177 lines
19 KiB
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177 lines
19 KiB
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<document id="ABB558D09CDC2F82A62D38071CA25D9C" ID-CLB-Dataset="30595" ID-DOI="10.11646/zootaxa.4388.4.1" ID-GBIF-Dataset="a884362d-a9f8-4cee-8cd4-dcf59a73ede2" ID-ISSN="1175-5326" ID-Zenodo-Dep="1189072" ID-ZooBank="7A57C29B-6B16-445E-A8D7-750C736E6B42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520318467094" checkinUser="plazi" docAuthor="Cupello, Mario" docDate="2018" docId="5C282C04FFC4FFE6FF2D55741AFAFC7B" docLanguage="en" docName="zootaxa.4388.4.1.pdf" docOrigin="Zootaxa 4388 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Canthidium (Canthidium) ardens Bates 1887" docType="treatment" docVersion="10" lastPageNumber="456" masterDocId="A011547CFFC1FFE3FFBA541B1E1AFFFB" masterDocTitle="On the types species of the New World dung beetle genus Canthidium Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae), with an annotated checklist of species" masterLastPageNumber="486" masterPageNumber="451" pageNumber="456" updateTime="1698528354521" updateUser="ExternalLinkService">
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<mods:title id="C95F9B096BE8FBA23B162CBCC3043269">On the types species of the New World dung beetle genus Canthidium Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae), with an annotated checklist of species</mods:title>
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<mods:namePart id="2FBEEED85D41272589857CA7D5AD040D">Cupello, Mario</mods:namePart>
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<mods:title id="158856623DCD1834D9F4DE94E759D8AE">Zootaxa</mods:title>
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<mods:date id="1DD36DAC5825B8137FF705346C769EB5">2018</mods:date>
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<mods:number id="A374162F9426BA362B1BAE183AEBE96C">2018-03-06</mods:number>
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<mods:number id="4CAB6AFA8A6E31FA0443A1DE28C5083B">4388</mods:number>
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<mods:identifier id="03B40851D58B4ABDC6F039C811847B0B" type="DOI">10.11646/zootaxa.4388.4.1</mods:identifier>
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<treatment id="5C282C04FFC4FFE6FF2D55741AFAFC7B" ID-DOI="http://doi.org/10.5281/zenodo.5994523" ID-GBIF-Taxon="141280861" ID-Zenodo-Dep="5994523" LSID="urn:lsid:plazi:treatment:5C282C04FFC4FFE6FF2D55741AFAFC7B" httpUri="http://treatment.plazi.org/id/5C282C04FFC4FFE6FF2D55741AFAFC7B" lastPageNumber="456" pageId="5" pageNumber="456">
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<subSubSection id="9C9BCE99FFC4FFE6FF2D55741CB3FE73" box="[151,681,367,392]" pageId="5" pageNumber="456" type="nomenclature">
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<paragraph id="D43E9D12FFC4FFE6FF2D55741CB3FE73" blockId="5.[151,1437,367,896]" box="[151,681,367,392]" pageId="5" pageNumber="456">
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<heading id="8F762A7EFFC4FFE6FF2D55741CB3FE73" bold="true" box="[151,681,367,392]" fontSize="10" level="2" pageId="5" pageNumber="456" reason="2">
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<taxonomicName id="1381E691FFC4FFE6FF2D55741CB3FE73" ID-CoL="QN8W" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[151,681,367,392]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="ardens" subGenus="Canthidium">
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<emphasis id="E6F54100FFC4FFE6FF2D55741CB3FE73" bold="true" box="[151,681,367,392]" pageId="5" pageNumber="456">
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<emphasis id="E6F54100FFC4FFE6FF2D55741F38FE73" bold="true" box="[151,290,367,392]" italics="true" pageId="5" pageNumber="456">Canthidium</emphasis>
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(
