nov. Grey area and heavy line arcs show mouth cone and placids, respectively. The table lists the scalid arrangement by sector." figureDoi="http://doi.org/10.5281/zenodo.10659231" httpUri="https://zenodo.org/record/10659231/files/figure.png" pageId="11" pageNumber="12">11</figureCitation>
). Midventral placid broadest (ca. 17 μm wide at base, ca. 12 μm wide at tip); remaining placids with similar size (ca. 10 μm wide at base, ca. 4 μm wide at tip).
</paragraph>
<paragraph id="F62036B2FFA3D24FFF3923ACFB4FFB79" blockId="11.[136,738,253,1974]" lastBlockId="11.[813,1415,253,1975]" pageId="11" pageNumber="12">
Segment 1 consists of complete cuticular ring (
<figureCitation id="6EA42A37FFA3D24FFF73238CFECEFCF8" box="[143,293,1725,1750]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFEC8238CFEB7FCF8" box="[308,348,1725,1750]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9A</figureCitation>
). Non-bracteate cuticular hairs densely cover entire segment. Paired, rounded subdorsal and laterodorsal sensory spots located close to anterior margin of segment (
<figureCitation id="6EA42A37FFA3D24FFDC7222CFD48FD18" box="[571,675,1821,1846]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFD53222CFD3EFD19" box="[687,725,1821,1847]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9C</figureCitation>
). Rounded ventromedial sensory spots centered between anterior and posterior margins (
<figureCitation id="6EA42A37FFA3D24FFD5C226CFF45FDB8" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFF41224CFF1EFDB8" box="[189,245,1917,1942]" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">10D</figureCitation>
). Type-1 glandular cell outlets situated anteriorly in middorsal and lateroventral positions (
<figureCitation id="6EA42A37FFA3D24FFCC825CCFC24FB38" box="[820,975,253,278]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFC1C25CCFBDBFB39" box="[992,1072,253,279]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9A, C</figureCitation>
). Primary pectinate fringe conspicuous, with longer fringe tips, especially on ventral side (
<figureCitation id="6EA42A37FFA3D24FFC3F240CFBA0FB78" box="[963,1099,317,342]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFBAA240CFB8EFB79" box="[1110,1125,317,343]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9</figureCitation>
A-C).
</paragraph>
<paragraph id="F62036B2FFA3D24FFC95246CFAB2F9D9" blockId="11.[813,1415,253,1975]" pageId="11" pageNumber="12">
Segment 2 also with complete cuticular ring (
<figureCitation id="6EA42A37FFA3D24FFCC8244CFC23FBB8" box="[820,968,381,406]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
), with thick pachycyclus at anterior margin (
<figureCitation id="6EA42A37FFA3D24FFC6124ACFBABFB98" box="[925,1088,413,438]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFBAE24ACFB88FB99" box="[1106,1123,413,439]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9</figureCitation>
A-C). Entire cuticular surface, except anterior area, covered with bracteate cuticular hairs on this and following eight segments. Oval sensory spots in middorsal, laterodorsal, midlateral, and ventromedial positions (
<figureCitation id="6EA42A37FFA3D24FFC5A270CFBDAF878" box="[934,1073,573,598]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFBC2270CFBA7F879" box="[1086,1100,573,599]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9</figureCitation>
A-C, 10D). Unpaired type- 1 glandular cell outlet present in middorsal position (
<figureCitation id="6EA42A37FFA3D24FFCC9274CFC77F8B8" box="[821,924,637,662]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Fig. 8A</figureCitation>
). Paired type-1 glandular cell outlets close to anterior margin in ventromedial position on this and following eight segments (
<figureCitation id="6EA42A37FFA3D24FFB09278CFA69F8F8" box="[1269,1410,701,726]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8B, D</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFCD127ECFC9FF8D8" box="[813,884,733,758]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9A, B</figureCitation>
). Type-2 glandular cell outlets in subdorsal, laterodorsal, and ventrolateral positions (
<figureCitation id="6EA42A37FFA3D24FFAEB27CCFCA9F918" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFCB2262CFCB6F919" box="[846,861,797,823]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9</figureCitation>
A-C). Type-2 glandular cell outlets show single large pore in LM and
<collectionCode id="908EAE77FFA3D24FFBCB260CFB9AF979" box="[1079,1137,829,855]" pageId="11" pageNumber="12">SEM</collectionCode>
(
<figureCitation id="6EA42A37FFA3D24FFB7E260CFAFAF979" box="[1154,1297,829,855]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">Figs. 9B, C</figureCitation>
; see also figures 12A, B for type-2 glandular cell outlet on segments 6 and 8). Posterior margin of segment with primary pectinate fringe having long tips, with ventromedial tips obliquely orientated, pointing towards the midventral line (
<figureCitation id="6EA42A37FFA3D24FFB8426ECFAEBF9D8" box="[1144,1280,989,1014]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFAF726ECFAF1F9D9" box="[1291,1306,989,1015]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9</figureCitation>
A-C).
</paragraph>
<paragraph id="F62036B2FFA3D24FFC9526CCFC9BFFB9" blockId="11.[813,1415,253,1975]" pageId="11" pageNumber="12">
Segment 3 and following eight segments consist of one tergal and two sternal plates (
<figureCitation id="6EA42A37FFA3D24FFAB5212CFC9EFE78" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFC78210CFC40FE78" box="[900,939,1085,1110]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9A</figureCitation>
). Each plate with thicker pachycycli in anterior area (
<figureCitation id="6EA42A37FFA3D24FFC26216CFB82FE58" box="[986,1129,1117,1142]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFB84216CFB2AFE58" box="[1144,1217,1117,1142]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9A, B</figureCitation>
). Sensory spots in subdorsal, midlateral, and sublateral positions (
<figureCitation id="6EA42A37FFA3D24FFCC821ACFC43FE98" box="[820,936,1181,1206]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFC4421ACFC0BFE98" box="[952,992,1181,1206]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9A</figureCitation>
). Two unpaired type-1 glandular cell outlets present middorsally (
<figureCitation id="6EA42A37FFA3D24FFB49218CFAFFFEF8" box="[1205,1300,1213,1238]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Fig. 8A</figureCitation>
). Primary pectinate fringes on this and following two segments as on segment 2. Secondary pectinate fringes, consisting of single belt of minute teeth, visible on anterior dorsal plate in
<collectionCode id="908EAE77FFA3D24FFB4F200CFB06FF79" box="[1203,1261,1341,1367]" pageId="11" pageNumber="12">SEM</collectionCode>
observation, at least on this and following five segments (
<figureCitation id="6EA42A37FFA3D24FFAAB206CFC8FFFB9" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">Fig. 10G</figureCitation>
).
</paragraph>
<paragraph id="F62036B2FFA3D24FFC9520ACFC98FCB8" blockId="11.[813,1415,253,1975]" pageId="11" pageNumber="12">
Segment 4 with short middorsal acicular spine (
<figureCitation id="6EA42A37FFA3D24FFC80208CFC0DFFF8" box="[892,998,1469,1494]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFC0E208CFBC1FFF9" box="[1010,1066,1469,1495]" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">10G</figureCitation>
). Paired laterodorsal sensory spots and paired subdorsal type-2 glandular cell outlets present (
<figureCitation id="6EA42A37FFA3D24FFBFD20CCFB9FFC38" box="[1025,1140,1533,1558]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFB7820CCFB54FC39" box="[1156,1215,1533,1559]" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">10G</figureCitation>
). Paired type-1 glandular cell outlets present paradorsally on this and following six segments. Fringe tips on dorsal, lateral, and ventral sides similar in length and width.
</paragraph>
<paragraph id="F62036B2FFA3D24FFC9523ACFAA3FD18" blockId="11.[813,1415,253,1975]" pageId="11" pageNumber="12">
Segment 5 with lateroventral tubules (
<figureCitation id="6EA42A37FFA3D24FFABB23ACFCB9FCF8" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFC9F238CFC61FCF8" box="[867,906,1725,1750]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9D</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFC66238CFC18FCF8" box="[922,1011,1725,1750]" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">10D, F</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFBFF238CFBD0FCF8" box="[1027,1083,1725,1750]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[211,1335,250,1816]" captionTargetId="figure-99@17.[211,1335,249,1817]" captionTargetPageId="17" captionText="Fig. 12. Echinoderes serratulus sp. nov., scanning electron micrographs. A, segments 5 and 6, lateral view; B, segments 8 and 9, lateroventral view; C, segment 6, sublateral view; D, segment 9, sublateral view; E, segments 9-11 in female, dorsal view; F, segments 10 and 11 in male, lateral view; G, segments 10 and 11 in female, lateral view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; pe, penile spine; po, pore; si, sieve area; sls, sublateral acicular spine." figureDoi="http://doi.org/10.5281/zenodo.10659233" httpUri="https://zenodo.org/record/10659233/files/figure.png" pageId="11" pageNumber="12">12A</figureCitation>
). Paired sensory spots in subdorsal, midlateral, and ventromedial positions (
<figureCitation id="6EA42A37FFA3D24FFCC823CCFC2AFD38" box="[820,961,1789,1814]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFC2C23CCFC1EFD38" box="[976,1013,1789,1814]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9A</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFBFE23CCFBCAFD38" box="[1026,1057,1789,1814]" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">10</figureCitation>
D-F). Pair of type-2 glandular cell outlets in midlateral position (
<figureCitation id="6EA42A37FFA3D24FFB4F222CFAD0FD18" box="[1203,1339,1821,1846]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
).
</paragraph>
<paragraph id="F62036B2FFA3D249FC95220CFE4BFB18" blockId="11.[813,1415,253,1975]" lastBlockId="13.[136,737,253,406]" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
Segment 6 similar to segment 5 except for absence of lateroventral tubules and presence of short sublateral acicular spines (
<figureCitation id="6EA42A37FFA3D24FFB11224CFA6AFDB8" box="[1261,1409,1917,1942]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="11" pageNumber="12">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFCD122ACFC9DFD98" box="[813,886,1949,1974]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="11" pageNumber="12">9A, D</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFC7822ACFC33FD98" box="[900,984,1949,1974]" captionStart="Fig" captionStartId="15.[136,172,1840,1860]" captionTargetBox="[184,1366,429,1814]" captionTargetId="figure-89@15.[183,1366,428,1815]" captionTargetPageId="15" captionText="Fig. 10. Echinoderes serratulus sp. nov., scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe." figureDoi="http://doi.org/10.5281/zenodo.10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="11" pageNumber="12">10D, F</figureCitation>
,
<figureCitation id="6EA42A37FFA3D24FFC1A22ACFBABFD99" box="[998,1088,1949,1975]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[211,1335,250,1816]" captionTargetId="figure-99@17.[211,1335,249,1817]" captionTargetPageId="17" captionText="Fig. 12. Echinoderes serratulus sp. nov., scanning electron micrographs. A, segments 5 and 6, lateral view; B, segments 8 and 9, lateroventral view; C, segment 6, sublateral view; D, segment 9, sublateral view; E, segments 9-11 in female, dorsal view; F, segments 10 and 11 in male, lateral view; G, segments 10 and 11 in female, lateral view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; pe, penile spine; po, pore; si, sieve area; sls, sublateral acicular spine." figureDoi="http://doi.org/10.5281/zenodo.10659233" httpUri="https://zenodo.org/record/10659233/files/figure.png" pageId="11" pageNumber="12">12A, C</figureCitation>
). Primary pectinate fringes of this and following three segments with long, conspicuous fringe tips.
