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<emphasis id="C8D8EAA8317BD91944B26973FE121E06" bold="true" box="[151,398,1053,1080]" italics="true" pageId="6" pageNumber="407">Hypoxys quadridens</emphasis>
(Fabricius, 1803)
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(
<figureCitation id="62972A3F317BD91944BA692FFF711E65" box="[159,237,1089,1115]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="6" pageNumber="407">Figs. 1</figureCitation>
A–G; 18 A–B; 21)
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<paragraph id="FA1336BA317BD91944B269E8FED01EA0" blockId="6.[151,1437,1157,1642]" box="[151,332,1157,1182]" pageId="6" pageNumber="407">-- Stoll, Pun.: 135</paragraph>
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<paragraph id="FA1336BA317BD91944B269CAFE381EC4" blockId="6.[151,1437,1157,1642]" pageId="6" pageNumber="407">
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<taxonomicName id="3DAC4D39317BD91944B269CAFDA71E82" authority="Fabricius, 1803: 148" authorityName="Fabricius" authorityPageNumber="148" authorityYear="1803" box="[151,571,1187,1212]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="C8D8EAA8317BD91944B269CAFEC61E82" box="[151,346,1188,1212]" italics="true" pageId="6" pageNumber="407">Edessa quadridens</emphasis>
Fabricius, 1803: 148
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; 
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<bibRefCitation id="9E3D4B4B317BD919466D69CDFC9F1E82" author="Dallas, W. S." box="[584,771,1187,1212]" pageId="6" pageNumber="407" refId="ref24638" refString="Dallas, W. S. (1851) List of the specimens of hemipterous insects in the collection of the British Museum. British Museum, London, 390 pp., 15 pls." type="book" year="1851">Dallas, 1851: 329</bibRefCitation>
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; Stål, 1868: 36; Walker, 1868: 425; 
<treatmentCitation id="7B0D10AB317BD91940A569CDFA891E82" author="Stal, C." box="[1152,1301,1187,1212]" page="55" pageId="6" pageNumber="407" year="1872">
<bibRefCitation id="9E3D4B4B317BD91940A569CDFA891E82" author="Stal, C." box="[1152,1301,1187,1212]" pageId="6" pageNumber="407" pagination="1 - 159" refId="ref25463" refString="Stal, C. (1872) Enumeratio Hemipterorum. Bidrag till em forteckning ofver alla hittils kanda Hemiptera, Jemte systematiska meddelanden. 2. Kongliga Svenska Vetenskaps-Akademien Handlingar, 10 (4), 1 - 159. https: // doi. org / 10.5962 / bhl. title. 12549" type="journal article" year="1872">Stål, 1872: 55</bibRefCitation>
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; Lethierry &amp; Severin, 1893: 193 (catalog); 
<treatmentCitation id="7B0D10AB317BD91945D169ACFD5A1EE5" author="Breddin, G." box="[500,710,1218,1243]" page="140" pageId="6" pageNumber="407" year="1904">
<bibRefCitation id="9E3D4B4B317BD91945D169ACFD5A1EE5" author="Breddin, G." box="[500,710,1218,1243]" pageId="6" pageNumber="407" pagination="135 - 153" refId="ref24439" refString="Breddin, G. (1904 a) Beitrage zur Systematik der Rhynchoten. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1904, 135 - 153." type="journal article" year="1904">Breddin, 1904a: 140</bibRefCitation>
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; Osborn, 1904: 197; 
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<bibRefCitation id="9E3D4B4B317BD91947BF69ACFBFC1EE5" author="Breddin, G." box="[922,1120,1218,1243]" pageId="6" pageNumber="407" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907: 327</bibRefCitation>
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; 
<treatmentCitation id="7B0D10AB317BD919404E69ACFAA91EE5" author="Kirkaldy, G. W." box="[1131,1333,1218,1243]" page="163" pageId="6" pageNumber="407" year="1909">
<bibRefCitation id="9E3D4B4B317BD919404E69ACFAA91EE5" author="Kirkaldy, G. W." box="[1131,1333,1218,1243]" pageId="6" pageNumber="407" refId="ref25016" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera). Vol. 1. Cimicidae. Felix L. Dames ed., Berlin, XL + 392 pp." type="book" year="1909">Kirkaldy, 1909: 163</bibRefCitation>
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(catalog); Froeschner, 1981: 66.
