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<document id="995C6CF3B75F2F624E3C280448F444D0" ID-CLB-Dataset="27316" ID-DOI="10.11646/zootaxa.4583.1.1" ID-GBIF-Dataset="027ed27d-5035-4646-93a6-c23b565f3d20" ID-ISSN="1175-5326" ID-Zenodo-Dep="2637603" ID-ZooBank="27F24995-359E-46F6-AB22-75568BACFDCF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555054242242" checkinUser="plazi" docAuthor="Zhang, - H." docDate="2019" docId="724987F6FF812417FF7EB970FF393EA8" docLanguage="en" docName="zootaxa.4583.1.1.pdf" docOrigin="Zootaxa 4583 (1)" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Eulodrobia eulo" docType="treatment" docVersion="15" lastPageNumber="11" masterDocId="8E70FF8EFF86241DFFE9BE79FFE73928" masterDocTitle="New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages" masterLastPageNumber="67" masterPageNumber="1" pageNumber="8" updateTime="1698720421338" updateUser="ExternalLinkService">
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<mods:title id="C60C7A0F1E5BB9356553710AB7E93353">New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="9715605A22B2232723A0182EA081234F">Author</mods:roleTerm>
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<mods:namePart id="543059A7D499BA3FE188B3A196D2F8E6">Zhang, - H.</mods:namePart>
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<mods:typeOfResource id="01E8E00EA5973C8AEA5DD3CFF1158687">text</mods:typeOfResource>
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<mods:title id="B4EDF2B2DE5647280DDF76C5AFCBDEA9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="00EAC11228381FFF48D310B898345DD2">
<mods:date id="09543FCEFEF8D166FCDF742021A052D8">2019</mods:date>
<mods:detail id="994D7633AA072DB95F379C06EA5DA0E6" type="pubDate">
<mods:number id="5D40A3BC8FBFFC59BCAA7070D8D5C35E">2019-04-10</mods:number>
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<mods:detail id="6252EAD9932439C6B5F7C72C0B4A4F11" type="volume">
<mods:number id="147A3B2E6F7EB81B6043C175A120FDEC">4583</mods:number>
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<mods:number id="46451EEA1B6C0765AB870CCFF140B857">1</mods:number>
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<mods:classification id="7C5F1AD3EE9D87BB9B2D0B4387807B4B">journal article</mods:classification>
<mods:identifier id="B02C47CC5EC577F9CCC0F3F110E9C407" type="CLB-Dataset">27316</mods:identifier>
<mods:identifier id="81C096F1EACD8E3E672F9B07FABF64E1" type="DOI">10.11646/zootaxa.4583.1.1</mods:identifier>
<mods:identifier id="AA8509CC7D26D6DAACCB5B8C0BC1CEB8" type="GBIF-Dataset">027ed27d-5035-4646-93a6-c23b565f3d20</mods:identifier>
<mods:identifier id="8B661C68B26D305C1436C464740D412A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="737859A0829D442B265423C93010D760" type="Zenodo-Dep">2637603</mods:identifier>
<mods:identifier id="E3D7D7ED775F2A552E5A92348FC13F95" type="ZooBank">27F24995-359E-46F6-AB22-75568BACFDCF</mods:identifier>
</mods:mods>
<treatment id="724987F6FF812417FF7EB970FF393EA8" ID-GBIF-Taxon="156198451" LSID="urn:lsid:plazi:treatment:724987F6FF812417FF7EB970FF393EA8" httpUri="http://treatment.plazi.org/id/724987F6FF812417FF7EB970FF393EA8" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">
<subSubSection id="B2FA656BFF81241AFF7EB970FC5E3E0B" box="[151,953,1801,1827]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="FA5F36E0FF81241AFF7EB970FC5E3E0B" blockId="7.[151,953,1801,1827]" box="[151,953,1801,1827]" pageId="7" pageNumber="8">
<heading id="A117818CFF81241AFF7EB970FC5E3E0B" bold="true" box="[151,953,1801,1827]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="C894EAF2FF81241AFF7EB970FC5E3E0B" bold="true" box="[151,953,1801,1827]" pageId="7" pageNumber="8">
<taxonomicName id="3DE04D63FF81241AFF7EB970FD6D3E0B" ID-CoL="7R9CV" authority="(Ponder &amp; Clark, 1990)" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[151,650,1801,1827]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF81241AFF7EB970FEBB3E0B" bold="true" box="[151,348,1801,1827]" italics="true" pageId="7" pageNumber="8">Eulodrobia eulo</emphasis>
(
<bibRefCitation id="9E714B11FF81241AFE83B970FD653E0B" author="Ponder, W. F. &amp; Clark, G. A." box="[362,642,1801,1827]" pageId="7" pageNumber="8" pagination="301 - 363" refId="ref35763" refString="Ponder, W. F. &amp; Clark, G. A. (1990) A radiation of hydrobiid snails in threatened artesian springs in Western Queensland. Records of the Australian Museum, 42, 301 - 363. https: // doi. org / 10.3853 / j. 0067 - 1975.42.1990.119" type="journal article" year="1990">Ponder &amp; Clark, 1990</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="3DE04D63FF81241AFD21B970FC553E0B" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[712,946,1801,1827]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF81241AFD21B970FCB63E0B" bold="true" box="[712,849,1801,1827]" italics="true" pageId="7" pageNumber="8">Eulodrobia</emphasis>
cf.
<emphasis id="C894EAF2FF81241AFC97B970FC553E0B" bold="true" box="[894,946,1801,1827]" italics="true" pageId="7" pageNumber="8">eulo</emphasis>
</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF81241AFF7EB929FEE53E8B" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="FA5F36E0FF81241AFF7EB929FEE53E8B" blockId="7.[151,1435,1872,1955]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="DAF011CEFF81241AFF7EB929FEE53E8B" pageId="7" pageNumber="8">
<taxonomicName id="3DE04D63FF81241AFF7EB929FDAC3E4E" ID-CoL="7R9CV" authority="Ponder &amp; Clark, 1990: 342" authorityName="Ponder &amp; Clark" authorityPageNumber="342" authorityYear="1990" box="[151,587,1872,1894]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF81241AFF7EB929FED73E4E" box="[151,304,1872,1894]" italics="true" pageId="7" pageNumber="8">Jardinella eulo</emphasis>
<treatmentCitation id="7B4110F1FF81241AFEDFB929FDAC3E4E" author="Ponder, W. F. &amp; Clark, G. A." box="[310,587,1872,1894]" page="342" pageId="7" pageNumber="8" year="1990">
<bibRefCitation id="9E714B11FF81241AFEDFB929FDFE3E4E" author="Ponder, W. F. &amp; Clark, G. A." box="[310,537,1872,1894]" pageId="7" pageNumber="8" pagination="301 - 363" refId="ref35763" refString="Ponder, W. F. &amp; Clark, G. A. (1990) A radiation of hydrobiid snails in threatened artesian springs in Western Queensland. Records of the Australian Museum, 42, 301 - 363. https: // doi. org / 10.3853 / j. 0067 - 1975.42.1990.119" type="journal article" year="1990">Ponder &amp; Clark, 1990</bibRefCitation>
: 342
</treatmentCitation>
</taxonomicName>
, figs. 6C, 31A, B, 32, A, B, 33C, D, 34D, F.
<taxonomicName id="3DE04D63FF81241AFF7EB917FDC13EAC" ID-CoL="7R9CV" authority="Perez et al. 2005: 547" authorityName="Perez" authorityPageNumber="547" authorityYear="2005" box="[151,550,1902,1925]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF81241AFF7EB917FED43EAC" box="[151,307,1902,1924]" italics="true" pageId="7" pageNumber="8">Jardinella eulo</emphasis>
<treatmentCitation id="7B4110F1FF81241AFED5B916FDC13EAC" author="Perez, K. E. &amp; Ponder, W. F. &amp; Colgan, D. J. &amp; Clark, S. A. &amp; Lydeard, C." box="[316,550,1902,1925]" page="547" pageId="7" pageNumber="8" year="2005">
<bibRefCitation id="9E714B11FF81241AFED5B916FE143EAC" author="Perez, K. E. &amp; Ponder, W. F. &amp; Colgan, D. J. &amp; Clark, S. A. &amp; Lydeard, C." box="[316,499,1902,1925]" pageId="7" pageNumber="8" pagination="545 - 556" refId="ref35351" refString="Perez, K. E., Ponder, W. F., Colgan, D. J., Clark, S. A. &amp; Lydeard, C. (2005) Molecular phylogeny and biogeography of springassociated hydrobiid snails of the Great Artesian Basin, Australia. Molecular Phylogenetics and Evolution, 34, 545 - 556. https: // doi. org / 10.1016 / j. ympev. 2004.11.020" type="journal article" year="2005">
Perez
<emphasis id="C894EAF2FF81241AFE97B909FE533EAC" box="[382,436,1902,1925]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
2005
</bibRefCitation>
: 547
</treatmentCitation>
</taxonomicName>
(C.400135);
<emphasis id="C894EAF2FF81241AFD5DB917FCFC3EAC" box="[692,795,1902,1924]" italics="true" pageId="7" pageNumber="8">Jardinella</emphasis>
sp.