</paragraph>
<caption id="A2E0663AFFA4D248FF5225CCFD07FB17" pageId="12" pageNumber="13" startId="12.[174,242,253,278]" targetType="table">
<paragraph id="F62036B2FFA4D248FF5225CCFD07FB17" blockId="12.[174,1452,253,313]" pageId="12" pageNumber="13">
<emphasis id="C4EBEAA0FFA4D248FF5225CCFEFBFB38" bold="true" box="[174,272,253,278]" pageId="12" pageNumber="13">Table 3.</emphasis>
Measurements for adult
<taxonomicName id="319F4D31FFA4D248FDB825CCFCBDFB38" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[580,854,253,278]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="12" pageNumber="13" phylum="Kinorhyncha" rank="species" species="serratulus" status="sp. nov.">
<emphasis id="C4EBEAA0FFA4D248FDB825CCFCBDFB38" box="[580,854,253,278]" italics="true" pageId="12" pageNumber="13">Echinoderes serratulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFD857DBFFA4D248FCA225CCFC57FB38" box="[862,956,253,278]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
(in micrometers). Columns N and SD are sample size and standard deviation, respectively
</paragraph>
</caption>
<paragraph id="F62036B2FFA4D248FF4E2450FAA2FE97" pageId="12" pageNumber="13">
<table id="849FC412FFA42DBBFF4E2450FAA1FE97" box="[178,1354,353,1209]" gridcols="5" gridrows="29" pageId="12" pageNumber="13">
<tr id="48AF34F0FFA42DBBFF4E2450FAA1FB5B" box="[178,1354,353,373]" gridrow="0" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E2450FEDBFB5B" box="[178,304,353,373]" gridcol="0" gridrow="0" pageId="12" pageNumber="13">Character</th>
<th id="0B7E5D8CFFA42DBBFDD92450FDDFFB5B" box="[549,564,353,373]" gridcol="1" gridrow="0" pageId="12" pageNumber="13">N</th>
<th id="0B7E5D8CFFA42DBBFD0A2450FC88FB5B" box="[758,867,353,373]" gridcol="2" gridrow="0" pageId="12" pageNumber="13">Range</th>
<th id="0B7E5D8CFFA42DBBFBF22450FBA0FB5B" box="[1038,1099,353,373]" gridcol="3" gridrow="0" pageId="12" pageNumber="13">Mean</th>
<th id="0B7E5D8CFFA42DBBFAF12450FAA1FB5B" box="[1293,1354,353,373]" gridcol="4" gridrow="0" pageId="12" pageNumber="13">SD</th>
</tr>
<tr id="48AF34F0FFA42DBBFF4E24A8FAA1FB83" box="[178,1354,409,429]" gridrow="1" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E24A8FEDBFB83" box="[178,304,409,429]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">TL</th>
<td id="0B7E5D8CFFA42DBBFDD924A8FDDFFB83" box="[549,564,409,429]" gridcol="1" gridrow="1" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A24A8FC88FB83" box="[758,867,409,429]" gridcol="2" gridrow="1" pageId="12" pageNumber="13">321-359</td>
<td id="0B7E5D8CFFA42DBBFBF224A8FBA0FB83" box="[1038,1099,409,429]" gridcol="3" gridrow="1" pageId="12" pageNumber="13">348</td>
<td id="0B7E5D8CFFA42DBBFAF124A8FAA1FB83" box="[1293,1354,409,429]" gridcol="4" gridrow="1" pageId="12" pageNumber="13">15.32</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2487FAA1FBE4" box="[178,1354,438,458]" gridrow="2" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E2487FEDBFBE4" box="[178,304,438,458]" gridcol="0" gridrow="2" pageId="12" pageNumber="13">MSW-6</th>
<td id="0B7E5D8CFFA42DBBFDD92487FDDFFBE4" box="[549,564,438,458]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A2487FC88FBE4" box="[758,867,438,458]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">62-69</td>
<td id="0B7E5D8CFFA42DBBFBF22487FBA0FBE4" box="[1038,1099,438,458]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">66</td>
<td id="0B7E5D8CFFA42DBBFAF12487FAA1FBE4" box="[1293,1354,438,458]" gridcol="4" gridrow="2" pageId="12" pageNumber="13">3.09</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E24E2FAA1FBC9" box="[178,1354,467,487]" gridrow="3" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E24E2FEDBFBC9" box="[178,304,467,487]" gridcol="0" gridrow="3" pageId="12" pageNumber="13">MSW-6/TL</th>
<td id="0B7E5D8CFFA42DBBFDD924E2FDDFFBC9" box="[549,564,467,487]" gridcol="1" gridrow="3" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A24E2FC88FBC9" box="[758,867,467,487]" gridcol="2" gridrow="3" pageId="12" pageNumber="13">17.9-19.9%</td>
<td id="0B7E5D8CFFA42DBBFBF224E2FBA0FBC9" box="[1038,1099,467,487]" gridcol="3" gridrow="3" pageId="12" pageNumber="13">18.9%</td>
<td id="0B7E5D8CFFA42DBBFAF124E2FAA1FBC9" box="[1293,1354,467,487]" gridcol="4" gridrow="3" pageId="12" pageNumber="13">1.00%</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E24C1FAA1F82A" box="[178,1354,496,516]" gridrow="4" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E24C1FEDBF82A" box="[178,304,496,516]" gridcol="0" gridrow="4" pageId="12" pageNumber="13">SW-10</th>
<td id="0B7E5D8CFFA42DBBFDD924C1FDDFF82A" box="[549,564,496,516]" gridcol="1" gridrow="4" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A24C1FC88F82A" box="[758,867,496,516]" gridcol="2" gridrow="4" pageId="12" pageNumber="13">54-59</td>
<td id="0B7E5D8CFFA42DBBFBF224C1FBA0F82A" box="[1038,1099,496,516]" gridcol="3" gridrow="4" pageId="12" pageNumber="13">56</td>
<td id="0B7E5D8CFFA42DBBFAF124C1FAA1F82A" box="[1293,1354,496,516]" gridcol="4" gridrow="4" pageId="12" pageNumber="13">1.76</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E273CFAA1F80F" box="[178,1354,525,545]" gridrow="5" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E273CFEDBF80F" box="[178,304,525,545]" gridcol="0" gridrow="5" pageId="12" pageNumber="13">SW-10/TL</th>
<td id="0B7E5D8CFFA42DBBFDD9273CFDDFF80F" box="[549,564,525,545]" gridcol="1" gridrow="5" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A273CFC88F80F" box="[758,867,525,545]" gridcol="2" gridrow="5" pageId="12" pageNumber="13">15.1-17.0%</td>
<td id="0B7E5D8CFFA42DBBFBF2273CFBA0F80F" box="[1038,1099,525,545]" gridcol="3" gridrow="5" pageId="12" pageNumber="13">16.1%</td>
<td id="0B7E5D8CFFA42DBBFAF1273CFAA1F80F" box="[1293,1354,525,545]" gridcol="4" gridrow="5" pageId="12" pageNumber="13">0.81%</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E271BFAA1F810" box="[178,1354,554,574]" gridrow="6" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E271BFEDBF810" box="[178,304,554,574]" gridcol="0" gridrow="6" pageId="12" pageNumber="13">S1</th>
<td id="0B7E5D8CFFA42DBBFDD9271BFDDFF810" box="[549,564,554,574]" gridcol="1" gridrow="6" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A271BFC88F810" box="[758,867,554,574]" gridcol="2" gridrow="6" pageId="12" pageNumber="13">36-39</td>
<td id="0B7E5D8CFFA42DBBFBF2271BFBA0F810" box="[1038,1099,554,574]" gridcol="3" gridrow="6" pageId="12" pageNumber="13">37</td>
<td id="0B7E5D8CFFA42DBBFAF1271BFAA1F810" box="[1293,1354,554,574]" gridcol="4" gridrow="6" pageId="12" pageNumber="13">1.25</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2777FAA1F875" box="[178,1354,582,603]" gridrow="7" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E2777FEDBF875" box="[178,304,582,603]" gridcol="0" gridrow="7" pageId="12" pageNumber="13">S2</th>
<td id="0B7E5D8CFFA42DBBFDD92777FDDFF875" box="[549,564,582,603]" gridcol="1" gridrow="7" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A2777FC88F875" box="[758,867,582,603]" gridcol="2" gridrow="7" pageId="12" pageNumber="13">33-37</td>
<td id="0B7E5D8CFFA42DBBFBF22777FBA0F875" box="[1038,1099,582,603]" gridcol="3" gridrow="7" pageId="12" pageNumber="13">35</td>
<td id="0B7E5D8CFFA42DBBFAF12777FAA1F875" box="[1293,1354,582,603]" gridcol="4" gridrow="7" pageId="12" pageNumber="13">1.82</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2752FAA1F856" box="[178,1354,611,632]" gridrow="8" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E2752FEDBF856" box="[178,304,611,632]" gridcol="0" gridrow="8" pageId="12" pageNumber="13">S3</th>
<td id="0B7E5D8CFFA42DBBFDD92752FDDFF856" box="[549,564,611,632]" gridcol="1" gridrow="8" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A2752FC88F856" box="[758,867,611,632]" gridcol="2" gridrow="8" pageId="12" pageNumber="13">28-31</td>
<td id="0B7E5D8CFFA42DBBFBF22752FBA0F856" box="[1038,1099,611,632]" gridcol="3" gridrow="8" pageId="12" pageNumber="13">30</td>
<td id="0B7E5D8CFFA42DBBFAF12752FAA1F856" box="[1293,1354,611,632]" gridcol="4" gridrow="8" pageId="12" pageNumber="13">1.21</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E27B1FAA1F8BA" box="[178,1354,640,660]" gridrow="9" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E27B1FEDBF8BA" box="[178,304,640,660]" gridcol="0" gridrow="9" pageId="12" pageNumber="13">S4</th>
<td id="0B7E5D8CFFA42DBBFDD927B1FDDFF8BA" box="[549,564,640,660]" gridcol="1" gridrow="9" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A27B1FC88F8BA" box="[758,867,640,660]" gridcol="2" gridrow="9" pageId="12" pageNumber="13">31-34</td>
<td id="0B7E5D8CFFA42DBBFBF227B1FBA0F8BA" box="[1038,1099,640,660]" gridcol="3" gridrow="9" pageId="12" pageNumber="13">33</td>
<td id="0B7E5D8CFFA42DBBFAF127B1FAA1F8BA" box="[1293,1354,640,660]" gridcol="4" gridrow="9" pageId="12" pageNumber="13">1.28</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E27ACFAA1F89F" box="[178,1354,669,689]" gridrow="10" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E27ACFEDBF89F" box="[178,304,669,689]" gridcol="0" gridrow="10" pageId="12" pageNumber="13">S5</th>
<td id="0B7E5D8CFFA42DBBFDD927ACFDDFF89F" box="[549,564,669,689]" gridcol="1" gridrow="10" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A27ACFC88F89F" box="[758,867,669,689]" gridcol="2" gridrow="10" pageId="12" pageNumber="13">34-38</td>
<td id="0B7E5D8CFFA42DBBFBF227ACFBA0F89F" box="[1038,1099,669,689]" gridcol="3" gridrow="10" pageId="12" pageNumber="13">37</td>
<td id="0B7E5D8CFFA42DBBFAF127ACFAA1F89F" box="[1293,1354,669,689]" gridcol="4" gridrow="10" pageId="12" pageNumber="13">1.68</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E278BFAA1F8E0" box="[178,1354,698,718]" gridrow="11" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E278BFEDBF8E0" box="[178,304,698,718]" gridcol="0" gridrow="11" pageId="12" pageNumber="13">S6</th>