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<paragraph id="FA1336BA317BD91944B2686EFEBD1F09" blockId="6.[151,1437,1157,1642]" pageId="6" pageNumber="407">
<treatmentCitationGroup id="DABC1194317BD91944B2686EFEBD1F09" pageId="6" pageNumber="407">
<taxonomicName id="3DAC4D39317BD91944B2686EFDAD1F26" authority="Fabricius, 1803: 148" authorityName="Fabricius" authorityPageNumber="148" authorityYear="1803" box="[151,561,1279,1304]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="brunnipes">
<emphasis id="C8D8EAA8317BD91944B2686EFED11F26" box="[151,333,1280,1304]" italics="true" pageId="6" pageNumber="407">Edessa brunnipes</emphasis>
Fabricius, 1803: 148
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; Walker, 1868: 427; Lethierry &amp; Severin, 1893: 193 (catalog); 
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<bibRefCitation id="9E3D4B4B317BD91940EE6991FA001F26" author="Kirkaldy, G. W." box="[1227,1436,1279,1304]" pageId="6" pageNumber="407" refId="ref25016" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera). Vol. 1. Cimicidae. Felix L. Dames ed., Berlin, XL + 392 pp." type="book" year="1909">Kirkaldy, 1909: 163</bibRefCitation>
</treatmentCitation>
(catalog)
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</paragraph>
<paragraph id="FA1336BA317BD91944B26850FF6C1F4A" blockId="6.[151,1437,1157,1642]" pageId="6" pageNumber="407">
<treatmentCitationGroup id="DABC1194317BD91944B26850FF6C1F4A" pageId="6" pageNumber="407">
<taxonomicName id="3DAC4D39317BD91944B26850FD541F68" authority="Lepeletier &amp; Serville, 1825: 55" authorityName="Lepeletier &amp; Serville" authorityPageNumber="55" authorityYear="1825" box="[151,712,1341,1366]" class="Insecta" family="Acanthosomatidae" genus="Pentatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="luteicornis">
<emphasis id="C8D8EAA8317BD91944B26850FEE51F6B" box="[151,377,1341,1365]" italics="true" pageId="6" pageNumber="407">Pentatoma luteicornis</emphasis>
Lepeletier &amp; Serville, 1825: 55
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; Lethierry &amp; Severin, 1893: 193 (catalog); 
<treatmentCitation id="7B0D10AB317BD91940A96853FAC61F68" author="Kirkaldy, G. W." box="[1164,1370,1341,1366]" page="163" pageId="6" pageNumber="407" year="1909">
<bibRefCitation id="9E3D4B4B317BD91940A96853FAC61F68" author="Kirkaldy, G. W." box="[1164,1370,1341,1366]" pageId="6" pageNumber="407" refId="ref25016" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera). Vol. 1. Cimicidae. Felix L. Dames ed., Berlin, XL + 392 pp." type="book" year="1909">Kirkaldy, 1909: 163</bibRefCitation>
</treatmentCitation>
(catalog)
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</paragraph>
<paragraph id="FA1336BA317BD91944B26815FB421FAD" blockId="6.[151,1437,1157,1642]" box="[151,1246,1402,1427]" pageId="6" pageNumber="407">
<treatmentCitationGroup id="DABC1194317BD91944B26815FB421FAD" box="[151,1246,1402,1427]" pageId="6" pageNumber="407">
<taxonomicName id="3DAC4D39317BD91944B26815FE601FAD" authority="Hahn, 1831: 127" authorityName="Hahn" authorityPageNumber="127" authorityYear="1831" box="[151,508,1402,1427]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="schaefferi">
<emphasis id="C8D8EAA8317BD91944B26815FED51FAD" box="[151,329,1403,1427]" italics="true" pageId="6" pageNumber="407">Edessa schaefferi</emphasis>
Hahn, 1831: 127
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; Lethierry &amp; Severin, 1893: 193 (catalog); 
<treatmentCitation id="7B0D10AB317BD91947966814FBE21FAD" author="Kirkaldy, G. W." box="[947,1150,1402,1427]" page="163" pageId="6" pageNumber="407" year="1909">
<bibRefCitation id="9E3D4B4B317BD91947966814FBE21FAD" author="Kirkaldy, G. W." box="[947,1150,1402,1427]" pageId="6" pageNumber="407" refId="ref25016" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera). Vol. 1. Cimicidae. Felix L. Dames ed., Berlin, XL + 392 pp." type="book" year="1909">Kirkaldy, 1909: 163</bibRefCitation>
</treatmentCitation>
(catalog)
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</paragraph>
<paragraph id="FA1336BA317BD91944B268F4FD061F8C" blockId="6.[151,1437,1157,1642]" box="[151,666,1433,1458]" pageId="6" pageNumber="407">
<treatmentCitationGroup id="DABC1194317BD91944B268F4FD061F8C" box="[151,666,1433,1458]" pageId="6" pageNumber="407">
<treatmentCitation id="7B0D10AB317BD91944B268F4FD061F8C" author="Amyot, C. J. B. &amp; Serville, A." box="[151,666,1433,1458]" page="157" pageId="6" pageNumber="407" year="1843">
<taxonomicName id="3DAC4D39317BD91944B268F4FD061F8C" authority=", Amyot &amp; Serville, 1843: 157" authorityName=", Amyot &amp; Serville" authorityYear="1843" box="[151,666,1433,1458]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="C8D8EAA8317BD91944B268F4FEFB1F8F" box="[151,359,1433,1457]" italics="true" pageId="6" pageNumber="407">Hypoxys quadridens</emphasis>
, 
<bibRefCitation id="9E3D4B4B317BD919455768F7FD061F8C" author="Amyot, C. J. B. &amp; Serville, A." box="[370,666,1433,1458]" pageId="6" pageNumber="407" refId="ref24336" refString="Amyot, C. J. B. &amp; Serville, A. (1843) Historie naturelle des insects. Hemipteres. Cuyrage accompagne de planches. Librairie encyclopedique de Roret, Paris, lxxvi + 675 pp., 12 pls." type="book" year="1843">Amyot &amp; Serville, 1843: 157</bibRefCitation>
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<paragraph id="FA1336BA317BD91944B268D6FC581FEE" blockId="6.[151,1437,1157,1642]" box="[151,964,1463,1488]" pageId="6" pageNumber="407">
<treatmentCitationGroup id="DABC1194317BD91944B268D6FC581FEE" box="[151,964,1463,1488]" pageId="6" pageNumber="407">
<treatmentCitation id="7B0D10AB317BD91944B268D6FE6B1FEE" author="Breddin, G." box="[151,503,1463,1488]" page="327" pageId="6" pageNumber="407" year="1907">
<taxonomicName id="3DAC4D39317BD91944B268D6FE6B1FEE" authority="Breddin, 1907: 327" authorityName="Breddin" authorityPageNumber="327" authorityYear="1907" box="[151,503,1463,1488]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="jugalis">