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<emphasis id="E6F54100FFC4FFE6FE8855741FA7FE73" bold="true" box="[306,445,367,392]" italics="true" pageId="5" pageNumber="456">Canthidium</emphasis>
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)
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<emphasis id="E6F54100FFC4FFE6FE7655741C00FE73" bold="true" box="[460,538,367,392]" italics="true" pageId="5" pageNumber="456">ardens</emphasis>
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<bibRefCitation id="B010E0E3FFC4FFE6FD9B55741CB3FE73" box="[545,681,367,392]" pageId="5" pageNumber="456" refId="ref25529" refString="Bates, H. W. (1887) Tribe Lamellicornia. In: Bates, H. W. (1886 - 1890) Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. Taylor and Francis, London, pp. 25 - 64, pls. 2 - 3. [publication date taken from Bousquet 2016]" type="journal article">Bates, 1887</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9C9BCE99FFC4FFE6FF7D558F1AFAFC7B" pageId="5" pageNumber="456" type="reference_group">
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<paragraph id="D43E9D12FFC4FFE6FF7D558F1C34FDE3" blockId="5.[151,1437,367,896]" pageId="5" pageNumber="456">
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<treatmentCitationGroup id="F491BA3CFFC4FFE6FF7D558F1C34FDE3" infered="true" pageId="5" pageNumber="456">
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<taxonomicName id="1381E691FFC4FFE6FF7D558F1C4DFE57" ID-CoL="QN8W" authority="Bates, 1887: 48" authorityName="Bates" authorityPageNumber="48" authorityYear="1887" box="[199,599,404,428]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="ardens">
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<emphasis id="E6F54100FFC4FFE6FF7D558F1F84FE57" box="[199,414,404,428]" italics="true" pageId="5" pageNumber="456">Canthidium ardens</emphasis>
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<treatmentCitation id="5520BB03FFC4FFE6FE1C558E1C4DFE57" author="" box="[422,599,404,428]" page="48" pageId="5" pageNumber="456">
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<bibRefCitation id="B010E0E3FFC4FFE6FE1C558E1C35FE57" box="[422,559,404,428]" pageId="5" pageNumber="456" refId="ref25529" refString="Bates, H. W. (1887) Tribe Lamellicornia. In: Bates, H. W. (1886 - 1890) Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. Taylor and Francis, London, pp. 25 - 64, pls. 2 - 3. [publication date taken from Bousquet 2016]" type="journal article">Bates, 1887</bibRefCitation>
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: 48
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</treatmentCitation>
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</taxonomicName>
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[here transferred to the subgenus
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<taxonomicName id="1381E691FFC4FFE6FC60558F1A44FE57" ID-CoL="3GVY" authorityName="Erichson" authorityYear="1847" box="[986,1118,404,428]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus">
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<emphasis id="E6F54100FFC4FFE6FC60558F1A44FE57" box="[986,1118,404,428]" italics="true" pageId="5" pageNumber="456">Canthidium</emphasis>
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</taxonomicName>
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].
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<emphasis id="E6F54100FFC4FFE6FBCC558F1B03FE56" bold="true" box="[1142,1305,404,429]" pageId="5" pageNumber="456">Type locality:</emphasis>
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Nicaragua: Chontales, and Panama: Chiriquí: Bugaba: Volcán (“Nicaragua, Chontales [...]; Panama, Bugaba, Volcan de Chiriqui”).
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<emphasis id="E6F54100FFC4FFE6FEA355C71FD0FE0E" bold="true" box="[281,458,476,501]" pageId="5" pageNumber="456">Type material:</emphasis>
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Unknown number of syntypes of both sexes (BMNH).
