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<mods:title id="C5A12160F8CBAA81CCFE7FC9C0954997">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<taxonomicName id="23946389517B1816FF0EFE26ED1C1D94" ID-CoL="363C" authority="Ballantyne 2013" authorityName="Ballantyne in Ballantyne &amp; Lambkin" authorityYear="2013" box="[151,548,441,468]" class="Insecta" family="Lampyridae" genus="Australoluciola" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C418517B1816FF0EFE26EE6A1D93" bold="true" box="[151,338,441,468]" italics="true" pageId="64" pageNumber="65">Australoluciola</emphasis>
<bibRefCitation id="800565FB517B1816FEC3FE26ED1C1D94" author="Ballantyne" box="[346,548,441,467]" firstAuthor="Ballantyne" pageId="64" pageNumber="65" pagination="1 - 48" refId="ref114209" refString="Ballantyne, L., Fu, X. - H., Lambkin, C., Jeng, M. - L., Faust, L., Wijekoon, W. M. C. D., Li, D. - Q. &amp; Zhu, T. - F. (2013) Studies on South-east Asian fireflies: Abscondita, a new genus with details of life history, flashing patterns and behaviour of Abs. chinensis (L.) and Abs. terminalis (Olivier) (Coleoptera: Lampyridae: Luciolinae). Zootaxa, 3721 (1), 1 - 48. https: // doi. org / 10.11646 / zootaxa. 3721.1.1" type="journal article" year="2013">Ballantyne 2013</bibRefCitation>
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<paragraph id="E42B180A517B1816FF0EFE42EE761DB0" blockId="64.[151,548,441,503]" box="[151,334,477,503]" pageId="64" pageNumber="65">
<figureCitation id="7CAF048F517B1816FF0EFE42EFC21DB0" box="[151,250,477,503]" captionStart="FIGURES 1−8" captionStartId="11.[151,264,1682,1707]" captionTargetBox="[152,1435,197,1617]" captionTargetId="figure@11.[151,1435,197,1619]" captionTargetPageId="11" captionText="FIGURES 1−8. Key to genera characters 1. 1, 2 Australoluciola anthracina type male (MNHN); 3, 4 Aus. japenensis paratype male; 5, 6 Atyphella scabra male; 7, 8 Aty. leucura type male (MNHN). 1, 2, 5, 6 aedeagus ventral (1, 5), and dorsal (2, 6) (lateral lobes are arrowed). 3, 4, 7, 8 aedeagal sheath dorsal (3, 7), and ventral (4, 8). Oblique arrows in anterior section indicate point where tergite arms join the lateral margins of the sheath sternite; vertical arrow pointing to foot of page in 3 indicates anterior margin of sheath tergite; vertical and horizontal lines in 4 indicate length (vertical) and width (horizontal) of sheath sternite; lower paired arrows in 8 indicate emarginated margin of sheath sternite." figureDoi="http://doi.org/10.5281/zenodo.3508102" httpUri="https://zenodo.org/record/3508102/files/figure.png" pageId="64" pageNumber="65">Figs 3, 4</figureCitation>
, 
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<taxonomicName id="23946389517B1816FF0EFDBEEC2D1E7E" ID-CoL="363C" authority="Ballantyne in Ballantyne &amp; Lambkin 2013: 43" authorityName="Ballantyne in Ballantyne &amp; Lambkin" authorityPageNumber="43" authorityYear="2013" box="[151,789,544,569]" class="Insecta" family="Lampyridae" genus="Australoluciola" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418517B1816FF0EFDBEEE0D1E7E" box="[151,309,545,569]" italics="true" pageId="64" pageNumber="65">Australoluciola</emphasis>
Ballantyne in 
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<bibRefCitation id="800565FB517B1816FE50FDBFEC2D1E7E" author="Ballantyne &amp; Lambkin" box="[457,789,544,569]" firstAuthor="Ballantyne" pageId="64" pageNumber="65" pagination="1 - 162" refId="ref114597" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2013) Systematics and phylogenetics of Indo-Pacific Luciolinae fireflies (Coleoptera: Lampyridae) and the description of new genera. Zootaxa, 3653 (1), 1 - 162. https: // doi. org / 10.11646 / zootaxa. 3653.1.1" type="journal article" year="2013">Ballantyne &amp; Lambkin 2013: 43</bibRefCitation>
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. 