<emphasis id="C8D8EAA8317BD91944B268D6FEB61FEE" box="[151,298,1464,1488]" italics="true" pageId="6" pageNumber="407">Edessa jugalis</emphasis>
<bibRefCitation id="9E3D4B4B317BD919451468D9FE6B1FEE" author="Breddin, G." box="[305,503,1463,1488]" pageId="6" pageNumber="407" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907: 327</bibRefCitation>
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, 329; 
<treatmentCitation id="7B0D10AB317BD919461068D9FC9C1FEE" author="Kirkaldy, G. W." box="[565,768,1463,1488]" page="368" pageId="6" pageNumber="407" year="1909">
<bibRefCitation id="9E3D4B4B317BD919461068D9FC9C1FEE" author="Kirkaldy, G. W." box="[565,768,1463,1488]" pageId="6" pageNumber="407" refId="ref25016" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera). Vol. 1. Cimicidae. Felix L. Dames ed., Berlin, XL + 392 pp." type="book" year="1909">Kirkaldy, 1909: 368</bibRefCitation>
</treatmentCitation>
(catalog) 
<emphasis id="C8D8EAA8317BD919474268D6FC581FEE" bold="true" box="[871,964,1464,1488]" pageId="6" pageNumber="407">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="FA1336BA317BD91944B268B9FAA41FD1" blockId="6.[151,1437,1157,1642]" box="[151,1336,1494,1519]" pageId="6" pageNumber="407">
<materialsCitation id="4AC43CE7317BD91944B268B9FAA41FD1" ID-GBIF-Occurrence="2570056118" box="[151,1336,1494,1519]" collectingDate="86-10-20" collectionCode="ZMUC" pageId="6" pageNumber="407" specimenCount="1" typeStatus="lectotype">
<typeStatus id="25178818317BD91944B268B9FF601FD1" box="[151,252,1495,1519]" pageId="6" pageNumber="407" type="lectotype">Lectotype</typeStatus>
male 
<taxonomicName id="3DAC4D39317BD919451F68B9FE661FD1" authorityName="Fabricius" authorityYear="1803" box="[314,506,1495,1519]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="C8D8EAA8317BD919451F68B9FE661FD1" box="[314,506,1495,1519]" italics="true" pageId="6" pageNumber="407">Edessa quadridens</emphasis>
</taxonomicName>
. America Meridionalis, ZMUC 00 
<date id="8E12107A317BD919474468B8FC311FD1" box="[865,941,1494,1519]" pageId="6" pageNumber="407" value="86-10-20">
<collectingDate id="9E56E992317BD919474468B8FC311FD1" box="[865,941,1494,1519]" pageId="6" pageNumber="407" value="86-10-20">102086</collectingDate>
</date>
(
<collectionCode id="9CBDAE7F317BD919479F68B9FB951FD1" box="[954,1033,1495,1519]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="6" pageNumber="407">ZMUC</collectionCode>
). Examined. Here designated.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B2B66531317BD91944B26898FBD11C54" pageId="6" pageNumber="407" type="description">
<paragraph id="FA1336BA317BD91944B26898FDFF1C75" blockId="6.[151,1437,1157,1642]" pageId="6" pageNumber="407">
<materialsCitation id="4AC43CE7317BD91944B26898FDFF1C75" ID-GBIF-Occurrence="2570056063" collectingDate="86-10-20" collectionCode="SDEI" country="Suriname" location="Museum of Strasburg" municipality="We" pageId="6" pageNumber="407" specimenCount="2" typeStatus="lectotype">
<typeStatus id="25178818317BD91944B26898FF601C30" box="[151,252,1526,1550]" pageId="6" pageNumber="407" type="lectotype">Lectotype</typeStatus>
male 
<taxonomicName id="3DAC4D39317BD919451D689BFE561C33" authorityName="Breddin" authorityYear="1907" box="[312,458,1525,1549]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="jugalis">
<emphasis id="C8D8EAA8317BD919451D689BFE561C33" box="[312,458,1525,1549]" italics="true" pageId="6" pageNumber="407">Edessa jugalis</emphasis>
</taxonomicName>
. 
<collectingCountry id="82BB762A317BD91945F0689BFDAA1C30" box="[469,566,1525,1550]" name="Suriname" pageId="6" pageNumber="407">Suriname</collectingCountry>
(
<collectionCode id="9CBDAE7F317BD9194666689BFD1D1C33" box="[579,641,1525,1549]" pageId="6" pageNumber="407">SDEI</collectionCode>
). Designated by 
<bibRefCitation id="9E3D4B4B317BD9194703689BFC7A1C33" author="Gaedike, H." box="[806,998,1525,1550]" pageId="6" pageNumber="407" pagination="79 - 159" refId="ref24984" refString="Gaedike, H. (1971) Katalog der in Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen- V. Heteroptera. Beitrage zur Entomologie, 21, 79 - 159." type="journal article" year="1971">Gaedike (1971:89)</bibRefCitation>
. Examined. Gaedike also indicated 
<typeStatus id="25178818317BD919416E689BFE8C1C12" pageId="6" pageNumber="407" type="paralectotype">paralectotypes</typeStatus>
in SDEI and 
<location id="FF736061317BD91945B36B7AFDEE1C12" LSID="urn:lsid:plazi:treatment:720587AC317BD91844B26973FE5A1E65:FF736061317BD91945B36B7AFDEE1C12" box="[406,626,1555,1580]" country="Suriname" municipality="We" name="Museum of Strasburg" pageId="6" pageNumber="407">Museum of Strasburg</location>
, but did not mention quantity or sex of them. 
<collectingMunicipality id="1A77ACC0317BD91940126B7AFBC41C15" box="[1079,1112,1556,1579]" pageId="6" pageNumber="407">We</collectingMunicipality>
have examined only one female with the same data as lectotype in SDEI.
</materialsCitation>
</paragraph>
<paragraph id="FA1336BA317BD91944B26B3CFBD11C54" blockId="6.[151,1437,1157,1642]" box="[151,1101,1617,1642]" pageId="6" pageNumber="407">
<materialsCitation id="4AC43CE7317BD91944B26B3CFBD11C54" ID-GBIF-Occurrence="2570056086" box="[151,1101,1617,1642]" collectionCode="ZMUC" pageId="6" pageNumber="407" specimenCount="1" typeStatus="lectotype">
<typeStatus id="25178818317BD91944B26B3CFF601C54" box="[151,252,1618,1642]" pageId="6" pageNumber="407" type="lectotype">Lectotype</typeStatus>
female 
<emphasis id="C8D8EAA8317BD919456B6B3FFD951C57" box="[334,521,1617,1641]" italics="true" pageId="6" pageNumber="407">
<taxonomicName id="3DAC4D39317BD919456B6B3FFD991C57" authorityName="Fabricius" authorityYear="1803" box="[334,517,1617,1641]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="brunnipes">Edessa brunnipes</taxonomicName>
.
