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<document id="7D61F5C7AAF8EF066B20B7791F4C3919" ID-CLB-Dataset="161516" ID-DOI="10.11646/zootaxa.5195.1.5" ID-GBIF-Dataset="94f5f3e5-eb0a-4cb0-8f6c-9e1391430140" ID-ISSN="1175-5326" ID-Zenodo-Dep="7180668" ID-ZooBank="7BCF5C20-88D0-4740-8698-87919D9F8D58" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665395377019" checkinUser="plazi" docAuthor="Fanti, Fabrizio &amp; Pankowski, Maximilian G." docDate="2022" docId="561C87D12230DD43FF64FC85FBE8FD9D" docLanguage="en" docName="zootaxa.5195.1.5.pdf" docOrigin="Zootaxa 5195 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Silis (Silis) boninoi FANTI &amp; M. G. PANKOWSKI 2022, sp. nov." docType="treatment" docVersion="5" lastPageNumber="90" masterDocId="AA25FFA92231DD40FFF3FFB2FFE2FF84" masterDocTitle="A new fossil Silis Charpentier, 1825 from Baltic amber (Cantharidae, Silinae, Silini)" masterLastPageNumber="91" masterPageNumber="87" pageNumber="88" updateTime="1699407079626" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="4B1493F49733597E5F1AF95E0B0D57D6">
<mods:title id="474C0C6435507BBF7192230EBB555C73">A new fossil Silis Charpentier, 1825 from Baltic amber (Cantharidae, Silinae, Silini)</mods:title>
</mods:titleInfo>
<mods:name id="62D1C40F338FEDF8B57F0347985C3E94" type="personal">
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<mods:namePart id="6CA54CFB6D878432E8090BA1E1305DC2">Fanti, Fabrizio</mods:namePart>
<mods:affiliation id="3B3AE08D4F5FBDA00590DE05B97BC095">Via del Tamburino 69, I- 53040 Piazze (SI), Italy.</mods:affiliation>
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<mods:namePart id="3DE8E621C56A46FA10F4C2D4A26CA23B">Pankowski, Maximilian G.</mods:namePart>
<mods:affiliation id="B0EE026092430637FD0478D805E2242F">16405 Fox Valley Terrace, Rockville, Maryland 20853, USA.</mods:affiliation>
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<mods:title id="9B4AC93FC452F1C271D03D052D2F3131">Zootaxa</mods:title>
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<mods:part id="9C69937CEC45694ABE656E6D1AF1F029">
<mods:date id="A8CB26527C91D7F1FC6D44F3A394F9E0">2022</mods:date>
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<mods:number id="2B4BCE4B53F8BE3D9FF47C0C3411C034">2022-10-10</mods:number>
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<mods:number id="47D327FC9E837D10EB3979EFFE7486A8">5195</mods:number>
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<mods:identifier id="3A1131F852FAEC9B0DF5436BF97CC655" type="CLB-Dataset">161516</mods:identifier>
<mods:identifier id="C154DF475DB936A49D357DB6F659460E" type="DOI">10.11646/zootaxa.5195.1.5</mods:identifier>
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<treatment id="561C87D12230DD43FF64FC85FBE8FD9D" ID-DOI="http://doi.org/10.5281/zenodo.7180662" ID-GBIF-Taxon="202401688" ID-Zenodo-Dep="7180662" LSID="urn:lsid:plazi:treatment:561C87D12230DD43FF64FC85FBE8FD9D" httpUri="http://treatment.plazi.org/id/561C87D12230DD43FF64FC85FBE8FD9D" lastPageId="3" lastPageNumber="90" pageId="1" pageNumber="88">
<subSubSection id="96AF654C2230DD41FF64FC85FC91FCD6" box="[151,883,823,850]" pageId="1" pageNumber="88" type="nomenclature">
<paragraph id="DE0A36C72230DD41FF64FC85FC91FCD6" blockId="1.[151,883,823,886]" box="[151,883,823,850]" pageId="1" pageNumber="88">
<heading id="854281AB2230DD41FF64FC85FC91FCD6" bold="true" box="[151,883,823,850]" fontSize="11" level="1" pageId="1" pageNumber="88" reason="1">
<emphasis id="ECC1EAD52230DD41FF64FC85FC91FCD6" bold="true" box="[151,883,823,850]" pageId="1" pageNumber="88">
<taxonomicName id="19B54D442230DD41FF64FC85FCECFCD6" authority="FANTI &amp; M. G. PANKOWSKI" authorityName="FANTI &amp; M. G. PANKOWSKI" authorityYear="2022" box="[151,782,823,850]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="boninoi" status="sp. nov." subGenus="Silis">
