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<document id="87C65001500E586DA7DDA70687927E37" ID-DOI="10.5281/zenodo.174318" ID-GBIF-Dataset="b8813703-00e8-48ff-bb05-667752b9a7f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="174318" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459795368271" checkinUser="plazi" docAuthor="Hogan, James E." docDate="2006" docId="517C8791FFCF6846FEA212562A888E19" docLanguage="en" docName="zt01337p068.pdf" docOrigin="Zootaxa 1337" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Psammocoryza vanemdeni Hogan, new species" docType="treatment" docVersion="10" lastPageNumber="67" masterDocId="AD45FFE9FFCC6840FFAA100A280A890A" masterDocTitle="Psammocoryza vanemdeni, a new genus and species of scaritine beetle from the Atlantic coast of Brazil (Coleoptera: Carabidae: Clivinini)" masterLastPageNumber="68" masterPageNumber="61" pageNumber="64" updateTime="1698219676329" updateUser="plazi">
<mods:mods id="2FEF1D09B9551995DEBF9C60952FEF7A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="6EC4E810CE9D367ACF719C1B9F94EE92">Psammocoryza vanemdeni, a new genus and species of scaritine beetle from the Atlantic coast of Brazil (Coleoptera: Carabidae: Clivinini)</mods:title>
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<mods:name id="6E8B2C0679830844AA0DFA049C3F7D2E" type="personal">
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<mods:roleTerm id="EFB75DF8E851C99E8B60DC8FBB6748C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FA6B464249AC3B936F91EF684E4D94EB">Hogan, James E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5803EA6C621515BF0A393AA24B8BFC55">text</mods:typeOfResource>
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<mods:title id="22C2C8CCF958F3D1A9F1ED3574EC2477">Zootaxa</mods:title>
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<mods:part id="B9E58E04F4118B5DB245D7EE64338147">
<mods:date id="F37BCF8097811531B114186524C4B893">2006</mods:date>
<mods:detail id="18BC7F9BDCE31C89BA9C9300AEA8D43B" type="volume">
<mods:number id="83830D3D0D82640050B0B92E0A7B671A">1337</mods:number>
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<mods:classification id="02ABB6F2699083C596032CB1AF3928B9">journal article</mods:classification>
<mods:identifier id="1DA4D2D344CF302BC600ABEDB70DF8DF" type="DOI">10.5281/zenodo.174318</mods:identifier>
<mods:identifier id="C76D3520C262549D5BC55484256C04A0" type="GBIF-Dataset">b8813703-00e8-48ff-bb05-667752b9a7f2</mods:identifier>
<mods:identifier id="43C8B19CF8334A78723BEA3134D46649" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F11BACAB079832875C691B7FD8635830" type="Zenodo-Dep">174318</mods:identifier>
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<treatment id="517C8791FFCF6846FEA212562A888E19" ID-DOI="http://doi.org/10.5281/zenodo.5670924" ID-GBIF-Taxon="119351022" ID-Zenodo-Dep="5670924" LSID="urn:lsid:plazi:treatment:517C8791FFCF6846FEA212562A888E19" httpUri="http://treatment.plazi.org/id/517C8791FFCF6846FEA212562A888E19" lastPageId="6" lastPageNumber="67" pageId="3" pageNumber="64">
<subSubSection id="91CF650CFFCF6843FEA2125629828B93" pageId="3" pageNumber="64" type="nomenclature">
<paragraph id="D96A3687FFCF6843FEA212562B498B7F" blockId="3.[264,835,603,665]" box="[264,835,603,630]" pageId="3" pageNumber="64">
<heading id="822281EBFFCF6843FEA212562B498B7F" bold="true" box="[264,835,603,630]" fontSize="11" level="1" pageId="3" pageNumber="64" reason="1">
<emphasis id="EBA1EA95FFCF6843FEA212562B498B7F" bold="true" box="[264,835,603,630]" pageId="3" pageNumber="64">
<taxonomicName id="1ED54D04FFCF6843FEA212562AAC8B7F" ID-CoL="4NC6Q" authority="Hogan" authorityName="Hogan" box="[264,678,603,630]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="vanemdeni" status="sp. nov.">