<td id="0B7E5D8CFFA42DBBFDD9278BFDDFF8E0" box="[549,564,698,718]" gridcol="1" gridrow="11" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A278BFC88F8E0" box="[758,867,698,718]" gridcol="2" gridrow="11" pageId="12" pageNumber="13">38-42</td>
<td id="0B7E5D8CFFA42DBBFBF2278BFBA0F8E0" box="[1038,1099,698,718]" gridcol="3" gridrow="11" pageId="12" pageNumber="13">40</td>
<td id="0B7E5D8CFFA42DBBFAF1278BFAA1F8E0" box="[1293,1354,698,718]" gridcol="4" gridrow="11" pageId="12" pageNumber="13">1.51</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E27E6FAA1F8C5" box="[178,1354,727,747]" gridrow="12" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E27E6FEDBF8C5" box="[178,304,727,747]" gridcol="0" gridrow="12" pageId="12" pageNumber="13">S7</th>
<td id="0B7E5D8CFFA42DBBFDD927E6FDDFF8C5" box="[549,564,727,747]" gridcol="1" gridrow="12" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A27E6FC88F8C5" box="[758,867,727,747]" gridcol="2" gridrow="12" pageId="12" pageNumber="13">40-48</td>
<td id="0B7E5D8CFFA42DBBFBF227E6FBA0F8C5" box="[1038,1099,727,747]" gridcol="3" gridrow="12" pageId="12" pageNumber="13">45</td>
<td id="0B7E5D8CFFA42DBBFAF127E6FAA1F8C5" box="[1293,1354,727,747]" gridcol="4" gridrow="12" pageId="12" pageNumber="13">2.86</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E27C5FAA1F926" box="[178,1354,756,776]" gridrow="13" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E27C5FEDBF926" box="[178,304,756,776]" gridcol="0" gridrow="13" pageId="12" pageNumber="13">S8</th>
<td id="0B7E5D8CFFA42DBBFDD927C5FDDFF926" box="[549,564,756,776]" gridcol="1" gridrow="13" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A27C5FC88F926" box="[758,867,756,776]" gridcol="2" gridrow="13" pageId="12" pageNumber="13">46-53</td>
<td id="0B7E5D8CFFA42DBBFBF227C5FBA0F926" box="[1038,1099,756,776]" gridcol="3" gridrow="13" pageId="12" pageNumber="13">50</td>
<td id="0B7E5D8CFFA42DBBFAF127C5FAA1F926" box="[1293,1354,756,776]" gridcol="4" gridrow="13" pageId="12" pageNumber="13">2.67</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2620FAA1F90B" box="[178,1354,785,805]" gridrow="14" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E2620FEDBF90B" box="[178,304,785,805]" gridcol="0" gridrow="14" pageId="12" pageNumber="13">S9</th>
<td id="0B7E5D8CFFA42DBBFDD92620FDDFF90B" box="[549,564,785,805]" gridcol="1" gridrow="14" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A2620FC88F90B" box="[758,867,785,805]" gridcol="2" gridrow="14" pageId="12" pageNumber="13">47-53</td>
<td id="0B7E5D8CFFA42DBBFBF22620FBA0F90B" box="[1038,1099,785,805]" gridcol="3" gridrow="14" pageId="12" pageNumber="13">50</td>
<td id="0B7E5D8CFFA42DBBFAF12620FAA1F90B" box="[1293,1354,785,805]" gridcol="4" gridrow="14" pageId="12" pageNumber="13">2.37</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E261FFAA1F96C" box="[178,1354,814,834]" gridrow="15" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E261FFEDBF96C" box="[178,304,814,834]" gridcol="0" gridrow="15" pageId="12" pageNumber="13">S10</th>
<td id="0B7E5D8CFFA42DBBFDD9261FFDDFF96C" box="[549,564,814,834]" gridcol="1" gridrow="15" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A261FFC88F96C" box="[758,867,814,834]" gridcol="2" gridrow="15" pageId="12" pageNumber="13">44-46</td>
<td id="0B7E5D8CFFA42DBBFBF2261FFBA0F96C" box="[1038,1099,814,834]" gridcol="3" gridrow="15" pageId="12" pageNumber="13">45</td>
<td id="0B7E5D8CFFA42DBBFAF1261FFAA1F96C" box="[1293,1354,814,834]" gridcol="4" gridrow="15" pageId="12" pageNumber="13">0.66</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E267BFAA1F971" box="[178,1354,842,863]" gridrow="16" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E267BFEDBF971" box="[178,304,842,863]" gridcol="0" gridrow="16" pageId="12" pageNumber="13">S11</th>
<td id="0B7E5D8CFFA42DBBFDD9267BFDDFF971" box="[549,564,842,863]" gridcol="1" gridrow="16" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A267BFC88F971" box="[758,867,842,863]" gridcol="2" gridrow="16" pageId="12" pageNumber="13">44-48</td>
<td id="0B7E5D8CFFA42DBBFBF2267BFBA0F971" box="[1038,1099,842,863]" gridcol="3" gridrow="16" pageId="12" pageNumber="13">46</td>
<td id="0B7E5D8CFFA42DBBFAF1267BFAA1F971" box="[1293,1354,842,863]" gridcol="4" gridrow="16" pageId="12" pageNumber="13">1.75</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2659FAA1F952" box="[178,1354,872,892]" gridrow="17" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E2659FEDBF952" box="[178,304,872,892]" gridcol="0" gridrow="17" pageId="12" pageNumber="13">MD4 (ac)</th>
<td id="0B7E5D8CFFA42DBBFDD92659FDDFF952" box="[549,564,872,892]" gridcol="1" gridrow="17" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A2659FC88F952" box="[758,867,872,892]" gridcol="2" gridrow="17" pageId="12" pageNumber="13">10-14</td>
<td id="0B7E5D8CFFA42DBBFBF22659FBA0F952" box="[1038,1099,872,892]" gridcol="3" gridrow="17" pageId="12" pageNumber="13">12</td>
<td id="0B7E5D8CFFA42DBBFAF12659FAA1F952" box="[1293,1354,872,892]" gridcol="4" gridrow="17" pageId="12" pageNumber="13">1.68</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E26B5FAA1F9B6" box="[178,1354,900,920]" gridrow="18" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E26B5FEDBF9B6" box="[178,304,900,920]" gridcol="0" gridrow="18" pageId="12" pageNumber="13">LV5 (tu)</th>
<td id="0B7E5D8CFFA42DBBFDD926B5FDDFF9B6" box="[549,564,900,920]" gridcol="1" gridrow="18" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A26B5FC88F9B6" box="[758,867,900,920]" gridcol="2" gridrow="18" pageId="12" pageNumber="13">11-13</td>
<td id="0B7E5D8CFFA42DBBFBF226B5FBA0F9B6" box="[1038,1099,900,920]" gridcol="3" gridrow="18" pageId="12" pageNumber="13">12</td>
<td id="0B7E5D8CFFA42DBBFAF126B5FAA1F9B6" box="[1293,1354,900,920]" gridcol="4" gridrow="18" pageId="12" pageNumber="13">1.14</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2690FAA1F99B" box="[178,1354,929,949]" gridrow="19" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E2690FEDBF99B" box="[178,304,929,949]" gridcol="0" gridrow="19" pageId="12" pageNumber="13">SL6 (ac)</th>
<td id="0B7E5D8CFFA42DBBFDD92690FDDFF99B" box="[549,564,929,949]" gridcol="1" gridrow="19" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A2690FC88F99B" box="[758,867,929,949]" gridcol="2" gridrow="19" pageId="12" pageNumber="13">9-12</td>
<td id="0B7E5D8CFFA42DBBFBF22690FBA0F99B" box="[1038,1099,929,949]" gridcol="3" gridrow="19" pageId="12" pageNumber="13">10</td>
<td id="0B7E5D8CFFA42DBBFAF12690FAA1F99B" box="[1293,1354,929,949]" gridcol="4" gridrow="19" pageId="12" pageNumber="13">1.53</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E268FFAA1F9FC" box="[178,1354,958,978]" gridrow="20" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E268FFEDBF9FC" box="[178,304,958,978]" gridcol="0" gridrow="20" pageId="12" pageNumber="13">SL7 (ac)</th>
<td id="0B7E5D8CFFA42DBBFDD9268FFDDFF9FC" box="[549,564,958,978]" gridcol="1" gridrow="20" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A268FFC88F9FC" box="[758,867,958,978]" gridcol="2" gridrow="20" pageId="12" pageNumber="13">9-13</td>
<td id="0B7E5D8CFFA42DBBFBF2268FFBA0F9FC" box="[1038,1099,958,978]" gridcol="3" gridrow="20" pageId="12" pageNumber="13">11</td>
<td id="0B7E5D8CFFA42DBBFAF1268FFAA1F9FC" box="[1293,1354,958,978]" gridcol="4" gridrow="20" pageId="12" pageNumber="13">1.71</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E26EAFAA1F9C1" box="[178,1354,987,1007]" gridrow="21" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E26EAFEDBF9C1" box="[178,304,987,1007]" gridcol="0" gridrow="21" pageId="12" pageNumber="13">LV8 (tu)</th>
<td id="0B7E5D8CFFA42DBBFDD926EAFDDFF9C1" box="[549,564,987,1007]" gridcol="1" gridrow="21" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A26EAFC88F9C1" box="[758,867,987,1007]" gridcol="2" gridrow="21" pageId="12" pageNumber="13">9-13</td>
<td id="0B7E5D8CFFA42DBBFBF226EAFBA0F9C1" box="[1038,1099,987,1007]" gridcol="3" gridrow="21" pageId="12" pageNumber="13">12</td>
<td id="0B7E5D8CFFA42DBBFAF126EAFAA1F9C1" box="[1293,1354,987,1007]" gridcol="4" gridrow="21" pageId="12" pageNumber="13">1.59</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E26C9FAA1FE22" box="[178,1354,1016,1036]" gridrow="22" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E26C9FEDBFE22" box="[178,304,1016,1036]" gridcol="0" gridrow="22" pageId="12" pageNumber="13">ML9 (tu)</th>
<td id="0B7E5D8CFFA42DBBFDD926C9FDDFFE22" box="[549,564,1016,1036]" gridcol="1" gridrow="22" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A26C9FC88FE22" box="[758,867,1016,1036]" gridcol="2" gridrow="22" pageId="12" pageNumber="13">16-21</td>
<td id="0B7E5D8CFFA42DBBFBF226C9FBA0FE22" box="[1038,1099,1016,1036]" gridcol="3" gridrow="22" pageId="12" pageNumber="13">19</td>
<td id="0B7E5D8CFFA42DBBFAF126C9FAA1FE22" box="[1293,1354,1016,1036]" gridcol="4" gridrow="22" pageId="12" pageNumber="13">1.96</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2124FAA1FE07" box="[178,1354,1045,1065]" gridrow="23" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E2124FEDBFE07" box="[178,304,1045,1065]" gridcol="0" gridrow="23" pageId="12" pageNumber="13">LD10 (tu) (m)</th>
<td id="0B7E5D8CFFA42DBBFDD92124FDDFFE07" box="[549,564,1045,1065]" gridcol="1" gridrow="23" pageId="12" pageNumber="13">3</td>
<td id="0B7E5D8CFFA42DBBFD0A2124FC88FE07" box="[758,867,1045,1065]" gridcol="2" gridrow="23" pageId="12" pageNumber="13">39-41</td>
<td id="0B7E5D8CFFA42DBBFBF22124FBA0FE07" box="[1038,1099,1045,1065]" gridcol="3" gridrow="23" pageId="12" pageNumber="13">40</td>