<emphasis id="ECC1EAD52230DD41FF64FC85FF29FCD6" bold="true" box="[151,203,823,850]" italics="true" pageId="1" pageNumber="88">Silis</emphasis>
(
<emphasis id="ECC1EAD52230DD41FF2FFC85FEF2FCD6" bold="true" box="[220,272,823,850]" italics="true" pageId="1" pageNumber="88">Silis</emphasis>
)
<emphasis id="ECC1EAD52230DD41FED1FC85FE9DFCD6" bold="true" box="[290,383,823,850]" italics="true" pageId="1" pageNumber="88">boninoi</emphasis>
FANTI &amp; M. G. PANKOWSKI
</taxonomicName>
<taxonomicNameLabel id="F7F257AE2230DD41FCE6FC8AFC91FCD6" box="[789,883,824,850]" pageId="1" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="96AF654C2230DD41FF64FCEEFEF5F82A" pageId="1" pageNumber="88" type="description">
<paragraph id="DE0A36C72230DD41FF64FCEEFF09FCF2" blockId="1.[151,883,823,886]" box="[151,235,860,886]" pageId="1" pageNumber="88">
(
<figureCitation id="468E2A422230DD41FF6CFCEEFF02FCF2" box="[159,224,860,886]" captionStart="FIGURE 1" captionStartId="2.[151,250,1329,1353]" captionTargetBox="[151,1436,179,1304]" captionTargetId="figure-22@2.[151,1436,179,1304]" captionTargetPageId="2" captionText="FIGURE 1. Silis (Silis) boninoi sp. nov. in Baltic amber. A: Holotype, dorsal view. B: Holotype, ventral view. C: Holotype, detail of prolegs. D: Holotype, detail of pronotum." figureDoi="http://doi.org/10.5281/zenodo.7180670" httpUri="https://zenodo.org/record/7180670/files/figure.png" pageId="1" pageNumber="88">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="DE0A36C72230DD41FF64FC16FDC8FC66" blockId="1.[151,1437,932,2039]" pageId="1" pageNumber="88">
<emphasis id="ECC1EAD52230DD41FF64FC16FECBFC3A" bold="true" box="[151,297,932,958]" pageId="1" pageNumber="88">Description.</emphasis>
Adult, winged, male. Body length:
<quantity id="194D9B222230DD41FD4CFC16FCFBFC3B" box="[703,793,932,959]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="88" unit="mm" value="2.7">2.7 mm</quantity>
; elytra: approximately
<quantity id="194D9B222230DD41FBEEFC16FB96FC3B" box="[1053,1140,932,959]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="88" unit="mm" value="2.4">2.4 mm</quantity>
long, and 1.0 mm wide at apex. Entirely blackish-dark brown.
</paragraph>
<paragraph id="DE0A36C72230DD41FF34FC5EFCC0F89A" blockId="1.[151,1437,932,2039]" pageId="1" pageNumber="88">Head large, slightly narrower than pronotum, smooth, equipped with several long setae, largely covered by pronotum. Eyes rounded and protruded, wide, inserted in upper and lateral part of the head. Maxillary palpi 4- segmented, unequal in length with last palpomere very elongated and securiform. Labial palps 3-segmented with last palpomere securiform. Antennal insertions and eyes not juxtaposed, antennae surpassing two-thirds of the elytra, 11-segmented, filiform, each antennomere covered with several setae; scape elongated, slightly club-shaped; antennomere II (pedicel) short, about 2.1 times shorter than scape; antennomeres III–V subequal in length, longer than antennomere II, rather robust; antennomeres VI–IX subequal in length, slightly longer than antennomeres III–V; antennomere X very slightly shorter and more slender than previous ones; antennomere XI filiform, oblong with extremely thin and pointed apex. Pronotum strongly transverse, surface undulating with concavities (pores) and equipped with sparse long setae, apparently without punctuation; anterior margin curved and protruding in the middle; sides strongly sinuous, with two very short processes on each side: Posterior processes are thicker and wider than anterior ones and rounded apically; anterior processes are shorter than posterior ones and rounded apically; posterior margin irregular and sinuous with two expansions in the middle. Scutellum triangular with rounded apex. Elytra wider than pronotum, elongated and surpassing last abdominal segments, enlarged at humeri and apex, restricted in