<emphasis id="EBA1EA95FFCF6843FEA212562A438B7C" bold="true" box="[264,585,604,630]" italics="true" pageId="3" pageNumber="64">Psammocoryza vanemdeni</emphasis>
Hogan
</taxonomicName>
,
<taxonomicNameLabel id="F09257EEFFCF6843FD1912512B498B7F" box="[691,835,603,629]" pageId="3" pageNumber="64" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D96A3687FFCF6843FEA2127529828B93" blockId="3.[264,835,603,665]" box="[264,392,639,665]" pageId="3" pageNumber="64">
(
<figureCitation id="41EE2A02FFCF6843FEBB127529758B93" box="[273,383,639,665]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 7" captionStartId-0="2.[264,382,1571,1595]" captionStartId-1="4.[264,382,1012,1036]" captionTargetBox-0="[264,1323,414,1531]" captionTargetBox-1="[287,1302,616,987]" captionTargetId-0="figure@2.[264,1323,406,1547]" captionTargetId-1="figure@4.[264,1323,598,988]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 4. Psammocoryza vanemdeni Hogan, new genus and species, holotype, female. 1, habitus, latero-dorsal aspect (total length of specimen 3.64 mm); 2, labium, ventral aspect, showing palpi and ligula; 3, left maxilla, ventral aspect; 4, ovipositor styli, dorsal aspect, Scale bar = 0.2 mm." captionText-1="FIGURES 5 – 7. Psammocoryza vanemdeni Hogan, new genus and species. 5, holotype, female; 6 – 7, paratype, male. 5, left fore leg, right lateral aspect; 6, parameres, dorsal aspect; 7, phallus, with endophallus inverted, left lateral aspect. Scale bar = 0.2 mm." httpUri-0="https://zenodo.org/record/174319/files/figure.png" httpUri-1="https://zenodo.org/record/174320/files/figure.png" pageId="3" pageNumber="64">Figs. 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="91CF650CFFCF6843FEA212C42ACE8D23" pageId="3" pageNumber="64" type="materials_examined">
<paragraph id="D96A3687FFCF6843FEA212C42ACE8D23" blockId="3.[264,1323,718,1905]" pageId="3" pageNumber="64">
<emphasis id="EBA1EA95FFCF6843FEA212C429C98BE2" bold="true" box="[264,451,718,744]" pageId="3" pageNumber="64">
<typeStatus id="066E8825FFCF6843FEA212C429498BE2" box="[264,323,718,744]" pageId="3" pageNumber="64">Type</typeStatus>
material:
</emphasis>
<typeStatus id="066E8825FFCF6843FE7A12C52A488BE3" box="[464,578,719,745]" pageId="3" pageNumber="64" type="holotype">Holotype</typeStatus>
: Female, deposited in the Natural History Museum, London,
<collectingCountry id="A1C27617FFCF6843FEA212FD293C8A1B" box="[264,310,759,785]" name="United Kingdom" pageId="3" pageNumber="64">UK</collectingCountry>
. “Barra de Serinhaem, Pernambuco,
<date id="AD6B1047FFCF6843FD5912FD2B638A1B" box="[755,873,759,785]" pageId="3" pageNumber="64" value="1937-05-25">25.v.1937</date>
, O. Schubart, Br.867”, “Nordstrand”, “gen.n. S2, van Emden det. 1949”, “F.van Emden Bequest B.M. 1960–129”, “gen. nr.
<taxonomicName id="1ED54D04FFCF6843FEA2134D29AE8A6B" box="[264,420,839,865]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">Schizogenius</taxonomicName>
det. S.W.Nichols 1984”. In good condition; articles 2–5 of both hind tarsi missing; left antenna with articles 8–11 missing. Head darkened in colour due to maceration in KOH.
<typeStatus id="066E8825FFCF6843FDBB139D2A768ABB" box="[529,636,919,945]" pageId="3" pageNumber="64" type="paratype">Paratype</typeStatus>
: Male, also deposited in the Natural History Museum, London. “Barra de Serinhaem, Pernambuco,
<date id="AD6B1047FFCF6843FC2F13B52BF18AD3" box="[901,1019,959,985]" pageId="3" pageNumber="64" value="1937-05-25">25.v.1937</date>
, O.Schubart, Br.864”, “Nordstrand”, “F.van Emden Bequest B.M. 1960–129”. Specimen incomplete: head, pterothorax and right elytron missing.