<td id="0B7E5D8CFFA42DBBFAF12124FAA1FE07" box="[1293,1354,1045,1065]" gridcol="4" gridrow="23" pageId="12" pageNumber="13">1.11</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2103FAA1FE68" box="[178,1354,1074,1094]" gridrow="24" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E2103FEDBFE68" box="[178,304,1074,1094]" gridcol="0" gridrow="24" pageId="12" pageNumber="13">LD10 (tu) (f)</th>
<td id="0B7E5D8CFFA42DBBFDD92103FDDFFE68" box="[549,564,1074,1094]" gridcol="1" gridrow="24" pageId="12" pageNumber="13">2</td>
<td id="0B7E5D8CFFA42DBBFD0A2103FC88FE68" box="[758,867,1074,1094]" gridcol="2" gridrow="24" pageId="12" pageNumber="13">16-17</td>
<td id="0B7E5D8CFFA42DBBFBF22103FBA0FE68" box="[1038,1099,1074,1094]" gridcol="3" gridrow="24" pageId="12" pageNumber="13">17</td>
<td id="0B7E5D8CFFA42DBBFAF12103FAA1FE68" box="[1293,1354,1074,1094]" gridcol="4" gridrow="24" pageId="12" pageNumber="13">0.96</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E217FFAA1FE4D" box="[178,1354,1102,1123]" gridrow="25" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E217FFEDBFE4D" box="[178,304,1102,1123]" gridcol="0" gridrow="25" pageId="12" pageNumber="13">LTS</th>
<td id="0B7E5D8CFFA42DBBFDD9217FFDDFFE4D" box="[549,564,1102,1123]" gridcol="1" gridrow="25" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A217FFC88FE4D" box="[758,867,1102,1123]" gridcol="2" gridrow="25" pageId="12" pageNumber="13">44-49</td>
<td id="0B7E5D8CFFA42DBBFBF2217FFBA0FE4D" box="[1038,1099,1102,1123]" gridcol="3" gridrow="25" pageId="12" pageNumber="13">46</td>
<td id="0B7E5D8CFFA42DBBFAF1217FFAA1FE4D" box="[1293,1354,1102,1123]" gridcol="4" gridrow="25" pageId="12" pageNumber="13">1.65</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E215AFAA1FEAE" box="[178,1354,1131,1152]" gridrow="26" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E215AFEDBFEAE" box="[178,304,1131,1152]" gridcol="0" gridrow="26" pageId="12" pageNumber="13">LTAS</th>
<td id="0B7E5D8CFFA42DBBFDD9215AFDDFFEAE" box="[549,564,1131,1152]" gridcol="1" gridrow="26" pageId="12" pageNumber="13">2</td>
<td id="0B7E5D8CFFA42DBBFD0A215AFC88FEAE" box="[758,867,1131,1152]" gridcol="2" gridrow="26" pageId="12" pageNumber="13">20-21</td>
<td id="0B7E5D8CFFA42DBBFBF2215AFBA0FEAE" box="[1038,1099,1131,1152]" gridcol="3" gridrow="26" pageId="12" pageNumber="13">20</td>
<td id="0B7E5D8CFFA42DBBFAF1215AFAA1FEAE" box="[1293,1354,1131,1152]" gridcol="4" gridrow="26" pageId="12" pageNumber="13">0.66</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E21B9FAA1FEB2" box="[178,1354,1160,1180]" gridrow="27" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E21B9FEDBFEB2" box="[178,304,1160,1180]" gridcol="0" gridrow="27" pageId="12" pageNumber="13">LTS/TL</th>
<td id="0B7E5D8CFFA42DBBFDD921B9FDDFFEB2" box="[549,564,1160,1180]" gridcol="1" gridrow="27" pageId="12" pageNumber="13">5</td>
<td id="0B7E5D8CFFA42DBBFD0A21B9FC88FEB2" box="[758,867,1160,1180]" gridcol="2" gridrow="27" pageId="12" pageNumber="13">12.4-14.7%</td>
<td id="0B7E5D8CFFA42DBBFBF221B9FBA0FEB2" box="[1038,1099,1160,1180]" gridcol="3" gridrow="27" pageId="12" pageNumber="13">13.4%</td>
<td id="0B7E5D8CFFA42DBBFAF121B9FAA1FEB2" box="[1293,1354,1160,1180]" gridcol="4" gridrow="27" pageId="12" pageNumber="13">0.86%</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2194FAA1FE97" box="[178,1354,1189,1209]" gridrow="28" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E2194FEDBFE97" box="[178,304,1189,1209]" gridcol="0" gridrow="28" pageId="12" pageNumber="13">LTAS/TL</th>
<td id="0B7E5D8CFFA42DBBFDD92194FDDFFE97" box="[549,564,1189,1209]" gridcol="1" gridrow="28" pageId="12" pageNumber="13">2</td>
<td id="0B7E5D8CFFA42DBBFD0A2194FC88FE97" box="[758,867,1189,1209]" gridcol="2" gridrow="28" pageId="12" pageNumber="13">5.6-5.8%</td>
<td id="0B7E5D8CFFA42DBBFBF22194FBA0FE97" box="[1038,1099,1189,1209]" gridcol="3" gridrow="28" pageId="12" pageNumber="13">5.7%</td>
<td id="0B7E5D8CFFA42DBBFAF12194FAA1FE97" box="[1293,1354,1189,1209]" gridcol="4" gridrow="28" pageId="12" pageNumber="13">0.11%</td>
</tr>
</table>
</paragraph>
<paragraph id="F62036B2FFA4D248FF5221E2FE7CFF11" blockId="12.[174,1453,1234,1343]" pageId="12" pageNumber="13">
<tableNote id="0B79373CFFA4D248FF5221E2FE7CFF11" pageId="12" pageNumber="13" targetBox="[178,1354,353,1209]" targetPageId="12">Abbreviations: (ac), acicular spine; (f), female; LD, length of laterodorsal tubule; LTAS, length of lateral terminal accessory spine; LTS, length of lateral terminal spine; LV, length of lateroventral tubule; (m), male; MD: length of middorsal acicular spine; ML, length of midlateral tubule; MSW, maximum sternal width; S, segment length; SL, length of sublateral acicular spine; SW, standard width; TL, trunk length; (tu), tubule.</tableNote>
</paragraph>
<caption id="A2E0663AFFA4D248FF5220B0FA49FFB4" box="[174,1442,1409,1435]" pageId="12" pageNumber="13" startId="12.[174,242,1409,1434]" targetType="table">
<paragraph id="F62036B2FFA4D248FF5220B0FA49FFB4" blockId="12.[174,1442,1409,1435]" box="[174,1442,1409,1435]" pageId="12" pageNumber="13">
<emphasis id="C4EBEAA0FFA4D248FF5220B0FEE4FFB4" bold="true" box="[174,271,1409,1434]" pageId="12" pageNumber="13">Table 4.</emphasis>
Summary of location of cuticular structures, tubules, and spines in
<taxonomicName id="319F4D31FFA4D248FBD220B0FAD4FFB4" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1070,1343,1409,1434]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="12" pageNumber="13" phylum="Kinorhyncha" rank="species" species="serratulus" status="sp. nov.">
<emphasis id="C4EBEAA0FFA4D248FBD220B0FAD4FFB4" box="[1070,1343,1409,1434]" italics="true" pageId="12" pageNumber="13">Echinoderes serratulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFD857DBFFA4D248FABA20B0FA49FFB4" box="[1350,1442,1409,1434]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<table id="849FC412FFA42DBBFF4E2084FA7DFD1E" box="[178,1430,1461,1840]" gridcols="10" gridrows="12" pageId="12" pageNumber="13">
<paragraph id="F62036B2FFA4D248FEF02084FEB3FFE7" box="[268,344,1461,1481]" pageId="12" pageNumber="13">Position</paragraph>
<paragraph id="F62036B2FFA4D248FE7420F2FB6CFD1E" pageId="12" pageNumber="13">
<tr id="48AF34F0FFA42DBBFF4E20F2FA7DFFC8" box="[178,1430,1475,1510]" gridrow="0" pageId="12" pageNumber="13">
<th id="0B7E5D8CFFA42DBBFF4E20F2FE27FFC8" box="[178,460,1475,1510]" colspan="2" colspanRight="1" gridcol="0" gridrow="0" pageId="12" pageNumber="13">MD segment</th>
<th id="0B7E5D8CFFA42DBBFE0620F2FDC3FFC8" box="[506,552,1475,1510]" gridcol="2" gridrow="0" pageId="12" pageNumber="13">PD</th>
<th id="0B7E5D8CFFA42DBBFD8F20F2FD4AFFC8" box="[627,673,1475,1510]" gridcol="3" gridrow="0" pageId="12" pageNumber="13">SD</th>
<th id="0B7E5D8CFFA42DBBFD2720F2FCC0FFC8" box="[731,811,1475,1510]" gridcol="4" gridrow="0" pageId="12" pageNumber="13">LD</th>
<th id="0B7E5D8CFFA42DBBFCA820F2FC4FFFC8" box="[852,932,1475,1510]" gridcol="5" gridrow="0" pageId="12" pageNumber="13">ML</th>
<th id="0B7E5D8CFFA42DBBFC4020F2FBD6FFC8" box="[956,1085,1475,1510]" gridcol="6" gridrow="0" pageId="12" pageNumber="13">SL</th>
<th id="0B7E5D8CFFA42DBBFB9A20F2FB7FFFC8" box="[1126,1172,1475,1510]" gridcol="7" gridrow="0" pageId="12" pageNumber="13">LV</th>
<th id="0B7E5D8CFFA42DBBFB2320F2FAE6FFC8" box="[1247,1293,1475,1510]" gridcol="8" gridrow="0" pageId="12" pageNumber="13">VL</th>
<th id="0B7E5D8CFFA42DBBFABB20F2FA7DFFC8" box="[1351,1430,1475,1510]" gridcol="9" gridrow="0" pageId="12" pageNumber="13">VM</th>
</tr>
<tr id="48AF34F0FFA42DBBFF4E20CAFA7DFC21" box="[178,1430,1531,1551]" gridrow="1" pageId="12" pageNumber="13" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-8="1">
<th id="0B7E5D8CFFA42DBBFF4E20CAFF21FC21" box="[178,202,1531,1551]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">1</th>
<td id="0B7E5D8CFFA42DBBFE9820CAFE27FC21" box="[356,460,1531,1551]" gridcol="1" gridrow="1" pageId="12" pageNumber="13">gco1</td>
<td id="0B7E5D8CFFA42DBBFD8F20CAFD4AFC21" box="[627,673,1531,1551]" gridcol="3" gridrow="1" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFD2720CAFCC0FC21" box="[731,811,1531,1551]" gridcol="4" gridrow="1" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFB9A20CAFB7FFC21" box="[1126,1172,1531,1551]" gridcol="7" gridrow="1" pageId="12" pageNumber="13">gco1</td>
<td id="0B7E5D8CFFA42DBBFABB20CAFA7DFC21" box="[1351,1430,1531,1551]" gridcol="9" gridrow="1" pageId="12" pageNumber="13">ss</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2329FA7DFC02" box="[178,1430,1560,1580]" gridrow="2" pageId="12" pageNumber="13" rowspan-2="1" rowspan-6="1" rowspan-7="1">
<th id="0B7E5D8CFFA42DBBFF4E2329FF21FC02" box="[178,202,1560,1580]" gridcol="0" gridrow="2" pageId="12" pageNumber="13">2</th>
<td id="0B7E5D8CFFA42DBBFE982329FE27FC02" box="[356,460,1560,1580]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">ss, gco1</td>
<td id="0B7E5D8CFFA42DBBFD8F2329FD4AFC02" box="[627,673,1560,1580]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">gco2</td>
<td id="0B7E5D8CFFA42DBBFD272329FCC0FC02" box="[731,811,1560,1580]" gridcol="4" gridrow="2" pageId="12" pageNumber="13">ss, gco2</td>
<td id="0B7E5D8CFFA42DBBFCA82329FC4FFC02" box="[852,932,1560,1580]" gridcol="5" gridrow="2" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFB232329FAE6FC02" box="[1247,1293,1560,1580]" gridcol="8" gridrow="2" pageId="12" pageNumber="13">gco2</td>
<td id="0B7E5D8CFFA42DBBFABB2329FA7DFC02" box="[1351,1430,1560,1580]" gridcol="9" gridrow="2" pageId="12" pageNumber="13">ss, gco1</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2304FA7DFC67" box="[178,1430,1589,1609]" gridrow="3" pageId="12" pageNumber="13" rowspan-2="1" rowspan-4="1" rowspan-7="1" rowspan-8="1">
<th id="0B7E5D8CFFA42DBBFF4E2304FF21FC67" box="[178,202,1589,1609]" gridcol="0" gridrow="3" pageId="12" pageNumber="13">3</th>
