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<document id="51F6E3BDFD81CD7C1F5BABBE42275E1B" ID-DOI="10.11646/zootaxa.3934.1.1" ID-GBIF-Dataset="0abffe3e-a2c6-4623-b7c6-c8b26cee4a2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="236066" ID-ZooBank="11C2A2CB-30B3-4694-B379-AE9D47332F0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461012229557" checkinUser="plazi" docAuthor="Winterbottom, Richard &amp; Hoese, Douglass F." docDate="2015" docId="5519879AB822F366FF1FFDBE6F76EE2F" docLanguage="en" docName="zt03934p102.pdf" docOrigin="Zootaxa 3934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Trimma hoesei Winterbottom 1984" docType="treatment" docVersion="11" lastPageNumber="36" masterDocId="A920FFE2B803F345FF88FFF66B4DEB78" masterDocTitle="A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species" masterLastPageNumber="102" masterPageNumber="1" pageNumber="34" updateTime="1698567702668" updateUser="plazi">
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<mods:title id="E96F9FBDBF157234226B17C9FE65AD3F">A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species</mods:title>
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<mods:namePart id="B36C6C9B46A2F4EA319F38D0B40A5048">Winterbottom, Richard</mods:namePart>
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<mods:namePart id="9FE56956D7D1F9AC29E7E6DB0826EEBD">Hoese, Douglass F.</mods:namePart>
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<mods:date id="448F02FCFD05BC504EA0AD0001C624B7">2015</mods:date>
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<mods:number id="A75DEBCF854AA89EF8BD137134050EE1">3934</mods:number>
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<treatment id="5519879AB822F366FF1FFDBE6F76EE2F" ID-DOI="http://doi.org/10.5281/zenodo.5621512" ID-GBIF-Taxon="119600478" ID-Zenodo-Dep="5621512" LSID="urn:lsid:plazi:treatment:5519879AB822F366FF1FFDBE6F76EE2F" httpUri="http://treatment.plazi.org/id/5519879AB822F366FF1FFDBE6F76EE2F" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
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<taxonomicName id="1AB04D0FB822F364FF1FFDBE6902E91A" ID-CoL="58V66" authority="Winterbottom, 1984" authorityName="Winterbottom" authorityYear="1984" box="[151,591,584,610]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="hoesei">
<emphasis id="EFC4EA9EB822F364FF1FFDBE6902E91A" bold="true" box="[151,591,584,610]" pageId="33" pageNumber="34">
<emphasis id="EFC4EA9EB822F364FF1FFDBE6A01E91A" bold="true" box="[151,332,584,610]" italics="true" pageId="33" pageNumber="34">Trimma hoesei</emphasis>
<bibRefCitation id="B9214B7DB822F364FEDBFDBE6902E91A" author="Winterbottom" box="[339,591,584,610]" pageId="33" pageNumber="34" refString="Winterbottom, R. (1984) A review of the gobiid fish genus Trimma from the Chagos Archipelago, central Indian Ocean, with the description of seven new species. Canadian Journal of Zoology, 62, 695 - 715." type="journal article" year="1984">Winterbottom, 1984</bibRefCitation>
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<paragraph id="DD0F368CB822F364FF1FFD9C6A30E9FB" blockId="33.[151,591,584,679]" box="[151,381,618,643]" pageId="33" pageNumber="34">Forktail Pygmygoby</paragraph>
<paragraph id="DD0F368CB822F364FF1FFD786AE0E9DE" blockId="33.[151,591,584,679]" box="[151,429,654,679]" pageId="33" pageNumber="34">
<figureCitation id="458B2A09B822F364FF1FFD786BAEE9DF" box="[151,227,654,679]" captionStart="FIGURE 1" captionStartId="7.[151,250,738,760]" captionTargetBox="[151,1435,195,713]" captionTargetId="figure@7.[151,1435,195,713]" captionTargetPageId="7" captionText="FIGURE 1. Dorsal view of heads of A: Trimma caesiura, (26.3 mm SL male, ROM 64124, New Caledonia) to show fleshy interorbital width and interorbital and postorbital trenches; B: T. hoesei (21.7 mm SL female, ROM 83303, Palau) to show wide fleshy interorbital and fleshy median interorbital ridge. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/236067/files/figure.png" pageId="33" pageNumber="34">Figs. 1</figureCitation>
B, 15–16, Pl. 1 H
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B822F364FF1FFD2E69ABE830" pageId="33" pageNumber="34" type="reference_group">
<paragraph id="DD0F368CB822F364FF1FFD2E6919E873" blockId="33.[151,1436,726,840]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="FDA011A2B822F364FF1FFD2E6919E873" pageId="33" pageNumber="34">
<taxonomicName id="1AB04D0FB822F364FF1FFD2E6961E994" ID-CoL="58V66" authority="Winterbottom, 1984:704" authorityName="Winterbottom" authorityPageNumber="704" authorityYear="1984" box="[151,556,726,749]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="hoesei">
<emphasis id="EFC4EA9EB822F364FF1FFD2E6A60E994" box="[151,301,726,749]" italics="true" pageId="33" pageNumber="34">Trimma hoesei</emphasis>
<treatmentCitation id="5C11109DB822F364FEBAFD206961E994" author="Winterbottom" box="[306,556,726,748]" page="704" pageId="33" pageNumber="34" year="1984">
<bibRefCitation id="B9214B7DB822F364FEBAFD20694DE994" author="Winterbottom" box="[306,512,726,748]" pageId="33" pageNumber="34" refString="Winterbottom, R. (1984) A review of the gobiid fish genus Trimma from the Chagos Archipelago, central Indian Ocean, with the description of seven new species. Canadian Journal of Zoology, 62, 695 - 715." type="journal article" year="1984">Winterbottom, 1984</bibRefCitation>
:704
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(Chagos Archipelago, central Indian ocean);
<treatmentCitation id="5C11109DB822F364FC7FFD206F9AE994" author="Allen" box="[1015,1239,726,749]" page="331" pageId="33" pageNumber="34" year="2004">
<bibRefCitation id="B9214B7DB822F364FC7FFD206FE1E994" author="Allen" box="[1015,1196,726,749]" pageId="33" pageNumber="34" refString="Allen, G., Steene, R., Humann, P. &amp; DeLoach, N. (2004) Reef Fish Identification. Tropical Pacific. New World Publications Ltd., Florida, 457 pp." type="book" year="2004">
Allen
<emphasis id="EFC4EA9EB822F364FBBDFD2E6F25E994" box="[1077,1128,726,749]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
, 2004
</bibRefCitation>
:331
</treatmentCitation>
;
<treatmentCitation id="5C11109DB822F364FB69FD206A6AE873" author="Allen" page="940" pageId="33" pageNumber="34" year="2012">
<bibRefCitation id="B9214B7DB822F364FB69FD206BB1E873" author="Allen" pageId="33" pageNumber="34" refString="Allen, G. R. &amp; Erdmann, M. V. (2012) Reef fishes of the East Indies. Vol. III. Tropical Reef Research, Perth, pp. 857 - 1292." type="book chapter" year="2012">Allen &amp; Erdmann, 2012</bibRefCitation>
:940
</treatmentCitation>
(Cocos (Keeling) Ids to
<collectingCountry id="A5A7761CB822F364FDA9FD036904E873" box="[545,585,757,779]" name="Fiji" pageId="33" pageNumber="34">Fiji</collectingCountry>
).