<collectingCountry id="82F77670FF81241AFCA3B917FB933EAC" box="[842,1140,1902,1924]" name="Australia" pageId="7" pageNumber="8">
Commonwealth of
<treatmentCitation id="7B4110F1FF81241AFBFDB917FB933EAC" author="Commonwealth of Australia" box="[1044,1140,1902,1924]" page="37" pageId="7" pageNumber="8" year="2014">
<bibRefCitation id="9E714B11FF81241AFBFDB917FB933EAC" author="Commonwealth of Australia" box="[1044,1140,1902,1924]" pageId="7" pageNumber="8" refId="ref33702" refString="Commonwealth of Australia (2014) Ecological and hydrogeological survey of the Great Artesian Basin springs-Springsure, Eulo, Bourke and Bogan River Supergroups. Fol. 2. Hydrogeological profiles, Knowledge report, prepared by UniQuest for the Department of the Environment, Commonwealth of Australia, Canberra. Department of the Environment, Public Affairs, Canberra, xiii + 307 pp." type="book" year="2014">Australia</bibRefCitation>
</treatmentCitation>
</collectingCountry>
, 2014: 37 (=
<taxonomicName id="3DE04D63FF81241AFAEFB917FF143E8B" ID-CoL="7R9CV" authority="Ponder &amp; Clark, 1990" authorityName="Ponder &amp; Clark" authorityYear="1990" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF81241AFAEFB917FA9E3EAC" box="[1286,1401,1902,1924]" italics="true" pageId="7" pageNumber="8">Eulodrobia</emphasis>
cf.
<emphasis id="C894EAF2FF81241AFF2EB9F4FF143E8B" box="[199,243,1933,1955]" italics="true" pageId="7" pageNumber="8">eulo</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF812415FF7EB9A9FBA53885" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="FA5F36E0FF81241AFF7EB9A9FA7B3EC1" blockId="7.[151,1436,1999,2025]" box="[151,1436,1999,2025]" pageId="7" pageNumber="8">
<emphasis id="C894EAF2FF81241AFF7EB9A9FE093EC0" bold="true" box="[151,494,1999,2025]" pageId="7" pageNumber="8">
Material examined.
<taxonomicName id="3DE04D63FF81241AFE63B9A9FE093EC0" box="[394,494,1999,2025]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF81241AFE63B9A9FE4B3ECF" bold="true" box="[394,428,2000,2023]" italics="true" pageId="7" pageNumber="8">Eu</emphasis>
.
<emphasis id="C894EAF2FF81241AFE55B9B6FE093EC0" box="[444,494,1999,2024]" italics="true" pageId="7" pageNumber="8">eulo</emphasis>
</taxonomicName>
</emphasis>
.
<collectingRegion id="3824F802FF81241AFE14B9A9FD633EC1" box="[509,644,2000,2025]" country="Australia" name="Queensland" pageId="7" pageNumber="8">Queensland</collectingRegion>
, Massey Spring (=Rocky Springs), Granites Springs, near Mt Francis
</paragraph>
<paragraph id="FA5F36E0FF8E2415FF7EBEE1FEEB39D0" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="9">
ca
<quantity id="3D189B05FF8E2415FF51BEEEFEE73998" box="[184,256,151,176]" metricMagnitude="4" metricUnit="m" metricValue="5.2" pageId="8" pageNumber="9" unit="km" value="52.0">52 km</quantity>
SW of Eulo,
<geoCoordinate id="9FD45027FF8E2415FE76BEE1FDDF3998" box="[415,568,152,176]" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="15" value="-28.3">28° 18' 00&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FF8E2415FDAFBEE1FD163987" box="[582,753,152,176]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="15" value="144.53334">144° 32' 00&quot; E</geoCoordinate>
, on small-leaved plants, W.F. Ponder &amp; P.H. Colman,
<date id="8E5E1020FF8E2415FAB5BEE1FF2839FC" pageId="8" pageNumber="9" value="1984-09-07">7 Sep 1984</date>
(
<typeStatus id="255B8842FF8E2415FF01BEC5FEA939FD" box="[232,334,188,213]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
, C.156778, and
<typeStatus id="255B8842FF8E2415FDCABEC4FD7639FD" box="[547,657,189,213]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
, C.156779). Additional material from same locality, C.410724, C.479953.
</paragraph>
<paragraph id="FA5F36E0FF8E2415FF2EBF7DFBA53885" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="9">
<taxonomicName id="3DE04D63FF8E2415FF2EBF7DFEA93834" authority="Unnamed" authorityName="Unnamed" box="[199,334,259,285]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF8E2415FF2EBF7DFEA93834" bold="true" box="[199,334,259,285]" pageId="8" pageNumber="9">
<emphasis id="C894EAF2FF8E2415FF2EBF7DFF0E3833" bold="true" box="[199,233,260,283]" italics="true" pageId="8" pageNumber="9">Eu</emphasis>
. cf.
<emphasis id="C894EAF2FF8E2415FEF5BF7AFEA93834" bold="true" box="[284,334,259,284]" italics="true" pageId="8" pageNumber="9">eulo</emphasis>
</emphasis>
</taxonomicName>
‘Unnamed spring’ (=Tunga Spring) at Tunga Bore,
<collectingRegion id="3824F802FF8E2415FC76BF7DFBC13835" box="[927,1062,260,285]" country="Australia" name="Queensland" pageId="8" pageNumber="9">Queensland</collectingRegion>
,
<geoCoordinate id="9FD45027FF8E2415FBDBBF7DFB2C3834" box="[1074,1227,260,284]" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="15" value="-28.216667">28° 13' 00&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FF8E2415FB31BF7DFA653834" box="[1240,1410,260,284]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="15" value="144.63333">144° 38' 00&quot; E</geoCoordinate>
,
<date id="8E5E1020FF8E2415FA67BF7DFEE43868" pageId="8" pageNumber="9" value="1984-09-06">6 Sep 1984</date>
, W.F. Ponder &amp; P.H. Colman, spring flows into boulder strewn creek, C.156780 (designated as
<typeStatus id="255B8842FF8E2415FAD8BF51FA7B3868" box="[1329,1436,296,320]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="3DE04D63FF8E2415FF5DBF34FEF6384D" box="[180,273,332,357]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF8E2415FF5DBF34FEF6384D" box="[180,273,332,357]" italics="true" pageId="8" pageNumber="9">Eu. eulo</emphasis>
</taxonomicName>
). Other material from same locality
<date id="8E5E1020FF8E2415FD40BF35FCDA384C" box="[681,829,332,356]" pageId="8" pageNumber="9" value="2000-08-28">28 Aug 2000</date>
, R. J. Fensham, artesian spring with flow enhanced by bore head, on mud and plant surfaces, C.410643 (SEM material), 20+; C.479952, 1;
<date id="8E5E1020FF8E2415FBAABF09FB2A38A0" box="[1091,1229,368,392]" pageId="8" pageNumber="9" value="2001-05-02">2 May 2001</date>
, W.F. Ponder &amp; C. Lydeard, in shallow muddy habitat at head of spring, C.400135 (molecular sample).