<td id="0B7E5D8CFFA42DBBFE982304FE27FC67" box="[356,460,1589,1609]" gridcol="1" gridrow="3" pageId="12" pageNumber="13">gco1, gco1</td>
<td id="0B7E5D8CFFA42DBBFD8F2304FD4AFC67" box="[627,673,1589,1609]" gridcol="3" gridrow="3" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFCA82304FC4FFC67" box="[852,932,1589,1609]" gridcol="5" gridrow="3" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFC402304FBD6FC67" box="[956,1085,1589,1609]" gridcol="6" gridrow="3" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFABB2304FA7DFC67" box="[1351,1430,1589,1609]" gridcol="9" gridrow="3" pageId="12" pageNumber="13">gco1</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2363FA7DFC48" box="[178,1430,1618,1638]" gridrow="4" pageId="12" pageNumber="13" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="0B7E5D8CFFA42DBBFF4E2363FF21FC48" box="[178,202,1618,1638]" gridcol="0" gridrow="4" pageId="12" pageNumber="13">4</th>
<td id="0B7E5D8CFFA42DBBFE982363FE27FC48" box="[356,460,1618,1638]" gridcol="1" gridrow="4" pageId="12" pageNumber="13">ac</td>
<td id="0B7E5D8CFFA42DBBFE062363FDC3FC48" box="[506,552,1618,1638]" gridcol="2" gridrow="4" pageId="12" pageNumber="13">gco1</td>
<td id="0B7E5D8CFFA42DBBFD8F2363FD4AFC48" box="[627,673,1618,1638]" gridcol="3" gridrow="4" pageId="12" pageNumber="13">gco2</td>
<td id="0B7E5D8CFFA42DBBFD272363FCC0FC48" box="[731,811,1618,1638]" gridcol="4" gridrow="4" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFABB2363FA7DFC48" box="[1351,1430,1618,1638]" gridcol="9" gridrow="4" pageId="12" pageNumber="13">gco1</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E235EFA7DFCAD" box="[178,1430,1647,1667]" gridrow="5" pageId="12" pageNumber="13" rowspan-1="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<th id="0B7E5D8CFFA42DBBFF4E235EFF21FCAD" box="[178,202,1647,1667]" gridcol="0" gridrow="5" pageId="12" pageNumber="13">5</th>
<td id="0B7E5D8CFFA42DBBFE06235EFDC3FCAD" box="[506,552,1647,1667]" gridcol="2" gridrow="5" pageId="12" pageNumber="13">gco1</td>
<td id="0B7E5D8CFFA42DBBFD8F235EFD4AFCAD" box="[627,673,1647,1667]" gridcol="3" gridrow="5" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFCA8235EFC4FFCAD" box="[852,932,1647,1667]" gridcol="5" gridrow="5" pageId="12" pageNumber="13">ss, gco2</td>
<td id="0B7E5D8CFFA42DBBFB9A235EFB7FFCAD" box="[1126,1172,1647,1667]" gridcol="7" gridrow="5" pageId="12" pageNumber="13">tu</td>
<td id="0B7E5D8CFFA42DBBFABB235EFA7DFCAD" box="[1351,1430,1647,1667]" gridcol="9" gridrow="5" pageId="12" pageNumber="13">ss, gco1</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E23BDFA7DFC8E" box="[178,1430,1676,1696]" gridrow="6" pageId="12" pageNumber="13" rowspan-1="1" rowspan-4="1" rowspan-7="1" rowspan-8="1">
<th id="0B7E5D8CFFA42DBBFF4E23BDFF21FC8E" box="[178,202,1676,1696]" gridcol="0" gridrow="6" pageId="12" pageNumber="13">6</th>
<td id="0B7E5D8CFFA42DBBFE0623BDFDC3FC8E" box="[506,552,1676,1696]" gridcol="2" gridrow="6" pageId="12" pageNumber="13">gco1</td>
<td id="0B7E5D8CFFA42DBBFD8F23BDFD4AFC8E" box="[627,673,1676,1696]" gridcol="3" gridrow="6" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFCA823BDFC4FFC8E" box="[852,932,1676,1696]" gridcol="5" gridrow="6" pageId="12" pageNumber="13">ss, gco2</td>
<td id="0B7E5D8CFFA42DBBFC4023BDFBD6FC8E" box="[956,1085,1676,1696]" gridcol="6" gridrow="6" pageId="12" pageNumber="13">ac</td>
<td id="0B7E5D8CFFA42DBBFABB23BDFA7DFC8E" box="[1351,1430,1676,1696]" gridcol="9" gridrow="6" pageId="12" pageNumber="13">ss, gco1</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E2398FA7DFC93" box="[178,1430,1705,1725]" gridrow="7" pageId="12" pageNumber="13" rowspan-1="1" rowspan-4="1" rowspan-7="1" rowspan-8="1">
<th id="0B7E5D8CFFA42DBBFF4E2398FF21FC93" box="[178,202,1705,1725]" gridcol="0" gridrow="7" pageId="12" pageNumber="13">7</th>
<td id="0B7E5D8CFFA42DBBFE062398FDC3FC93" box="[506,552,1705,1725]" gridcol="2" gridrow="7" pageId="12" pageNumber="13">gco1</td>
<td id="0B7E5D8CFFA42DBBFD8F2398FD4AFC93" box="[627,673,1705,1725]" gridcol="3" gridrow="7" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFCA82398FC4FFC93" box="[852,932,1705,1725]" gridcol="5" gridrow="7" pageId="12" pageNumber="13">ss, gco2</td>
<td id="0B7E5D8CFFA42DBBFC402398FBD6FC93" box="[956,1085,1705,1725]" gridcol="6" gridrow="7" pageId="12" pageNumber="13">ac</td>
<td id="0B7E5D8CFFA42DBBFABB2398FA7DFC93" box="[1351,1430,1705,1725]" gridcol="9" gridrow="7" pageId="12" pageNumber="13">ss, gco1</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E23F4FA7DFCF7" box="[178,1430,1733,1753]" gridrow="8" pageId="12" pageNumber="13" rowspan-1="1" rowspan-6="1" rowspan-8="1">
<th id="0B7E5D8CFFA42DBBFF4E23F4FF21FCF7" box="[178,202,1733,1753]" gridcol="0" gridrow="8" pageId="12" pageNumber="13">8</th>
<td id="0B7E5D8CFFA42DBBFE0623F4FDC3FCF7" box="[506,552,1733,1753]" gridcol="2" gridrow="8" pageId="12" pageNumber="13">gco1</td>
<td id="0B7E5D8CFFA42DBBFD8F23F4FD4AFCF7" box="[627,673,1733,1753]" gridcol="3" gridrow="8" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFD2723F4FCC0FCF7" box="[731,811,1733,1753]" gridcol="4" gridrow="8" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFCA823F4FC4FFCF7" box="[852,932,1733,1753]" gridcol="5" gridrow="8" pageId="12" pageNumber="13">gco2</td>
<td id="0B7E5D8CFFA42DBBFB9A23F4FB7FFCF7" box="[1126,1172,1733,1753]" gridcol="7" gridrow="8" pageId="12" pageNumber="13">tu</td>
<td id="0B7E5D8CFFA42DBBFABB23F4FA7DFCF7" box="[1351,1430,1733,1753]" gridcol="9" gridrow="8" pageId="12" pageNumber="13">gco1</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E23D3FA7DFCD8" box="[178,1430,1762,1782]" gridrow="9" pageId="12" pageNumber="13" rowspan-1="1" rowspan-7="1">
<th id="0B7E5D8CFFA42DBBFF4E23D3FF21FCD8" box="[178,202,1762,1782]" gridcol="0" gridrow="9" pageId="12" pageNumber="13">9</th>
<td id="0B7E5D8CFFA42DBBFE0623D3FDC3FCD8" box="[506,552,1762,1782]" gridcol="2" gridrow="9" pageId="12" pageNumber="13">gco1</td>
<td id="0B7E5D8CFFA42DBBFD8F23D3FD4AFCD8" box="[627,673,1762,1782]" gridcol="3" gridrow="9" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFD2723D3FCC0FCD8" box="[731,811,1762,1782]" gridcol="4" gridrow="9" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFCA823D3FC4FFCD8" box="[852,932,1762,1782]" gridcol="5" gridrow="9" pageId="12" pageNumber="13">tu</td>
<td id="0B7E5D8CFFA42DBBFC4023D3FBD6FCD8" box="[956,1085,1762,1782]" gridcol="6" gridrow="9" pageId="12" pageNumber="13">si</td>
<td id="0B7E5D8CFFA42DBBFB2323D3FAE6FCD8" box="[1247,1293,1762,1782]" gridcol="8" gridrow="9" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFABB23D3FA7DFCD8" box="[1351,1430,1762,1782]" gridcol="9" gridrow="9" pageId="12" pageNumber="13">gco1</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E23CEFA7DFD3D" box="[178,1430,1791,1811]" gridrow="10" pageId="12" pageNumber="13" rowspan-1="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="0B7E5D8CFFA42DBBFF4E23CEFF21FD3D" box="[178,202,1791,1811]" gridcol="0" gridrow="10" pageId="12" pageNumber="13">10</th>
<td id="0B7E5D8CFFA42DBBFE0623CEFDC3FD3D" box="[506,552,1791,1811]" gridcol="2" gridrow="10" pageId="12" pageNumber="13">gco1</td>
<td id="0B7E5D8CFFA42DBBFD8F23CEFD4AFD3D" box="[627,673,1791,1811]" gridcol="3" gridrow="10" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFD2723CEFCC0FD3D" box="[731,811,1791,1811]" gridcol="4" gridrow="10" pageId="12" pageNumber="13">tu</td>
<td id="0B7E5D8CFFA42DBBFB2323CEFAE6FD3D" box="[1247,1293,1791,1811]" gridcol="8" gridrow="10" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFABB23CEFA7DFD3D" box="[1351,1430,1791,1811]" gridcol="9" gridrow="10" pageId="12" pageNumber="13">gco1</td>
</tr>
<tr id="48AF34F0FFA42DBBFF4E222DFA7DFD1E" box="[178,1430,1820,1840]" gridrow="11" pageId="12" pageNumber="13" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-8="1" rowspan-9="1">
<th id="0B7E5D8CFFA42DBBFF4E222DFF21FD1E" box="[178,202,1820,1840]" gridcol="0" gridrow="11" pageId="12" pageNumber="13">11</th>
<td id="0B7E5D8CFFA42DBBFD8F222DFD4AFD1E" box="[627,673,1820,1840]" gridcol="3" gridrow="11" pageId="12" pageNumber="13">ss</td>
<td id="0B7E5D8CFFA42DBBFC40222DFBD6FD1E" box="[956,1085,1820,1840]" gridcol="6" gridrow="11" pageId="12" pageNumber="13">ltas (f), ps (m)</td>
<td id="0B7E5D8CFFA42DBBFB9A222DFB7FFD1E" box="[1126,1172,1820,1840]" gridcol="7" gridrow="11" pageId="12" pageNumber="13">lts</td>
</tr>
</paragraph>
</table>
<paragraph id="F62036B2FFA4D248FF522278FC3EFD9B" blockId="12.[174,1452,1865,1973]" pageId="12" pageNumber="13">
<tableNote id="0B79373CFFA4D248FF522278FC3EFD9B" pageId="12" pageNumber="13" targetBox="[178,1430,1461,1840]" targetPageId="12">Abbreviations: ac, acicular spine; (f), female condition of sexually dimorphic character; gco1, type-1 glandular cell outlet; gco2, type- 2 glandular cell outlet; LD, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; LV, lateroventral; (m), male condition of sexually dimorphic character; MD, middorsal; ML, midlateral; PD, paradorsal; ps, penile spine; SD, subdorsal; si, sieve plate; SL, sublateral; ss, sensory spot; tu, tubule; VL, ventrolateral; VM, ventromedial.</tableNote>
</paragraph>
<paragraph id="F62036B2FFA5D249FF39240CFDA8FB78" blockId="13.[136,737,253,406]" box="[197,579,317,343]" pageId="13" pageNumber="14">Segment 7 similar to segment 6.</paragraph>
<paragraph id="F62036B2FFA5D249FF39246CFC56FB78" blockId="13.[136,737,253,406]" lastBlockId="13.[813,1414,253,407]" pageId="13" pageNumber="14">
Segment 8 with lateroventral tubules (
<figureCitation id="6EA42A37FFA5D249FD5F246CFF47FBB8" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="13" pageNumber="14">Figs. 8B</figureCitation>
,