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<bibRefCitation id="800565FB517B1816FC86FDBEEB121E7E" author="Sartsanga" box="[799,1066,545,569]" etAl="et al." firstAuthor="Sartsanga" pageId="64" pageNumber="65" pagination="727 - 732" refId="ref121784" refString="Sartsanga, C., Chanchay, P., Chaovalit, S., Ballantyne, L. A., Swatdipong, A. &amp; Sriboonlert, A. (2017) Australoluciola thailandica Sriboonlert, Swatdipong and Sartsanga, new species (Coleoptera: Lampyridae: Luciolinae) from Thailand: First Record of the genus Australoluciola Ballantyne from Southeast Asia. The Coleopterists Bulletin, 71 (4), 727 - 732. https: // doi. org / 10.1649 / 0010 - 065 X- 71.4.727" type="journal article" year="2017">
Sartsanga 
<emphasis id="D6E0C418517B1816FC11FDBDEC831E7E" box="[904,955,545,569]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2017: 727
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.
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<emphasis id="D6E0C418517B1816FF0EFDDFEE261E1F" bold="true" box="[151,286,575,600]" pageId="64" pageNumber="65">
<typeStatus id="3B2FA6A8517B1816FF0EFDDFEFF41E1F" box="[151,204,576,600]" pageId="64" pageNumber="65">Type</typeStatus>
species
</emphasis>
. 
<taxonomicName id="23946389517B1816FEB3FDDEED751E1F" authority="Fabricius" authorityName="Fabricius" box="[298,589,575,600]" class="Insecta" family="Lampyridae" genus="Lampyris" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="D6E0C418517B1816FEB3FDDEEED11E1F" box="[298,489,576,600]" italics="true" pageId="64" pageNumber="65">Lampyris australis</emphasis>
Fabricius
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<paragraph id="E42B180A517B1816FF0EFD1AEE3A1F3F" blockId="64.[151,1437,645,1537]" pageId="64" pageNumber="65">
<emphasis id="D6E0C418517B1816FF0EFD1AEE321EE7" bold="true" box="[151,266,645,672]" pageId="64" pageNumber="65">Diagnosis</emphasis>
. Repeated with some modifications from 
<bibRefCitation id="800565FB517B1816FD47FD19EB661EE7" author="Ballantyne &amp; Lambkin" box="[734,1118,646,672]" firstAuthor="Ballantyne" pageId="64" pageNumber="65" pagination="1 - 162" refId="ref114597" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2013) Systematics and phylogenetics of Indo-Pacific Luciolinae fireflies (Coleoptera: Lampyridae) and the description of new genera. Zootaxa, 3653 (1), 1 - 162. https: // doi. org / 10.11646 / zootaxa. 3653.1.1" type="journal article" year="2013">Ballantyne &amp; Lambkin (2013: 39)</bibRefCitation>
. 
<taxonomicName id="23946389517B1816FBF1FD19EA2E1EE7" authorityName="Ballantyne in Ballantyne &amp; Lambkin" authorityYear="2013" box="[1128,1302,646,672]" class="Insecta" family="Lampyridae" genus="Australoluciola" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418517B1816FBF1FD19EA2E1EE7" box="[1128,1302,646,672]" italics="true" pageId="64" pageNumber="65">Australoluciola</emphasis>
</taxonomicName>
is predominantly an Australian and New Guinean genus. It is found across the northern Australian coastline to Darwin, and along the eastern coast of 
<collectingCountry id="9C83589A517B1816FE2DFD51ED231EAF" box="[436,539,718,744]" name="Australia" pageId="64" pageNumber="65">Australia</collectingCountry>
<materialsCitation id="54FC1257517B1816FDB8FD51EFC61F3F" ID-GBIF-Occurrence="2427485703" country="Thailand" location="Brisbane area" pageId="64" pageNumber="65" specimenCount="1" typeStatus="holotype">
either in mangroves or in more open forest south to the 
<location id="E14B4ED1517B1816FB1DFD51EA271EAE" LSID="urn:lsid:plazi:treatment:6C3DA91C517B1816FF0EFE26EABF1A47:E14B4ED1517B1816FB1DFD51EA271EAE" box="[1156,1311,718,745]" country="Thailand" name="Brisbane area" pageId="64" pageNumber="65">Brisbane area</location>
, where one species occurs in relict rain forest in suburban areas of Brisbane. Species are slim bodied and usually have orange pronota and black elytra. Females are macropterous and capable of flight. Larvae are slim bodied and terrestrial with laterotergites visible at the sides of the body when viewed from above. One species has been reported from 
<collectingCountry id="9C83589A517B1816FF0EFCC1EFC61F3F" box="[151,254,862,888]" name="Thailand" pageId="64" pageNumber="65">Thailand</collectingCountry>
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.