</emphasis>
America Meridionalis, Smidt (
<collectionCode id="9CBDAE7F317BD91947676B3FFC0D1C57" box="[834,913,1617,1641]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="6" pageNumber="407">ZMUC</collectionCode>
). Here designated.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B2B66531317BD91944B26BF9FDED1DC8" pageId="6" pageNumber="407" type="materials_examined">
<paragraph id="FA1336BA317BD91944B26BF9FDED1DC8" blockId="6.[151,1437,1687,2038]" pageId="6" pageNumber="407">
<emphasis id="C8D8EAA8317BD91944B26BF9FD221C8C" bold="true" box="[151,702,1687,1714]" pageId="6" pageNumber="407">
Material examined. 
<collectingCountry id="82BB762A317BD91945AF6BF9FDA61C8C" box="[394,570,1687,1714]" name="Venezuela" pageId="6" pageNumber="407">VENEZUELA</collectingCountry>
. 
<collectingRegion id="3868F858317BD919466D6BF6FD251C8C" box="[584,697,1688,1714]" country="Venezuela" name="Monagas" pageId="6" pageNumber="407">Monagas</collectingRegion>
:
</emphasis>
<specimenCount id="ECAAFD33317BD91946E26BF6FD401C8C" box="[711,732,1688,1714]" pageId="6" pageNumber="407" type="female">♀</specimenCount>
La Esperanza, 
<date id="8E12107A317BD91947A86BF9FB891C8C" box="[909,1045,1687,1714]" pageId="6" pageNumber="407" value="1965-10-25">25–X–1965</date>
, Inst. Zool., Agricola—Fac. Agro- nomia Univ. Central, F. Fernande. Y. C. J. Rosales (
<collectionCode id="9CBDAE7F317BD91946C26BD2FCB11CE8" box="[743,813,1724,1750]" pageId="6" pageNumber="407">IZAV</collectionCode>
); 
<specimenCount id="ECAAFD33317BD91947646BD2FCCA1CE8" box="[833,854,1724,1750]" pageId="6" pageNumber="407" type="male">♂</specimenCount>
La Pica, 3–X–65, Inst. Zool., Agricola–Fac. Agro- nomia Univ. Central, C. J. Rosales, F. Fernandes Y. 
<taxonomicName id="3DAC4D39317BD91946C06B8EFBB01CC4" authority="F. Comp." authorityName="F. Comp." box="[741,1068,1760,1786]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="C8D8EAA8317BD91946C06B8EFC271CC4" box="[741,955,1760,1786]" italics="true" pageId="6" pageNumber="407">Edessa quadridens</emphasis>
F. Comp.
</taxonomicName>
w. 
<typeStatus id="25178818317BD91940716B8EFB061CC4" box="[1108,1178,1760,1786]" pageId="6" pageNumber="407">TYPE</typeStatus>
Fernandes, JAM 2011 (
<collectionCode id="9CBDAE7F317BD91944BA6A6AFF791D20" box="[159,229,1796,1822]" pageId="6" pageNumber="407">IZAV</collectionCode>
); 
<emphasis id="C8D8EAA8317BD91944DF6A6DFEC01D20" bold="true" box="[250,348,1795,1822]" pageId="6" pageNumber="407">
<collectingRegion id="3868F858317BD91944DF6A6DFECB1D20" box="[250,343,1795,1822]" country="Venezuela" name="Bolivar" pageId="6" pageNumber="407">Bolivar</collectingRegion>
:
</emphasis>
<specimenCount id="ECAAFD33317BD91945406A6AFEE61D20" box="[357,378,1796,1822]" pageId="6" pageNumber="407" type="female">♀</specimenCount>
km 107, El Dorado 
<collectingCountry id="82BB762A317BD919464D6A6AFD6E1D20" box="[616,754,1796,1822]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="6" pageNumber="407">Santa Elena</collectingCountry>
, 
<specimenCount id="ECAAFD33317BD91946DB6A6DFCD21D20" box="[766,846,1795,1822]" pageId="6" pageNumber="407" type="generic">
<quantity id="3D549B5F317BD91946DB6A6DFCD61D20" box="[766,842,1795,1822]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="6" pageNumber="407" unit="m" value="520.0">520 m</quantity>
.
</specimenCount>
18–VIII–57, Inst. Zool, Agricola—Fac. Agronomia Univ. Central. F. Fernandez Y. C.J. Rosales (
<collectionCode id="9CBDAE7F317BD91946AF6A46FD4C1D7C" box="[650,720,1832,1858]" pageId="6" pageNumber="407">IZAV</collectionCode>
); 
<emphasis id="C8D8EAA8317BD91946C66A49FC451D7C" bold="true" box="[739,985,1831,1858]" pageId="6" pageNumber="407">
<collectingCountry id="82BB762A317BD91946C66A49FC481D7C" box="[739,980,1831,1858]" name="French Guiana" pageId="6" pageNumber="407">FRENCH GUIANA</collectingCountry>
.
</emphasis>
<specimenCount id="ECAAFD33317BD91947C56A46FC691D7C" box="[992,1013,1832,1858]" pageId="6" pageNumber="407" type="male">♂</specimenCount>
Otovalo, 
<collectingCountry id="82BB762A317BD91940406A46FA901D7C" box="[1125,1292,1832,1858]" name="Guyana" pageId="6" pageNumber="407">British Guiana</collectingCountry>
, 
<date id="8E12107A317BD91941326A46FA0B1D7C" box="[1303,1431,1832,1858]" pageId="6" pageNumber="407" value="1927-01-21">1–21–1927</date>
, J. Murray (
<collectionCode id="9CBDAE7F317BD91945376A22FEF51D58" box="[274,361,1868,1894]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="407" type="Museum">USNM</collectionCode>
); 
<emphasis id="C8D8EAA8317BD91945596A25FD801D58" bold="true" box="[380,540,1867,1894]" pageId="6" pageNumber="407">
<collectingCountry id="82BB762A317BD91945596A25FD8B1D58" box="[380,535,1867,1894]" name="Suriname" pageId="6" pageNumber="407">SURINAME</collectingCountry>
.