<figureCitation id="6EA42A37FFA5D249FF44244CFF17FBB8" box="[184,252,381,406]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="13" pageNumber="14">9A, E</figureCitation>
,
<figureCitation id="6EA42A37FFA5D249FEF4244CFED6FBB8" box="[264,317,381,406]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[211,1335,250,1816]" captionTargetId="figure-99@17.[211,1335,249,1817]" captionTargetPageId="17" captionText="Fig. 12. Echinoderes serratulus sp. nov., scanning electron micrographs. A, segments 5 and 6, lateral view; B, segments 8 and 9, lateroventral view; C, segment 6, sublateral view; D, segment 9, sublateral view; E, segments 9-11 in female, dorsal view; F, segments 10 and 11 in male, lateral view; G, segments 10 and 11 in female, lateral view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; pe, penile spine; po, pore; si, sieve area; sls, sublateral acicular spine." figureDoi="http://doi.org/10.5281/zenodo.10659233" httpUri="https://zenodo.org/record/10659233/files/figure.png" pageId="13" pageNumber="14">12B</figureCitation>
). Paired sensory spots in subdorsal and laterodorsal positions (
<figureCitation id="6EA42A37FFA5D249FB8725CCFB31FB38" box="[1147,1242,253,278]" captionStart="Fig" captionStartId="9.[136,170,1842,1862]" captionTargetBox="[159,1392,386,1817]" captionTargetId="figure-129@9.[158,1392,385,1817]" captionTargetPageId="9" captionText="Fig. 7. Echinoderes regina sp. nov., scanning electron micrographs. A, segments 4-6, laterodorsal view; B, segments 5-7, ventrolateral view; C, segments 8-10, ventral view; D, sublateral acicular spine on segment 6; E, segments 10 and 11 in female, dorsal view; F, segments 10 and 11 in male, laterodorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal acicular spine; lts lateral terminal spine; lvt, lateroventral tubule; ps, penile spine; sls, sublateral acicular spine; slt, sublateral tubule." figureDoi="http://doi.org/10.5281/zenodo.10659223" httpUri="https://zenodo.org/record/10659223/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
). Paired type- 2 glandular cell outlets in midlateral position (
<figureCitation id="6EA42A37FFA5D249FAB6242CFC9AFB78" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="13" pageNumber="14">Figs. 8A, B</figureCitation>
,
<figureCitation id="6EA42A37FFA5D249FC81240CFC5AFB78" box="[893,945,317,342]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[211,1335,250,1816]" captionTargetId="figure-99@17.[211,1335,249,1817]" captionTargetPageId="17" captionText="Fig. 12. Echinoderes serratulus sp. nov., scanning electron micrographs. A, segments 5 and 6, lateral view; B, segments 8 and 9, lateroventral view; C, segment 6, sublateral view; D, segment 9, sublateral view; E, segments 9-11 in female, dorsal view; F, segments 10 and 11 in male, lateral view; G, segments 10 and 11 in female, lateral view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; pe, penile spine; po, pore; si, sieve area; sls, sublateral acicular spine." figureDoi="http://doi.org/10.5281/zenodo.10659233" httpUri="https://zenodo.org/record/10659233/files/figure.png" pageId="13" pageNumber="14">12B</figureCitation>
).
</paragraph>
<paragraph id="F62036B2FFA5D24BFC95246CFE8EFB58" blockId="13.[813,1414,253,407]" lastBlockId="15.[136,737,253,374]" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
Segment 9 with midlateral tubules (
<figureCitation id="6EA42A37FFA5D249FAEB246CFC83FBB8" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="13" pageNumber="14">Figs. 8A, C, D</figureCitation>
,
<figureCitation id="6EA42A37FFA5D249FC8A244CFC7DFBB8" box="[886,918,381,406]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="13" pageNumber="14">9F</figureCitation>
,
<figureCitation id="6EA42A37FFA5D249FC58244CFC16FBB8" box="[932,1021,381,406]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[211,1335,250,1816]" captionTargetId="figure-99@17.[211,1335,249,1817]" captionTargetPageId="17" captionText="Fig. 12. Echinoderes serratulus sp. nov., scanning electron micrographs. A, segments 5 and 6, lateral view; B, segments 8 and 9, lateroventral view; C, segment 6, sublateral view; D, segment 9, sublateral view; E, segments 9-11 in female, dorsal view; F, segments 10 and 11 in male, lateral view; G, segments 10 and 11 in female, lateral view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; pe, penile spine; po, pore; si, sieve area; sls, sublateral acicular spine." figureDoi="http://doi.org/10.5281/zenodo.10659233" httpUri="https://zenodo.org/record/10659233/files/figure.png" pageId="13" pageNumber="14">12B, D</figureCitation>
). Paired subdorsal, laterodorsal, and ventrolateral sensory spots present (
<figureCitation id="6EA42A37FFA7D24BFD8F25CCFD2EFB38" box="[627,709,253,278]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="15" pageNumber="16">Figs. 8</figureCitation>
A- D, 9A, E). Sieve plates with oval sieve area and single posterior pore in sublateral position (
<figureCitation id="6EA42A37FFA7D24BFD5F240CFF7CFB58" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="15" pageNumber="16">Figs. 8</figureCitation>
B-D, 9E, 12B, D).
</paragraph>
<caption id="A2E0663AFFA5D249FF742224FE53FD9B" ID-DOI="http://doi.org/10.5281/zenodo.10659225" ID-Zenodo-Dep="10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="13" pageNumber="14" startId="13.[136,170,1813,1833]" targetBox="[160,1385,462,1787]" targetPageId="13" targetType="figure">
<paragraph id="F62036B2FFA5D249FF742224FE53FD9B" blockId="13.[136,1414,1813,1973]" pageId="13" pageNumber="14">
<emphasis id="C4EBEAA0FFA5D249FF742224FF2DFD07" bold="true" box="[136,198,1813,1833]" pageId="13" pageNumber="14">Fig. 8.</emphasis>
<taxonomicName id="319F4D31FFA5D249FF2F2224FE45FD07" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[211,430,1813,1833]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="13" pageNumber="14" phylum="Kinorhyncha" rank="species" species="serratulus" status="sp. nov.">
<emphasis id="C4EBEAA0FFA5D249FF2F2224FE45FD07" box="[211,430,1813,1833]" italics="true" pageId="13" pageNumber="14">Echinoderes serratulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFD857DBFFA5D249FE482224FDE9FD07" box="[436,514,1813,1833]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
, camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot.
</paragraph>
</caption>
<caption id="A2E0663AFFA6D24AFF522224FCDAFD9B" ID-DOI="http://doi.org/10.5281/zenodo.10659227" ID-Zenodo-Dep="10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="14" pageNumber="15" startId="14.[174,208,1813,1833]" targetBox="[291,1333,250,1785]" targetPageId="14" targetType="figure">
<paragraph id="F62036B2FFA6D24AFF522224FCDAFD9B" blockId="14.[174,1452,1813,1973]" pageId="14" pageNumber="15">
<emphasis id="C4EBEAA0FFA6D24AFF522224FF06FD07" bold="true" box="[174,237,1813,1833]" pageId="14" pageNumber="15">Fig. 9.</emphasis>
<taxonomicName id="319F4D31FFA6D24AFF012224FE31FD07" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[253,474,1813,1833]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="15" phylum="Kinorhyncha" rank="species" species="serratulus" status="sp. nov.">
<emphasis id="C4EBEAA0FFA6D24AFF012224FE31FD07" box="[253,474,1813,1833]" italics="true" pageId="14" pageNumber="15">Echinoderes serratulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFD857DBFFA6D24AFE1D2224FDDAFD07" box="[481,561,1813,1833]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
, Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate.
</paragraph>
</caption>
<paragraph id="F62036B2FFA7D254FC9525CCFF00FBB8" blockId="15.[813,1414,253,374]" lastBlockId="16.[174,775,253,919]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
Segment 10 with long laterodorsal tubules in males, short laterodorsal tubules in females (
<figureCitation id="6EA42A37FFA7D24BFAB6242CFC9EFB79" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="15" pageNumber="16">Figs. 8A, C</figureCitation>
,
<figureCitation id="6EA42A37FFA7D24BFC7E240CFC49FB78" box="[898,930,317,342]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="15" pageNumber="16">9F</figureCitation>
,
<figureCitation id="6EA42A37FFA7D24BFC53240CFBECFB79" box="[943,1031,317,343]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[211,1335,250,1816]" captionTargetId="figure-99@17.[211,1335,249,1817]" captionTargetPageId="17" captionText="Fig. 12. Echinoderes serratulus sp. nov., scanning electron micrographs. A, segments 5 and 6, lateral view; B, segments 8 and 9, lateroventral view; C, segment 6, sublateral view; D, segment 9, sublateral view; E, segments 9-11 in female, dorsal view; F, segments 10 and 11 in male, lateral view; G, segments 10 and 11 in female, lateral view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; pe, penile spine; po, pore; si, sieve area; sls, sublateral acicular spine." figureDoi="http://doi.org/10.5281/zenodo.10659233" httpUri="https://zenodo.org/record/10659233/files/figure.png" pageId="15" pageNumber="16">12E, G</figureCitation>
;
<tableCitation id="BB1D0309FFA7D24BFBE8240CFB81FB78" box="[1044,1130,317,342]" captionStart="Table 1" captionStartId="3.[136,205,253,278]" captionText="Table 1. Measurements for adult Echinoderes regina sp. nov. (in micrometers). Columns N and SD are sample size and standard deviation, respectively" pageId="15" pageNumber="16">Table 1</tableCitation>
). Paired subdorsal and ventrolateral sensory spots present (
<figureCitation id="6EA42A37FFA7D24BFB0A246CFAA5FB58" box="[1270,1358,349,374]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="15" pageNumber="16">Figs. 8</figureCitation>
A-D, 12E, G). Primary pectinate fringe with shorter and thinner tips than preceding segments in middorsal to ventrolateral areas, and with tips similar in length to those of preceding segments in ventromedial area.
