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<document id="16778CBA7A8122AE061C2E7E3909C174" ID-CLB-Dataset="24286" ID-DOI="10.11646/zootaxa.4728.2.2" ID-GBIF-Dataset="a1c40b73-0c11-4863-a188-412c33523e02" ID-ISSN="1175-5326" ID-Zenodo-Dep="3621117" ID-ZooBank="AE4F8089-7D2E-4ABA-AA09-CD504705788A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1579677041620" checkinUser="plazi" docAuthor="Wagner, Rüdiger &amp; Withers, Phil" docDate="2020" docId="784D878B601B21028FF10120FF598BE2" docLanguage="en" docName="zootaxa.4728.2.2.pdf" docOrigin="Zootaxa 4728 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tonnoiriella graeca Wagner 1993" docType="treatment" docVersion="6" lastPageNumber="206" masterDocId="8474FFF3600C21158F66006DFFB28F20" masterDocTitle="The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)" masterLastPageNumber="210" masterPageNumber="183" pageNumber="206" updateTime="1698778572713" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8F1F00999A746A5672584DCA8B49F90B">The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)</mods:title>
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<mods:namePart id="2F4C449D340601539F1A62C1E635F73E">Wagner, Rüdiger</mods:namePart>
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<mods:namePart id="4EC1ACFD0D91B7BA49C1D97CCBCD557A">Withers, Phil</mods:namePart>
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<mods:title id="45A9FD354F769C9B64D1FF2C8CEF5A5D">Zootaxa</mods:title>
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<mods:date id="DB15F79E375A5DE734A004AE6A0A4F83">2020</mods:date>
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<treatment id="784D878B601B21028FF10120FF598BE2" ID-GBIF-Taxon="161827859" LSID="urn:lsid:plazi:treatment:784D878B601B21028FF10120FF598BE2" httpUri="http://treatment.plazi.org/id/784D878B601B21028FF10120FF598BE2" lastPageNumber="206" pageId="23" pageNumber="206">
<subSubSection id="B8FE6516601B21028FF10120FD848E47" box="[151,566,333,360]" pageId="23" pageNumber="206" type="nomenclature">
<paragraph id="F05B369D601B21028FF10120FD848E47" blockId="23.[151,566,333,395]" box="[151,566,333,360]" pageId="23" pageNumber="206">
<heading id="AB1381F1601B21028FF10120FD848E47" bold="true" box="[151,566,333,360]" fontSize="11" level="1" pageId="23" pageNumber="206" reason="1">
<taxonomicName id="37E44D1E601B21028FF10120FD848E47" ID-CoL="7CHTH" authority="Wagner, 1993" authorityName="Wagner" authorityYear="1993" box="[151,566,333,360]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="206" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="C290EA8F601B21028FF10120FD848E47" bold="true" box="[151,566,333,360]" pageId="23" pageNumber="206">
<emphasis id="C290EA8F601B21028FF10120FE368E47" bold="true" box="[151,388,333,360]" italics="true" pageId="23" pageNumber="206">Tonnoiriella graeca</emphasis>
<bibRefCitation id="94754B6C601B21028EEA0120FD848E47" author="Wagner, R." box="[396,566,333,359]" pageId="23" pageNumber="206" pagination="401 - 416" refId="ref20909" refString="Wagner, R. (1993) Einige neue europaische Psychodiden. Entomofauna, 14 (24), 401 - 416." type="journal article" year="1993">Wagner, 1993</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601B21028FF1011CFE9A8EAB" box="[151,296,368,395]" pageId="23" pageNumber="206" type="description">
<paragraph id="F05B369D601B21028FF1011CFE9A8EAB" blockId="23.[151,566,333,395]" box="[151,296,368,395]" pageId="23" pageNumber="206">
(