</paragraph>
</subSubSection>
<subSubSection id="91CF650CFFCF6845FE92143C2A558F0D" lastPageId="5" lastPageNumber="66" pageId="3" pageNumber="64" type="description">
<paragraph id="D96A3687FFCF6843FE92143C29CE8DAB" blockId="3.[264,1323,718,1905]" pageId="3" pageNumber="64">
<emphasis id="EBA1EA95FFCF6843FE92143C29DA8D5A" bold="true" box="[312,464,1078,1104]" pageId="3" pageNumber="64">Description.</emphasis>
Due to the poor condition of the male
<typeStatus id="066E8825FFCF6843FC0C143D2C018D5B" box="[934,1035,1079,1105]" pageId="3" pageNumber="64" type="paratype">paratype</typeStatus>
the description is based on the female
<typeStatus id="066E8825FFCF6843FE1F14552A118D73" box="[437,539,1119,1145]" pageId="3" pageNumber="64" type="holotype">holotype</typeStatus>
unless otherwise stated. Small clivinine carabid beetle (length of elytra
<quantity id="1E2D9B62FFCF6843FEF8148D29B28DAB" box="[338,440,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="3" pageNumber="64" unit="mm" value="1.92">1.92mm</quantity>
).
</paragraph>
<paragraph id="D96A3687FFCF6843FE9214A42BC48DC3" blockId="3.[264,1323,718,1905]" box="[312,974,1198,1225]" pageId="3" pageNumber="64">
<emphasis id="EBA1EA95FFCF6843FE9214A4299A8DC2" bold="true" box="[312,400,1198,1224]" pageId="3" pageNumber="64">Colour</emphasis>
uniform pale testaceous (pale brownish-yellow).
</paragraph>
<paragraph id="D96A3687FFCF6844FE9214DC2C3F88F4" blockId="3.[264,1323,718,1905]" lastBlockId="4.[264,1323,284,550]" lastPageId="4" lastPageNumber="65" pageId="3" pageNumber="64">
<emphasis id="EBA1EA95FFCF6843FE9214DC2AA58DFA" bold="true" box="[312,687,1238,1264]" pageId="3" pageNumber="64">Head capsule, dorsal surface:</emphasis>
large, slightly narrower than pronotum. Surface dull due to coarse isodiametric microsculpture.
<emphasis id="EBA1EA95FFCF6843FCB514F42B8E8C12" bold="true" box="[799,900,1278,1304]" pageId="3" pageNumber="64">Clypeus</emphasis>
tri-tuberculate at anterior margin, central tubercle extending a small distance posteriorly onto clypeus as a carina, lateral tubercles also extended posteriorly onto clypeus as carinae but longer, almost reaching clypeal suture. Antero-lateral angles of clypeus sharp and carinate, almost reaching level of anterior tubercles. Single central pair of clypeal carinae almost continuous with second pair of carinae of frons, slightly converging anteriorly, terminating just before anterior margin of clypeus either side of central tubercle. One pair of clypeal setae. Clypeal suture deeply impressed lateraly, absent medially. Supra-antennal plate carinate laterally, rounded.
<emphasis id="EBA1EA95FFCF6843FEDD163429CA8F52" bold="true" box="[375,448,1598,1624]" pageId="3" pageNumber="64">Frons</emphasis>
with seven pairs of paired longitudinal carinae; first, fourth and fifth pairs nearly parallel; second pair weakly developed, stronger basally; third pair convergent anteriorly, continued almost unbroken onto clypeus; fourth pair weakly developed, narrower and lower than carinae one, three and five; fifth pair interrupted for a short distance at level of eye, then continued to clypeal suture; sixth pair shorter than carinae one to five, curved inwards around insertion of anterior supra-orbital seta and terminating a short distance beyond; seventh pair also short, touching the dorsal margin of the eye then continued to clypeal suture. On the right hand side only the fifth carina is interrupted for a short distance. The form of the carinae may therefore show some individual variation. Two pairs of supra-orbital setae; anterior pair located close to eye, adjacent to carina six; posterior pair at base of carina five.
<emphasis id="EBA1EA95FFC86844FD4F11612B2F888F" bold="true" box="[741,805,363,389]" pageId="4" pageNumber="65">Neck</emphasis>
marked dorsaly with coarse longitudinal rugosity.