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF8E2417FF2EBFCEFA723DDF" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="FA5F36E0FF8E2415FF2EBFCEFD123BAD" blockId="8.[151,1437,151,645]" pageId="8" pageNumber="9">
<emphasis id="C894EAF2FF8E2415FF2EBFCEFEE438F8" bold="true" box="[199,259,439,464]" pageId="8" pageNumber="9">Shell</emphasis>
(
<figureCitation id="62DB2A65FF8E2415FEFABFCEFE6838F8" box="[275,399,439,464]" captionStart="FIGURE 2" captionStartId="8.[151,250,1857,1879]" captionTargetBox="[185,1418,742,1819]" captionTargetId="figure@8.[150,1436,697,1836]" captionTargetPageId="8" captionText="FIGURE 2. Shells of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp., holotype, spring at Town Common, Eulo (C.479950). B. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). C. Eulodrobia cf. eulo (Ponder &amp; Clark, 1990), Tunga Spring (C.410643). D. Eulodrobia ovata n. sp., holotype, Yowah Springs, Bundoona Station (C.479944). E. Eulodrobia bundoona n. sp., holotype, Yowah Springs, Bundoona Station (C.479943). F. Eulodrobia carinata n. sp., holotype, Yowah Springs, Bundoona Station (C.479942). G–I. Eulodrobia spirula n. sp., Yowah Springs, Bundoona Station. G. dorsal view of holotype (479945). H, I. apertural view of paratypes (C.400133). Scale bars 1 mm." httpUri="https://zenodo.org/record/2637607/files/figure.png" pageId="8" pageNumber="9">Fig. 2B, C</figureCitation>
,
<figureCitation id="62DB2A65FF8E2415FE74BFC1FE2738F8" box="[413,448,440,464]" captionStart="FIGURE 3" captionStartId="9.[151,250,1476,1498]" captionTargetBox="[221,1370,184,1448]" captionTargetId="figure@9.[204,1382,169,1464]" captionTargetPageId="9" captionText="FIGURE 3. Protoconchs and protoconch microsculpture of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). B. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). C. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). D. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). E. Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131). F. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133). Scale bars 100 µm (lowpowered images [left]), 10 µm (high-powered images [right])." httpUri="https://zenodo.org/record/2637609/files/figure.png" pageId="8" pageNumber="9">3B</figureCitation>
). Ovate, spire outline slightly convex, normally coiled, opaque. Length
<quantity id="3D189B05FF8E2415FB11BFC1FA7A38E7" box="[1272,1437,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.935" metricValueMax="2.43" metricValueMin="1.44" pageId="8" pageNumber="9" unit="mm" value="1.935" valueMax="2.43" valueMin="1.44">1.44–2.43 mm</quantity>
(mean
<quantity id="3D189B05FF8E2415FF0DBFA5FED938DC" box="[228,318,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="9" unit="mm" value="2.4">2.4 mm</quantity>
), width
<quantity id="3D189B05FF8E2415FE7EBFA5FDF838DC" box="[407,543,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="8" pageNumber="9" unit="mm" value="1.9" valueMax="2.3" valueMin="1.5">1.5–2.3 mm</quantity>
(mean
<quantity id="3D189B05FF8E2415FD9BBFA5FD2B38DC" box="[626,716,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="9" unit="mm" value="1.9">1.9 mm</quantity>
). Protoconch of 1.5 whorls, minutely and irregularly pitted (
<figureCitation id="62DB2A65FF8E2415FA86BFA5FF5D3B30" captionStart="FIGURE 3" captionStartId="9.[151,250,1476,1498]" captionTargetBox="[221,1370,184,1448]" captionTargetId="figure@9.[204,1382,169,1464]" captionTargetPageId="9" captionText="FIGURE 3. Protoconchs and protoconch microsculpture of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). B. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). C. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). D. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). E. Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131). F. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133). Scale bars 100 µm (lowpowered images [left]), 10 µm (high-powered images [right])." httpUri="https://zenodo.org/record/2637609/files/figure.png" pageId="8" pageNumber="9">Fig. 3B</figureCitation>
). Teleoconch whorls moderately convex, evenly rounded, total number 3.5–4.3 (mean 3.6). Umbilicus moderate. Sculpture of close growth lines, often raised into weak riblets. Aperture ovate, inner lip narrow, thick, slightly separated along whole length of parietal wall, outer lip medium thickness. Periostracum moderately developed, yellow-brown to reddish or orange-brown.
</paragraph>
<caption id="AE9F6668FF8E2415FF7EB938FB033ED8" httpUri="https://zenodo.org/record/2637607/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1857,1879]" targetBox="[185,1418,742,1819]" targetPageId="8">
<paragraph id="FA5F36E0FF8E2415FF7EB938FB033ED8" blockId="8.[151,1436,1857,2032]" pageId="8" pageNumber="9">
<emphasis id="C894EAF2FF8E2415FF7EB938FEF33E7F" bold="true" box="[151,276,1857,1879]" pageId="8" pageNumber="9">FIGURE 2.</emphasis>
Shells of species of
<taxonomicName id="3DE04D63FF8E2415FE03B938FDBA3E7F" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[490,605,1857,1879]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="C894EAF2FF8E2415FE03B938FDBA3E7F" box="[490,605,1857,1879]" italics="true" pageId="8" pageNumber="9">Eulodrobia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8E2415FD8DB938FD4B3E7F" box="[612,684,1857,1879]" pageId="8" pageNumber="9" rank="genus">
<emphasis id="C894EAF2FF8E2415FD8DB938FD4B3E7F" bold="true" box="[612,684,1857,1879]" pageId="8" pageNumber="9">n. gen.</emphasis>
</taxonomicNameLabel>
A.
<taxonomicName id="3DE04D63FF8E2415FD3DB938FC4E3E7F" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[724,937,1857,1879]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="fenshami" status="sp. nov.">
<emphasis id="C894EAF2FF8E2415FD3DB938FC4E3E7F" box="[724,937,1857,1879]" italics="true" pageId="8" pageNumber="9">Eulodrobia fenshami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8E2415FC59B938FC0C3E7F" box="[944,1003,1857,1879]" pageId="8" pageNumber="9" rank="species">
<emphasis id="C894EAF2FF8E2415FC59B938FC0C3E7F" bold="true" box="[944,1003,1857,1879]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, spring at Town Common, Eulo (C.479950). B.
<taxonomicName id="3DE04D63FF8E2415FEDCB919FD333E5E" authority="(Ponder &amp; Clark, 1990)" authorityName="Ponder &amp; Clark" authorityYear="1990" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[309,724,1888,1910]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF8E2415FEDCB919FE3D3E5E" box="[309,474,1888,1910]" italics="true" pageId="8" pageNumber="9">Eulodrobia eulo</emphasis>
(Ponder &amp; Clark, 1990)
</taxonomicName>
, paratype, Massey Spring (C.156779). C.
<taxonomicName id="3DE04D63FF8E2415FB69B919FEA53EBC" authority="(Ponder &amp; Clark, 1990)" authorityName="Ponder &amp; Clark" authorityYear="1990" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF8E2415FB69B919FB153E5E" box="[1152,1266,1888,1910]" italics="true" pageId="8" pageNumber="9">Eulodrobia</emphasis>
cf.
<emphasis id="C894EAF2FF8E2415FAF3B919FAA13E5E" box="[1306,1350,1888,1910]" italics="true" pageId="8" pageNumber="9">eulo</emphasis>
(Ponder &amp; Clark, 1990)
</taxonomicName>
, Tunga Spring (C.410643). D.
<taxonomicName id="3DE04D63FF8E2415FD48B907FCBD3EBD" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[673,858,1918,1941]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="ovata" status="sp. nov.">
<emphasis id="C894EAF2FF8E2415FD48B907FCBD3EBD" box="[673,858,1918,1941]" italics="true" pageId="8" pageNumber="9">Eulodrobia ovata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8E2415FC8EB907FC4F3EBC" box="[871,936,1918,1940]" pageId="8" pageNumber="9" rank="species">
<emphasis id="C894EAF2FF8E2415FC8EB907FC4F3EBC" bold="true" box="[871,936,1918,1940]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, Yowah Springs, Bundoona Station (C.479944). E.
<taxonomicName id="3DE04D63FF8E2415FEDCB9E4FDF33E9B" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[309,532,1949,1971]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="bundoona" status="sp. nov.">
<emphasis id="C894EAF2FF8E2415FEDCB9E4FDF33E9B" box="[309,532,1949,1971]" italics="true" pageId="8" pageNumber="9">Eulodrobia bundoona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8E2415FDF5B9E4FDB13E9B" box="[540,598,1949,1971]" pageId="8" pageNumber="9" rank="species">
<emphasis id="C894EAF2FF8E2415FDF5B9E4FDB13E9B" bold="true" box="[540,598,1949,1971]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, Yowah Springs, Bundoona Station (C.479943). F.
<taxonomicName id="3DE04D63FF8E2415FB25B9E4FA7B3E9B" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[1228,1436,1949,1971]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="carinata" status="sp. nov.">
<emphasis id="C894EAF2FF8E2415FB25B9E4FA7B3E9B" box="[1228,1436,1949,1971]" italics="true" pageId="8" pageNumber="9">Eulodrobia carinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8E2415FF7EB9C5FF353EFA" box="[151,210,1980,2002]" pageId="8" pageNumber="9" rank="species">
<emphasis id="C894EAF2FF8E2415FF7EB9C5FF353EFA" bold="true" box="[151,210,1980,2002]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, Yowah Springs, Bundoona Station (C.479942). G–I.
<taxonomicName id="3DE04D63FF8E2415FC98B9C5FBD33EFA" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[881,1076,1980,2002]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="spirula" status="sp. nov.">
<emphasis id="C894EAF2FF8E2415FC98B9C5FBD33EFA" box="[881,1076,1980,2002]" italics="true" pageId="8" pageNumber="9">Eulodrobia spirula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8E2415FBD4B9C5FB9E3EFA" box="[1085,1145,1980,2002]" pageId="8" pageNumber="9" rank="species">
<emphasis id="C894EAF2FF8E2415FBD4B9C5FB9E3EFA" bold="true" box="[1085,1145,1980,2002]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
, Yowah Springs, Bundoona Station. G. dorsal view of holotype (479945). H, I. apertural view of paratypes (C.400133). Scale bars 1 mm.