<figureCitation id="68DF2A18601B21028FF9011CFEAD8EAB" box="[159,287,368,395]" captionStart="FIGURES 50–53" captionStartId="24.[151,264,1201,1226]" captionTargetBox="[169,1415,360,1162]" captionTargetId="figure@24.[151,1436,345,1176]" captionTargetPageId="24" captionText="FIGURES 50–53: 50–51 T. graeca 50 hypandrium, gonopods, aedeagus, 51 epandrial processes and hypoproct; 52–53 T. hatayensis 52 gonocoxite and gonostylus, 53 aedeagus oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621131" httpUri="https://zenodo.org/record/3621131/files/figure.png" pageId="23" pageNumber="206">Figs 50, 51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601B21028FF101D8FCA18EED" box="[151,787,436,462]" pageId="23" pageNumber="206" type="reference_group">
<paragraph id="F05B369D601B21028FF101D8FCA18EED" blockId="23.[151,787,436,462]" box="[151,787,436,462]" pageId="23" pageNumber="206">
<taxonomicName id="37E44D1E601B21028FF101D8FE448EED" ID-CoL="7CHTH" authority="Wagner, 1993" authorityName="Wagner" authorityYear="1993" box="[151,502,437,462]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="206" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="C290EA8F601B21028FF101D8FED28EED" box="[151,352,437,461]" italics="true" pageId="23" pageNumber="206">Tonnoiriella graeca</emphasis>
<bibRefCitation id="94754B6C601B21028E0001DBFE448EED" author="Wagner, R." box="[358,502,437,462]" pageId="23" pageNumber="206" pagination="401 - 416" refId="ref20909" refString="Wagner, R. (1993) Einige neue europaische Psychodiden. Entomofauna, 14 (24), 401 - 416." type="journal article" year="1993">Wagner, 1993</bibRefCitation>
</taxonomicName>
: Entomofauna 14 (24): 406.
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601B21028FF10196FE198D1A" pageId="23" pageNumber="206" type="materials_examined">
<paragraph id="F05B369D601B21028FF10196FE198D1A" blockId="23.[151,1437,507,1218]" pageId="23" pageNumber="206">
<emphasis id="C290EA8F601B21028FF10196FF4D8D36" bold="true" box="[151,255,507,534]" pageId="23" pageNumber="206">Material</emphasis>
:
<materialsCitation id="408C3CC0601B21028E680191FE198D1A" ID-GBIF-Occurrence="2556505413" collectingDate="1984-04-17" collectionCode="RW" country="Greece" elevation="170" latitude="37.866665" location="Andros island" longLatPrecision="1178" longitude="24.833334" municipality="Remmara" pageId="23" pageNumber="206" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="E6E2FD14601B21028E680191FE838D36" box="[270,305,508,534]" pageId="23" pageNumber="206" type="male">1♂</specimenCount>
<typeStatus id="2F5F883F601B21028E5E0196FE2F8D35" box="[312,413,507,533]" pageId="23" pageNumber="206" type="holotype">holotype</typeStatus>
,
<collectingCountry id="88F3760D601B21028ECE0191FE488D36" box="[424,506,508,534]" name="Greece" pageId="23" pageNumber="206">Greece</collectingCountry>
,
<location id="F53B6046601B21028D620196FD168D35" LSID="urn:lsid:plazi:treatment:784D878B601B21028FF10120FF598BE2:F53B6046601B21028D620196FD168D35" box="[516,676,507,533]" country="Greece" latitude="37.866665" longLatPrecision="1178" longitude="24.833334" municipality="Remmara" name="Andros island" pageId="23" pageNumber="206">Andros island</location>
,
<collectingMunicipality id="103FACE7601B21028DC90191FCAE8D36" box="[687,796,508,534]" pageId="23" pageNumber="206">Remmara</collectingMunicipality>
(
<geoCoordinate id="95D0505A601B21028C4D0196FC3F8D36" box="[811,909,507,534]" degrees="37" direction="north" minutes="52" orientation="latitude" pageId="23" pageNumber="206" precision="925" value="37.866665">37°52’N</geoCoordinate>
/
<geoCoordinate id="95D0505A601B21028CF00196FC478D36" box="[918,1013,507,534]" degrees="24" direction="east" minutes="50" orientation="longitude" pageId="23" pageNumber="206" precision="925" value="24.833334">24°50’E</geoCoordinate>