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</paragraph>
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<treatmentCitationGroup id="FDA011A2B822F364FF1FFCE3691FE852" box="[151,594,788,810]" pageId="33" pageNumber="34">
<taxonomicName id="1AB04D0FB822F364FF1FFCE36BABE852" ID-CoL="827C" box="[151,230,789,810]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB822F364FF1FFCE36BABE852" box="[151,230,789,810]" italics="true" pageId="33" pageNumber="34">Trimma</emphasis>
</taxonomicName>
sp. 4:
<treatmentCitation id="5C11109DB822F364FEA1FCE26A98E852" author="Kuiter" box="[297,469,788,810]" page="204" pageId="33" pageNumber="34" year="1998">
<bibRefCitation id="B9214B7DB822F364FEA1FCE26AE4E852" author="Kuiter" box="[297,425,788,810]" pageId="33" pageNumber="34" refString="Kuiter, R. H. (1998) Photo Guide to the Fishes of the Maldives. Atoll Editions, Victoria, 257 pp." type="book" year="1998">Kuiter, 1998</bibRefCitation>
:204
</treatmentCitation>
(
<collectingCountry id="A5A7761CB822F364FE6AFCE2690BE852" box="[482,582,788,810]" name="Maldives" pageId="33" pageNumber="34">Maldives</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD0F368CB822F364FF1FFCC569ABE830" blockId="33.[151,1436,726,840]" box="[151,742,818,840]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="FDA011A2B822F364FF1FFCC569ABE830" box="[151,742,818,840]" pageId="33" pageNumber="34">
<taxonomicName id="1AB04D0FB822F364FF1FFCC56BABE830" ID-CoL="827C" box="[151,230,819,840]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB822F364FF1FFCC56BABE830" box="[151,230,819,840]" italics="true" pageId="33" pageNumber="34">Trimma</emphasis>
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sp. 11:
<treatmentCitation id="5C11109DB822F364FEBDFCC4692BE830" author="Kuiter" box="[309,614,818,840]" page="706" pageId="33" pageNumber="34" year="2004">
<bibRefCitation id="B9214B7DB822F364FEBDFCC46976E830" author="Kuiter" box="[309,571,818,840]" pageId="33" pageNumber="34" refString="Kuiter, R. H. &amp; Tonozuka, T. (2004) Pictorial Guide to Indonesian Reef Fishes. Part 3. PT Dive &amp; Dive's, Bali, Indonesia, iv + 623 - 893 pp." type="book chapter" year="2004">Kuiter &amp; Tonozuka, 2004</bibRefCitation>
:706
</treatmentCitation>
(
<collectingCountry id="A5A7761CB822F364FDFBFCC46996E830" box="[627,731,818,840]" name="Indonesia" pageId="33" pageNumber="34">Indonesia</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B822F364FF1FFC836F02EE61" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="DD0F368CB822F364FF1FFC836EDCE8F6" blockId="33.[151,1437,885,2026]" box="[151,1425,885,910]" pageId="33" pageNumber="34">
<emphasis id="EFC4EA9EB822F364FF1FFC836A4BE8F6" bold="true" box="[151,262,885,910]" pageId="33" pageNumber="34">Material.</emphasis>
The
<typeStatus id="020B882EB822F364FEB7FC806A22E8F6" box="[319,367,886,910]" pageId="33" pageNumber="34">type</typeStatus>
material (
<specimenCount id="CBB6FD05B822F364FE6AFC8069C6E8F6" box="[482,651,885,910]" pageId="33" pageNumber="34" type="generic">128 specimens</specimenCount>
, 9.0–
<quantity id="1A489B69B822F364FD4FFC806866E8F5" box="[711,811,886,910]" metricMagnitude="-2" metricUnit="m" metricValue="2.42" pageId="33" pageNumber="34" unit="mm" value="24.2">24.2 mm</quantity>
SL) from the Chagos Archipelago, plus the following:
</paragraph>
<paragraph id="DD0F368CB822F364FF4FFC6F6965E8AD" blockId="33.[151,1437,885,2026]" pageId="33" pageNumber="34">
<emphasis id="EFC4EA9EB822F364FF4FFC6F69E3E8CA" bold="true" box="[199,686,921,946]" pageId="33" pageNumber="34">Australian material. Great Barrier Reef:</emphasis>
One Tree Id:
<collectionCode id="BBA1AE49B822F364FCC2FC6C68CAE8CA" box="[842,903,922,946]" pageId="33" pageNumber="34">AMS</collectionCode>
I.20577-008, (23),
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<quantity id="1A489B69B822F364FBEAFC6C6FD3E8C9" box="[1122,1182,922,946]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="34" unit="m" value="30.0">30 m</quantity>
.
</specimenCount>
<emphasis id="EFC4EA9EB822F364FB21FC6F6ED6E8CA" bold="true" box="[1193,1435,921,946]" pageId="33" pageNumber="34">Cocos (Keeling) Ids:</emphasis>
<collectionCode id="BBA1AE49B822F364FF1FFC486B91E8AE" LSID="urn:lsid:biocol.org:col:34981" box="[151,220,958,982]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="33" pageNumber="34">WAM</collectionCode>
P. 29906.017, (15),
<quantity id="1A489B69B822F364FE36FC486969E8AD" box="[446,548,958,982]" metricMagnitude="1" metricUnit="m" metricValue="2.55" metricValueMax="3.1" metricValueMin="2.0" pageId="33" pageNumber="34" unit="m" value="25.5" valueMax="31.0" valueMin="20.0">
20–
<specimenCount id="CBB6FD05B822F364FE62FC486969E8AD" box="[490,548,958,982]" pageId="33" pageNumber="34" type="generic">31 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="DD0F368CB822F364FF4FFC176F02EE61" blockId="33.[151,1437,885,2026]" pageId="33" pageNumber="34">
<emphasis id="EFC4EA9EB822F364FF4FFC176954E882" bold="true" box="[199,537,993,1018]" pageId="33" pageNumber="34">
Other Material.
<collectingCountry id="A5A7761CB822F364FE13FC176959E882" box="[411,532,993,1018]" name="Indonesia" pageId="33" pageNumber="34">Indonesia</collectingCountry>
:
</emphasis>
Bone Rate Id:
<collectionCode id="BBA1AE49B822F364FD69FC14687FE881" LSID="urn:lsid:biocol.org:col:34697" box="[737,818,994,1017]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="33" pageNumber="34">BPBM</collectionCode>
.