</paragraph>
</caption>
<caption id="AE9F6668FF8F2414FF7EBBBDFCED3F5B" httpUri="https://zenodo.org/record/2637609/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1476,1498]" targetBox="[221,1370,184,1448]" targetPageId="9">
<paragraph id="FA5F36E0FF8F2414FF7EBBBDFCED3F5B" blockId="9.[151,1436,1476,1651]" pageId="9" pageNumber="10">
<emphasis id="C894EAF2FF8F2414FF7EBBBDFEF13CF2" bold="true" box="[151,278,1476,1498]" pageId="9" pageNumber="10">FIGURE 3.</emphasis>
Protoconchs and protoconch microsculpture of species of
<taxonomicName id="3DE04D63FF8F2414FC6DBBBDFC103CF2" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[900,1015,1476,1498]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="C894EAF2FF8F2414FC6DBBBDFC103CF2" box="[900,1015,1476,1498]" italics="true" pageId="9" pageNumber="10">Eulodrobia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8F2414FBE9BBBDFBAC3CF2" box="[1024,1099,1476,1498]" pageId="9" pageNumber="10" rank="genus">
<emphasis id="C894EAF2FF8F2414FBE9BBBDFBAC3CF2" bold="true" box="[1024,1099,1476,1498]" pageId="9" pageNumber="10">n. gen.</emphasis>
</taxonomicNameLabel>
A.
<taxonomicName id="3DE04D63FF8F2414FB9EBBBDFAA83CF2" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[1143,1359,1476,1498]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="fenshami" status="sp. nov.">
<emphasis id="C894EAF2FF8F2414FB9EBBBDFAA83CF2" box="[1143,1359,1476,1498]" italics="true" pageId="9" pageNumber="10">Eulodrobia fenshami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8F2414FAB0BBBDFA713CF2" box="[1369,1430,1476,1498]" pageId="9" pageNumber="10" rank="species">
<emphasis id="C894EAF2FF8F2414FAB0BBBDFA713CF2" bold="true" box="[1369,1430,1476,1498]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, spring at Town Common, Eulo (C.410721). B.
<taxonomicName id="3DE04D63FF8F2414FD01BB9BFB683CD0" authority="(Ponder &amp; Clark, 1990)" authorityName="Ponder &amp; Clark" authorityYear="1990" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[744,1167,1506,1528]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF8F2414FD01BB9BFC683CD0" box="[744,911,1506,1528]" italics="true" pageId="9" pageNumber="10">Eulodrobia eulo</emphasis>
(Ponder &amp; Clark, 1990)
</taxonomicName>
, paratype, Massey Spring (C.156779). C.
<taxonomicName id="3DE04D63FF8F2414FEDFB878FE003F3F" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[310,487,1537,1559]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="ovata" status="sp. nov.">
<emphasis id="C894EAF2FF8F2414FEDFB878FE003F3F" box="[310,487,1537,1559]" italics="true" pageId="9" pageNumber="10">Eulodrobia ovata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8F2414FE07B878FDC03F3F" box="[494,551,1537,1559]" pageId="9" pageNumber="10" rank="species">
<emphasis id="C894EAF2FF8F2414FE07B878FDC03F3F" bold="true" box="[494,551,1537,1559]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, Yowah Springs, Bundoona Stn (C.400130). D.
<taxonomicName id="3DE04D63FF8F2414FB9EB878FAB23F3F" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[1143,1365,1537,1559]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="bundoona" status="sp. nov.">
<emphasis id="C894EAF2FF8F2414FB9EB878FAB23F3F" box="[1143,1365,1537,1559]" italics="true" pageId="9" pageNumber="10">Eulodrobia bundoona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8F2414FAB5B878FA713F3F" box="[1372,1430,1537,1559]" pageId="9" pageNumber="10" rank="species">
<emphasis id="C894EAF2FF8F2414FAB5B878FA713F3F" bold="true" box="[1372,1430,1537,1559]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, Yowah Springs, Bundoona Stn (C.400132). E.
<taxonomicName id="3DE04D63FF8F2414FD34B859FC4A3F1E" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[733,941,1568,1590]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="carinata" status="sp. nov.">
<emphasis id="C894EAF2FF8F2414FD34B859FC4A3F1E" box="[733,941,1568,1590]" italics="true" pageId="9" pageNumber="10">Eulodrobia carinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8F2414FC5DB859FC083F1E" box="[948,1007,1568,1590]" pageId="9" pageNumber="10" rank="species">
<emphasis id="C894EAF2FF8F2414FC5DB859FC083F1E" bold="true" box="[948,1007,1568,1590]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, Yowah Springs, Bundoona Stn (C.400131). F.
<taxonomicName id="3DE04D63FF8F2414FED0B847FE193F7C" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[313,510,1598,1620]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="spirula" status="sp. nov.">
<emphasis id="C894EAF2FF8F2414FED0B847FE193F7C" box="[313,510,1598,1620]" italics="true" pageId="9" pageNumber="10">Eulodrobia spirula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8F2414FDE1B847FDA13F7C" box="[520,582,1598,1620]" pageId="9" pageNumber="10" rank="species">
<emphasis id="C894EAF2FF8F2414FDE1B847FDA13F7C" bold="true" box="[520,582,1598,1620]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, Yowah Springs, Bundoona Stn (C.400133). Scale bars 100 µm (lowpowered images [left]), 10 µm (high-powered images [right]).
</paragraph>
</caption>
<paragraph id="FA5F36E0FF8F2414FF2EB8CFFC983FDB" blockId="9.[151,1437,1718,2031]" pageId="9" pageNumber="10">
<emphasis id="C894EAF2FF8F2414FF2EB8CFFEA83FE7" bold="true" box="[199,335,1718,1743]" pageId="9" pageNumber="10">Operculum</emphasis>
(
<figureCitation id="62DB2A65FF8F2414FE88B8CFFE053FE7" box="[353,482,1718,1743]" captionStart="FIGURE 4" captionStartId="12.[151,250,1521,1543]" captionTargetBox="[196,1406,256,1477]" captionTargetId="figure@12.[151,1435,229,1500]" captionTargetPageId="12" captionText="FIGURE 4. Opercula of species of Eulodrobia n. gen., external and internal views of each species. A, B. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). C, D. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). E, F. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). G, H. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). I, J. Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131). K, L. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133). Scale bars 500 µm." httpUri="https://zenodo.org/record/2637611/files/figure.png" pageId="9" pageNumber="10">Fig. 4C, D</figureCitation>
). Translucent, yellow-brown, slightly concave, nucleus acentric. Inner side lacking white smear, simple except for low rounded thickening at nucleus.
</paragraph>
<paragraph id="FA5F36E0FF8F2414FF2EB887FD0E3E13" blockId="9.[151,1437,1718,2031]" pageId="9" pageNumber="10">
<emphasis id="C894EAF2FF8F2414FF2EB887FDC13E3F" bold="true" box="[199,550,1790,1815]" pageId="9" pageNumber="10">Head-foot and external body.</emphasis>
Snout black, tentacles dark grey; dorsal and lateral parts of foot and opercular lobes pigmented, mantle roof and visceral coil black.
</paragraph>
<paragraph id="FA5F36E0FF8F2414FF2EB93FFA673EC7" blockId="9.[151,1437,1718,2031]" pageId="9" pageNumber="10">
<emphasis id="C894EAF2FF8F2414FF2EB93FFE9A3E77" bold="true" box="[199,381,1862,1887]" pageId="9" pageNumber="10">Mantle cavity.</emphasis>
Ctenidium well-developed, filaments 24–28, broadly triangular, apex towards right. Osphradium narrowly oval, towards posterior end of ctenidium, length relative to gill 0.29–0.31. Hypobranchial gland thin. Rectum with U-shaped bend, with longitudinal to oblique pellets, anus behind mantle collar. Kidney extends for about one third to one half of length into mantle cavity roof. Renal gland longitudinal. Pericardium extends for about one third to one half of length into mantle cavity roof, overlapping posterior end of ctenidium.