middle, apex strongly rounded, surface with very superficial punctation and several very long setae. Posterior wings transparent, completely covered by elytra but just barely shorter than them. Metasternum very elongated with a strongly rounded posterior margin, abdominal segments transverse and pubescent. Legs relatively short, robust and strongly pubescent; coxae massive; trochanters elongated and triangular-shaped with rounded apex; femora enlarged and slightly curved; tibiae cylindrical, with one apical and robust spur, protibiae as long as profemora, mesotibiae slightly shorter than mesofemora, metatibiae just slightly shorter than metafemora. Tarsal formula 5-5-5; first tarsomere elongated; second tarsomere about 1.2 times shorter than first; third tarsomere triangular and shorter than tarsomere II; fourth tarsomere deeply bilobed at sides; fifth tarsomere very elongated, slender, flat and curved; claws simple without basal tooth.</paragraph>
<paragraph id="DE0A36C72230DD41FF34F89BFEF5F82A" blockId="1.[151,1437,932,2039]" pageId="1" pageNumber="88">
Note. Female unknown. The specimen is a male, defined on the basis of long antennae and lateral sides of pronotum toothed. Sexual dimorphism is supposed because females of the genus
<taxonomicName id="19B54D442230DD41FBAFF8FEFB6EF8E2" authorityName="Charpentier" authorityYear="1825" box="[1116,1164,1868,1894]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="ECC1EAD52230DD41FBAFF8FEFB6EF8E2" box="[1116,1164,1868,1894]" italics="true" pageId="1" pageNumber="88">Silis</emphasis>
</taxonomicName>
are characterized by a pronotum with sides without appendages or with very small teeth, and have shorter antennae and a wider abdomen than males.
</paragraph>
</subSubSection>
<subSubSection id="96AF654C2230DD41FF34F80AFE34F872" pageId="1" pageNumber="88" type="etymology">
<paragraph id="DE0A36C72230DD41FF34F80AFE34F872" blockId="1.[151,1437,932,2039]" pageId="1" pageNumber="88">
<emphasis id="ECC1EAD52230DD41FF34F80AFEAAF856" bold="true" box="[199,328,1976,2002]" pageId="1" pageNumber="88">Etymology</emphasis>
. Named in honor of Enrico Bonino for donating the described specimen, photographing it and kindly giving us access to it.
</paragraph>
</subSubSection>
<caption id="8ACA664F2233DD42FF64FA83FD75FAE9" ID-DOI="http://doi.org/10.5281/zenodo.7180670" ID-Zenodo-Dep="7180670" httpUri="https://zenodo.org/record/7180670/files/figure.png" pageId="2" pageNumber="89" startId="2.[151,250,1329,1353]" targetBox="[151,1436,179,1304]" targetPageId="2">
<paragraph id="DE0A36C72233DD42FF64FA83FD75FAE9" blockId="2.[151,1437,1329,1390]" pageId="2" pageNumber="89">
<emphasis id="ECC1EAD52233DD42FF64FA83FEF7FACD" bold="true" box="[151,277,1329,1353]" pageId="2" pageNumber="89">FIGURE 1.</emphasis>
<taxonomicName id="19B54D442233DD42FEEFFA80FE03FACE" authority="Fanti &amp; Pankowski, 2022" authorityName="FANTI &amp; M. G. PANKOWSKI" authorityYear="2022" box="[284,481,1330,1354]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="boninoi" status="sp. nov." subGenus="Silis">
<emphasis id="ECC1EAD52233DD42FEEFFA80FEAAFACE" box="[284,328,1330,1354]" italics="true" pageId="2" pageNumber="89">Silis</emphasis>
(
<emphasis id="ECC1EAD52233DD42FEABFA80FE66FACE" box="[344,388,1330,1354]" italics="true" pageId="2" pageNumber="89">Silis</emphasis>
)
<emphasis id="ECC1EAD52233DD42FE67FA80FE03FACE" box="[404,481,1330,1354]" italics="true" pageId="2" pageNumber="89">boninoi</emphasis>
</taxonomicName>
<emphasis id="ECC1EAD52233DD42FE19FA80FDDEFACE" bold="true" box="[490,572,1330,1354]" pageId="2" pageNumber="89">
<taxonomicNameLabel id="F7F257AE2233DD42FE19FA80FDDEFACE" box="[490,572,1330,1354]" pageId="2" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Baltic amber. A: Holotype, dorsal view. B: Holotype, ventral view. C: Holotype, detail of prolegs. D: Holotype, detail of pronotum.
