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<mods:title id="C6C43D197E4B5EA05EC84BDFC15301D0">Reclassification of the Sack-bearer Moths (Lepidoptera, Mimallonoidea, Mimallonidae)</mods:title>
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<mods:namePart id="FD76FBB1F76CE9180A4F9F84A031CCE3">Laurent, Ryan A. St</mods:namePart>
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<mods:namePart id="1C3526685B88CBA1C808274A67F472BA">Kawahara, Akito Y.</mods:namePart>
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<subSubSection id="376FF72F049A7952B622977ACDDB8616" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="2CD78D394301B13AE13DB43A4241C880" pageId="11" pageNumber="12">
<taxonomicName id="50F58C06106F43A510939AF163602A6C" ID-CoL="927NN" authority="Schaus, 1928" authorityName="Schaus" authorityYear="1928" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Eadmuna Schaus, 1928</taxonomicName>
Figs 8, 45, 85, 86
</paragraph>
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<subSubSection id="B296EBF3C2333403972538C2FA7609A6" pageId="12" pageNumber="13" type="type species">
<paragraph id="1641D38662B22C5B5FA19BAD1DA931F4" pageId="12" pageNumber="13">
<pageBreakToken id="AC2932149D1B290DE2C58898E7D96E84" pageId="12" pageNumber="13" start="start">Type</pageBreakToken>
species.
</paragraph>
<paragraph id="5F20B44B7E8D74872874AF5A8F93A787" pageId="12" pageNumber="13">
<taxonomicName id="ACF287F514863163648C11D6665C3794" class="Insecta" family="Mimallonidae" genus="Cicinnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cicinnus esperans" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="esperans">Cicinnus esperans</taxonomicName>
Schaus, 1905.
</paragraph>
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<paragraph id="FAE523A648705298F2C172E02700EA5E" pageId="12" pageNumber="13">Diagnosis.</paragraph>
<paragraph id="4F7548628C519CC862D9973DA27024C4" pageId="12" pageNumber="13">Combination of the following characters: silvery-gray or brown ground color, forewing with hyaline or sub-hyaline patch bisected by the M2 vein; dorsal postmedial line incomplete, formed by brown crescents between veins; the presence of smooth wing margins without a sharply falcate forewing apex.</paragraph>
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<subSubSection id="D1E3E6F05B7D75A791DD997A9CB48E2F" pageId="12" pageNumber="13" type="apomorphies">
<paragraph id="5996632E5C33FE4CF8CB655AC6044FDC" pageId="12" pageNumber="13">Apomorphies.</paragraph>
<paragraph id="4E44AD14E6EB96CA637DEFBDCDF5C94A" pageId="12" pageNumber="13">Combination of the following characters: (1) Uncus clearly differentiated from tegumen as separate smaller structure (Fig. 8a) compared to single elongate triangular uncus + tegumen with little differentiation between these parts; (2) Triangular transtilla or valva apodeme (of uncertain homology) extensions finely wrinkled along their length (Fig. 8b); (3) Vesica with distinct cornutus (which itself may be formed by many cornuti fused into one) or scobinate patch (Fig. 8c).</paragraph>
</subSubSection>
<subSubSection id="8ABA3F0C8FD7251C3AED16B61989D30A" pageId="12" pageNumber="13" type="remarks">
<paragraph id="CAD0960449C02E181B12EFE35B20900E" pageId="12" pageNumber="13">Remarks.</paragraph>
<paragraph id="B57B1C4C434F495DE5689E0A4D4F767F" pageId="12" pageNumber="13">
<bibRefCitation id="D406C035DFB2B950C11C06F8119E7916" author="St Laurent, RA" journalOrPublisher="Systematic Entomology" pageId="76" pageNumber="77" pagination="729 - 761" title="Museum specimens provide phylogenomic data to resolve relationships of sack-bearer moths (Lepidoptera, Mimallonoidea, Mimallonidae)." url="https://doi.org/10.1111/syen.12301" volume="43" year="2018 a">St Laurent et al. (2018a)</bibRefCitation>
did not include this genus in their phylogeny and instead placed this genus in
<taxonomicName id="076BAF6D6BAD3015B9D502A7F42479FC" lsidName="" pageId="12" pageNumber="13" rank="subfamily" subfamily="Mimalloninae">Mimalloninae</taxonomicName>
based on morphology, due to clear similarities to the genus
<taxonomicName id="E255D129C2698A5A7EBC40C7B8FFE84C" class="Insecta" family="Mimallonidae" genus="Macessoga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macessoga" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Macessoga</taxonomicName>
, a genus that was included. Our ongoing molecular work, which includes broadened taxon coverage of
<taxonomicName id="7E7D9F8A09BCD588E819BAE20D988BB1" family="Mimallonidae" lsidName="" pageId="12" pageNumber="13" rank="family">Mimallonidae</taxonomicName>
, including
<taxonomicName id="D5E68FFC2A093C70E8DEBBFCD2100356" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
, fully supports this placement (St Laurent et al. in prep.). Additionally, all of our morphological phylogenetic results consistently place
<taxonomicName id="5892F25DA83DF98BED758AFA759479C4" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
as sister to
<taxonomicName id="E9C26513F52DBA2A7F418EF30FFA7C52" class="Insecta" family="Mimallonidae" genus="Macessoga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macessoga" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Macessoga</taxonomicName>
(Fig. 1, see also Suppl. materials 5-7), therefore confirming the placement of this genus in
<taxonomicName id="A63CBA1915FD82ACD11F0482F4657396" lsidName="" pageId="12" pageNumber="13" rank="subfamily" subfamily="Mimalloninae">Mimalloninae</taxonomicName>
.
</paragraph>
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