</paragraph>
<paragraph id="E42B180A517B1816FF5EFC1DEB2E196F" blockId="64.[151,1437,645,1537]" pageId="64" pageNumber="65">
The genus belongs in a group of genera characterized by: pronotal width less than width across elytral humeri, parallel-sided elytra, no MFC, an elongate slender aedeagus with LL largely concealed behind the ML when viewed from beneath (
<figureCitation id="7CAF048F517B1816FEA5FC55EE9A1FA3" box="[316,418,970,996]" captionStart="FIGURES 1−8" captionStartId="11.[151,264,1682,1707]" captionTargetBox="[152,1435,197,1617]" captionTargetId="figure@11.[151,1435,197,1619]" captionTargetPageId="11" captionText="FIGURES 1−8. Key to genera characters 1. 1, 2 Australoluciola anthracina type male (MNHN); 3, 4 Aus. japenensis paratype male; 5, 6 Atyphella scabra male; 7, 8 Aty. leucura type male (MNHN). 1, 2, 5, 6 aedeagus ventral (1, 5), and dorsal (2, 6) (lateral lobes are arrowed). 3, 4, 7, 8 aedeagal sheath dorsal (3, 7), and ventral (4, 8). Oblique arrows in anterior section indicate point where tergite arms join the lateral margins of the sheath sternite; vertical arrow pointing to foot of page in 3 indicates anterior margin of sheath tergite; vertical and horizontal lines in 4 indicate length (vertical) and width (horizontal) of sheath sternite; lower paired arrows in 8 indicate emarginated margin of sheath sternite." figureDoi="http://doi.org/10.5281/zenodo.3508102" httpUri="https://zenodo.org/record/3508102/files/figure.png" pageId="64" pageNumber="65">Figs 1, 2</figureCitation>
); aedeagal sheath elongate slender, widest across middle, without bulbous paraprocts, and with both sides of posterior half of sheath sternite tapering evenly towards a narrow entire apex (
<figureCitation id="7CAF048F517B1816FB45FC71EA7B184F" box="[1244,1347,1006,1032]" captionStart="FIGURES 1−8" captionStartId="11.[151,264,1682,1707]" captionTargetBox="[152,1435,197,1617]" captionTargetId="figure@11.[151,1435,197,1619]" captionTargetPageId="11" captionText="FIGURES 1−8. Key to genera characters 1. 1, 2 Australoluciola anthracina type male (MNHN); 3, 4 Aus. japenensis paratype male; 5, 6 Atyphella scabra male; 7, 8 Aty. leucura type male (MNHN). 1, 2, 5, 6 aedeagus ventral (1, 5), and dorsal (2, 6) (lateral lobes are arrowed). 3, 4, 7, 8 aedeagal sheath dorsal (3, 7), and ventral (4, 8). Oblique arrows in anterior section indicate point where tergite arms join the lateral margins of the sheath sternite; vertical arrow pointing to foot of page in 3 indicates anterior margin of sheath tergite; vertical and horizontal lines in 4 indicate length (vertical) and width (horizontal) of sheath sternite; lower paired arrows in 8 indicate emarginated margin of sheath sternite." figureDoi="http://doi.org/10.5281/zenodo.3508102" httpUri="https://zenodo.org/record/3508102/files/figure.png" pageId="64" pageNumber="65">Figs 3, 4</figureCitation>
). Males distinguished from 
<taxonomicName id="23946389517B1816FEECFB8DEECB186B" authorityName="Motschulsky" authorityYear="1854" box="[373,499,1042,1068]" class="Insecta" family="Lampyridae" genus="Colophotia" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418517B1816FEECFB8DEECB186B" box="[373,499,1042,1068]" italics="true" pageId="64" pageNumber="65">Colophotia</emphasis>
</taxonomicName>
in having no median carina on 
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, no expanded and oblique PLP and entire, not bipartite, LOs in 
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; from 
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<emphasis id="D6E0C418517B1816FE27FBA8ED121817" box="[446,554,1079,1104]" italics="true" pageId="64" pageNumber="65">Pteroptyx</emphasis>
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in having no MFC, no deflexed elytral apices, no bulbous aedeagal sheath paraprocts and entire, not bipartite, LOs in 
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; from those 
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<emphasis id="D6E0C418517B1816FD6BFBC4EC661833" box="[754,862,1115,1140]" italics="true" pageId="64" pageNumber="65">Pteroptyx</emphasis>
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with entire LO in 
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by the absence of a MFC, and from 
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<emphasis id="D6E0C418517B1816FF4DFBE0EE7318DF" box="[212,331,1151,1176]" italics="true" pageId="64" pageNumber="65">P. testacea</emphasis>