</paragraph>
</caption>
<subSubSection id="96AF654C2233DD42FF34FA2EFCDFF80E" pageId="2" pageNumber="89" type="materials_examined">
<paragraph id="DE0A36C72233DD42FF34FA2EFEA0FA5E" blockId="2.[151,1437,1436,2039]" pageId="2" pageNumber="89">
<materialsCitation id="6EDD3C9A2233DD42FF34FA2EFEA0FA5E" ID-GBIF-Occurrence="3922661301" country="Mexico" county="Cancun" location="Male" municipality="Past Museum" pageId="2" pageNumber="89" specimenCount="1" typeStatus="holotype">
<emphasis id="ECC1EAD52233DD42FF34FA2EFED9FA32" bold="true" box="[199,315,1436,1462]" pageId="2" pageNumber="89">
<typeStatus id="010E88652233DD42FF34FA2EFED4FA32" box="[199,310,1436,1462]" pageId="2" pageNumber="89" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="DB6A601C2233DD42FEB0FA2EFE9DFA32" LSID="urn:lsid:plazi:treatment:561C87D12230DD43FF64FC85FBE8FD9D:DB6A601C2233DD42FEB0FA2EFE9DFA32" box="[323,383,1436,1462]" country="Mexico" county="Cancun" municipality="Past Museum" name="Male" pageId="2" pageNumber="89">Male</location>
, in
<location id="DB6A601C2233DD42FE59FA2EFE0FFA32" LSID="urn:lsid:plazi:treatment:561C87D12230DD43FF64FC85FBE8FD9D:DB6A601C2233DD42FE59FA2EFE0FFA32" box="[426,493,1436,1462]" country="Mexico" county="Cancun" municipality="Past Museum" name="Baltic" pageId="2" pageNumber="89">Baltic</location>
amber, deposited at the
<location id="DB6A601C2233DD42FCF0FA2EFCDFFA32" LSID="urn:lsid:plazi:treatment:561C87D12230DD43FF64FC85FBE8FD9D:DB6A601C2233DD42FCF0FA2EFCDFFA32" box="[771,829,1436,1462]" country="Mexico" county="Cancun" municipality="Past Museum" name="Back" pageId="2" pageNumber="89">Back</location>
to the
<collectingMunicipality id="3E6EACBD2233DD42FC7EFA2EFBCBFA32" box="[909,1065,1436,1462]" pageId="2" pageNumber="89">Past Museum</collectingMunicipality>
,
<collectingCounty id="376B4E4B2233DD42FBC6FA2EFB6FFA32" box="[1077,1165,1436,1462]" pageId="2" pageNumber="89">Cancún</collectingCounty>
,
<collectingCountry id="A6A276572233DD42FB6AFA2EFB0DFA32" box="[1177,1263,1436,1462]" name="Mexico" pageId="2" pageNumber="89">Mexico</collectingCountry>
with accession no. BPM-1111.
</materialsCitation>
</paragraph>
<paragraph id="DE0A36C72233DD42FF34FA56FBA3FA7A" blockId="2.[151,1437,1436,2039]" box="[199,1089,1508,1534]" pageId="2" pageNumber="89">
<emphasis id="ECC1EAD52233DD42FF34FA56FE84FA7A" bold="true" box="[199,358,1508,1534]" pageId="2" pageNumber="89">Type locality.</emphasis>
Yantarny mine, Sambian Peninsula,
<collectingRegion id="1C71F8252233DD42FCF0FA56FC01FA7A" box="[771,995,1508,1534]" country="Russia" name="Kaliningrad" pageId="2" pageNumber="89">Kaliningrad Region</collectingRegion>
,
<collectingCountry id="A6A276572233DD42FC1DFA56FBDEFA7A" box="[1006,1084,1508,1534]" name="Russia" pageId="2" pageNumber="89">Russia</collectingCountry>
.