</emphasis>
<specimenCount id="ECAAFD33317BD91946076A22FDAB1D58" box="[546,567,1868,1894]" pageId="6" pageNumber="407" type="male">♂</specimenCount>
Tibiti 
<date id="8E12107A317BD91946A46A25FD731D5B" box="[641,751,1867,1893]" pageId="6" pageNumber="407" value="1949-01-06">6–I–1949</date>
, In. bostussenpaal 2–3, Museum Leiden 
<collectingCountry id="82BB762A317BD91940906A22FABC1D58" box="[1205,1312,1868,1894]" name="Suriname" pageId="6" pageNumber="407">Suriname</collectingCountry>
exp 
<date id="8E12107A317BD91941736A22FF4E1DB4" pageId="6" pageNumber="407" value="1948" valueMax="1949">1948– 1949</date>
, 
<taxonomicName id="3DAC4D39317BD91944FA6A1EFD1D1DB4" authority="Breddin, 1907 Comp." authorityName="Breddin" authorityYear="1907" box="[223,641,1904,1930]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="jugalis">
<emphasis id="C8D8EAA8317BD91944FA6A1EFE1E1DB4" box="[223,386,1904,1930]" italics="true" pageId="6" pageNumber="407">Edessa jugalis</emphasis>
<bibRefCitation id="9E3D4B4B317BD91945AE6A1EFDB11DB4" author="Breddin, G." box="[395,557,1904,1930]" pageId="6" pageNumber="407" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907</bibRefCitation>
Comp.
</taxonomicName>
w. 
<typeStatus id="25178818317BD919468E6A1EFD6D1DB4" box="[683,753,1904,1930]" pageId="6" pageNumber="407">TYPE</typeStatus>
Fernandes, JAM 1999 (
<collectionCode id="9CBDAE7F317BD91940216A1EFBC21DB4" box="[1028,1118,1904,1930]" country="Netherlands" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="407" type="Museum">RMNH</collectionCode>
); 
<emphasis id="C8D8EAA8317BD91940566A01FAE31DB4" bold="true" box="[1139,1407,1903,1930]" pageId="6" pageNumber="407">
<collectingCountry id="82BB762A317BD91940566A01FA881DB4" box="[1139,1300,1903,1930]" name="Colombia" pageId="6" pageNumber="407">COLOMBIA</collectingCountry>
. 
<collectingRegion id="3868F858317BD91941046A01FAE51DB4" box="[1313,1401,1903,1930]" country="Colombia" name="Bogota" pageId="6" pageNumber="407">Bogotá</collectingRegion>
:
</emphasis>
<specimenCount id="ECAAFD33317BD91941A26A1EFA001DB4" box="[1415,1436,1904,1930]" pageId="6" pageNumber="407" type="female">♀</specimenCount>
<date id="8E12107A317BD91944B26AFDFEB41D90" box="[151,296,1939,1966]" pageId="6" pageNumber="407" value="1929-06-25">25–VI–1929</date>
, M. H. Nicefero (
<collectionCode id="9CBDAE7F317BD91945CC6AFAFDDC1D90" box="[489,576,1940,1966]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="407" type="Museum">USNM</collectionCode>
); 
<emphasis id="C8D8EAA8317BD91946776AFDFCD31D90" bold="true" box="[594,847,1939,1966]" pageId="6" pageNumber="407">
<collectingCountry id="82BB762A317BD91946776AFDFD231D90" box="[594,703,1939,1966]" name="Brazil" pageId="6" pageNumber="407">BRAZIL</collectingCountry>
. 
<collectingRegion id="3868F858317BD91946EC6AFDFCD61D90" box="[713,842,1939,1966]" country="Brazil" name="Amazonas" pageId="6" pageNumber="407">Amazonas</collectingRegion>
:
</emphasis>
<specimenCount id="ECAAFD33317BD91947706AFAFCF61D90" box="[853,874,1940,1966]" pageId="6" pageNumber="407" type="male">♂</specimenCount>
Benjamin Constant, 3–1943, A. Parko (
<collectionCode id="9CBDAE7F317BD91941016AFAFAE91D90" box="[1316,1397,1940,1966]" country="Brazil" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="407">MNRJ</collectionCode>
); 
<specimenCount id="ECAAFD33317BD91941A26AFAFA001D90" box="[1415,1436,1940,1966]" pageId="6" pageNumber="407" type="female">♀</specimenCount>
Colkoen 
<collectingCountry id="82BB762A317BD91944DB6AD6FED81DEC" box="[254,324,1976,2002]" name="Brazil" pageId="6" pageNumber="407">Brasil</collectingCountry>
, Museum Leiden, 
<taxonomicName id="3DAC4D39317BD91946326AD6FC261DEC" authority="F. Cat." authorityName="F. Cat." box="[535,954,1976,2002]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="quadridens" subGenus="Hypoxys">
<emphasis id="C8D8EAA8317BD91946326AD6FDFA1DEC" box="[535,614,1976,2002]" italics="true" pageId="6" pageNumber="407">Edessa</emphasis>
(
<emphasis id="C8D8EAA8317BD919465D6AD6FD441DEF" box="[632,728,1976,2001]" italics="true" pageId="6" pageNumber="407">Hypoxys</emphasis>
) 
<emphasis id="C8D8EAA8317BD91946CC6AD6FCFB1DEC" box="[745,871,1976,2002]" italics="true" pageId="6" pageNumber="407">quadridens</emphasis>
F. Cat.
</taxonomicName>
Nº1, 
<taxonomicName id="3DAC4D39317BD91947D96AD6FA011DEC" authority="Breddin, 1907 Comp." authorityName="Breddin" authorityYear="1907" box="[1020,1437,1976,2002]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="jugalis">
<emphasis id="C8D8EAA8317BD91947D96AD6FB031DEC" box="[1020,1183,1976,2002]" italics="true" pageId="6" pageNumber="407">Edessa jugalis</emphasis>
<bibRefCitation id="9E3D4B4B317BD91940826AD6FAD51DEC" author="Breddin, G." box="[1191,1353,1976,2002]" pageId="6" pageNumber="407" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907</bibRefCitation>
Comp.