<collectingCountry id="A5A7761CB822F364FCCCFC1768FBE882" box="[836,950,993,1018]" name="Indonesia" pageId="33" pageNumber="34">Moluccas</collectingCountry>
:
<collectionCode id="BBA1AE49B822F364FC40FC146F54E881" LSID="urn:lsid:biocol.org:col:34697" box="[968,1049,994,1017]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="33" pageNumber="34">BPBM</collectionCode>
. Raja Ampat Ids:
<collectionCode id="BBA1AE49B822F364FA89FC146E0CE882" LSID="urn:lsid:biocol.org:col:34778" box="[1281,1345,994,1018]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="33" pageNumber="34">ROM</collectionCode>
85205, 4(20–26),
<quantity id="1A489B69B822F364FE85FBF06A39EF65" box="[269,372,1030,1054]" metricMagnitude="1" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="33" pageNumber="34" unit="m" value="20.5" valueMax="23.0" valueMin="18.0">18–23 m</quantity>
;
<collectionCode id="BBA1AE49B822F364FE09FBF06A8CEF66" LSID="urn:lsid:biocol.org:col:34778" box="[385,449,1030,1054]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="33" pageNumber="34">ROM</collectionCode>
84892, (29),
<quantity id="1A489B69B822F364FDD4FBF369D4EF65" box="[604,665,1029,1054]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="33" pageNumber="34" unit="m" value="56.0">56 m</quantity>
;
<collectionCode id="BBA1AE49B822F364FD2EFBF069ABEF66" LSID="urn:lsid:biocol.org:col:34778" box="[678,742,1030,1054]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="33" pageNumber="34">ROM</collectionCode>
85331, 11(11–22),
<quantity id="1A489B69B822F364FC4EFBF36F4EEF65" box="[966,1027,1029,1054]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="33" pageNumber="34" unit="m" value="66.0">66 m</quantity>
;
<collectionCode id="BBA1AE49B822F364FB98FBF06F1DEF66" LSID="urn:lsid:biocol.org:col:34778" box="[1040,1104,1030,1054]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="33" pageNumber="34">ROM</collectionCode>
85342, 27(18–28),
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20–
<specimenCount id="CBB6FD05B822F364FAD4FBF06ED5EF65" box="[1372,1432,1030,1054]" pageId="33" pageNumber="34" type="generic">27 m</specimenCount>
</quantity>
.
<collectionCode id="BBA1AE49B822F364FF1FFBDC6B91EF3A" LSID="urn:lsid:biocol.org:col:34981" box="[151,220,1066,1090]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="33" pageNumber="34">WAM</collectionCode>
P. 32925.005, (24),
<quantity id="1A489B69B822F364FE5AFBDF6971EF39" box="[466,572,1065,1090]" metricMagnitude="1" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="33" pageNumber="34" unit="m" value="45.0" valueMax="50.0" valueMin="40.0">
40–
<specimenCount id="CBB6FD05B822F364FE74FBDF6971EF39" box="[508,572,1065,1090]" pageId="33" pageNumber="34" type="generic">50 m</specimenCount>
</quantity>
.
<emphasis id="EFC4EA9EB822F364FDC4FBDF69D7EF3A" bold="true" box="[588,666,1065,1090]" pageId="33" pageNumber="34">
<collectingCountry id="A5A7761CB822F364FDC4FBDF69D8EF3A" box="[588,661,1065,1090]" name="Palau" pageId="33" pageNumber="34">Palau</collectingCountry>
:
</emphasis>
Koror Id:
<collectionCode id="BBA1AE49B822F364FC97FBDC6812EF3A" box="[799,863,1066,1090]" pageId="33" pageNumber="34">AMS</collectionCode>
;
<collectionCode id="BBA1AE49B822F364FCE7FBDC68E2EF3A" LSID="urn:lsid:biocol.org:col:34778" box="[879,943,1066,1090]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="33" pageNumber="34">ROM</collectionCode>
74773, 2(24.1–26.0),
<quantity id="1A489B69B822F364FB3FFBDF6E01EF39" box="[1207,1356,1065,1090]" metricMagnitude="1" metricUnit="m" metricValue="2.085" metricValueMax="2.65" metricValueMin="1.52" pageId="33" pageNumber="34" unit="m" value="20.85" valueMax="26.5" valueMin="15.2">15.2–26.5 m</quantity>
;
<collectionCode id="BBA1AE49B822F364FAD4FBDC6ED1EF3A" LSID="urn:lsid:biocol.org:col:34778" box="[1372,1436,1066,1090]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="33" pageNumber="34">ROM</collectionCode>
80356, (23.3),
<quantity id="1A489B69B822F364FECAFBB86ACCEF1D" box="[322,385,1102,1126]" metricMagnitude="1" metricUnit="m" metricValue="7.3" pageId="33" pageNumber="34" unit="m" value="73.0">73 m</quantity>
;
<collectionCode id="BBA1AE49B822F364FE07FBB86A82EF1E" LSID="urn:lsid:biocol.org:col:34778" box="[399,463,1102,1126]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="33" pageNumber="34">ROM</collectionCode>
80366, 8(18.6–27.1),
<quantity id="1A489B69B822F364FD59FBB86868EF1D" box="[721,805,1102,1126]" metricMagnitude="1" metricUnit="m" metricValue="1.7399999999999998" pageId="33" pageNumber="34" unit="m" value="17.4">17.4 m</quantity>
;
<collectionCode id="BBA1AE49B822F364FCBCFBB86839EF1E" LSID="urn:lsid:biocol.org:col:34778" box="[820,884,1102,1126]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="33" pageNumber="34">ROM</collectionCode>
80394, (21.0),
<quantity id="1A489B69B822F364FBA0FBB86FF6EF1D" box="[1064,1211,1102,1126]" metricMagnitude="1" metricUnit="m" metricValue="2.36" metricValueMax="2.7399999999999998" metricValueMin="1.98" pageId="33" pageNumber="34" unit="m" value="23.6" valueMax="27.4" valueMin="19.8">19.8–27.4 m</quantity>
. Helen Reef:
<collectionCode id="BBA1AE49B822F364FAD4FBB86ED1EF1E" LSID="urn:lsid:biocol.org:col:34778" box="[1372,1436,1102,1126]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="33" pageNumber="34">ROM</collectionCode>
83276, 25(12.6–26.2),
<quantity id="1A489B69B822F364FE11FB84694DEFF1" box="[409,512,1138,1162]" metricMagnitude="1" metricUnit="m" metricValue="2.8" metricValueMax="3.4" metricValueMin="2.2" pageId="33" pageNumber="34" unit="m" value="28.0" valueMax="34.0" valueMin="22.0">22–34 m</quantity>
;
<collectionCode id="BBA1AE49B822F364FD84FB846901EFF2" LSID="urn:lsid:biocol.org:col:34778" box="[524,588,1138,1162]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="33" pageNumber="34">ROM</collectionCode>
83303, 60(21.1–23.6),
<specimenCount id="CBB6FD05B822F364FCDEFB8468C2EFF2" box="[854,911,1138,1162]" pageId="33" pageNumber="34" type="generic">
<quantity id="1A489B69B822F364FCDEFB8468C6EFF2" box="[854,907,1138,1162]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="33" pageNumber="34" unit="m" value="27.0">27m</quantity>
.