</paragraph>
<caption id="A2E0663AFFA7D24BFF742201FB60FD9A" ID-DOI="http://doi.org/10.5281/zenodo.10659229" ID-Zenodo-Dep="10659229" httpUri="https://zenodo.org/record/10659229/files/figure.png" pageId="15" pageNumber="16" startId="15.[136,172,1840,1860]" targetBox="[184,1366,429,1814]" targetPageId="15" targetType="figure">
<paragraph id="F62036B2FFA7D24BFF742201FB60FD9A" blockId="15.[136,1415,1840,1972]" pageId="15" pageNumber="16">
<emphasis id="C4EBEAA0FFA7D24BFF742201FF3CFD6A" bold="true" box="[136,215,1840,1860]" pageId="15" pageNumber="16">Fig. 10.</emphasis>
<taxonomicName id="319F4D31FFA7D24BFF152201FE3BFD6A" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[233,464,1840,1860]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="15" pageNumber="16" phylum="Kinorhyncha" rank="species" species="serratulus" status="sp. nov.">
<emphasis id="C4EBEAA0FFA7D24BFF152201FE3BFD6A" box="[233,464,1840,1860]" italics="true" pageId="15" pageNumber="16">Echinoderes serratulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFD857DBFFA7D24BFE252201FDC6FD6A" box="[473,557,1840,1860]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
, scanning electron micrographs. A, entire animal, lateroventral view; B, mouth cone, lateroventral view; C, introvert, ventrolateral view; D, segments 1-7, ventral view; E, segment 5, ventromedial view; F, segments 3-6, ventrolateral view; G, segments 3-5, dorsal view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ha, hair; int, introvert; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; oos, outer oral style; ppf, primary pectinate fringe; psc, primary spinoscalid; sec, sector; seg, segment; spf, secondary pectinate fringe.
</paragraph>
</caption>
<paragraph id="F62036B2FFB8D254FF1624ACFDE1F8D8" blockId="16.[174,775,253,919]" pageId="16" pageNumber="17">
Segment 11 with short, plump lateral terminal spines (
<figureCitation id="6EA42A37FFB8D254FEEC248CFE83FBF8" box="[272,360,445,470]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="16" pageNumber="17">Figs. 8</figureCitation>
A-D, 9A, F, G, 10A, 12E-G). Pair of short lateral terminal accessory spines present in females (
<figureCitation id="6EA42A37FFB8D254FEB724CCFE2AF838" box="[331,449,509,534]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="16" pageNumber="17">Figs. 8B</figureCitation>
,
<figureCitation id="6EA42A37FFB8D254FE2D24CCFE17F839" box="[465,508,509,535]" captionStart="Fig" captionStartId="14.[174,208,1813,1833]" captionTargetBox="[291,1333,250,1785]" captionTargetId="figure-108@14.[291,1333,249,1786]" captionTargetPageId="14" captionText="Fig. 9. Echinoderes serratulus sp. nov., Nomarski photomicrographs of holotype female (ZIHU-5053). A, entire animal, ventral view; B, segments 2 and 3, ventral view; C, segments 1 and 2, dorsal view; D, segments 5 and 6, lateroventral view; E, segments 8 and 9, lateroventral view; F, segments 9-11, laterodorsal view; G, segments 10 and 11, ventrolateral view. Dashed circles indicate sensory spots. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; mvp, midventral placid; ppf, primary pectinate fringe; si, sieve plate; sls, sublateral acicular spine; sp, sternal plate." figureDoi="http://doi.org/10.5281/zenodo.10659227" httpUri="https://zenodo.org/record/10659227/files/figure.png" pageId="16" pageNumber="17">9G</figureCitation>
,
<figureCitation id="6EA42A37FFB8D254FDF024CCFD84F839" box="[524,623,509,535]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[211,1335,250,1816]" captionTargetId="figure-99@17.[211,1335,249,1817]" captionTargetPageId="17" captionText="Fig. 12. Echinoderes serratulus sp. nov., scanning electron micrographs. A, segments 5 and 6, lateral view; B, segments 8 and 9, lateroventral view; C, segment 6, sublateral view; D, segment 9, sublateral view; E, segments 9-11 in female, dorsal view; F, segments 10 and 11 in male, lateral view; G, segments 10 and 11 in female, lateral view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; pe, penile spine; po, pore; si, sieve area; sls, sublateral acicular spine." figureDoi="http://doi.org/10.5281/zenodo.10659233" httpUri="https://zenodo.org/record/10659233/files/figure.png" pageId="16" pageNumber="17">12E, G</figureCitation>
), and three pairs of penile spines present in males (
<figureCitation id="6EA42A37FFB8D254FD39272CFF3FF878" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="16" pageNumber="17">Figs. 8D</figureCitation>
,
<figureCitation id="6EA42A37FFB8D254FF1F270CFEFCF878" box="[227,279,573,598]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[211,1335,250,1816]" captionTargetId="figure-99@17.[211,1335,249,1817]" captionTargetPageId="17" captionText="Fig. 12. Echinoderes serratulus sp. nov., scanning electron micrographs. A, segments 5 and 6, lateral view; B, segments 8 and 9, lateroventral view; C, segment 6, sublateral view; D, segment 9, sublateral view; E, segments 9-11 in female, dorsal view; F, segments 10 and 11 in male, lateral view; G, segments 10 and 11 in female, lateral view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; pe, penile spine; po, pore; si, sieve area; sls, sublateral acicular spine." figureDoi="http://doi.org/10.5281/zenodo.10659233" httpUri="https://zenodo.org/record/10659233/files/figure.png" pageId="16" pageNumber="17">12F</figureCitation>
). Cuticular hairs absent. Paired sensory spots in subdorsal position (
<figureCitation id="6EA42A37FFB8D254FDE7276CFD5FF859" box="[539,692,605,631]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="16" pageNumber="17">Figs. 8A, C</figureCitation>
,
<figureCitation id="6EA42A37FFB8D254FD3F276CFD11F858" box="[707,762,605,630]" captionStart="Fig" captionStartId="17.[136,171,1841,1861]" captionTargetBox="[211,1335,250,1816]" captionTargetId="figure-99@17.[211,1335,249,1817]" captionTargetPageId="17" captionText="Fig. 12. Echinoderes serratulus sp. nov., scanning electron micrographs. A, segments 5 and 6, lateral view; B, segments 8 and 9, lateroventral view; C, segment 6, sublateral view; D, segment 9, sublateral view; E, segments 9-11 in female, dorsal view; F, segments 10 and 11 in male, lateral view; G, segments 10 and 11 in female, lateral view. Dashed circles indicate sensory spots. Abbreviations: gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mlt, midlateral tubule; pe, penile spine; po, pore; si, sieve area; sls, sublateral acicular spine." figureDoi="http://doi.org/10.5281/zenodo.10659233" httpUri="https://zenodo.org/record/10659233/files/figure.png" pageId="16" pageNumber="17">12E</figureCitation>
). Tergal plate partially divided at middorsal line from central to posterior part. Posterior edge of tergal plate projects laterally and ends in pointed tergal extensions (
<figureCitation id="6EA42A37FFB8D254FEC027ECFE65F8D8" box="[316,398,733,758]" captionStart="Fig" captionStartId="13.[136,170,1813,1833]" captionTargetBox="[160,1385,462,1787]" captionTargetId="figure-162@13.[160,1385,462,1787]" captionTargetPageId="13" captionText="Fig. 8. Echinoderes serratulus sp. nov., camera lucida drawings. A, B, Holotype, female (ZIHU-5053), entire animal, dorsal and ventral views, respectively; C, D, allotype, male (ZIHU-5054), segments 9-11, dorsal and ventral views, respectively. Double, gray, and black circles indicate sensory spots, type-1 glandular cell outlets, and type-2 glandular cell outlets, respectively. Abbreviations: gco1, type-1 glandular cell outlet; gco2, type-2 glandular cell outlet; ldt, laterodorsal tubule; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvt, lateroventral tubule; mds, middorsal acicular spine; mlt, midlateral tubule; ps, penile spine; si, sieve plate; sls, sublateral acicular spine; ss, sensory spot." figureDoi="http://doi.org/10.5281/zenodo.10659225" httpUri="https://zenodo.org/record/10659225/files/figure.png" pageId="16" pageNumber="17">Figs. 8</figureCitation>
A-D, 12E).
</paragraph>
<paragraph id="F62036B2FFB8D254FF1627CCFB79F878" blockId="16.[174,775,253,919]" lastBlockId="16.[850,1452,253,918]" pageId="16" pageNumber="17">
<emphasis id="C4EBEAA0FFB8D254FF1627CCFEB2F938" box="[234,345,765,790]" italics="true" pageId="16" pageNumber="17">Remarks</emphasis>
: The combination of the arrangement of spines and tubules and the presence of large sieve plates assigns
<taxonomicName id="319F4D31FFB8D254FE0E260CFCECF978" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[498,775,829,854]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="16" pageNumber="17" phylum="Kinorhyncha" rank="species" species="serratulus" status="sp. nov.">
<emphasis id="C4EBEAA0FFB8D254FE0E260CFCECF978" box="[498,775,829,854]" italics="true" pageId="16" pageNumber="17">Echinoderes serratulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFD857DBFFB8D254FF52266CFEFAF958" box="[174,273,861,886]" pageId="16" pageNumber="17" rank="species">sp. nov</taxonomicNameLabel>
to be a member of the
<emphasis id="C4EBEAA0FFB8D254FD9D266CFF16F9B8" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="319F4D31FFB8D254FD9D266CFCEEF958" authorityName="Claparede" authorityYear="1863" box="[609,773,861,886]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="16" pageNumber="17" phylum="Kinorhyncha" rank="genus">Echinoderes</taxonomicName>
coulli-
</emphasis>
group (see also DISCUSSION). Within the group,
<taxonomicName id="319F4D31FFB8D254FC1A25CCFAFCFB38" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[998,1303,253,278]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="16" pageNumber="17" phylum="Kinorhyncha" rank="species" species="serratulus" status="sp. nov.">
<emphasis id="C4EBEAA0FFB8D254FC1A25CCFAFCFB38" box="[998,1303,253,278]" italics="true" pageId="16" pageNumber="17">Echinoderes serratulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFD857DBFFB8D254FADF25CCFA60FB38" box="[1315,1419,253,278]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="319F4D31FFB8D254FC25242CFBBAFB18" authorityName="Sorensen, Gasiorowski, Randso, Sanchez &amp; Neves" authorityYear="2016" box="[985,1105,285,310]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="16" pageNumber="17" phylum="Kinorhyncha" rank="species" species="annae">
<emphasis id="C4EBEAA0FFB8D254FC25242CFC00FB18" box="[985,1003,285,310]" italics="true" pageId="16" pageNumber="17">E</emphasis>
.