<emphasis id="EBA1EA95FFC86844FEDE11992AE588A7" bold="true" box="[372,751,403,429]" pageId="4" pageNumber="65">Head capsule, ventral surface:</emphasis>
Buccal fissure wide and extended to below level of submentum. Gula narrow, at narrowest point 0.2 times maximum width of mentum; surface smooth and shining, in contrast to the microsculptured genae.
</paragraph>
<paragraph id="D96A3687FFC86844FE9212012B758B2C" blockId="4.[264,1323,284,550]" box="[312,895,523,550]" pageId="4" pageNumber="65">
<emphasis id="EBA1EA95FFC86844FE92120129788B2F" bold="true" box="[312,370,523,549]" pageId="4" pageNumber="65">Eyes</emphasis>
large and prominent, not enclosed by genae.
</paragraph>
<caption id="8DAA660FFFC86844FEA213FE2B838D5B" httpUri="https://zenodo.org/record/174320/files/figure.png" pageId="4" pageNumber="65" targetBox="[287,1302,616,987]" targetPageId="4">
<paragraph id="D96A3687FFC86844FEA213FE2B838D5B" blockId="4.[264,1323,1012,1105]" pageId="4" pageNumber="65">
<emphasis id="EBA1EA95FFC86844FEA213FE29BD8D06" bold="true" box="[264,439,1012,1036]" pageId="4" pageNumber="65">FIGURES 5–7.</emphasis>
<taxonomicName id="1ED54D04FFC86844FE6B13FE2B3F8D01" authority="Hogan" authorityName="Hogan" box="[449,821,1012,1035]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="vanemdeni" status="gen. nov.">
<emphasis id="EBA1EA95FFC86844FE6B13FE2AE88D01" box="[449,738,1012,1035]" italics="true" pageId="4" pageNumber="65">Psammocoryza vanemdeni</emphasis>
Hogan
</taxonomicName>
,
<taxonomicNameLabel id="F09257EEFFC86844FCE913FE2BBD8D01" box="[835,951,1012,1035]" pageId="4" pageNumber="65" rank="genus">new genus</taxonomicNameLabel>
and species. 5, holotype, female; 6–7, paratype, male. 5, left fore leg, right lateral aspect; 6, parameres, dorsal aspect; 7, phallus, with endophallus inverted, left lateral aspect. Scale bar = 0.2mm.
</paragraph>
</caption>
<paragraph id="D96A3687FFC86844FE92148E2AFF8DE4" blockId="4.[264,1323,1156,1902]" pageId="4" pageNumber="65">
<emphasis id="EBA1EA95FFC86844FE92148E29BA8D94" bold="true" box="[312,432,1156,1182]" pageId="4" pageNumber="65">Antennae</emphasis>
(
<figureCitation id="41EE2A02FFC86844FE60148E2A0D8D94" box="[458,519,1156,1182]" captionStart="FIGURES 1 – 4" captionStartId="2.[264,382,1571,1595]" captionTargetBox="[264,1323,414,1531]" captionTargetId="figure@2.[264,1323,406,1547]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Psammocoryza vanemdeni Hogan, new genus and species, holotype, female. 1, habitus, latero-dorsal aspect (total length of specimen 3.64 mm); 2, labium, ventral aspect, showing palpi and ligula; 3, left maxilla, ventral aspect; 4, ovipositor styli, dorsal aspect, Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/174319/files/figure.png" pageId="4" pageNumber="65">Fig.1</figureCitation>
) pubescent from article three; articles four to ten moniliform. Antennal scape slightly longer than articles two and three combined, with long, sub-apical dorsal seta. Pedicel with long ventral seta.
</paragraph>
<paragraph id="D96A3687FFC86844FE9214F629C98FC4" blockId="4.[264,1323,1156,1902]" pageId="4" pageNumber="65">
<emphasis id="EBA1EA95FFC86844FE9214F62A408C1C" bold="true" box="[312,586,1276,1302]" pageId="4" pageNumber="65">Mouthparts. Labrum</emphasis>
bilobed, slightly emarginate at middle; dorsal surface with seven setae, lateral margins with six pairs of curved setae, reflexed forwards.