</specimenCount>
<emphasis id="EFC4EA9EB822F364FC1EFB876F7FEFF2" bold="true" box="[918,1074,1137,1162]" pageId="33" pageNumber="34">
New
<collectingCountry id="A5A7761CB822F364FC5BFB876F60EFF2" box="[979,1069,1137,1162]" name="United Kingdom" pageId="33" pageNumber="34">Britain</collectingCountry>
:
</emphasis>
<collectionCode id="BBA1AE49B822F364FBB1FB846F33EFF2" LSID="urn:lsid:biocol.org:col:34981" box="[1081,1150,1138,1162]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="33" pageNumber="34">WAM</collectionCode>
P. 28178.012, 2 (16–21),
<quantity id="1A489B69B822F364FF1FFB606A4FEFD5" box="[151,258,1174,1198]" metricMagnitude="1" metricUnit="m" metricValue="3.65" metricValueMax="4.0" metricValueMin="3.3" pageId="33" pageNumber="34" unit="m" value="36.5" valueMax="40.0" valueMin="33.0">
33–
<specimenCount id="CBB6FD05B822F364FF48FB606A4FEFD5" box="[192,258,1174,1198]" pageId="33" pageNumber="34" type="generic">40 m</specimenCount>
</quantity>
.
<emphasis id="EFC4EA9EB822F364FE99FB636946EFD6" bold="true" box="[273,523,1173,1198]" pageId="33" pageNumber="34">
<collectingCountry id="A5A7761CB822F364FE99FB63694BEFD6" box="[273,518,1173,1198]" name="Papua New Guinea" pageId="33" pageNumber="34">Papua New Guinea</collectingCountry>
:
</emphasis>
Manus Id:
<collectionCode id="BBA1AE49B822F364FD12FB606993EFD6" LSID="urn:lsid:biocol.org:col:34981" box="[666,734,1174,1198]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="33" pageNumber="34">WAM</collectionCode>
P. 27826.068, 7 (19–20),
<quantity id="1A489B69B822F364FB91FB636FC9EFD5" box="[1049,1156,1173,1198]" metricMagnitude="1" metricUnit="m" metricValue="3.8" metricValueMax="4.1" metricValueMin="3.5" pageId="33" pageNumber="34" unit="m" value="38.0" valueMax="41.0" valueMin="35.0">35–41 m</quantity>
;
<collectionCode id="BBA1AE49B822F364FB1CFB606F94EFD6" LSID="urn:lsid:biocol.org:col:34981" box="[1172,1241,1174,1198]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="33" pageNumber="34">WAM</collectionCode>
P. 27826.093, 6 (18–20),
<quantity id="1A489B69B822F364FE88FB4F6A25EFA9" box="[256,360,1209,1234]" metricMagnitude="1" metricUnit="m" metricValue="3.8" metricValueMax="4.1" metricValueMin="3.5" pageId="33" pageNumber="34" unit="m" value="38.0" valueMax="41.0" valueMin="35.0">35–41 m</quantity>
;
<collectionCode id="BBA1AE49B822F364FEFEFB4C6AF6EFAA" LSID="urn:lsid:biocol.org:col:34981" box="[374,443,1210,1234]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="33" pageNumber="34">WAM</collectionCode>
P. 28178.012, 2 (16–21),
<quantity id="1A489B69B822F364FD62FB4C681FEFA9" box="[746,850,1210,1234]" metricMagnitude="1" metricUnit="m" metricValue="3.65" metricValueMax="4.0" metricValueMin="3.3" pageId="33" pageNumber="34" unit="m" value="36.5" valueMax="40.0" valueMin="33.0">
33–
<specimenCount id="CBB6FD05B822F364FC9BFB4C681FEFA9" box="[787,850,1210,1234]" pageId="33" pageNumber="34" type="generic">40 m</specimenCount>
</quantity>
. Ninigo Id:
<collectionCode id="BBA1AE49B822F364FC57FB4C6F7DEFAA" LSID="urn:lsid:biocol.org:col:34871" box="[991,1072,1210,1234]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="34">USNM</collectionCode>
313556,
<quantity id="1A489B69B822F364FB16FB4F6E4DEFA9" box="[1182,1280,1209,1234]" metricMagnitude="0" metricUnit="m" metricValue="2.3" metricValueMax="4.6" metricValueMin="0.0" pageId="33" pageNumber="34" unit="m" value="2.3" valueMax="4.6" valueMin="0.0">0–4.6 m</quantity>
.
<emphasis id="EFC4EA9EB822F364FA85FB4F6ED1EFAA" bold="true" box="[1293,1436,1209,1234]" pageId="33" pageNumber="34">
<collectingCountry id="A5A7761CB822F364FA85FB4F6EDAEFAA" box="[1293,1431,1209,1234]" name="Philippines" pageId="33" pageNumber="34">Philippines</collectingCountry>
:
</emphasis>
Balicasag Id:
<collectionCode id="BBA1AE49B822F364FEBEFB286ACAEF8E" LSID="urn:lsid:biocol.org:col:34871" box="[310,391,1246,1270]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="34">USNM</collectionCode>
313544, 6(15–24),
<quantity id="1A489B69B822F364FDE5FB28698AEF8D" box="[621,711,1246,1270]" metricMagnitude="1" metricUnit="m" metricValue="1.2" metricValueMax="2.4" metricValueMin="0.0" pageId="33" pageNumber="34" unit="m" value="12.0" valueMax="24.0" valueMin="0.0">
0–
<specimenCount id="CBB6FD05B822F364FD01FB28698AEF8D" box="[649,711,1246,1270]" pageId="33" pageNumber="34" type="generic">24 m</specimenCount>
</quantity>
. Southern Cebu:
<collectionCode id="BBA1AE49B822F364FC18FB2868ACEF8E" LSID="urn:lsid:biocol.org:col:34871" box="[912,993,1246,1270]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="34">USNM</collectionCode>
263539, 13(15–22),
<quantity id="1A489B69B822F364FB5CFB286E70EF8D" box="[1236,1341,1246,1270]" metricMagnitude="1" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="33" pageNumber="34" unit="m" value="21.0" valueMax="24.0" valueMin="18.0">18–24 m</quantity>
;
<collectionCode id="BBA1AE49B822F364FAC3FB286ED1EF8E" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,1246,1270]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="34">USNM</collectionCode>
333312, (18),
<quantity id="1A489B69B822F364FEBDFAF46AD1EE61" box="[309,412,1282,1306]" metricMagnitude="1" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="33" pageNumber="34" unit="m" value="21.0" valueMax="24.0" valueMin="18.0">
18–
<specimenCount id="CBB6FD05B822F364FED7FAF46AD1EE61" box="[351,412,1282,1306]" pageId="33" pageNumber="34" type="generic">24 m</specimenCount>
</quantity>
.