</taxonomicName>
w. 
<typeStatus id="25178818317BD91944926AB2FF611DC8" box="[183,253,2012,2038]" pageId="6" pageNumber="407">TYPE</typeStatus>
Fernandes, JAM 1999 (
<collectionCode id="9CBDAE7F317BD919462E6AB2FDFA1DC8" box="[523,614,2012,2038]" country="Netherlands" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="407" type="Museum">RMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="B2B66531317AD91844E26DF7FAD61AC5" pageId="7" pageNumber="408" type="description">
<paragraph id="FA1336BA317AD91844E26DF7FAD61AC5" blockId="7.[151,1437,152,1115]" pageId="7" pageNumber="408">
<emphasis id="C8D8EAA8317AD91844E26DF7FEE21A8D" bold="true" box="[199,382,153,179]" pageId="7" pageNumber="408">Measurements.</emphasis>
total length: 16.2–19.0; head length: 2.1–2.4; head width: 3.2–3.7; pronotal width: 10.0–11.6; abdominal width: 9.0–10.2; length of antennal segments. I: 0.8–1.0; II: 1.6–2.2; III: 1.1–1.7; IV: 4.1–5.6; V: 5.0–6.2; interocular distance: 1.4–1.7; scutellum length: 7.3–9.3; scutellum width: 5.7–6.8; pronotal length: 3.3–3.9.
</paragraph>
</subSubSection>
<subSubSection id="B2B66531317AD91844E26C6AFDDA185C" pageId="7" pageNumber="408" type="diagnosis">
<paragraph id="FA1336BA317AD91844E26C6AFDDA185C" blockId="7.[151,1437,152,1115]" pageId="7" pageNumber="408">
<emphasis id="C8D8EAA8317AD91844E26C6AFEDE1B21" bold="true" box="[199,322,260,287]" pageId="7" pageNumber="408">Diagnosis.</emphasis>
Anterolateral margin of pronotum with solid black stripe reaching all or part of its extension with remaining part smoky black and narrow. Cicatrices of pronotum with concolorous punctures delimiting anterior sulci (
<figureCitation id="62972A3F317AD91844F86C22FED41B59" box="[221,328,332,359]" captionStart="FIGURE 18" captionStartId="34.[151,250,1983,2008]" captionTargetBox="[244,1330,184,1951]" captionTargetId="figure@34.[190,1398,181,1957]" captionTargetPageId="34" captionText="FIGURE 18. Dorsal and ventral view. H. quadridens A–B; H. boerneri C–D; H. claricolor E–F; H. dolosus G–H; H. eburatulus I–J; H. necopinatus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677905" httpUri="https://zenodo.org/record/3677905/files/figure.png" pageId="7" pageNumber="408">Fig. 18A</figureCitation>
). Anterolateral margin not punctured. Male (
<figureCitation id="62972A3F317AD918476D6C22FC1B1B59" box="[840,903,332,359]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="7" pageNumber="408">Fig. 1</figureCitation>
A–C). Pygophore with dorsal rim sinuous and shallowly excavated (
<figureCitation id="62972A3F317AD91845A96C1EFE7A1BB5" box="[396,486,368,395]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="7" pageNumber="408">Fig. 1A</figureCitation>
). Superior process of genital cup elongate, elliptic with a ventral notch and adjacent to dorsal rim (
<figureCitation id="62972A3F317AD91845136CFAFE101B91" box="[310,396,404,431]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="7" pageNumber="408">Fig. 1B</figureCitation>
). Paramere elongate, spatulate, broad distally, anteriorly curved touching superior process of genital cup (
<figureCitation id="62972A3F317AD91845056CD6FEEA1BED" box="[288,374,440,467]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="7" pageNumber="408">Fig. 1B</figureCitation>
). Proctiger with dorsolateral tufts of setae separated by tumid ridge; posterior face subrectangular and slightly tumid (
<figureCitation id="62972A3F317AD91845B06CB2FE771BC9" box="[405,491,476,503]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="7" pageNumber="408">Fig. 1B</figureCitation>
). Ventral rim with expansion acuminate, flat, surpassing level of posterolateral angle in ventral view (
<figureCitation id="62972A3F317AD91845756F6EFE3B1825" box="[336,423,512,539]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="7" pageNumber="408">Fig. 1C</figureCitation>
); base of each expansion with a small transverse carina. Female (
<figureCitation id="62972A3F317AD91840AF6F6EFB781825" box="[1162,1252,512,539]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="7" pageNumber="408">Fig. 1D</figureCitation>
). Gonocoxites 8 posterior margin slightly sinuous, inner angle projecting over gonocoxites 9, sutural border slightly divergent distally. Gonapophyses 8 barely exposed.
</paragraph>
</subSubSection>
<subSubSection id="B2B66531317AD91844E26F02FAF61E2D" pageId="7" pageNumber="408" type="discussion">
<paragraph id="FA1336BA317AD91844E26F02FAF61E2D" blockId="7.[151,1437,152,1115]" pageId="7" pageNumber="408">
<emphasis id="C8D8EAA8317AD91844E26F02FED318B9" bold="true" box="[199,335,620,647]" pageId="7" pageNumber="408">Comments.</emphasis>
The type series of 
<taxonomicName id="3DAC4D39317AD91846146F03FD4C18B8" box="[561,720,620,646]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="C8D8EAA8317AD91846146F03FD4C18B8" box="[561,720,620,646]" italics="true" pageId="7" pageNumber="408">E. quadridens</emphasis>
</taxonomicName>
is formed by four specimens (all in Copenhagen, but part belonging to Kiel collection), three males and one female. Male without head presented in this paper (
<figureCitation id="62972A3F317AD918412E6FFEFAC31895" box="[1291,1375,656,683]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="7" pageNumber="408">Fig. 1F</figureCitation>
) was photographed by Pieter van Doesburg as the 
<typeStatus id="25178818317AD91846A26FDAFD7218F0" box="[647,750,692,718]" pageId="7" pageNumber="408" type="lectotype">lectotype</typeStatus>
of the species several years ago and we are confirming his selection. 