</paragraph>
<caption id="AE9F6668FF8C2417FF7EBEE1FB4D39E7" ID-Table-UUID="AE9F6668FF8C2417FF7EBEE1FB4D39E7" httpUri="http://table.plazi.org/id/AE9F6668FF8C2417FF7EBEE1FB4D39E7" pageId="10" pageNumber="11" targetBox="[159,1337,234,711]" targetIsTable="true" targetPageId="10">
<paragraph id="FA5F36E0FF8C2417FF7EBEE1FB4D39E7" blockId="10.[151,1435,151,207]" pageId="10" pageNumber="11">
<emphasis id="C894EAF2FF8C2417FF7EBEE1FEED3987" bold="true" box="[151,266,152,175]" pageId="10" pageNumber="11">TABLE 1.</emphasis>
Shell measurements (in mm) of the type material of
<taxonomicName id="3DE04D63FF8C2417FCD4BEEEFB093987" authority="(Ponder &amp; Clark, 1990)" authorityName="Ponder &amp; Clark" authorityYear="1990" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[829,1262,151,175]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF8C2417FCD4BEEEFC0D3987" box="[829,1002,151,175]" italics="true" pageId="10" pageNumber="11">Eulodrobia eulo</emphasis>
(Ponder &amp; Clark, 1990)
</taxonomicName>
(C.410724) and
<taxonomicName id="3DE04D63FF8C2417FF7EBECEFE8339E7" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[151,356,183,207]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF8C2417FF7EBECEFE8339E7" box="[151,356,183,207]" italics="true" pageId="10" pageNumber="11">Eulodrobia cf. eulo</emphasis>
</taxonomicName>
from Tunga Spring (C.410643). See Materials and Methods for abbreviations.
</paragraph>
</caption>
<paragraph id="FA5F36E0FF8C2417FF76BE92FADF3BEF" pageId="10" pageNumber="11">
<table id="88E0C440FF8CDBE2FF76BE93FADE3BEF" box="[159,1337,234,711]" gridcols="7" gridrows="12" pageId="10" pageNumber="11">
<tr id="44D034A2FF8CDBE2FF76BE93FADE3828" box="[159,1337,234,256]" gridrow="0" pageId="10" pageNumber="11">
<th id="07015DDEFF8CDBE2FF76BE93FF1A3828" box="[159,253,234,256]" gridcol="0" gridrow="0" pageId="10" pageNumber="11">
<taxonomicName id="3DE04D63FF8C2417FF76BE92FF133828" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[159,244,234,256]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="eulo">Eu. eulo</taxonomicName>
</th>
<th id="07015DDEFF8CDBE2FE39BE93FE1B3828" box="[464,508,234,256]" gridcol="1" gridrow="0" pageId="10" pageNumber="11">SH</th>
<th id="07015DDEFF8CDBE2FD9DBE93FD423828" box="[628,677,234,256]" gridcol="2" gridrow="0" pageId="10" pageNumber="11">SW</th>
<th id="07015DDEFF8CDBE2FCFEBE93FCAD3828" box="[791,842,234,256]" gridcol="3" gridrow="0" pageId="10" pageNumber="11">AH</th>
<th id="07015DDEFF8CDBE2FC52BE93FC003828" box="[955,999,234,256]" gridcol="4" gridrow="0" pageId="10" pageNumber="11">AW</th>
<th id="07015DDEFF8CDBE2FBB7BE93FB773828" box="[1118,1168,234,256]" gridcol="5" gridrow="0" pageId="10" pageNumber="11">BW</th>
<th id="07015DDEFF8CDBE2FAEBBE93FADE3828" box="[1282,1337,234,256]" gridcol="6" gridrow="0" pageId="10" pageNumber="11">TW</th>
</tr>
<tr id="44D034A2FF8CDBE2FF76BF6DFADE3802" box="[159,1337,276,298]" gridrow="1" pageId="10" pageNumber="11">
<th id="07015DDEFF8CDBE2FF76BF6DFF1A3802" box="[159,253,276,298]" gridcol="0" gridrow="1" pageId="10" pageNumber="11">Holotype</th>
<td id="07015DDEFF8CDBE2FE39BF6DFE1B3802" box="[464,508,276,298]" gridcol="1" gridrow="1" pageId="10" pageNumber="11">2.55</td>
<td id="07015DDEFF8CDBE2FD9DBF6DFD423802" box="[628,677,276,298]" gridcol="2" gridrow="1" pageId="10" pageNumber="11">2.64</td>
<td id="07015DDEFF8CDBE2FCFEBF6DFCAD3802" box="[791,842,276,298]" gridcol="3" gridrow="1" pageId="10" pageNumber="11">1.55</td>
<td id="07015DDEFF8CDBE2FC52BF6DFC003802" box="[955,999,276,298]" gridcol="4" gridrow="1" pageId="10" pageNumber="11">1.04</td>
<td id="07015DDEFF8CDBE2FBB7BF6DFB773802" box="[1118,1168,276,298]" gridcol="5" gridrow="1" pageId="10" pageNumber="11">1.75</td>
<td id="07015DDEFF8CDBE2FAEBBF6DFADE3802" box="[1282,1337,276,298]" gridcol="6" gridrow="1" pageId="10" pageNumber="11">3.5</td>
</tr>
<tr id="44D034A2FF8CDBE2FF76BF44FADE387B" box="[159,1337,317,339]" gridrow="2" pageId="10" pageNumber="11">
<th id="07015DDEFF8CDBE2FF76BF44FADE387B" box="[159,1337,317,339]" colspan="7" colspanRight="6" gridcol="0" gridrow="2" pageId="10" pageNumber="11">Paratypes (20)</th>
</tr>
<tr id="44D034A2FF8CDBE2FF76BF1FFADE3854" box="[159,1337,358,380]" gridrow="3" pageId="10" pageNumber="11">
<th id="07015DDEFF8CDBE2FF76BF1FFF1A3854" box="[159,253,358,380]" gridcol="0" gridrow="3" pageId="10" pageNumber="11">Min</th>
<td id="07015DDEFF8CDBE2FE39BF1FFE1B3854" box="[464,508,358,380]" gridcol="1" gridrow="3" pageId="10" pageNumber="11">1.44</td>
<td id="07015DDEFF8CDBE2FD9DBF1FFD423854" box="[628,677,358,380]" gridcol="2" gridrow="3" pageId="10" pageNumber="11">1.53</td>
<td id="07015DDEFF8CDBE2FCFEBF1FFCAD3854" box="[791,842,358,380]" gridcol="3" gridrow="3" pageId="10" pageNumber="11">0.97</td>
<td id="07015DDEFF8CDBE2FC52BF1FFC003854" box="[955,999,358,380]" gridcol="4" gridrow="3" pageId="10" pageNumber="11">0.86</td>
<td id="07015DDEFF8CDBE2FBB7BF1FFB773854" box="[1118,1168,358,380]" gridcol="5" gridrow="3" pageId="10" pageNumber="11">1.25</td>
<td id="07015DDEFF8CDBE2FAEBBF1FFADE3854" box="[1282,1337,358,380]" gridcol="6" gridrow="3" pageId="10" pageNumber="11">3.6</td>
</tr>
<tr id="44D034A2FF8CDBE2FF76BFE9FADE388E" box="[159,1337,400,422]" gridrow="4" pageId="10" pageNumber="11">
<th id="07015DDEFF8CDBE2FF76BFE9FF1A388E" box="[159,253,400,422]" gridcol="0" gridrow="4" pageId="10" pageNumber="11">Max</th>
<td id="07015DDEFF8CDBE2FE39BFE9FE1B388E" box="[464,508,400,422]" gridcol="1" gridrow="4" pageId="10" pageNumber="11">2.43</td>
<td id="07015DDEFF8CDBE2FD9DBFE9FD42388E" box="[628,677,400,422]" gridcol="2" gridrow="4" pageId="10" pageNumber="11">2.32</td>
<td id="07015DDEFF8CDBE2FCFEBFE9FCAD388E" box="[791,842,400,422]" gridcol="3" gridrow="4" pageId="10" pageNumber="11">1.42</td>
<td id="07015DDEFF8CDBE2FC52BFE9FC00388E" box="[955,999,400,422]" gridcol="4" gridrow="4" pageId="10" pageNumber="11">1.34</td>
<td id="07015DDEFF8CDBE2FBB7BFE9FB77388E" box="[1118,1168,400,422]" gridcol="5" gridrow="4" pageId="10" pageNumber="11">1.98</td>
<td id="07015DDEFF8CDBE2FAEBBFE9FADE388E" box="[1282,1337,400,422]" gridcol="6" gridrow="4" pageId="10" pageNumber="11">4.25</td>
</tr>
<tr id="44D034A2FF8CDBE2FF76BFC0FADE38E7" box="[159,1337,441,463]" gridrow="5" pageId="10" pageNumber="11">