<emphasis id="EFC4EA9EB822F364FE2EFAF76909EE62" bold="true" box="[422,580,1281,1306]" pageId="33" pageNumber="34">Solomon Ids:</emphasis>
Florida Id:
<collectionCode id="BBA1AE49B822F364FD41FAF4684BEE62" box="[713,774,1282,1306]" pageId="33" pageNumber="34">AMS</collectionCode>
I.17500–027, (20),
<quantity id="1A489B69B822F364FC6DFAF46F06EE61" box="[997,1099,1282,1306]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="33" pageNumber="34" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">
20–
<specimenCount id="CBB6FD05B822F364FB98FAF46F06EE61" box="[1040,1099,1282,1306]" pageId="33" pageNumber="34" type="generic">30 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B822F364FF4FFAD36928ED6E" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="DD0F368CB822F364FF4FFAD36928ED6E" blockId="33.[151,1437,885,2026]" pageId="33" pageNumber="34">
<emphasis id="EFC4EA9EB822F364FF4FFAD36A0CEE46" bold="true" box="[199,321,1317,1342]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
A species of
<taxonomicName id="1AB04D0FB822F364FE69FAD0697AEE45" box="[481,567,1318,1341]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB822F364FE69FAD0697AEE45" box="[481,567,1318,1341]" italics="true" pageId="33" pageNumber="34">Trimma</emphasis>
</taxonomicName>
with the posterior one quarter or less of the predorsal midline scaled, these scales usually overlapping rows from either side rather than centred on the midline; a relatively broad bony interorbital (about pupil diameter in width); 14–15 pectoral-fin rays with the upper three and lower 2–3 unbranched; multiple, sequential branches to the first four pelvic-fin rays (up to five branch points, six branch tips); 3–4 + 14–15 = 17–19 gill rakers on the outer face of the first gill arch; a forked caudal fin (
<figureCitation id="458B2A09B822F364FB1DFA436FA5EEB6" box="[1173,1256,1461,1486]" captionStart="FIGURE 16" captionStartId="34.[151,250,1094,1116]" captionTargetBox="[206,1385,760,1015]" captionTargetId="figure@34.[151,1435,697,1073]" captionTargetPageId="34" captionText="FIGURE 16. Trimma hoesei, 24.1 mm SL female, ROM 83276, Helen Reef, Palau. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/236083/files/figure.png" pageId="33" pageNumber="34">Fig. 16</figureCitation>
); ventral half of body translucent yellow-orange to red in life, pale above; body uniformly straw-coloured in preservative with darker diffuse saddles across the dorsum.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B822F366FF4FF9D768C2EAF0" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34" type="description">
<paragraph id="DD0F368CB822F364FF4FF9D76E59EDFA" blockId="33.[151,1437,885,2026]" pageId="33" pageNumber="34">
<emphasis id="EFC4EA9EB822F364FF4FF9D76A14ED42" bold="true" box="[199,345,1569,1594]" pageId="33" pageNumber="34">Description.</emphasis>
The following description is taken mostly from
<bibRefCitation id="B9214B7DB822F364FCF4F9D76F20ED42" author="Winterbottom" box="[892,1133,1569,1594]" pageId="33" pageNumber="34" refString="Winterbottom, R. (1984) A review of the gobiid fish genus Trimma from the Chagos Archipelago, central Indian Ocean, with the description of seven new species. Canadian Journal of Zoology, 62, 695 - 715." type="journal article" year="1984">Winterbottom (1984)</bibRefCitation>
, with corrections made for differences in methodology, omissions and errors in text and figure derived from 10 (of 60) specimens from Helen Reef,
<collectingCountry id="A5A7761CB822F364FF52F99F6A55EDFA" box="[218,280,1641,1666]" name="Palau" pageId="33" pageNumber="34">Palau</collectingCountry>
(ROM 83303, 10(21.1–23.6), because of the paucity of Australian material of this species.
</paragraph>
<paragraph id="DD0F368CB822F367FF4FF97B69EFE9FC" blockId="33.[151,1437,885,2026]" lastBlockId="34.[151,1437,151,645]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
Dorsal fin VI +
<date id="A90E104CB822F364FEF3F9786A89EDDE" box="[379,452,1678,1702]" pageId="33" pageNumber="34" value="2010-01-09">I 9–10</date>
(usually 10), second and third spines longest with second reaching between the base of third ray of second dorsal fin and anterior region of the peduncle, third spine may reach posteriorly as far as base of fourth fin ray, first ray of second dorsal fin usually branched, last few rays increasingly elongated so that last reaches posteriorly to, or almost to, dorsal procurrent caudal-fin rays, last ray consisting of two unbranched rami; anal fin
<date id="A90E104CB822F364FF79F8E86A75EC4E" box="[241,312,1822,1846]" pageId="33" pageNumber="34" value="2011-01-09">I 9–11</date>
(usually 10), first ray branched, last few rays increasingly elongated so that last reaches to, or almost to, ventral procurrent caudal-fin rays, anterior ramus of last ray branched, posterior ramus unbranched; pectoral fin 14–16 (usually 15), with three dorsal and 2–3 ventral rays unbranched, others branched once dichotomously; pelvic fin
<date id="A90E104CB822F364FF48F87C6BAEECDA" box="[192,227,1929,1954]" pageId="33" pageNumber="34">I 5</date>
, fifth ray branched once dichotomously reaching posteriorly to urogenital papilla and 78–92% of fourth, which reaches posteriorly to region of first few elements of anal fin, rays 1–4 with 3–5 sequential branches (i.e. 4–6 branch tips per ray), no fraenum, basal membrane 7–12% fourth ray; caudal fin forked. Scales on sides of nape extend to above middle of opercle. Lateral scales 23–24; transverse scales 8; predorsal scales 1–5 (
<quantity id="1A489B69B821F367FB6FFF6E6E63EBC8" box="[1255,1326,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="34" pageNumber="35" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">1–3 in</quantity>
the
<collectingCountry id="A5A7761CB821F367FAD6FF616ED1EBC8" box="[1374,1436,151,176]" name="Palau" pageId="34" pageNumber="35">Palau</collectingCountry>
material), count includes scales from left and right sides that overlap in the midline, confined to area just in front of origin of first dorsal fin; body scales ctenoid; scales anterior to a line between pectoral-fin base and first dorsal-fin origin cycloid or weakly ctenoid; scales in belly midline cycloid anteriorly becoming ctenoid posteriorly; pectoralfin base with 4–5 vertical rows of cycloid scales; 5–6 scales in irregular rows across midline anterior to pelvic fin; no scales on cheek or opercle. Outer row of upper jaw and inner and outer rows of lower jaw teeth consisting of curved, evenly spaced, enlarged canines, other teeth short, conical and in irregular rows. Tongue broadly to moderately pointed, three-quarters pupil diameter in width. Gill opening extending anteroventrally to below midpupil; first gill arch with 3–4 + 14–15 outer rakers. Anterior nares in a short, slim tube; posterior opening pore-like with a slightly raised rim; nasal sac slightly elevated; nasal apparatus confined to middle half of snout. Interorbital broad (~ pupil diameter), with two shallow troughs on either side of a median ridge, interorbital region forming a flattened, rounded 'W' in cross section (
<figureCitation id="458B2A09B821F367FDC6FDD269DDE944" box="[590,656,548,573]" captionStart="FIGURE 1" captionStartId="7.[151,250,738,760]" captionTargetBox="[151,1435,195,713]" captionTargetId="figure@7.[151,1435,195,713]" captionTargetPageId="7" captionText="FIGURE 1. Dorsal view of heads of A: Trimma caesiura, (26.3 mm SL male, ROM 64124, New Caledonia) to show fleshy interorbital width and interorbital and postorbital trenches; B: T. hoesei (21.7 mm SL female, ROM 83303, Palau) to show wide fleshy interorbital and fleshy median interorbital ridge. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/236067/files/figure.png" pageId="34" pageNumber="35">Fig. 1</figureCitation>
B); epaxialis reaching anteriorly to a point perpendicular with middle of pupil, a dermal ridge in dorsal midline extending posteriorly to near anterior extent of predorsal scales. Abdominal/caudal vertebral transition
<typeStatus id="020B882EB821F367FDC0FD9B6932E9FD" box="[584,639,621,645]" pageId="34" pageNumber="35">Type</typeStatus>
A.