<emphasis id="C4EBEAA0FFB8D254FBFD242CFBBAFB18" box="[1025,1105,285,310]" italics="true" pageId="16" pageNumber="17">annae</emphasis>
</taxonomicName>
and
<taxonomicName id="319F4D31FFB8D254FB66242CFB3EFB78" authority="Yamasaki and Fujimoto (2014)" authorityName="Yamasaki and Fujimoto" authorityYear="2014" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="16" pageNumber="17" phylum="Kinorhyncha" rank="species" species="hwiizaa">
<emphasis id="C4EBEAA0FFB8D254FB66242CFA41FB18" box="[1178,1450,285,310]" italics="true" pageId="16" pageNumber="17">Echinoderes hwiizaa</emphasis>
<bibRefCitation id="920E4B43FFB8D254FCAE240CFB3EFB78" author="Yamasaki H &amp; Fujimoto S." box="[850,1237,317,342]" pageId="16" pageNumber="17" pagination="27 - 52" refId="ref9756" refString="Yamasaki H, Fujimoto S. 2014. Two new species in the Echinoderes coulli group (Echinoderidae, Cyclorhagida, Kinorhyncha) from the Ryukyu Islands, Japan. ZooKeys 382: 27 - 52. doi: 10.3897 / zookeys. 382.6761." type="journal article" year="2014">Yamasaki and Fujimoto (2014)</bibRefCitation>
</taxonomicName>
in having paired tubules on segment 9, and plump lateral terminal spines on segment 11 (
<bibRefCitation id="920E4B43FFB8D254FB82244CFC78FB98" author="Yamasaki H &amp; Fujimoto S." pageId="16" pageNumber="17" pagination="27 - 52" refId="ref9756" refString="Yamasaki H, Fujimoto S. 2014. Two new species in the Echinoderes coulli group (Echinoderidae, Cyclorhagida, Kinorhyncha) from the Ryukyu Islands, Japan. ZooKeys 382: 27 - 52. doi: 10.3897 / zookeys. 382.6761." type="journal article" year="2014">Yamasaki and Fujimoto 2014</bibRefCitation>
;
<bibRefCitation id="920E4B43FFB8D254FC5E24ADFB5BFB98" author="SOrensen MV &amp; Gasiorowski L &amp; RandsO PV &amp; Sanchez N &amp; Neves RC" box="[930,1200,412,438]" pageId="16" pageNumber="17" pagination="3 - 27" refId="ref9583" refString="SOrensen MV, Gasiorowski L, RandsO PV, Sanchez N, Neves RC. 2016. First report of kinorhynchs from Singapore, with the description of three new species. Raffles Bull Zool 64: 3 - 27." type="journal article" year="2016">SØrensen et al. 2016</bibRefCitation>
). It is also similar to
<taxonomicName id="319F4D31FFB8D254FCAE248CFB86FBF8" authorityName="Sorensen, Rho, Min, Kim &amp; Chang" authorityYear="2012" box="[850,1133,445,470]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="16" pageNumber="17" phylum="Kinorhyncha" rank="species" species="aspinosus">
<emphasis id="C4EBEAA0FFB8D254FCAE248CFB86FBF8" box="[850,1133,445,470]" italics="true" pageId="16" pageNumber="17">Echinoderes aspinosus</emphasis>
</taxonomicName>
<bibRefCitation id="920E4B43FFB8D254FB8B248DFA67FBF8" author="SOrensen MV &amp; Rho HS &amp; Min W &amp; Kim D &amp; Chang CY" box="[1143,1420,444,470]" pageId="16" pageNumber="17" pagination="161 - 196" refId="ref9699" refString="SOrensen MV, Rho HS, Min W, Kim D, Chang CY. 2012. An exploration of Echinoderes (Kinorhyncha: Cyclorhagida) in Korean and neighboring waters, with the description of four new species and a redescription of E. tchefouensis Lou, 1934. Zootaxa 3368: 161 - 196." type="journal article" year="2012">SØrensen et al. (2012)</bibRefCitation>
in having paired large sieve plates on segment 9, and the tips of the ventral primary pectinate fringes pointing towards the midventral line on segments 2-5 (
<bibRefCitation id="920E4B43FFB8D254FC7A270DFB6CF878" author="SOrensen MV &amp; Rho HS &amp; Min W &amp; Kim D &amp; Chang CY" box="[902,1159,572,598]" pageId="16" pageNumber="17" pagination="161 - 196" refId="ref9699" refString="SOrensen MV, Rho HS, Min W, Kim D, Chang CY. 2012. An exploration of Echinoderes (Kinorhyncha: Cyclorhagida) in Korean and neighboring waters, with the description of four new species and a redescription of E. tchefouensis Lou, 1934. Zootaxa 3368: 161 - 196." type="journal article" year="2012">SØrensen et al. 2012</bibRefCitation>
).
</paragraph>
<paragraph id="F62036B2FFB8D254FC73276CFA8FF898" blockId="16.[850,1452,253,918]" pageId="16" pageNumber="17">
<taxonomicName id="319F4D31FFB8D254FC73276CFB75F858" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[911,1182,605,630]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="16" pageNumber="17" phylum="Kinorhyncha" rank="species" species="serratulus" status="sp. nov.">
<emphasis id="C4EBEAA0FFB8D254FC73276CFB75F858" box="[911,1182,605,630]" italics="true" pageId="16" pageNumber="17">Echinoderes serratulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFD857DBFFB8D254FB59276CFAEAF858" box="[1189,1281,605,630]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="319F4D31FFB8D254FA6E276CFC77F8B8" authorityName="Sorensen, Gasiorowski, Randso, Sanchez &amp; Neves" authorityYear="2016" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="16" pageNumber="17" phylum="Kinorhyncha" rank="species" species="annae">
<emphasis id="C4EBEAA0FFB8D254FA6E276CFC77F8B8" italics="true" pageId="16" pageNumber="17">E. annae</emphasis>
</taxonomicName>
in having minute sublateral acicular spines on segments 6 and 7 (
<bibRefCitation id="920E4B43FFB8D254FBA527ADFAB1F898" author="SOrensen MV &amp; Gasiorowski L &amp; RandsO PV &amp; Sanchez N &amp; Neves RC" box="[1113,1370,668,694]" pageId="16" pageNumber="17" pagination="3 - 27" refId="ref9583" refString="SOrensen MV, Gasiorowski L, RandsO PV, Sanchez N, Neves RC. 2016. First report of kinorhynchs from Singapore, with the description of three new species. Raffles Bull Zool 64: 3 - 27." type="journal article" year="2016">SØrensen et al. 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BE856539FFB8D256FC73278CFE54FB58" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="description">
<paragraph id="F62036B2FFB8D254FC73278CFBABF958" blockId="16.[850,1452,253,918]" pageId="16" pageNumber="17">
<taxonomicName id="319F4D31FFB8D254FC73278CFB42F8F8" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[911,1193,701,726]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="16" pageNumber="17" phylum="Kinorhyncha" rank="species" species="serratulus" status="sp. nov.">
<emphasis id="C4EBEAA0FFB8D254FC73278CFB42F8F8" box="[911,1193,701,726]" italics="true" pageId="16" pageNumber="17">Echinoderes serratulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFD857DBFFB8D254FB49278CFAFDF8F8" box="[1205,1302,701,726]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="319F4D31FFB8D254FCAE27ECFC3EF8D8" authorityName="Yamasaki and Fujimoto" authorityYear="2014" box="[850,981,733,758]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="16" pageNumber="17" phylum="Kinorhyncha" rank="species" species="hwiizaa">
<emphasis id="C4EBEAA0FFB8D254FCAE27ECFC3EF8D8" box="[850,981,733,758]" italics="true" pageId="16" pageNumber="17">E. hwiizaa</emphasis>
</taxonomicName>
in having a middorsal acicular spine on segment 4 and a conspicuous ventral primary pectinate fringe on segments 1 and 2, and in lacking midlateral tubules on segment 8 (
<bibRefCitation id="920E4B43FFB8D254FAC9260CFBDDF958" author="Yamasaki H &amp; Fujimoto S." pageId="16" pageNumber="17" pagination="27 - 52" refId="ref9756" refString="Yamasaki H, Fujimoto S. 2014. Two new species in the Echinoderes coulli group (Echinoderidae, Cyclorhagida, Kinorhyncha) from the Ryukyu Islands, Japan. ZooKeys 382: 27 - 52. doi: 10.3897 / zookeys. 382.6761." type="journal article" year="2014">Yamasaki and Fujimoto 2014</bibRefCitation>
).
</paragraph>
<paragraph id="F62036B2FFB8D256FC73264CFE54FB58" blockId="16.[850,1452,253,918]" lastBlockId="18.[174,775,253,374]" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<taxonomicName id="319F4D31FFB8D254FC73264CFB49F9B8" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[911,1186,893,918]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="16" pageNumber="17" phylum="Kinorhyncha" rank="species" species="serratulus" status="sp. nov.">
<emphasis id="C4EBEAA0FFB8D254FC73264CFB49F9B8" box="[911,1186,893,918]" italics="true" pageId="16" pageNumber="17">Echinoderes serratulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFD857DBFFB8D254FB51264CFAE7F9B8" box="[1197,1292,893,918]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
easily differs from
<taxonomicName id="319F4D31FFBAD256FF1325CCFE7BFB38" authorityName="Sorensen, Rho, Min, Kim &amp; Chang" authorityYear="2012" box="[239,400,253,278]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="18" pageNumber="19" phylum="Kinorhyncha" rank="species" species="aspinosus">
<emphasis id="C4EBEAA0FFBAD256FF1325CCFE7BFB38" box="[239,400,253,278]" italics="true" pageId="18" pageNumber="19">E. aspinosus</emphasis>
</taxonomicName>
in having acicular spines and tubules on segments 4-10 (
<taxonomicName id="319F4D31FFBAD256FE09242CFD65FB18" authorityName="Sorensen, Rho, Min, Kim &amp; Chang" authorityYear="2012" box="[501,654,285,310]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="18" pageNumber="19" phylum="Kinorhyncha" rank="species" species="aspinosus">
<emphasis id="C4EBEAA0FFBAD256FE09242CFDECFB18" box="[501,519,285,310]" italics="true" pageId="18" pageNumber="19">E</emphasis>
.
<emphasis id="C4EBEAA0FFBAD256FDEA242CFD65FB18" box="[534,654,285,310]" italics="true" pageId="18" pageNumber="19">aspinosus</emphasis>
</taxonomicName>
lacks any acicular spines and tubules on segments 4-10) (
<bibRefCitation id="920E4B43FFBAD256FF48246DFE5EFB58" author="SOrensen MV &amp; Rho HS &amp; Min W &amp; Kim D &amp; Chang CY" box="[180,437,348,374]" pageId="18" pageNumber="19" pagination="161 - 196" refId="ref9699" refString="SOrensen MV, Rho HS, Min W, Kim D, Chang CY. 2012. An exploration of Echinoderes (Kinorhyncha: Cyclorhagida) in Korean and neighboring waters, with the description of four new species and a redescription of E. tchefouensis Lou, 1934. Zootaxa 3368: 161 - 196." type="journal article" year="2012">SØrensen et al. 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>