),
<quantity id="371C9B78601B21028B6C0191FBE38D36" box="[1034,1105,508,534]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="23" pageNumber="206" unit="m" value="170.0">
<elevation id="7BC9D1AE601B21028B6C0191FBE38D36" box="[1034,1105,508,534]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="23" pageNumber="206" unit="m" value="170.0">170 m</elevation>
</quantity>
a.s.l;
<date id="845A105D601B21028BF50191FA808D36" box="[1171,1330,507,534]" pageId="23" pageNumber="206" value="1984-04-17">
<collectingDate id="941EE9B5601B21028BF50191FA808D36" box="[1171,1330,507,534]" pageId="23" pageNumber="206" value="1984-04-17">17 April 1984</collectingDate>
</date>
, leg. Ma- licky. (in collection
<collectionCode id="96F5AE58601B21028E11024DFE178D1A" box="[375,421,544,570]" pageId="23" pageNumber="206">RW</collectionCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601B21028FA1022EFE6F8B76" pageId="23" pageNumber="206" type="description">
<paragraph id="F05B369D601B21028FA1022EFC6F8DCD" blockId="23.[151,1437,507,1218]" pageId="23" pageNumber="206">
<emphasis id="C290EA8F601B21028FA1022EFEE38D7E" bold="true" box="[199,337,579,606]" pageId="23" pageNumber="206">Description</emphasis>
: Head with eyebridge of 3 facet rows; distance between eyes 5 facet diameters; with a row of postocular bristles. Antenna with scape, pedicel and 13 flagellomeres, apical segment missing: absolute length 0.034- 0.054-0.044-0.048-0.040-0.036-0.038-0.036-0.036-0.036-0.036-0.028-0.026-0.022-0.018- mm; relative length 17- 27-22-24-20-18-19-18-18-18-
<date id="845A105D601B21028E8802DDFDC48DEA" box="[494,630,688,714]" pageId="23" pageNumber="206" value="2018-11-14" valueMax="2018-11-13" valueMin="2018-11-14">18-14-13-11</date>
-9-. Flagellomeres 5–10 (not clearly discernable might equally be 11) with a pair of digitiform, curved ascoids in the distal third. Palpus missing.
</paragraph>
<paragraph id="F05B369D601B21028FA1029AFC2D8C32" blockId="23.[151,1437,507,1218]" box="[199,927,759,786]" pageId="23" pageNumber="206">
Wing 3 times longer than wide; length
<quantity id="371C9B78601B21028D1A0295FD508C32" box="[636,738,760,786]" metricMagnitude="-3" metricUnit="m" metricValue="1.81" pageId="23" pageNumber="206" unit="mm" value="1.81">1.81 mm</quantity>
, width
<quantity id="371C9B78601B21028C52029AFC298C32" box="[820,923,759,786]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="23" pageNumber="206" unit="mm" value="0.61">0.61 mm</quantity>
.
</paragraph>
<paragraph id="F05B369D601B21028FA10376FCCB8C5E" blockId="23.[151,1437,507,1218]" pageId="23" pageNumber="206">
Male terminalia with an equally thin hypandrium. Gonocoxites about
<date id="845A105D601B21028CAA0371FC5A8C16" box="[972,1000,796,822]" pageId="23" pageNumber="206">2x</date>
longer than their median width, 1 short seta in the distal third, and 1 long and 1 short setae near the inner basis; condyles oval. Gonostyli 1.5 times longer than gonocoxites, almost straight with broad angular tip (
<figureCitation id="68DF2A18601B21028C7E030EFCDE8C5E" box="[792,876,867,894]" captionStart="FIGURES 50–53" captionStartId="24.[151,264,1201,1226]" captionTargetBox="[169,1415,360,1162]" captionTargetId="figure@24.[151,1436,345,1176]" captionTargetPageId="24" captionText="FIGURES 50–53: 50–51 T. graeca 50 hypandrium, gonopods, aedeagus, 51 epandrial processes and hypoproct; 52–53 T. hatayensis 52 gonocoxite and gonostylus, 53 aedeagus oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621131" httpUri="https://zenodo.org/record/3621131/files/figure.png" pageId="23" pageNumber="206">Fig. 50</figureCitation>
).