</paragraph>
<caption id="89CF6604B821F367FF1FFBB06E4CEF24" box="[151,1281,1094,1116]" httpUri="https://zenodo.org/record/236083/files/figure.png" pageId="34" pageNumber="35" targetBox="[206,1385,760,1015]" targetPageId="34">
<paragraph id="DD0F368CB821F367FF1FFBB06E4CEF24" blockId="34.[151,1281,1094,1116]" box="[151,1281,1094,1116]" pageId="34" pageNumber="35">
<emphasis id="EFC4EA9EB821F367FF1FFBB06A52EF24" bold="true" box="[151,287,1094,1116]" pageId="34" pageNumber="35">FIGURE 16.</emphasis>
<taxonomicName id="1AB04D0FB821F367FEADFBB16AF1EF24" box="[293,444,1094,1116]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="hoesei">
<emphasis id="EFC4EA9EB821F367FEADFBB16AF1EF24" box="[293,444,1094,1116]" italics="true" pageId="34" pageNumber="35">Trimma hoesei</emphasis>
</taxonomicName>
, 24.1 mm SL female, ROM 83276, Helen Reef, Palau. Photo by R. Winterbottom.
</paragraph>
</caption>
<paragraph id="DD0F368CB821F366FF4FFB726F3BEB8F" blockId="34.[151,1437,1156,2009]" lastBlockId="35.[151,1437,151,1367]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<emphasis id="EFC4EA9EB821F367FF4FFB72690EEFE5" box="[199,579,1156,1181]" italics="true" pageId="34" pageNumber="35">Colour pattern. Freshly collected</emphasis>
. A
<quantity id="1A489B69B821F367FDF9FB726995EFE4" box="[625,728,1156,1180]" metricMagnitude="-2" metricUnit="m" metricValue="2.41" pageId="34" pageNumber="35" unit="mm" value="24.1">24.1 mm</quantity>
SL female from Helen Reef,
<collectingCountry id="A5A7761CB821F367FBB9FB726F22EFE5" box="[1073,1135,1156,1181]" name="Palau" pageId="34" pageNumber="35">Palau</collectingCountry>
(
<figureCitation id="458B2A09B821F367FB09FB726F99EFE5" box="[1153,1236,1156,1181]" captionStart="FIGURE 16" captionStartId="34.[151,250,1094,1116]" captionTargetBox="[206,1385,760,1015]" captionTargetId="figure@34.[151,1435,697,1073]" captionTargetPageId="34" captionText="FIGURE 16. Trimma hoesei, 24.1 mm SL female, ROM 83276, Helen Reef, Palau. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/236083/files/figure.png" pageId="34" pageNumber="35">Fig. 16</figureCitation>
) with translucent dorsum, scale pockets thinly outlined in red and scattered, half pupil diameter yellow spots, usually below posterior margins of scales. Ventrum red, margined distally by diffuse, half pupil-diameter wide orange stripe just lateral to anal fin origin. Neural canal visible through body as diffuse dark grey stripe; dorsal surface of swimbladder visible as series of diffuse charcoal blotches. Ventral part of head off-white, cheek with scattered red chromatophores and diffuse, light yellow, oblique stripe, and opercle red with continuation of yellow cheek stripe. Snout reddish, with oblique yellow stripe from midregion of anterior margin of eye across upper jaw; anterior halves of both jaws red. Iris golden, margined dorsally and ventrally by dark purple stripe with similar stripe through middle. Nape with scattered red chromatophores and diffuse yellowish spots and blotches. Elements of dorsal fins red, accentuated at base with vague spots, and narrow (one-quarter pupil width) translucent proximal stripe with scattered red melanophores followed by diffuse, half-pupil width yellow stripe. Distally, first dorsal fin membranes translucent with scattered iridocytes and dark chromatophores, second dorsal fin similar, but with occasional diffuse yellow spots centred on fin rays and dark chromatophores concentrated along margin of fin. Anal fin yellow with half pupil diameter red basal stripe and dark chromatophores concentrated along distal edge. Caudal fin reddish, with four yellow stripes, two along rays forming elongated lobes of fin, and two arising from common source at middle base of fin over ural centrum, diverging and widening posteriorly. Distal margins of hypural outlined by thin red bar, interrupted at midline by yellow stripe. Pelvic fins hyaline with scattering of yellow and grey chromatophores. Freshly collected,
<quantity id="1A489B69B821F367FEEFF91E6A81EC78" box="[359,460,1768,1792]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="34" pageNumber="35" unit="mm" value="21.3">21.3 mm</quantity>
SL female from Chagos (see
<bibRefCitation id="B9214B7DB821F367FC97F9116F48EC78" author="Winterbottom" box="[799,1029,1767,1792]" pageId="34" pageNumber="35" refString="Winterbottom, R. (1984) A review of the gobiid fish genus Trimma from the Chagos Archipelago, central Indian Ocean, with the description of seven new species. Canadian Journal of Zoology, 62, 695 - 715." type="journal article" year="1984">Winterbottom, 1984</bibRefCitation>
, fig. 4) and two other photographed specimens from
<collectingCountry id="A5A7761CB821F367FEDAF8FA6ADDEC5D" box="[338,400,1804,1829]" name="Palau" pageId="34" pageNumber="35">Palau</collectingCountry>
(23.3 and 26.0 mm SL females) very similar. The latter have scale pockets on dorsum redbrown or grey; and caudal fin reddish-purple with faint, diffuse yellow stripes, outer pair (along fin rays forming upper and lower lobes of fin) about three times wider than inner pair. Live: specimen photographed in
<collectingCountry id="A5A7761CB821F367FA99F8A26E34EC15" box="[1297,1401,1876,1901]" name="Maldives" pageId="34" pageNumber="35">Maldives</collectingCountry>
by R. Kuiter. Translucent with red wash extending from abdomen to just behind eye, four light blotches on otherwise black nerve cord, several irregularly spaced blue dashes on head (including top of eye), and white tip to elongated second dorsal spine. Pair photographed spawning on alcyonarian by G. Allen at East Cape, Milne Bay,
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in about
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: pale cerise ventrally, with yellow nerve cord, dorsum semi-translucent purple-red. Several specimens from
<collectingCountry id="A5A7761CB820F366FF5EFF4A6A59EBAD" box="[214,276,188,213]" name="Palau" pageId="35" pageNumber="36">Palau</collectingCountry>
photographed by P. Colin (see Pl. 1 H): similar to
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specimen, but translucent dorsum tinged with red or yellow, some show diffuse or well demarcated pale stripe in middle of snout.