<bibRefCitation id="9E3D4B4B317AD91844D56FB6FE0718CD" author="Breddin, G." box="[240,411,728,755]" pageId="7" pageNumber="408" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin (1907)</bibRefCitation>
made it clear that he had not seen the type of 
<taxonomicName id="3DAC4D39317AD91847B46FB7FBB118CC" box="[913,1069,728,754]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="C8D8EAA8317AD91847B46FB7FBB118CC" box="[913,1069,728,754]" italics="true" pageId="7" pageNumber="408">E. quadridens</emphasis>
</taxonomicName>
, and postulated that the best candidate to be 
<taxonomicName id="3DAC4D39317AD918453A6F93FE201928" box="[287,444,764,790]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="C8D8EAA8317AD918453A6F93FE201928" box="[287,444,764,790]" italics="true" pageId="7" pageNumber="408">E. quadridens</emphasis>
</taxonomicName>
was 
<taxonomicName id="3DAC4D39317AD91845D06F93FDC31928" authorityName="Breddin" authorityYear="1907" box="[501,607,764,790]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="jugalis">
<emphasis id="C8D8EAA8317AD91845D06F93FDC31928" box="[501,607,764,790]" italics="true" pageId="7" pageNumber="408">E. jugalis</emphasis>
</taxonomicName>
. We confirm Breddin’s supposition considering 
<taxonomicName id="3DAC4D39317AD918405E6F93FB791928" authorityName="Breddin" authorityYear="1907" box="[1147,1253,764,791]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="jugalis">
<emphasis id="C8D8EAA8317AD918405E6F93FB101928" box="[1147,1164,765,790]" italics="true" pageId="7" pageNumber="408">E</emphasis>
. 
<emphasis id="C8D8EAA8317AD91840BC6F92FB791928" box="[1177,1253,764,790]" italics="true" pageId="7" pageNumber="408">jugalis</emphasis>
</taxonomicName>
junior synonym of 
<emphasis id="C8D8EAA8317AD91844906E4FFEC61904" box="[181,346,800,827]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="3DAC4D39317AD91844906E4FFED31905" box="[181,335,800,827]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="quadriden">
<emphasis id="C8D8EAA8317AD91844906E4FFED31905" bold="true" box="[181,335,800,827]" italics="true" pageId="7" pageNumber="408">H. quadriden</emphasis>
</taxonomicName>
s
</emphasis>
. The male of 
<taxonomicName id="3DAC4D39317AD91845D06E4FFD471904" authorityName=", Amyot &amp; Serville" authorityYear="1843" box="[501,731,800,826]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="C8D8EAA8317AD91845D06E4FFD471904" box="[501,731,800,826]" italics="true" pageId="7" pageNumber="408">Hypoxys quadridens</emphasis>
</taxonomicName>
shares with 
<taxonomicName id="3DAC4D39317AD91847426E4FFC671905" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[871,1019,801,827]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="offuscatus">
<emphasis id="C8D8EAA8317AD91847426E4FFC671905" box="[871,1019,801,827]" italics="true" pageId="7" pageNumber="408">H. offuscatus</emphasis>
</taxonomicName>
the paramere spatulate (
<figureCitation id="62972A3F317AD91841296E4EFAF81905" box="[1292,1380,800,827]" captionStart="FIGURE 7" captionStartId="16.[151,250,1746,1771]" captionTargetBox="[169,1410,207,1693]" captionTargetId="figure@16.[151,1436,181,1721]" captionTargetPageId="16" captionText="FIGURE 7. H. offuscatus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="7" pageNumber="408">Fig. 7B</figureCitation>
), but both species can be separated by the dorsal tumid ridge narrow of proctiger (
<figureCitation id="62972A3F317AD91847CA6E2AFBD91961" box="[1007,1093,836,863]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="7" pageNumber="408">Fig. 1B</figureCitation>
), and expansion of ventral rim of pygophore acuminated (
<figureCitation id="62972A3F317AD91845EC6E06FD8319BD" box="[457,543,872,899]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="7" pageNumber="408">Fig. 1C</figureCitation>
) in 
<emphasis id="C8D8EAA8317AD91846756E07FD6A19BC" box="[592,758,872,899]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="3DAC4D39317AD91846756E07FD7719BD" box="[592,747,872,899]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="quadriden">
<emphasis id="C8D8EAA8317AD91846756E07FD7719BD" bold="true" box="[592,747,872,899]" italics="true" pageId="7" pageNumber="408">H. quadriden</emphasis>
</taxonomicName>
s
</emphasis>
(dorsal ridge not tumid and wide, and expansion of ventral rim rounded and tumid in 
<taxonomicName id="3DAC4D39317AD91845986EE3FDCD1999" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[445,593,909,935]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="offuscatus">
<emphasis id="C8D8EAA8317AD91845986EE3FDCD1999" box="[445,593,909,935]" italics="true" pageId="7" pageNumber="408">H. offuscatus</emphasis>
</taxonomicName>
). The female of 
<emphasis id="C8D8EAA8317AD91847296EE3FC2D1998" box="[780,945,908,935]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="3DAC4D39317AD91847296EE3FC3A1999" box="[780,934,908,935]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="quadriden">
<emphasis id="C8D8EAA8317AD91847296EE3FC3A1999" bold="true" box="[780,934,908,935]" italics="true" pageId="7" pageNumber="408">H. quadriden</emphasis>
</taxonomicName>
s
</emphasis>
can be confused with 
<taxonomicName id="3DAC4D39317AD91840886EE3FAD91998" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[1197,1349,908,934]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="trabeculus">
<emphasis id="C8D8EAA8317AD91840886EE3FAD91998" box="[1197,1349,908,934]" italics="true" pageId="7" pageNumber="408">H. trabeculus</emphasis>
</taxonomicName>
(see 