<th id="07015DDEFF8CDBE2FF76BFC0FF1A38E7" box="[159,253,441,463]" gridcol="0" gridrow="5" pageId="10" pageNumber="11">Mean</th>
<td id="07015DDEFF8CDBE2FE39BFC0FE1B38E7" box="[464,508,441,463]" gridcol="1" gridrow="5" pageId="10" pageNumber="11">1.91</td>
<td id="07015DDEFF8CDBE2FD9DBFC0FD4238E7" box="[628,677,441,463]" gridcol="2" gridrow="5" pageId="10" pageNumber="11">1.92</td>
<td id="07015DDEFF8CDBE2FCFEBFC0FCAD38E7" box="[791,842,441,463]" gridcol="3" gridrow="5" pageId="10" pageNumber="11">1.21</td>
<td id="07015DDEFF8CDBE2FC52BFC0FC0038E7" box="[955,999,441,463]" gridcol="4" gridrow="5" pageId="10" pageNumber="11">1.09</td>
<td id="07015DDEFF8CDBE2FBB7BFC0FB7738E7" box="[1118,1168,441,463]" gridcol="5" gridrow="5" pageId="10" pageNumber="11">1.62</td>
<td id="07015DDEFF8CDBE2FAEBBFC0FADE38E7" box="[1282,1337,441,463]" gridcol="6" gridrow="5" pageId="10" pageNumber="11">3.82</td>
</tr>
<tr id="44D034A2FF8CDBE2FF76BF9BFADE38D0" box="[159,1337,482,504]" gridrow="6" pageId="10" pageNumber="11">
<th id="07015DDEFF8CDBE2FF76BF9BFF1A38D0" box="[159,253,482,504]" gridcol="0" gridrow="6" pageId="10" pageNumber="11">SD</th>
<td id="07015DDEFF8CDBE2FE39BF9BFE1B38D0" box="[464,508,482,504]" gridcol="1" gridrow="6" pageId="10" pageNumber="11">0.31</td>
<td id="07015DDEFF8CDBE2FD9DBF9BFD4238D0" box="[628,677,482,504]" gridcol="2" gridrow="6" pageId="10" pageNumber="11">0.23</td>
<td id="07015DDEFF8CDBE2FCFEBF9BFCAD38D0" box="[791,842,482,504]" gridcol="3" gridrow="6" pageId="10" pageNumber="11">0.13</td>
<td id="07015DDEFF8CDBE2FC52BF9BFC0038D0" box="[955,999,482,504]" gridcol="4" gridrow="6" pageId="10" pageNumber="11">0.13</td>
<td id="07015DDEFF8CDBE2FBB7BF9BFB7738D0" box="[1118,1168,482,504]" gridcol="5" gridrow="6" pageId="10" pageNumber="11">0.23</td>
<td id="07015DDEFF8CDBE2FAEBBF9BFADE38D0" box="[1282,1337,482,504]" gridcol="6" gridrow="6" pageId="10" pageNumber="11">0.20</td>
</tr>
<tr id="44D034A2FF8CDBE2FF76BC75FADE3B0A" box="[159,1337,524,546]" gridrow="7" pageId="10" pageNumber="11">
<th id="07015DDEFF8CDBE2FF76BC75FADE3B0A" box="[159,1337,524,546]" colspan="7" colspanRight="6" gridcol="0" gridrow="7" pageId="10" pageNumber="11">
<taxonomicName id="3DE04D63FF8C2417FF76BC74FEF33B0A" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[159,276,524,546]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF8C2417FF76BC74FF5C3B0A" box="[159,187,525,546]" italics="true" pageId="10" pageNumber="11">Eu</emphasis>
. cf.
<emphasis id="C894EAF2FF8C2417FF01BC75FEF33B0A" box="[232,276,524,546]" italics="true" pageId="10" pageNumber="11">eulo</emphasis>
</taxonomicName>
(20)
</th>
</tr>
<tr id="44D034A2FF8CDBE2FF76BC4CFADE3B63" box="[159,1337,565,587]" gridrow="8" pageId="10" pageNumber="11">
<th id="07015DDEFF8CDBE2FF76BC4CFF1A3B63" box="[159,253,565,587]" gridcol="0" gridrow="8" pageId="10" pageNumber="11">Min</th>
<td id="07015DDEFF8CDBE2FE39BC4CFE1B3B63" box="[464,508,565,587]" gridcol="1" gridrow="8" pageId="10" pageNumber="11">2.43</td>
<td id="07015DDEFF8CDBE2FD9DBC4CFD423B63" box="[628,677,565,587]" gridcol="2" gridrow="8" pageId="10" pageNumber="11">2.45</td>
<td id="07015DDEFF8CDBE2FCFEBC4CFCAD3B63" box="[791,842,565,587]" gridcol="3" gridrow="8" pageId="10" pageNumber="11">1.44</td>
<td id="07015DDEFF8CDBE2FC52BC4CFC003B63" box="[955,999,565,587]" gridcol="4" gridrow="8" pageId="10" pageNumber="11">1.44</td>
<td id="07015DDEFF8CDBE2FBB7BC4CFB773B63" box="[1118,1168,565,587]" gridcol="5" gridrow="8" pageId="10" pageNumber="11">2.10</td>
<td id="07015DDEFF8CDBE2FAEBBC4CFADE3B63" box="[1282,1337,565,587]" gridcol="6" gridrow="8" pageId="10" pageNumber="11">4.10</td>
</tr>
<tr id="44D034A2FF8CDBE2FF76BC27FADE3B5C" box="[159,1337,606,628]" gridrow="9" pageId="10" pageNumber="11">
<th id="07015DDEFF8CDBE2FF76BC27FF1A3B5C" box="[159,253,606,628]" gridcol="0" gridrow="9" pageId="10" pageNumber="11">Max</th>
<td id="07015DDEFF8CDBE2FE39BC27FE1B3B5C" box="[464,508,606,628]" gridcol="1" gridrow="9" pageId="10" pageNumber="11">3.00</td>
<td id="07015DDEFF8CDBE2FD9DBC27FD423B5C" box="[628,677,606,628]" gridcol="2" gridrow="9" pageId="10" pageNumber="11">3.01</td>
<td id="07015DDEFF8CDBE2FCFEBC27FCAD3B5C" box="[791,842,606,628]" gridcol="3" gridrow="9" pageId="10" pageNumber="11">1.86</td>
<td id="07015DDEFF8CDBE2FC52BC27FC003B5C" box="[955,999,606,628]" gridcol="4" gridrow="9" pageId="10" pageNumber="11">1.73</td>
<td id="07015DDEFF8CDBE2FBB7BC27FB773B5C" box="[1118,1168,606,628]" gridcol="5" gridrow="9" pageId="10" pageNumber="11">2.53</td>
<td id="07015DDEFF8CDBE2FAEBBC27FADE3B5C" box="[1282,1337,606,628]" gridcol="6" gridrow="9" pageId="10" pageNumber="11">4.40</td>
</tr>
<tr id="44D034A2FF8CDBE2FF76BCF1FADE3BB6" box="[159,1337,648,670]" gridrow="10" pageId="10" pageNumber="11">
<th id="07015DDEFF8CDBE2FF76BCF1FF1A3BB6" box="[159,253,648,670]" gridcol="0" gridrow="10" pageId="10" pageNumber="11">Mean</th>
<td id="07015DDEFF8CDBE2FE39BCF1FE1B3BB6" box="[464,508,648,670]" gridcol="1" gridrow="10" pageId="10" pageNumber="11">2.70</td>
<td id="07015DDEFF8CDBE2FD9DBCF1FD423BB6" box="[628,677,648,670]" gridcol="2" gridrow="10" pageId="10" pageNumber="11">2.79</td>
<td id="07015DDEFF8CDBE2FCFEBCF1FCAD3BB6" box="[791,842,648,670]" gridcol="3" gridrow="10" pageId="10" pageNumber="11">1.67</td>
<td id="07015DDEFF8CDBE2FC52BCF1FC003BB6" box="[955,999,648,670]" gridcol="4" gridrow="10" pageId="10" pageNumber="11">1.53</td>
<td id="07015DDEFF8CDBE2FBB7BCF1FB773BB6" box="[1118,1168,648,670]" gridcol="5" gridrow="10" pageId="10" pageNumber="11">2.31</td>
<td id="07015DDEFF8CDBE2FAEBBCF1FADE3BB6" box="[1282,1337,648,670]" gridcol="6" gridrow="10" pageId="10" pageNumber="11">4.24</td>
</tr>
<tr id="44D034A2FF8CDBE2FF76BCC8FADE3BEF" box="[159,1337,689,711]" gridrow="11" pageId="10" pageNumber="11">
<th id="07015DDEFF8CDBE2FF76BCC8FF1A3BEF" box="[159,253,689,711]" gridcol="0" gridrow="11" pageId="10" pageNumber="11">SD</th>
<td id="07015DDEFF8CDBE2FE39BCC8FE1B3BEF" box="[464,508,689,711]" gridcol="1" gridrow="11" pageId="10" pageNumber="11">0.18</td>
<td id="07015DDEFF8CDBE2FD9DBCC8FD423BEF" box="[628,677,689,711]" gridcol="2" gridrow="11" pageId="10" pageNumber="11">0.15</td>
<td id="07015DDEFF8CDBE2FCFEBCC8FCAD3BEF" box="[791,842,689,711]" gridcol="3" gridrow="11" pageId="10" pageNumber="11">0.11</td>
<td id="07015DDEFF8CDBE2FC52BCC8FC003BEF" box="[955,999,689,711]" gridcol="4" gridrow="11" pageId="10" pageNumber="11">0.07</td>