</paragraph>
<paragraph id="DE0A36C72233DD42FF34F9BAFC0AF9A6" blockId="2.[151,1437,1436,2039]" box="[199,1000,1544,1571]" pageId="2" pageNumber="89">
<emphasis id="ECC1EAD52233DD42FF34F9BAFE8EF9A6" bold="true" box="[199,364,1544,1570]" pageId="2" pageNumber="89">Type horizon.</emphasis>
Middle Eocene: Bartonian-Priabonian (45.0–38.0 Mya).
</paragraph>
<paragraph id="DE0A36C72233DD42FF34F99EFCD3F9C2" blockId="2.[151,1437,1436,2039]" box="[199,817,1580,1606]" pageId="2" pageNumber="89">
<emphasis id="ECC1EAD52233DD42FF34F99EFE90F9C2" bold="true" box="[199,370,1580,1606]" pageId="2" pageNumber="89">Syninclusions.</emphasis>
A few botanical fragments (trichomes).
</paragraph>
<paragraph id="DE0A36C72233DD42FF34F9E2FE89F97E" blockId="2.[151,1437,1436,2039]" pageId="2" pageNumber="89">
<emphasis id="ECC1EAD52233DD42FF34F9E2FE31F9EE" bold="true" box="[199,467,1616,1642]" pageId="2" pageNumber="89">Systematic placement.</emphasis>
The specimen is clearly a new species belonging to the subfamily
<taxonomicName id="19B54D442233DD42FB27F9E2FAC0F9EE" authorityName="Mulsant" authorityYear="1862" box="[1236,1314,1616,1642]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="subFamily" subFamily="Silinae">Silinae</taxonomicName>
and to the genus
<taxonomicName id="19B54D442233DD42FF17F9C6FEF6F90A" authorityName="Charpentier" authorityYear="1825" box="[228,276,1652,1678]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="ECC1EAD52233DD42FF17F9C6FEF6F90A" box="[228,276,1652,1678]" italics="true" pageId="2" pageNumber="89">Silis</emphasis>
</taxonomicName>
(
<bibRefCitation id="BA244B362233DD42FED4F9C6FE02F90A" author="Brancucci, M." box="[295,480,1652,1678]" pageId="2" pageNumber="89" pagination="215 - 388" refId="ref2465" refString="Brancucci, M. (1980) Morphologie comparee, evolution et systematique des Cantharidae (Insecta: Coleoptera). Entomologica Basiliensia, 5, 215 - 388." type="journal article" year="1980">Brancucci 1980</bibRefCitation>
;
<bibRefCitation id="BA244B362233DD42FE02F9C6FD4BF90A" author="Kazantsev, S. V." box="[497,681,1652,1678]" pageId="2" pageNumber="89" pagination="1229 - 1232" refId="ref3018" refString="Kazantsev, S. V. (1997) A New Species of the Genus Silis (Coleoptera, Cantharidae) from Siberia, with Notes on Zoogeography of the Genus. Zoologicheskii Zhurnal, 76 (10), 1214 - 1217. [in Russian with English summary, translated into English in 1997: Entomological Review, 77 (9), 1229 - 1232.]" type="journal article" year="1997">Kazantsev 1997</bibRefCitation>
) based on its maxillary palpi sub-equal in length with their last palpomere securiform, elytra elongated, tarsomeres IV wider than other tarsomeres, pores on the anterior part of the pronotum, the internal claw simple, tibial spurs conspicuous and the lateral section of its pronotum having two lobes on each side.