<taxonomicName id="3DAC4D39317AD91841A46EE3FE9119F4" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="trabeculus">
<emphasis id="C8D8EAA8317AD91841A46EE3FE9119F4" italics="true" pageId="7" pageNumber="408">H. trabeculus</emphasis>
</taxonomicName>
comments for differences between these two species). A few female specimens of 
<taxonomicName id="3DAC4D39317AD91840886EDFFAFF19F5" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[1197,1379,944,971]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="oxyacanthus">
<emphasis id="C8D8EAA8317AD91840886EDFFAFF19F5" bold="true" box="[1197,1379,944,971]" italics="true" pageId="7" pageNumber="408">H. oxyacanthus</emphasis>
</taxonomicName>
with solid black stripe on posterior part of pronotum can be confused with 
<emphasis id="C8D8EAA8317AD91847826EBBFBD019D0" box="[935,1100,980,1007]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="3DAC4D39317AD91847826EBBFBDD19D1" box="[935,1089,980,1007]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="quadriden">
<emphasis id="C8D8EAA8317AD91847826EBBFBDD19D1" bold="true" box="[935,1089,980,1007]" italics="true" pageId="7" pageNumber="408">H. quadriden</emphasis>
</taxonomicName>
s
</emphasis>
, but punctuation of pronotum is different (
<figureCitation id="62972A3F317AD91845056E96FE1F1E2D" box="[288,387,1016,1043]" captionStart="FIGURE 18" captionStartId="34.[151,250,1983,2008]" captionTargetBox="[244,1330,184,1951]" captionTargetId="figure@34.[190,1398,181,1957]" captionTargetPageId="34" captionText="FIGURE 18. Dorsal and ventral view. H. quadridens A–B; H. boerneri C–D; H. claricolor E–F; H. dolosus G–H; H. eburatulus I–J; H. necopinatus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677905" httpUri="https://zenodo.org/record/3677905/files/figure.png" pageId="7" pageNumber="408">Fig. 18A</figureCitation>
and 
<figureCitation id="62972A3F317AD918459F6E97FE771E2D" box="[442,491,1017,1043]" captionStart="FIGURE 19" captionStartId="35.[151,250,1983,2008]" captionTargetBox="[250,1354,187,1936]" captionTargetId="figure@35.[190,1398,181,1957]" captionTargetPageId="35" captionText="FIGURE 19. Dorsal and ventral view. H. offuscatus A–B; H. trabeculus C–D; H. triangularis E–F; H. capito G–H; H. subrastratus I–J; H. oxyacanthus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677907" httpUri="https://zenodo.org/record/3677907/files/figure.png" pageId="7" pageNumber="408">19K</figureCitation>
) and gonocoxites 8 of 
<emphasis id="C8D8EAA8317AD91846CE6E97FC0C1E2C" box="[747,912,1016,1043]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="3DAC4D39317AD91846CE6E97FC191E2D" box="[747,901,1016,1043]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="quadriden">
<emphasis id="C8D8EAA8317AD91846CE6E97FC191E2D" bold="true" box="[747,901,1016,1043]" italics="true" pageId="7" pageNumber="408">H. quadriden</emphasis>
</taxonomicName>
s
</emphasis>
are more convex than in 
<taxonomicName id="3DAC4D39317AD91840886E97FAFE1E2D" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[1197,1378,1016,1043]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="oxyacanthus">
<emphasis id="C8D8EAA8317AD91840886E97FAFE1E2D" bold="true" box="[1197,1378,1016,1043]" italics="true" pageId="7" pageNumber="408">H. oxyacanthus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B2B66531317AD91844B26972FE5A1E65" pageId="7" pageNumber="408" type="distribution">
<paragraph id="FA1336BA317AD91844B26972FE5A1E65" blockId="7.[151,1437,152,1115]" pageId="7" pageNumber="408">
<emphasis id="C8D8EAA8317AD91844B26972FE021E09" bold="true" box="[151,414,1052,1079]" pageId="7" pageNumber="408">
Distribution (
<figureCitation id="62972A3F317AD918451F6972FE131E09" box="[314,399,1052,1079]" captionStart="FIGURE 21" captionStartId="37.[151,250,1173,1198]" captionTargetBox="[159,1429,193,1137]" captionTargetId="figure@37.[151,1436,181,1148]" captionTargetPageId="37" captionText="FIGURE 21. Distributional map of the species of H. quadridens group." figureDoi="http://doi.org/10.5281/zenodo.3677911" httpUri="https://zenodo.org/record/3677911/files/figure.png" pageId="7" pageNumber="408">Fig. 21</figureCitation>
).
</emphasis>
<collectingCountry id="82BB762A317AD91845806973FDCC1E09" box="[421,592,1053,1079]" name="Venezuela" pageId="7" pageNumber="408">VENEZUELA</collectingCountry>
: 
<collectingRegion id="3868F858317AD91846796973FD591E09" box="[604,709,1053,1079]" country="Venezuela" name="Monagas" pageId="7" pageNumber="408">Monagas</collectingRegion>
, 
<collectingRegion id="3868F858317AD91846F46972FCB51E08" box="[721,809,1052,1078]" country="Venezuela" name="Bolivar" pageId="7" pageNumber="408">Bolivar</collectingRegion>
; 
<collectingCountry id="82BB762A317AD91847106973FB821E09" box="[821,1054,1053,1079]" name="French Guiana" pageId="7" pageNumber="408">FRENCH GUIANA</collectingCountry>
; 
<collectingCountry id="82BB762A317AD918400E6973FB5D1E09" box="[1067,1217,1053,1079]" name="Suriname" pageId="7" pageNumber="408">SURINAME</collectingCountry>
; 
<collectingCountry id="82BB762A317AD91840E86973FAFB1E09" box="[1229,1383,1053,1079]" name="Colombia" pageId="7" pageNumber="408">COLOMBIA</collectingCountry>
: 
<collectingRegion id="3868F858317AD91841576973FF571E65" country="Colombia" name="Bogota" pageId="7" pageNumber="408">Bogotá</collectingRegion>
; 
<collectingCountry id="82BB762A317AD91844F3692FFEDC1E65" box="[214,320,1089,1115]" name="Brazil" pageId="7" pageNumber="408">BRAZIL</collectingCountry>
: Amazonas.
</paragraph>
</subSubSection>
</treatment>
</document>