<td id="07015DDEFF8CDBE2FBB7BCC8FB773BEF" box="[1118,1168,689,711]" gridcol="5" gridrow="11" pageId="10" pageNumber="11">0.13</td>
<td id="07015DDEFF8CDBE2FAEBBCC8FADE3BEF" box="[1282,1337,689,711]" gridcol="6" gridrow="11" pageId="10" pageNumber="11">0.09</td>
</tr>
</table>
</paragraph>
<paragraph id="FA5F36E0FF8C2417FF2EBD56FDED3A9C" blockId="10.[151,1437,815,1920]" pageId="10" pageNumber="11">
<emphasis id="C894EAF2FF8C2417FF2EBD56FEF93A60" bold="true" box="[199,286,815,840]" pageId="10" pageNumber="11">Radula</emphasis>
(
<figureCitation id="62DB2A65FF8C2417FEC6BD56FE953A60" box="[303,370,815,840]" captionStart="FIGURE 5" captionStartId="13.[151,250,1812,1834]" captionTargetBox="[269,1322,207,1777]" captionTargetId="figure@13.[269,1322,193,1794]" captionTargetPageId="13" captionText="FIGURE 5. Radulae of Eulodrobia n. gen. A, B. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). A, half rows, B, detail of lateral tooth. C–E. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779), C, half rows, D detail of lateral tooth, E, detail of central tooth. F–H. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). F. Detail of central tooth. G. Detail of inner marginal tooth. H. Detail of outer lateral teeth. I–K. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). I. Part of tooth row. J. Detail of lateral teeth. K. Detail of inner marginal tooth. L.Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131), half rows. M. N. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133), half rows. Scale bars 10 µm." httpUri="https://zenodo.org/record/2637613/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
C–E). Central teeth with cusp formula 3+1+3, basal cusps 2+2; median cusp narrow, pointed, about 3 times longer than adjacent cusps (note: worn in
<figureCitation id="62DB2A65FF8C2417FCEEBD2DFCBB3A45" box="[775,860,852,877]" captionStart="FIGURE 5" captionStartId="13.[151,250,1812,1834]" captionTargetBox="[269,1322,207,1777]" captionTargetId="figure@13.[269,1322,193,1794]" captionTargetPageId="13" captionText="FIGURE 5. Radulae of Eulodrobia n. gen. A, B. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). A, half rows, B, detail of lateral tooth. C–E. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779), C, half rows, D detail of lateral tooth, E, detail of central tooth. F–H. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). F. Detail of central tooth. G. Detail of inner marginal tooth. H. Detail of outer lateral teeth. I–K. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). I. Part of tooth row. J. Detail of lateral teeth. K. Detail of inner marginal tooth. L.Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131), half rows. M. N. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133), half rows. Scale bars 10 µm." httpUri="https://zenodo.org/record/2637613/files/figure.png" pageId="10" pageNumber="11">Fig. 5E</figureCitation>
). Lateral teeth with cusp formula 2–3+1+2–3, main cusp narrow, blunt or pointed, about twice as long as adjacent cusps. Inner marginal teeth with 12–16 cusps. Outer marginal teeth with 17–21 cusps.
</paragraph>
<paragraph id="FA5F36E0FF8C2417FF2EBDC6FD313DA5" blockId="10.[151,1437,815,1920]" pageId="10" pageNumber="11">
<emphasis id="C894EAF2FF8C2417FF2EBDC6FDFF3AF0" bold="true" box="[199,536,959,984]" pageId="10" pageNumber="11">Female reproductive system</emphasis>
(
<figureCitation id="62DB2A65FF8C2417FDC1BDC6FD473AF0" box="[552,672,959,984]" captionStart="FIGURE 6" captionStartId="15.[151,250,1512,1534]" captionTargetBox="[181,1411,211,1442]" captionTargetId="figure@15.[150,1436,193,1490]" captionTargetPageId="15" captionText="FIGURE 6. Female reproductive system of species of Eulodrobia n. gen., shown from left side. A, B. Eulodrobia fenshami n. sp. (C. 410721). C, F. Eulodrobia eulo (Ponder &amp; Clark, 1990) (C. 410724). D, E. Eulodrobia ovata n. sp. (C. 400130). G, H. Eulodrobia bundoona n. sp. (C.400132). I, L. Eulodrobia spirula n. sp. (C.400133). J, K. Eulodrobia carinata n. sp. (C.400131). Abbreviations: ag—albumen gland; bc—bursa copulatrix; cg—capsule gland; ga—opening of oviduct; pwposterior wall of mantle cavity; ro—renal oviduct; sr—seminal receptacle; v—vestibule; vc—ventral channel. Scale bars 500 µm." httpUri="https://zenodo.org/record/2637615/files/figure.png" pageId="10" pageNumber="11">Fig. 6C, F</figureCitation>
). Ovary weakly lobed. Renal oviduct with single (
<figureCitation id="62DB2A65FF8C2417FB35BDC6FAC73AF0" box="[1244,1312,959,984]" captionStart="FIGURE 4" captionStartId="12.[151,250,1521,1543]" captionTargetBox="[196,1406,256,1477]" captionTargetId="figure@12.[151,1435,229,1500]" captionTargetPageId="12" captionText="FIGURE 4. Opercula of species of Eulodrobia n. gen., external and internal views of each species. A, B. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). C, D. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). E, F. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). G, H. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). I, J. Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131). K, L. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133). Scale bars 500 µm." httpUri="https://zenodo.org/record/2637611/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
) or double (
<bibRefCitation id="9E714B11FF8C2417FF77BD9DFE1B3AD4" author="Ponder, W. F. &amp; Clark, G. A." box="[158,508,996,1021]" pageId="10" pageNumber="11" pagination="301 - 363" refId="ref35763" refString="Ponder, W. F. &amp; Clark, G. A. (1990) A radiation of hydrobiid snails in threatened artesian springs in Western Queensland. Records of the Australian Museum, 42, 301 - 363. https: // doi. org / 10.3853 / j. 0067 - 1975.42.1990.119" type="journal article" year="1990">Ponder &amp; Clark 1990, fig. 34E</bibRefCitation>
) U-shaped arch. Seminal receptacle oval, duct very short. Bursa copulatrix behind albumen gland, pyriform to oval, shorter than albumen gland, bursal duct enters bursa anteroventrally, bursal duct joins coiled oviduct well behind posterior mantle cavity wall. Albumen gland partly in mantle cavity. Capsule gland with two distinct glandular zones apparent in some, medium thickness in cross section, slightly indented by rectum. Anterior vestibule small, opening terminal.