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<emphasis id="E6F54100FFC4FFE6FBFB55C71AC6FE0E" bold="true" box="[1089,1244,476,501]" pageId="5" pageNumber="456">Distribution:</emphasis>
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From Mexico to Panama (Kohlmann & Solís 2006a).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="D43E9D12FFC4FFE6FF7D563F1DD8FCC3" blockId="5.[151,1437,367,896]" pageId="5" pageNumber="456">
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<treatmentCitationGroup id="F491BA3CFFC4FFE6FF7D563F1DD8FCC3" infered="true" pageId="5" pageNumber="456">
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<taxonomicName id="1381E691FFC4FFE6FF7D563F1CDCFDC7" ID-CoL="5DQ9B" authority="Bates, 1887: 48" authorityName="Bates" authorityPageNumber="48" authorityYear="1887" box="[199,710,548,572]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="subSpecies" species="ardens" subSpecies="mutatum">
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<emphasis id="E6F54100FFC4FFE6FF7D563F1C13FDC7" box="[199,521,548,572]" italics="true" pageId="5" pageNumber="456">Canthidium ardens mutatum</emphasis>
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<treatmentCitation id="5520BB03FFC4FFE6FDAB563E1CDCFDC7" author="" box="[529,710,548,572]" page="48" pageId="5" pageNumber="456">
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<bibRefCitation id="B010E0E3FFC4FFE6FDAB563E1C81FDC7" box="[529,667,548,572]" pageId="5" pageNumber="456" refId="ref25529" refString="Bates, H. W. (1887) Tribe Lamellicornia. In: Bates, H. W. (1886 - 1890) Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. Taylor and Francis, London, pp. 25 - 64, pls. 2 - 3. [publication date taken from Bousquet 2016]" type="journal article">Bates, 1887</bibRefCitation>
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: 48
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</treatmentCitation>
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</taxonomicName>
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,
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<taxonomicNameLabel id="FDC6FC7BFFC4FFE6FD69563F1D9FFDC6" box="[723,901,548,573]" pageId="5" pageNumber="456">
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<emphasis id="E6F54100FFC4FFE6FD69563F1D9FFDC6" bold="true" box="[723,901,548,573]" pageId="5" pageNumber="456">new synonymy</emphasis>
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</taxonomicNameLabel>
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[except for
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<bibRefCitation id="B010E0E3FFC4FFE6FBAB563F1AEEFDC7" box="[1041,1268,548,573]" pageId="5" pageNumber="456" refId="ref25720" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2. United States National Museum Bulletin, 185, 189 - 341." type="journal article">Blackwelder (1944)</bibRefCitation>
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, this name has been overlooked by all authors since its establishment. Nonetheless, it is clearly available as Bates presented a proper description for it (“
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<taxonomicName id="1381E691FFC4FFE6FE6C56771C87FD7F" ID-CoL="5DQ9B" authorityName="Bates" authorityYear="1887" box="[470,669,620,644]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="subSpecies" species="ardens" subSpecies="mutatum">
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Var.
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<emphasis id="E6F54100FFC4FFE6FDA956771C87FD7F" box="[531,669,620,644]" italics="true" pageId="5" pageNumber="456">C. mutatum</emphasis>
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</taxonomicName>
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[under
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<taxonomicName id="1381E691FFC4FFE6FCBB56771DC4FD7F" ID-CoL="QN8W" authorityName="Bates" authorityYear="1887" box="[769,990,620,644]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="ardens">
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<emphasis id="E6F54100FFC4FFE6FCBB56771DC4FD7F" box="[769,990,620,644]" italics="true" pageId="5" pageNumber="456">Canthidium ardens</emphasis>
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</taxonomicName>
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]. Saturate viridi-aeneum,
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<taxonomicName id="1381E691FFC4FFE6FAAB56771B8DFD7E" ID-CoL="QND8" authorityName="Preudhomme de Borre" authorityYear="1886" box="[1297,1431,620,645]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="splendidum">splendidum</taxonomicName>
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; corpore subtus et pedibus concoloribus”) and treated the taxon as a “local variety” of
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<taxonomicName id="1381E691FFC4FFE6FBDD568B1ACEFD53" ID-CoL="QN8W" authorityName="Bates" authorityYear="1887" box="[1127,1236,656,680]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="ardens">
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<emphasis id="E6F54100FFC4FFE6FBDD568B1ACEFD53" box="[1127,1236,656,680]" italics="true" pageId="5" pageNumber="456">C. ardens</emphasis>
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</taxonomicName>