</paragraph>
<paragraph id="DE0A36C72233DD42FF23F8B6FCDFF80E" blockId="2.[151,1437,1436,2039]" pageId="2" pageNumber="89">
Genera that appear most similar to
<taxonomicName id="19B54D442233DD42FD92F8B6FD73F89A" authorityName="Charpentier" authorityYear="1825" box="[609,657,1796,1822]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="ECC1EAD52233DD42FD92F8B6FD73F89A" box="[609,657,1796,1822]" italics="true" pageId="2" pageNumber="89">Silis</emphasis>
</taxonomicName>
are distinguished by a few different characters. The genus
<taxonomicName id="19B54D442233DD42FAC7F8B6FA7EF89A" authorityName="Wittmer" authorityYear="1978" box="[1332,1436,1796,1822]" class="Insecta" family="Cantharidae" genus="Podosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="ECC1EAD52233DD42FAC7F8B6FA7EF89A" box="[1332,1436,1796,1822]" italics="true" pageId="2" pageNumber="89">Podosilis</emphasis>
</taxonomicName>
has a complex pronotal armature at the sides and a cleft outer claw in males (
<bibRefCitation id="BA244B362233DD42FC15F89AFB7FF8C6" author="Kazantsev, S. V." box="[998,1181,1832,1858]" pageId="2" pageNumber="89" pagination="1229 - 1232" refId="ref3018" refString="Kazantsev, S. V. (1997) A New Species of the Genus Silis (Coleoptera, Cantharidae) from Siberia, with Notes on Zoogeography of the Genus. Zoologicheskii Zhurnal, 76 (10), 1214 - 1217. [in Russian with English summary, translated into English in 1997: Entomological Review, 77 (9), 1229 - 1232.]" type="journal article" year="1997">Kazantsev 1997</bibRefCitation>
,
<bibRefCitation id="BA244B362233DD42FB54F89AFB00F8C6" author="Kazantsev, S. V." box="[1191,1250,1832,1858]" pageId="2" pageNumber="89" pagination="158 - 164" refId="ref3127" refString="Kazantsev, S. V. (2019) New species of Podosilis Wittmer, 1978 (Coleoptera: Cantharidae) from China and Indochina. Russian Entomological Journal, 28 (2), 158 - 164." type="journal article" year="2019">2019</bibRefCitation>
,
<bibRefCitation id="BA244B362233DD42FB1FF89AFACBF8C6" author="Kazantsev, S. V." box="[1260,1321,1832,1858]" pageId="2" pageNumber="89" pagination="260 - 268" refId="ref3166" refString="Kazantsev, S. V. (2020) New Baltic amber soldier beetles (Insecta: Coleoptera: Cantharidae) with some taxonomic notes. Palaeoentomology, 3 (3), 260 - 268. https: // doi. org / 10.11646 / palaeoentomology. 3.3.7" type="journal article" year="2020">2020</bibRefCitation>
).
<taxonomicName id="19B54D442233DD42FAC9F89AFA7EF8C6" authorityName="Kazantsev" authorityYear="2011" box="[1338,1436,1832,1858]" class="Insecta" family="Cantharidae" genus="Autosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="ECC1EAD52233DD42FAC9F89AFA7EF8C6" box="[1338,1436,1832,1858]" italics="true" pageId="2" pageNumber="89">Autosilis</emphasis>
</taxonomicName>
features the anterior half of its mesonotum smooth without clear pores, and the anal ring in the male has a tergite and sternite that are roughly equally wide (
<bibRefCitation id="BA244B362233DD42FD84F8C2FCD2F80E" author="Kazantsev, S. V." box="[631,816,1904,1930]" pageId="2" pageNumber="89" pagination="1229 - 1232" refId="ref3018" refString="Kazantsev, S. V. (1997) A New Species of the Genus Silis (Coleoptera, Cantharidae) from Siberia, with Notes on Zoogeography of the Genus. Zoologicheskii Zhurnal, 76 (10), 1214 - 1217. [in Russian with English summary, translated into English in 1997: Entomological Review, 77 (9), 1229 - 1232.]" type="journal article" year="1997">Kazantsev 1997</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="96AF654C2233DD43FF34F826FBD9FE0D" lastPageId="3" lastPageNumber="90" pageId="2" pageNumber="89" type="diagnosis">
<paragraph id="DE0A36C72233DD43FF34F826FD57FEC5" blockId="2.[151,1437,1436,2039]" lastBlockId="3.[151,1437,151,537]" lastPageId="3" lastPageNumber="90" pageId="2" pageNumber="89">
<emphasis id="ECC1EAD52233DD42FF34F826FE32F82A" bold="true" box="[199,464,1940,1966]" pageId="2" pageNumber="89">Differential diagnosis.</emphasis>