</paragraph>
<paragraph id="FA5F36E0FF8C2417FF2EBAEEFA723DDF" blockId="10.[151,1437,815,1920]" pageId="10" pageNumber="11">
<emphasis id="C894EAF2FF8C2417FF2EBAEEFE1A3D98" bold="true" box="[199,509,1175,1200]" pageId="10" pageNumber="11">Male reproductive system</emphasis>
(
<figureCitation id="62DB2A65FF8C2417FDE5BAEEFD883D87" box="[524,623,1175,1200]" captionStart="FIGURE 7" captionStartId="16.[151,250,1324,1346]" captionTargetBox="[170,1416,747,1275]" captionTargetId="figure@16.[150,1436,735,1299]" captionTargetPageId="16" captionText="FIGURE 7. Prostate and vas deferens of species of Eulodrobia n. gen. Each species is shown with anterior at left and posterior at right, with line indicating position of posterior pallial wall. A. Eulodrobia fenshami n. sp. (C. 410721). B. Eulodrobia eulo (Ponder &amp; Clark, 1990) (C. 410724). C. Eulodrobia ovata n. sp. (C.400130). D. Eulodrobia bundoona n. sp. (C.400132). E, Eulodrobia carinata n. sp. (C. 400131). F. Eulodrobia spirula n. sp. (C.400133). Abbreviations: avd—anterior vas deferens; pw—position of posterior wall of pallial cavity; ve—posterior vas deferens. Scale bars 200 µm." httpUri="https://zenodo.org/record/2637617/files/figure.png" pageId="10" pageNumber="11">Figs. 7B</figureCitation>
,
<figureCitation id="62DB2A65FF8C2417FD92BAE1FD783D98" box="[635,671,1176,1200]" captionStart="FIGURE 8" captionStartId="17.[151,250,1042,1064]" captionTargetBox="[170,1435,206,1008]" captionTargetId="figure@17.[151,1435,195,1020]" captionTargetPageId="17" captionText="FIGURE 8. Penis and vas deferens of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp. (C. 410721). B. Eulodrobia eulo (Ponder &amp; Clark, 1990) (C. 410724). C. Eulodrobia ovata n. sp. (C. 400130). D. Eulodrobia bundoona n. sp. (C.400132). E, Eulodrobia carinata n. sp. (C. 400131). F. Eulodrobia spirula n. sp. (C.400133). Abbreviations: ngl—non-glandular lobe; pd—penial duct; vd—vas deferens. Scale bars 200 µm." httpUri="https://zenodo.org/record/2637619/files/figure.png" pageId="10" pageNumber="11">8B</figureCitation>
). Prostate gland less than half in mantle roof, bean-shaped, medium in cross section. Posterior pallial vas deferens slightly undulating to (usually) coiled, anteriorly slightly undulating. Penis behind right eye well down neck, distal end broad, terminal papilla absent, two non-glandular lobes present.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF8C2417FF2EBB7DFB3B3CF8" pageId="10" pageNumber="11" type="distribution">
<paragraph id="FA5F36E0FF8C2417FF2EBB7DFB3B3CF8" blockId="10.[151,1437,815,1920]" pageId="10" pageNumber="11">
<emphasis id="C894EAF2FF8C2417FF2EBB7DFE1E3C35" bold="true" box="[199,505,1284,1309]" pageId="10" pageNumber="11">Distribution and habitat.</emphasis>
Restricted to Massey Spring, Granites Spring group, a small spring on Granite Springs Station, Eulo Supergroup (
<collectingCountry id="82F77670FF8C2417FDCCBB5EFC8D3C68" box="[549,874,1319,1344]" name="Australia" pageId="10" pageNumber="11">
Commonwealth of
<bibRefCitation id="9E714B11FF8C2417FCEABB5EFC8D3C68" author="Commonwealth of Australia" box="[771,874,1319,1344]" pageId="10" pageNumber="11" refId="ref33702" refString="Commonwealth of Australia (2014) Ecological and hydrogeological survey of the Great Artesian Basin springs-Springsure, Eulo, Bourke and Bogan River Supergroups. Fol. 2. Hydrogeological profiles, Knowledge report, prepared by UniQuest for the Department of the Environment, Commonwealth of Australia, Canberra. Department of the Environment, Public Affairs, Canberra, xiii + 307 pp." type="book" year="2014">Australia</bibRefCitation>
</collectingCountry>
2014) (
<figureCitation id="62DB2A65FF8C2417FC2FBB5EFBEE3C68" box="[966,1033,1319,1344]" captionStart="FIGURE 1" captionStartId="4.[151,250,1844,1866]" captionTargetBox="[151,1435,498,1802]" captionTargetId="figure@4.[151,1435,498,1802]" captionTargetPageId="4" captionText="FIGURE 1. Map of the spring supergroups associated with the Great Artesian Basin (GAB) in Queensland, showing the main non-town locations mentioned in the text (squares) and some of the towns in the area (ovals). The pale grey shading is the discharge part of the GAB, the dark grey the recharge part. The inset map of Australia shows the area illustrated. The approximate location of Conondale is indicated by a triangle and Nundle by a star. The spring supergroups are outlined with dotted lines and are as follows: A, Mulligan River; B, Springvale; C, Flinders River; D, Hughenden; E, Barcaldine (see inset at upper right for details); F, Springsure; G, Eulo." httpUri="https://zenodo.org/record/2637605/files/figure.png" pageId="10" pageNumber="11">Fig. 1</figureCitation>
). This species was absent in a few other nearby small springs which occur in the Granites Spring group. A similar taxon occurs in Tunga Spring (see Remarks). Massey Spring and Tunga Spring are about
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apart and on two different local drainage systems, although both are tributaries of the Paroo River system. These populations can be formally distinguished by referring to them as
<taxonomicName id="3DE04D63FF8C2417FE93BBCEFDC83CF8" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[378,559,1463,1488]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF8C2417FE93BBCEFDC83CF8" box="[378,559,1463,1488]" italics="true" pageId="10" pageNumber="11">Eulodrobia eulo</emphasis>
</taxonomicName>
(from Massey Spring) and
<taxonomicName id="3DE04D63FF8C2417FC8DBBC0FC023CF8" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[868,997,1463,1488]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF8C2417FC8DBBC0FC643CF8" box="[868,899,1465,1488]" italics="true" pageId="10" pageNumber="11">Eu</emphasis>
. cf.
<emphasis id="C894EAF2FF8C2417FC5CBBCEFC023CF8" box="[949,997,1463,1488]" italics="true" pageId="10" pageNumber="11">eulo</emphasis>
</taxonomicName>
(from Tunga Spring).
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF8C2417FF2EBBA5FF393EA8" pageId="10" pageNumber="11" type="discussion">
<paragraph id="FA5F36E0FF8C2417FF2EBBA5FD743FE5" blockId="10.[151,1437,815,1920]" pageId="10" pageNumber="11">
<emphasis id="C894EAF2FF8C2417FF2EBBA5FEDC3CDD" bold="true" box="[199,315,1500,1525]" pageId="10" pageNumber="11">Remarks.</emphasis>
This species was originally recorded from both Massey Spring (= Granite or Rocky Springs) and Tunga Spring (at Tunga Bore). Molecular data for both populations (see
<figureCitation id="62DB2A65FF8C2417FC16BB86FBA93F30" box="[1023,1102,1535,1560]" captionStart="FIGURE 31" captionStartId="61.[151,250,1731,1753]" captionTargetBox="[227,1355,208,1693]" captionTargetId="figure@61.[196,1395,194,1717]" captionTargetPageId="61" captionText="FIGURE 31. Phylogram of Australian tateid spring taxa and selected outgroups with Bayesian posterior probability values (shown above nodes) and RAxML bootstrap support values (shown below nodes). Posterior probability values below 0.85 and bootstrap support below 70% are not shown, and asterisks indicate optimal support. Two parallel diagonal lines on branches indicate those branches have been shortened. Bayesian tree is shown. The grey box indicates the clade encompassing all Queensland spring taxa, the monotypic Nundalia from New South Wales, and the coastal, riverine genus Jardinella from northern Queensland. All taxa included in this clade are illustrated to the right and where possible aligned with their respective placement in the phylogram. Abbreviations: C—Carnarvoncochlea; E. (B.)—Edgbastonia (Barcaldinia); E. (E.) -Edgbastonia (Edgbastonia); NQ—North Queensland (coastal); NSW—New South Wales; S—Springvalia. Names of imaged taxa are indicated, next to the illustrations, by the three first letters in their species (or subspecies) name, or, when identical, the four first letters. For details of superscript numbers 1–40, see Table 20." httpUri="https://zenodo.org/record/2637669/files/figure.png" pageId="10" pageNumber="11">Fig 31</figureCitation>
) shows some differentiation suggesting that separate species-group status may eventually be necessary. Although the Tunga Spring population has not been examined anatomically, its shell, opercular and radular characters are very similar to those of specimens from Massey Spring. Also, the operculum in both populations has the same raised nipple-like thickening at the nucleus. However, a discriminant function analysis separated most specimens of the two forms, as detailed below in the Remarks under the next species.
</paragraph>
<paragraph id="FA5F36E0FF8C2417FF2EB8AEFF393EA8" blockId="10.[151,1437,815,1920]" pageId="10" pageNumber="11">
The spring at the
<typeStatus id="255B8842FF8C2417FE67B8A1FE593FD8" box="[398,446,1752,1776]" pageId="10" pageNumber="11">type</typeStatus>
locality (Massey Spring) is a medium-sized spring (
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Commonwealth of
<bibRefCitation id="9E714B11FF8C2417FB05B8AEFAB53FD8" author="Commonwealth of Australia" box="[1260,1362,1751,1776]" pageId="10" pageNumber="11" refId="ref33702" refString="Commonwealth of Australia (2014) Ecological and hydrogeological survey of the Great Artesian Basin springs-Springsure, Eulo, Bourke and Bogan River Supergroups. Fol. 2. Hydrogeological profiles, Knowledge report, prepared by UniQuest for the Department of the Environment, Commonwealth of Australia, Canberra. Department of the Environment, Public Affairs, Canberra, xiii + 307 pp." type="book" year="2014">Australia</bibRefCitation>
</collectingCountry>
2014) about
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SW of the village of Eulo while the other known population lives in a tiny spring next to Tunga Bore, which is located near Mt Tunga, about
<quantity id="3D189B05FF8C2417FDBFB966FD793E10" box="[598,670,1823,1848]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="11" unit="km" value="45.0">45 km</quantity>
WSW of Eulo, south-western
<collectingRegion id="3824F802FF8C2417FC14B966FB623E10" box="[1021,1157,1823,1848]" country="Australia" name="Queensland" pageId="10" pageNumber="11">Queensland</collectingRegion>
. Other springs near both these springs have substantially decreased in flow (
<collectingCountry id="82F77670FF8C2417FD05B93DFBD23E75" box="[748,1077,1860,1885]" name="Australia" pageId="10" pageNumber="11">
Commonwealth of
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</collectingCountry>
2014), and none contain these snails.
</paragraph>
</subSubSection>
</treatment>
</document>
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