</paragraph>
<paragraph id="DD0F368CB820F366FF4FFEF268C2EAF0" blockId="35.[151,1437,151,1367]" pageId="35" pageNumber="36">
<emphasis id="EFC4EA9EB820F366FF4FFEF26A7AEA65" box="[199,311,260,285]" italics="true" pageId="35" pageNumber="36">Preserved</emphasis>
. Pale straw-yellow with an irregular, diffuse and variably-developed line of chromatophores in midline of snout and anterior interorbital region, continuous line of melanophores in midventral septum between base of last anal-fin ray and ventral procurrent caudal-fin rays; anal fin dusky, few scale pockets outlined with dark chromatophores dorsally on body, forming series of diffuse saddles.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B820F366FF4FFE626A31EAA8" pageId="35" pageNumber="36" type="etymology">
<paragraph id="DD0F368CB820F366FF4FFE626A31EAA8" blockId="35.[151,1437,151,1367]" pageId="35" pageNumber="36">
<emphasis id="EFC4EA9EB820F366FF4FFE626A03EAD5" bold="true" box="[199,334,404,429]" pageId="35" pageNumber="36">Etymology.</emphasis>
Named for Douglass F. Hoese, co-author of this paper, in recognition of his many contributions to gobioid systematics.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B820F366FF4FFE2A6F5BE8DD" pageId="35" pageNumber="36" type="distribution">
<paragraph id="DD0F368CB820F366FF4FFE2A6859E918" blockId="35.[151,1437,151,1367]" pageId="35" pageNumber="36">
<emphasis id="EFC4EA9EB820F366FF4FFE2A6A2DEA8D" bold="true" box="[199,352,476,501]" pageId="35" pageNumber="36">Distribution.</emphasis>
The species has been collected at Cocos (Keeling) Ids, and photographed at the
<collectingCountry id="A5A7761CB820F366FA8AFE2A6E20EA8D" box="[1282,1389,476,501]" name="Maldives" pageId="35" pageNumber="36">Maldives</collectingCountry>
, the only other Indian Ocean localities beside the
<typeStatus id="020B882EB820F366FD06FDF669F3E960" box="[654,702,512,536]" pageId="35" pageNumber="36">type</typeStatus>
locality at the Chagos Archipelago. We have examined preserved material from off Timor and the
<collectingCountry id="A5A7761CB820F366FD84FDD26931E945" box="[524,636,548,573]" name="Indonesia" pageId="35" pageNumber="36">Moluccas</collectingCountry>
, the
<collectingCountry id="A5A7761CB820F366FD3BFDD26879E945" box="[691,820,548,573]" name="Philippines" pageId="35" pageNumber="36">Philippines</collectingCountry>
,
<collectingCountry id="A5A7761CB820F366FCC9FDD268CFE945" box="[833,898,548,573]" name="Palau" pageId="35" pageNumber="36">Palau</collectingCountry>
, Raja Ampat,
<collectingCountry id="A5A7761CB820F366FBAEFDD36E48E945" box="[1062,1285,548,573]" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
, New
<collectingCountry id="A5A7761CB820F366FAC6FDD26ED1E945" box="[1358,1436,548,573]" name="United Kingdom" pageId="35" pageNumber="36">Britain</collectingCountry>
and the
<collectingCountry id="A5A7761CB820F366FF67FDB16AE6E918" box="[239,427,583,608]" name="Solomon Islands" pageId="35" pageNumber="36">Solomon Islands</collectingCountry>
in the western Pacific (
<figureCitation id="458B2A09B820F366FD3BFDB1684BE918" box="[691,774,583,608]" captionStart="FIGURE 15" captionStartId="32.[151,250,1940,1962]" captionTargetBox="[157,1427,1235,1908]" captionTargetId="figure@32.[151,1436,1226,1918]" captionTargetPageId="32" captionText="FIGURE 15. Map of the distributions of Trimma flavatrum (solid inverted triangles) and T. hoesei (solid pentagons). A solid square with a white center indicates localities at which both species occur." httpUri="https://zenodo.org/record/236081/files/figure.png" pageId="35" pageNumber="36">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="DD0F368CB820F366FF4FFD9A68A5E840" blockId="35.[151,1437,151,1367]" pageId="35" pageNumber="36">
<emphasis id="EFC4EA9EB820F366FF4FFD9A6A21E9FD" bold="true" box="[199,364,620,645]" pageId="35" pageNumber="36">Comparisons.</emphasis>
<taxonomicName id="1AB04D0FB820F366FEFCFD9B6951E9FD" box="[372,540,620,645]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="hoesei">
<emphasis id="EFC4EA9EB820F366FEFCFD9B6951E9FD" box="[372,540,620,645]" italics="true" pageId="35" pageNumber="36">Trimma hoesei</emphasis>
</taxonomicName>
is unique among members of the genus in possessing a forked caudal fin (
<figureCitation id="458B2A09B820F366FAE7FD9A6BFBE9D0" captionStart="FIGURE 16" captionStartId="34.[151,250,1094,1116]" captionTargetBox="[206,1385,760,1015]" captionTargetId="figure@34.[151,1435,697,1073]" captionTargetPageId="34" captionText="FIGURE 16. Trimma hoesei, 24.1 mm SL female, ROM 83276, Helen Reef, Palau. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/236083/files/figure.png" pageId="35" pageNumber="36">Fig. 16</figureCitation>
). It differs from all other species of
<taxonomicName id="1AB04D0FB820F366FDD9FD6769EAE9D0" box="[593,679,657,680]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="35" pageNumber="36" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB820F366FDD9FD6769EAE9D0" box="[593,679,657,680]" italics="true" pageId="35" pageNumber="36">Trimma</emphasis>
</taxonomicName>
with a broad, bony interorbital except
<taxonomicName id="1AB04D0FB820F366FBEBFD676F86E9D0" box="[1123,1227,655,680]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="taylori">
<emphasis id="EFC4EA9EB820F366FBEBFD676F3DE9D0" box="[1123,1136,657,680]" italics="true" pageId="35" pageNumber="36">T</emphasis>
.
<emphasis id="EFC4EA9EB820F366FB08FD796F86E9D0" box="[1152,1227,655,680]" italics="true" pageId="35" pageNumber="36">taylori</emphasis>
</taxonomicName>
in having the first four pelvic-fin rays with 3–4 sequential branches (as opposed to a single sequential branch). It differs further from
<taxonomicName id="1AB04D0FB820F366FF1FFD2F6A4CE988" box="[151,257,727,752]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="taylori">
<emphasis id="EFC4EA9EB820F366FF1FFD2F6BE9E988" box="[151,164,729,752]" italics="true" pageId="35" pageNumber="36">T</emphasis>
.