<taxonomicName id="19B54D442233DD42FE29F826FD88F82A" authority="Fanti &amp; Pankowski, 2022" authorityName="FANTI &amp; M. G. PANKOWSKI" authorityYear="2022" box="[474,618,1940,1966]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="boninoi" status="sp. nov.">
<emphasis id="ECC1EAD52233DD42FE29F826FD88F82A" box="[474,618,1940,1966]" italics="true" pageId="2" pageNumber="89">Silis boninoi</emphasis>
</taxonomicName>
<emphasis id="ECC1EAD52233DD42FD80F826FD2DF82A" bold="true" box="[627,719,1940,1966]" pageId="2" pageNumber="89">
<taxonomicNameLabel id="F7F257AE2233DD42FD80F826FD2DF82A" box="[627,719,1940,1966]" pageId="2" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="19B54D442233DD42FC98F826FAE4F82A" authority="Parisi &amp; Fanti, 2019" authorityName="Parisi &amp; Fanti" authorityYear="2019" box="[875,1286,1940,1966]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="lombardii">
<emphasis id="ECC1EAD52233DD42FC98F826FBF3F82A" box="[875,1041,1940,1966]" italics="true" pageId="2" pageNumber="89">Silis lombardii</emphasis>
<bibRefCitation id="BA244B362233DD42FBE8F826FAE4F82A" author="Parisi, F. &amp; Fanti, F." box="[1051,1286,1940,1966]" pageId="2" pageNumber="89" pagination="119 - 123" refId="ref3320" refString="Parisi, F. &amp; Fanti, F. (2019) On the first Silis Charpentier, 1825 from Baltic amber (Coleoptera, Cantharidae). Fossil Record, 22 (2), 119 - 123. https: // doi. org / 10.5194 / fr- 22 - 119 - 2019" type="journal article" year="2019">Parisi &amp; Fanti, 2019</bibRefCitation>
</taxonomicName>
(from Baltic amber) based on the new species’ pronotum that is much more transverse and with shorter pronotal lobes on its sides, and by the new species’ significantly smaller size:
<quantity id="194D9B222233DD42FCE1F86EFC85F872" box="[786,871,2012,2038]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="89" unit="mm" value="2.7">2.7 mm</quantity>
vs.
<quantity id="194D9B222233DD42FC67F86EFC08F872" box="[916,1002,2012,2039]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="89" unit="mm" value="5.2">5.2 mm</quantity>
for
<taxonomicName id="19B54D442233DD42FBEBF86EFA7BF872" authority="(Parisi &amp; Fanti 2019)" baseAuthorityName="Parisi &amp; Fanti" baseAuthorityYear="2019" box="[1048,1433,2012,2038]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="lombardii">
<emphasis id="ECC1EAD52233DD42FBEBF86EFB42F872" box="[1048,1184,2012,2038]" italics="true" pageId="2" pageNumber="89">S. lombardii</emphasis>
(
<bibRefCitation id="BA244B362233DD42FB5CF86EFA73F872" author="Parisi, F. &amp; Fanti, F." box="[1199,1425,2012,2038]" pageId="2" pageNumber="89" pagination="119 - 123" refId="ref3320" refString="Parisi, F. &amp; Fanti, F. (2019) On the first Silis Charpentier, 1825 from Baltic amber (Coleoptera, Cantharidae). Fossil Record, 22 (2), 119 - 123. https: // doi. org / 10.5194 / fr- 22 - 119 - 2019" type="journal article" year="2019">Parisi &amp; Fanti 2019</bibRefCitation>
)
</taxonomicName>
. In addition,
<taxonomicName id="19B54D442232DD43FEECFF25FD6FFF35" authority="Wittmer, 1963" authorityName="Wittmer" authorityYear="1963" box="[287,653,151,177]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="chiapasensis">
<emphasis id="ECC1EAD52232DD43FEECFF25FE04FF35" box="[287,486,151,177]" italics="true" pageId="3" pageNumber="90">Silis chiapasensis</emphasis>
<bibRefCitation id="BA244B362232DD43FE1FFF25FD6FFF35" author="Wittmer, W." box="[492,653,151,177]" pageId="3" pageNumber="90" pagination="59" refId="ref3704" refString="Wittmer, W. (1963) A new cantharid from the Chiapas amber of Mexico. University of California Publications in Entomology, 31, 53 + pl. 3 (p. 59)." type="journal article" year="1963">Wittmer, 1963</bibRefCitation>
</taxonomicName>
(found in Mexican
<collectingRegion id="1C71F8252232DD43FC9FFF25FC24FF35" box="[876,966,151,177]" country="Mexico" name="Chiapas" pageId="3" pageNumber="90">Chiapas</collectingRegion>
amber) differs based on its pronotum that is deeply incised at the sides and its lobes that are more curved and longer than on the new species (