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differing only by colour. Since Kohlmann & Solís (2004a), in their revision of the
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<taxonomicName id="1381E691FFC4FFE6FC3656AF1A0AFD37" ID-CoL="3GVY" authorityName="Erichson" authorityYear="1847" box="[908,1040,692,716]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus">
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<emphasis id="E6F54100FFC4FFE6FC3656AF1A0AFD37" box="[908,1040,692,716]" italics="true" pageId="5" pageNumber="456">Canthidium</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="AC96DD82FFC4FFE6FBE056AF1AC3FD36" box="[1114,1241,692,717]" name="Costa Rica" pageId="5" pageNumber="456">Costa Rica</collectingCountry>
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, considered both green and red specimens as conspecifics, and because the only difference noted by Bates concerns the green colouration of his specimen, I chose to treat
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<taxonomicName id="1381E691FFC4FFE6FD2B56E71D0FFCEF" ID-CoL="5DQ9B" box="[657,789,764,788]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="mutatum">
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<emphasis id="E6F54100FFC4FFE6FD2B56E71D0FFCEF" box="[657,789,764,788]" italics="true" pageId="5" pageNumber="456">C. mutatum</emphasis>
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</taxonomicName>
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as a junior synonym of
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<taxonomicName id="1381E691FFC4FFE6FB9156E71A82FCEF" ID-CoL="QN8W" authorityName="Bates" authorityYear="1887" box="[1067,1176,764,788]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="ardens">
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<emphasis id="E6F54100FFC4FFE6FB9156E71A82FCEF" box="[1067,1176,764,788]" italics="true" pageId="5" pageNumber="456">C. ardens</emphasis>
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</taxonomicName>
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].
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<emphasis id="E6F54100FFC4FFE6FB0B56E71B4EFCEE" bold="true" box="[1201,1364,764,789]" pageId="5" pageNumber="456">
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<typeStatus id="0B3A23B0FFC4FFE6FB0B56E71AF6FCEE" box="[1201,1260,764,789]" pageId="5" pageNumber="456">Type</typeStatus>
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locality:
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</emphasis>
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<collectingCountry id="AC96DD82FFC4FFE6FAE656E71ED4FCC3" name="Costa Rica" pageId="5" pageNumber="456">Costa Rica</collectingCountry>
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.
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<emphasis id="E6F54100FFC4FFE6FF6257041F92FCC3" bold="true" box="[216,392,799,824]" pageId="5" pageNumber="456">
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<typeStatus id="0B3A23B0FFC4FFE6FF6257041F09FCC3" box="[216,275,799,824]" pageId="5" pageNumber="456">Type</typeStatus>
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material:
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</emphasis>
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<typeStatus id="0B3A23B0FFC4FFE6FE3557041FEDFCC3" box="[399,503,799,824]" pageId="5" pageNumber="456" type="holotype">Holotype</typeStatus>
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unsexed (MNHN—ex Bates collection).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="D43E9D12FFC4FFE6FF7D575F1AFAFC7B" blockId="5.[151,1437,367,896]" pageId="5" pageNumber="456">
|
||
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|
<treatmentCitationGroup id="F491BA3CFFC4FFE6FF7D575F1AFAFC7B" pageId="5" pageNumber="456">
|
||
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|
<taxonomicName id="1381E691FFC4FFE6FF7D575F1C93FCA7" ID-CoL="5X9GL" authority="Bates, 1887: 48" authorityName="Bates" authorityPageNumber="48" authorityYear="1887" box="[199,649,836,860]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="rhodopus">
|
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<emphasis id="E6F54100FFC4FFE6FF7D575F1FDBFCA7" box="[199,449,836,860]" italics="true" pageId="5" pageNumber="456">Canthidium rhodopus</emphasis>
|
||
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|
<treatmentCitation id="5520BB03FFC4FFE6FE74575E1C93FCA7" author="" box="[462,649,836,860]" page="48" pageId="5" pageNumber="456">
|
||
|
|
<bibRefCitation id="B010E0E3FFC4FFE6FE74575E1C46FCA7" box="[462,604,836,860]" pageId="5" pageNumber="456" refId="ref25529" refString="Bates, H. W. (1887) Tribe Lamellicornia. In: Bates, H. W. (1886 - 1890) Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. Taylor and Francis, London, pp. 25 - 64, pls. 2 - 3. [publication date taken from Bousquet 2016]" type="journal article">Bates, 1887</bibRefCitation>
|
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: 48
|
||
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</treatmentCitation>
|
||
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</taxonomicName>
|
||
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|
[synonymized by Kohlmann & Solís 2006a: 244].