<emphasis id="EFC4EA9EB820F366FF3EFD216A4CE988" box="[182,257,727,752]" italics="true" pageId="35" pageNumber="36">taylori</emphasis>
</taxonomicName>
in that the second and third dorsal spines are elongated in both sexes (elongated only in males of
<taxonomicName id="1AB04D0FB820F366FA0FFD2F6BAFE86D" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="taylori">
<emphasis id="EFC4EA9EB820F366FA0FFD2F6ED9E988" box="[1415,1428,729,752]" italics="true" pageId="35" pageNumber="36">T</emphasis>
.
<emphasis id="EFC4EA9EB820F366FF1FFD0A6BAFE86D" box="[151,226,764,789]" italics="true" pageId="35" pageNumber="36">taylori</emphasis>
</taxonomicName>
), and in having only a single expanded haemal arch (vs. 4–5 expanded haemal arches—see
<bibRefCitation id="B9214B7DB820F366FB71FD0A6B9FE840" author="Winterbottom" pageId="35" pageNumber="36" refString="Winterbottom, R. (1984) A review of the gobiid fish genus Trimma from the Chagos Archipelago, central Indian Ocean, with the description of seven new species. Canadian Journal of Zoology, 62, 695 - 715." type="journal article" year="1984">Winterbottom, 1984</bibRefCitation>
, fig. 26). See Comparisons section of
<taxonomicName id="1AB04D0FB820F366FDF4FCD769AFE840" box="[636,738,799,824]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="taylori">
<emphasis id="EFC4EA9EB820F366FDF4FCD769C7E840" box="[636,650,801,824]" italics="true" pageId="35" pageNumber="36">T</emphasis>
.
<emphasis id="EFC4EA9EB820F366FD1FFCE969AFE840" box="[663,738,799,824]" italics="true" pageId="35" pageNumber="36">taylori</emphasis>
</taxonomicName>
for further differences.
</paragraph>
<paragraph id="DD0F368CB820F366FF4FFCB36F5BE8DD" blockId="35.[151,1437,151,1367]" pageId="35" pageNumber="36">
Four specimens (Helen Reef,
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, n = 2 and Raja Ampat,
<collectingCountry id="A5A7761CB820F366FCEBFCB2689EE825" box="[867,979,836,861]" name="Indonesia" pageId="35" pageNumber="36">Indonesia</collectingCountry>
, n = 2) were sampled for CO1, with only a minor difference between them (
<bibRefCitation id="B9214B7DB820F366FD94FC91680AE8F8" author="Winterbottom" box="[540,839,871,896]" pageId="35" pageNumber="36" refString="Winterbottom, R., Hanner, R. H., Burridge, M. &amp; Zur, M. (2014) A cornucopia of cryptic species-a DNA barcode analysis of the gobiid fish genus Trimma (Percomorpha, Gobiiformes). ZooKeys, 381, 79 - 111. http: // dx. doi. org / 10.3897 / zookeys. 381.6445" type="journal article" year="2014">
Winterbottom
<emphasis id="EFC4EA9EB820F366FD4AFC9F69B6E8F8" box="[706,763,871,896]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
, 2014
</bibRefCitation>
). One of the Helen Reef specimens was phenetically closer to the two Raja Ampat specimens than to the other Helen Reef specimen.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B820F366FF4FFC596F76EE2F" pageId="35" pageNumber="36" type="discussion">
<paragraph id="DD0F368CB820F366FF4FFC596A95EF23" blockId="35.[151,1437,151,1367]" pageId="35" pageNumber="36">
<emphasis id="EFC4EA9EB820F366FF4FFC596A01E8B0" bold="true" box="[199,332,943,968]" pageId="35" pageNumber="36">Discussion.</emphasis>
<bibRefCitation id="B9214B7DB820F366FEDBFC59693EE8B0" author="Winterbottom" box="[339,627,943,968]" pageId="35" pageNumber="36" refString="Winterbottom, R. (1984) A review of the gobiid fish genus Trimma from the Chagos Archipelago, central Indian Ocean, with the description of seven new species. Canadian Journal of Zoology, 62, 695 - 715." type="journal article" year="1984">Winterbottom (1984:704)</bibRefCitation>
stated the basal membrane of the pelvic fin was “... delicate, apparently about one-third of the length of the 5th ray”. The material from
<collectingCountry id="A5A7761CB820F366FCEAFC2068EDE897" box="[866,928,982,1007]" name="Palau" pageId="35" pageNumber="36">Palau</collectingCountry>
(ROM 83303) is in very good condition and allows a quantification of the membrane. It is ‘U’-shaped, with the apex directed anteriorly in the midline and 7–12% of the length of the fourth ray. The arms of the U taper towards the fifth ray, and attach to it at about onethird of the length of the ray.
</paragraph>
<paragraph id="DD0F368CB820F366FF4FFB906842EE77" blockId="35.[151,1437,151,1367]" pageId="35" pageNumber="36">
This species has been collected infrequently, and the largest lots came from well-developed caves in a vertical wall at about
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in the Chagos Archipelago and from similar caves at
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at Helen Reef,
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. G. R. Allen provided us with a photograph of a pair of
<taxonomicName id="1AB04D0FB820F366FDFDFB46699AEFBF" box="[629,727,1198,1223]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="hoesei">
<emphasis id="EFC4EA9EB820F366FDFDFB46699AEFBF" box="[629,727,1198,1223]" italics="true" pageId="35" pageNumber="36">T. hoesei</emphasis>
</taxonomicName>
spawning on an alcyonarian coral that was taken in Milne Bay,
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at a depth of
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.
</specimenCount>
The species is probably more widely distributed in the Indo-west Pacific (sensu Winterbottom
<emphasis id="EFC4EA9EB820F366FEB2FB0E6A3FEE77" box="[314,370,1270,1295]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
, 1989) than current records indicate.
</paragraph>
<paragraph id="DD0F368CB820F366FF4FFAEA6F76EE2F" blockId="35.[151,1437,151,1367]" pageId="35" pageNumber="36">
<taxonomicName id="1AB04D0FB820F366FF4FFAEA6A21EE4B" box="[199,364,1306,1331]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="hoesei">
<emphasis id="EFC4EA9EB820F366FF4FFAEA6A21EE4B" box="[199,364,1306,1331]" italics="true" pageId="35" pageNumber="36">Trimma hoesei</emphasis>
</taxonomicName>
is normally found on drop-offs in depths of
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20–
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</quantity>
. In
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, it is known only from single specimens from the southern Great Barrier Reef and from
<collectingCountry id="A5A7761CB820F366FCADFAC96F7AEE2F" box="[805,1079,1342,1367]" name="Cocos (Keeling) Islands" pageId="35" pageNumber="36">Cocos (Keeling) Islands</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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