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by the presence in 
<taxonomicName id="23946389517B1816FDBBFBE0ED4618DF" box="[546,638,1151,1176]" class="Insecta" family="Lampyridae" genus="Pteroptyx" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="D6E0C418517B1816FDBBFBE0ED4618DF" box="[546,638,1151,1176]" italics="true" pageId="64" pageNumber="65">testacea</emphasis>
</taxonomicName>
of bipartite LO in 
<date id="902A3ECA517B1816FCD6FBE0EC4918DE" box="[847,881,1151,1177]" pageId="64" pageNumber="65">V7</date>
(
<bibRefCitation id="800565FB517B1816FCE6FBE1EB0518DE" author="Jusoh" box="[895,1085,1150,1177]" etAl="et al." firstAuthor="Jusoh" pageId="64" pageNumber="65" pagination="1 - 71" refId="ref118381" refString="Jusoh, W. F. A., Ballantyne, L., Lambkin, C. L., Hashim, N. R. &amp; Wahlberg, N. (2018) The firefly genus Pteroptyx Olivier revisited (Coleoptera: Lampyridae: Luciolinae). Zootaxa, 4456 (1), 1 - 71. https: // doi. org / 10.11646 / zootaxa. 4456.1.1" type="journal article" year="2018">
Jusoh 
<emphasis id="D6E0C418517B1816FC5EFBE0ECC718DF" box="[967,1023,1150,1176]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2018
</bibRefCitation>
)from 
<taxonomicName id="23946389517B1816FB1DFBE1EA3118DF" authorityName="Olivier" authorityYear="1885" box="[1156,1289,1150,1176]" class="Insecta" family="Lampyridae" genus="Pyrophanes" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418517B1816FB1DFBE1EA3118DF" box="[1156,1289,1150,1176]" italics="true" pageId="64" pageNumber="65">Pyrophanes</emphasis>
</taxonomicName>
(which has a MFC) and 
<taxonomicName id="23946389517B1816FE88FB3CEEE718FB" box="[273,479,1187,1212]" class="Insecta" family="Lampyridae" genus="Pteroptyx" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="D6E0C418517B1816FE88FB3CEEE718FB" box="[273,479,1187,1212]" italics="true" pageId="64" pageNumber="65">Pteroptyx testacea</emphasis>
</taxonomicName>
(which has no MFC) in being without both incurving lobes along 
<date id="902A3ECA517B1816FB25FB3CEBE618FA" box="[1212,1246,1187,1213]" pageId="64" pageNumber="65">V7</date>
and bipartite LO in 
<date id="902A3ECA517B1816FF2AFB58EFE018A6" box="[179,216,1223,1249]" pageId="64" pageNumber="65">V7</date>
; from 
<taxonomicName id="23946389517B1816FEBBFB58EEAD18A7" authorityName="Ballantyne in Ballantyne &amp; Lambkin" authorityYear="2013" box="[290,405,1223,1248]" class="Insecta" family="Lampyridae" genus="Trisinuata" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418517B1816FEBBFB58EEAD18A7" box="[290,405,1223,1248]" italics="true" pageId="64" pageNumber="65">Trisinuata</emphasis>
</taxonomicName>
by the entire LOs in 
<date id="902A3ECA517B1816FD1BFB58ED9C18A6" box="[642,676,1223,1249]" pageId="64" pageNumber="65">V7</date>
(those of 
<taxonomicName id="23946389517B1816FC8CFB58ECB018A7" authorityName="Ballantyne in Ballantyne &amp; Lambkin" authorityYear="2013" box="[789,904,1223,1248]" class="Insecta" family="Lampyridae" genus="Trisinuata" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418517B1816FC8CFB58ECB018A7" box="[789,904,1223,1248]" italics="true" pageId="64" pageNumber="65">Trisinuata</emphasis>
</taxonomicName>
are bipartite); from most 
<taxonomicName id="23946389517B1816FB31FB59EA0218A7" authorityName="Ballantyne in Ballantyne &amp; Lambkin" authorityYear="2013" box="[1192,1338,1222,1248]" class="Insecta" family="Lampyridae" genus="Medeopteryx" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418517B1816FB31FB59EA0218A7" box="[1192,1338,1222,1248]" italics="true" pageId="64" pageNumber="65">Medeopteryx</emphasis>
</taxonomicName>
in being without deflexed elytral apices. It differs from 
<taxonomicName id="23946389517B1816FD02FB75EDDE1943" box="[667,742,1258,1284]" class="Insecta" family="Lampyridae" genus="Inflata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C418517B1816FD02FB75EDDE1943" box="[667,742,1258,1284]" italics="true" pageId="64" pageNumber="65">Inflata</emphasis>
</taxonomicName>
most obviously by the slender aedeagal ML with narrow bases of the LL; the ML of 
<emphasis id="D6E0C418517B1816FE1EFA91EED4196F" box="[391,492,1294,1320]" italics="true" pageId="64" pageNumber="65">
<taxonomicName id="23946389517B1816FE1EFA91EEEA196F" box="[391,466,1294,1320]" class="Insecta" family="Lampyridae" genus="Inflata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Inflata</taxonomicName>
is
</emphasis>
bulbous and the bases of the LL much expanded.