</paragraph>
<paragraph id="F05B369D601B21028FA103E5FC888B2D" blockId="23.[151,1437,507,1218]" pageId="23" pageNumber="206">Parameres similar to other species. Ventral bridge not discernable, middle extensions oval with tips converging. Aedeagus with basiphallus narrowing apically, outlines of aedeagus distal of hypandrium rectangular, 3.5 times longer than wide. Basiphallus bilobed, ventral lobe thin with ventral arm connected to the distiphallus; dorsal lobe thin as well but shorter, little sclerotized difficult to identify.</paragraph>
<paragraph id="F05B369D601B21028FA1047AFE6F8B76" blockId="23.[151,1437,507,1218]" pageId="23" pageNumber="206">
Epandrium rhomboid, epandrial processes short, straight with 4 tenacula, 2 long feathered, 2 smaller (
<figureCitation id="68DF2A18601B21028A5A047AFA3D8B12" box="[1340,1423,1047,1074]" captionStart="FIGURES 50–53" captionStartId="24.[151,264,1201,1226]" captionTargetBox="[169,1415,360,1162]" captionTargetId="figure@24.[151,1436,345,1176]" captionTargetPageId="24" captionText="FIGURES 50–53: 50–51 T. graeca 50 hypandrium, gonopods, aedeagus, 51 epandrial processes and hypoproct; 52–53 T. hatayensis 52 gonocoxite and gonostylus, 53 aedeagus oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621131" httpUri="https://zenodo.org/record/3621131/files/figure.png" pageId="23" pageNumber="206">Fig. 51</figureCitation>
). Hypoproct rhomboid, setose.
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601B21028FA10432FD4F8B5A" box="[199,765,1119,1146]" pageId="23" pageNumber="206" type="distribution">
<paragraph id="F05B369D601B21028FA10432FD4F8B5A" blockId="23.[151,1437,507,1218]" box="[199,765,1119,1146]" pageId="23" pageNumber="206">
<emphasis id="C290EA8F601B21028FA10432FEEB8B5A" bold="true" box="[199,345,1119,1146]" pageId="23" pageNumber="206">Distribution</emphasis>
:
<collectingCountry id="88F3760D601B21028E0E040DFE088B5A" box="[360,442,1120,1146]" name="Greece" pageId="23" pageNumber="206">Greece</collectingCountry>
, Andros island (Egean area).
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601B21028FA104E9FF598BE2" pageId="23" pageNumber="206" type="discussion">
<paragraph id="F05B369D601B21028FA104E9FF598BE2" blockId="23.[151,1437,507,1218]" pageId="23" pageNumber="206">
<emphasis id="C290EA8F601B21028FA104E9FE868BBE" bold="true" box="[199,308,1156,1182]" pageId="23" pageNumber="206">Remarks</emphasis>
: a
<taxonomicName id="37E44D1E601B21028E0604E9FE5B8BBE" authorityName="Vaillant" authorityYear="1971" box="[352,489,1156,1182]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="206" phylum="Arthropoda" rank="genus">
<emphasis id="C290EA8F601B21028E0604E9FE5B8BBE" box="[352,489,1156,1182]" italics="true" pageId="23" pageNumber="206">Tonnoiriella</emphasis>
</taxonomicName>
species presently difficult to group. No close relations to any of the mentioned groups.
</paragraph>
</subSubSection>
</treatment>
</document>
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