|
||
|
|
<emphasis id="E6F54100FFC4FFE6FB4E575F1B86FCA6" bold="true" box="[1268,1436,836,861]" pageId="5" pageNumber="456">Type locality:</emphasis>
|
||
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|
Guatemala:
|
||
|
|
<materialsCitation id="64E9974FFFC4FFE6FEA557731AC6FC7B" ID-GBIF-Occurrence="1828997597" box="[287,1244,871,896]" collectionCode="BMNH" country="Guatemala" county="Two" location="Zapote Bobal" municipality="Zapote" pageId="5" pageNumber="456" specimenCount="2" stateProvince="Peten" typeStatus="syntype">
|
||
|
|
<collectingRegion id="164553F0FFC4FFE6FEA557731F78FC7B" box="[287,354,872,896]" country="Guatemala" name="Peten" pageId="5" pageNumber="456">Petén</collectingRegion>
|
||
|
|
:
|
||
|
|
<location id="D15ECBC9FFC4FFE6FED657731C1EFC7B" LSID="urn:lsid:plazi:treatment:5C282C04FFC4FFE6FF2D55741AFAFC7B:D15ECBC9FFC4FFE6FED657731C1EFC7B" box="[364,516,871,896]" country="Guatemala" county="Two" municipality="Zapote" name="Zapote Bobal" pageId="5" pageNumber="456" stateProvince="Peten">Zapote Bobal</location>
|
||
|
|
(“
|
||
|
|
<collectingCountry id="AC96DD82FFC4FFE6FDA7577C1C87FC7B" box="[541,669,871,896]" name="Guatemala" pageId="5" pageNumber="456">Guatemala</collectingCountry>
|
||
|
|
,
|
||
|
|
<collectingMunicipality id="345A0768FFC4FFE6FD1257731CE7FC7B" box="[680,765,872,896]" pageId="5" pageNumber="456">Zapote</collectingMunicipality>
|
||
|
|
”).
|
||
|
|
<emphasis id="E6F54100FFC4FFE6FCA3577C1DD3FC7B" bold="true" box="[793,969,871,896]" pageId="5" pageNumber="456">
|
||
|
|
<typeStatus id="0B3A23B0FFC4FFE6FCA3577C1D4EFC7B" box="[793,852,871,896]" pageId="5" pageNumber="456">Type</typeStatus>
|
||
|
|
material:
|
||
|
|
</emphasis>
|
||
|
|
<collectingCounty id="3D5FE59EFFC4FFE6FC6A57731A1BFC84" box="[976,1025,872,895]" pageId="5" pageNumber="456">Two</collectingCounty>
|
||
|
|
<typeStatus id="0B3A23B0FFC4FFE6FBB257731A70FC7B" box="[1032,1130,872,896]" pageId="5" pageNumber="456" type="syntype">syntypes</typeStatus>
|
||
|
|
(
|
||
|
|
<collectionCode id="B29005D7FFC4FFE6FBC357731AC9FC7B" box="[1145,1235,872,896]" pageId="5" pageNumber="456">BMNH</collectionCode>
|
||
|
|
)
|
||
|
|
</materialsCitation>
|
||
|
|
.
|
||
|
|
</treatmentCitationGroup>
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
</treatment>
|
||
|
|
</document>
|