<bibRefCitation id="BA244B362232DD43FB1CFF09FA72FF51" author="Wittmer, W." box="[1263,1424,187,213]" pageId="3" pageNumber="90" pagination="59" refId="ref3704" refString="Wittmer, W. (1963) A new cantharid from the Chiapas amber of Mexico. University of California Publications in Entomology, 31, 53 + pl. 3 (p. 59)." type="journal article" year="1963">Wittmer 1963</bibRefCitation>
). Finally,
<taxonomicName id="19B54D442232DD43FF07FF6DFD08FF7D" authority="Fanti &amp; M. G. Pankowski 2021" authorityName="Fanti &amp; M. G. Pankowski" authorityYear="2021" box="[244,746,223,249]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="curleri">
<emphasis id="ECC1EAD52232DD43FF07FF6DFE9BFF7D" box="[244,377,223,249]" italics="true" pageId="3" pageNumber="90">Silis curleri</emphasis>
Fanti &amp; M. G. Pankowski 2021
</taxonomicName>
and
<taxonomicName id="19B54D442232DD43FCD7FF6DFAFBFF7D" authority="Fanti &amp; M. G. Pankowski 2021" authorityName="Fanti &amp; M. G. Pankowski" authorityYear="2021" box="[804,1305,223,249]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="hegnai">
<emphasis id="ECC1EAD52232DD43FCD7FF6DFC4BFF7D" box="[804,937,223,249]" italics="true" pageId="3" pageNumber="90">Silis hegnai</emphasis>
Fanti &amp; M. G. Pankowski 2021
</taxonomicName>
(both from Dominican amber) have a very different pronotal shape, with lobes that are longer and more curved, and a less transverse pronotum (
<bibRefCitation id="BA244B362232DD43FE78FE95FD4AFEC5" author="Fanti, F. &amp; Pankowski, M. G." box="[395,680,295,321]" pageId="3" pageNumber="90" pagination="163 - 170" refId="ref2905" refString="Fanti, F. &amp; Pankowski, M. G. (2021) Two new species of soldier beetles (Coleoptera, Cantharidae), the first from the tribe Silini in Dominican amber. Zootaxa, 4996 (1), 163 - 170. https: // doi. org / 10.11646 / zootaxa. 4996.1.7" type="journal article" year="2021">Fanti &amp; Pankowski 2021</bibRefCitation>
).
</paragraph>
<paragraph id="DE0A36C72232DD43FF34FEF9FBD9FE0D" blockId="3.[151,1437,151,537]" pageId="3" pageNumber="90">
The living and widely distributed species
<taxonomicName id="19B54D442232DD43FD6AFEF9FC1EFEE1" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[665,1020,330,357]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="ruficollis">
<emphasis id="ECC1EAD52232DD43FD6AFEF9FCD7FEE1" box="[665,821,331,357]" italics="true" pageId="3" pageNumber="90">Silis ruficollis</emphasis>
(Fabricius, 1775)
</taxonomicName>
has a pronotum with a lateral margin that is less toothed and indented compared to the new fossil species described here.
</paragraph>
</subSubSection>
<subSubSection id="96AF654C2232DD43FF34FE21FBE8FD9D" pageId="3" pageNumber="90" type="discussion">
<paragraph id="DE0A36C72232DD43FF34FE21FBE8FD9D" blockId="3.[151,1437,151,537]" pageId="3" pageNumber="90">
<emphasis id="ECC1EAD52232DD43FF34FE21FED9FE29" bold="true" box="[199,315,403,429]" pageId="3" pageNumber="90">Remarks.</emphasis>
The yellow amber piece has an elongated drop shape and measures approximately 22x
<quantity id="194D9B222232DD43FAE4FE21FA84FE29" box="[1303,1382,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="90" unit="mm" value="11.0">11 mm</quantity>
. The inclusion is complete and well visible. Once trapped, the specimen struggled to get out of the sticky resin and thus created characteristic ripples (“fringes”) in the amber. A photo of this new species is included in Enrico Bonino’s forthcoming book on amber (
<bibRefCitation id="BA244B362232DD43FE2CFE4DFD9FFD9D" author="Bonino, E." box="[479,637,511,537]" pageId="3" pageNumber="90" refId="ref2439" refString="Bonino, E. (2022) AmberArt - a journey between Science and Beauty. Self publishing, 300 pp. [In press]" type="book" year="2022">Bonino, 2022</bibRefCitation>
), expected to be published in 2022.
</paragraph>
</subSubSection>
</treatment>
</document>
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