</paragraph>
<paragraph id="E42B180A517B1816FF5EFAADEABF1A47" blockId="64.[151,1437,645,1537]" pageId="64" pageNumber="65">
All but three species have orange pronotum and dark brown elytra similar to many species of 
<emphasis id="D6E0C418517B1816FB55FAADEFDB1937" italics="true" pageId="64" pageNumber="65">
<taxonomicName id="23946389517B1816FB55FAADEA5A190B" authorityName="Ballantyne in Ballantyne &amp; Lambkin" authorityYear="2013" box="[1228,1378,1330,1356]" class="Insecta" family="Lampyridae" genus="Medeopteryx" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Medeopteryx</taxonomicName>
. 
<taxonomicName id="23946389517B1816FAF2FAACEFDB1937" pageId="64" pageNumber="65" rank="species" species="aspera">Aus. aspera</taxonomicName>
</emphasis>
has median brown pronotal markings, and in 
<taxonomicName id="23946389517B1816FD6AFAC8ECA31937" baseAuthorityName="MacLeay" baseAuthorityYear="1872" box="[755,923,1366,1393]" class="Insecta" family="Lampyridae" genus="Australoluciola" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="flavicollis">
<emphasis id="D6E0C418517B1816FD6AFAC8ECA31937" box="[755,923,1366,1393]" italics="true" pageId="64" pageNumber="65">Aus. flavicollis</emphasis>
</taxonomicName>
the pronotum is orange with the elytral base and/or margins may be pale. 
<taxonomicName id="23946389517B1816FE44FAE4EDB419D3" authorityName="Ballantyne" authorityYear="2013" box="[477,652,1403,1429]" class="Insecta" family="Lampyridae" genus="Lloydiella" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="japenensis">
<emphasis id="D6E0C418517B1816FE44FAE4EDB419D3" box="[477,652,1403,1429]" italics="true" pageId="64" pageNumber="65">Aus. japenensis</emphasis>
</taxonomicName>
alone has pale yellowish brown dorsal colour and dark elytral apices, a dorsal colour pattern characteristic of many S. E. Asian fireflies (
<bibRefCitation id="800565FB517B1816FC18FA01EB7119FE" author="McDermott" box="[897,1097,1438,1465]" firstAuthor="McDermott" pageId="64" pageNumber="65" pagination="1 - 149" refId="ref119380" refString="McDermott, F. A. (1966) Lampyridae. In: Steel, W. O. (Ed.), Coleopterorum Catalogus Supplementa. Pars 9. Editio Secunda. W. Junk, S'Gravenhage, pp. 1 - 149." type="book chapter" year="1966">McDermott 1966</bibRefCitation>
). 
<date id="902A3ECA517B1816FBC2FA00EB4519FE" box="[1115,1149,1439,1465]" pageId="64" pageNumber="65">V7</date>
may be trisinuate or with an MPP only, and the corners are rounded (there are no well-defined PLP); T8 has a well-defined ventral groove margined by ridges and with short, wide and apically rounded flanges in 
<taxonomicName id="23946389517B1816FC51FA78EB741A47" box="[968,1100,1511,1537]" pageId="64" pageNumber="65" rank="species" species="aspera">
<emphasis id="D6E0C418517B1816FC51FA78EB741A47" box="[968,1100,1511,1537]" italics="true" pageId="64" pageNumber="65">Aus. aspera</emphasis>
</taxonomicName>
and 
<taxonomicName id="23946389517B1816FB1AFA78EA711A47" box="[1155,1353,1510,1537]" pageId="64" pageNumber="65" rank="species" species="pharusaurea">
<emphasis id="D6E0C418517B1816FB1AFA78EA711A47" box="[1155,1353,1510,1537]" italics="true" pageId="64" pageNumber="65">Aus. pharusaurea</emphasis>
</taxonomicName>
only.
</paragraph>
</subSubSection>
</treatment>
</document>