<emphasis id="EBA1EA95FFC86844FB0D152E2D218C34" bold="true" box="[1191,1323,1316,1342]" pageId="4" pageNumber="65">Mandibles</emphasis>
prominent, hooked at apex; inner margin straight, inner margin of right mandible with small tooth, probably analagous to the anterior retinacular tooth of the right mandible of
<taxonomicName id="1ED54D04FFC86844FEA215972A6F8CBC" authority="Latreille (Ball 2001)" authorityName="Latreille (Ball" authorityYear="2001" box="[264,613,1436,1462]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFC86844FEA2159729548CBC" box="[264,350,1437,1462]" italics="true" pageId="4" pageNumber="65">Clivina</emphasis>
Latreille (
<bibRefCitation id="BD444B76FFC86844FE4A15962A568CBC" author="Ball" box="[480,604,1436,1462]" pageId="4" pageNumber="68" refString="Ball, G. E. (2001) The subgenera of Clivina Latreille in the Western Hemisphere, and a revision of subgenus Antroforceps Barr (new status), with notes about evolutionary aspects (Coleoptera: Carabidae: Clivinini). Special Publications, Japan Coleopterological Society, Osaka, 1, 129 - 156." type="journal article" year="2001">Ball 2001</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="EBA1EA95FFC86844FDDE15962ADE8CBC" bold="true" box="[628,724,1436,1462]" pageId="4" pageNumber="65">Maxilla</emphasis>
(
<figureCitation id="41EE2A02FFC86844FD4215962B2D8CBC" box="[744,807,1436,1462]" captionStart="FIGURES 1 – 4" captionStartId="2.[264,382,1571,1595]" captionTargetBox="[264,1323,414,1531]" captionTargetId="figure@2.[264,1323,406,1547]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Psammocoryza vanemdeni Hogan, new genus and species, holotype, female. 1, habitus, latero-dorsal aspect (total length of specimen 3.64 mm); 2, labium, ventral aspect, showing palpi and ligula; 3, left maxilla, ventral aspect; 4, ovipositor styli, dorsal aspect, Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/174319/files/figure.png" pageId="4" pageNumber="65">Fig.3</figureCitation>
) with lacinia setose along inner and outer margin, apex slightly hooked; terminal article of palpus elongate, swollen at base.
<emphasis id="EBA1EA95FFC86844FB6D15CE2D218CD4" bold="true" box="[1223,1323,1476,1502]" pageId="4" pageNumber="65">Labium</emphasis>
(
<figureCitation id="41EE2A02FFC86844FEBB15E629598F0C" box="[273,339,1516,1542]" captionStart="FIGURES 1 – 4" captionStartId="2.[264,382,1571,1595]" captionTargetBox="[264,1323,414,1531]" captionTargetId="figure@2.[264,1323,406,1547]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Psammocoryza vanemdeni Hogan, new genus and species, holotype, female. 1, habitus, latero-dorsal aspect (total length of specimen 3.64 mm); 2, labium, ventral aspect, showing palpi and ligula; 3, left maxilla, ventral aspect; 4, ovipositor styli, dorsal aspect, Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/174319/files/figure.png" pageId="4" pageNumber="65">Fig.2</figureCitation>
):
<emphasis id="EBA1EA95FFC86844FEC015E629DF8F0C" bold="true" box="[362,469,1516,1542]" pageId="4" pageNumber="65">Mentum</emphasis>
with triangular central tooth; one pair of paramedian anterior setae and one pair of paralateral posterior setae. Base of mentum with a pair of large ringed pores. Mentum and submentum separated by suture. Submentum with one pair of paramedian and one pair of paralateral setae.
<emphasis id="EBA1EA95FFC86844FD09166E2B4B8F74" bold="true" box="[675,833,1636,1662]" pageId="4" pageNumber="65">Prementum:</emphasis>
Glossal sclerite of ligula with two long apical setae; penultimate article of labial palpus with two setae; terminal article elongate, swollen at base.
</paragraph>
<paragraph id="D96A3687FFC86845FE9216D62C9088A7" blockId="4.[264,1323,1156,1902]" lastBlockId="5.[264,1323,284,1901]" lastPageId="5" lastPageNumber="66" pageId="4" pageNumber="65">
<emphasis id="EBA1EA95FFC86844FE9216D62A248FFC" bold="true" box="[312,558,1756,1782]" pageId="4" pageNumber="65">Thorax. Pronotum</emphasis>
transverse, surface dull and non-reflective due to coarse isodiametric microsculpture. Anterior angles sharp, projected slightly forwards; posterior angles rounded, each with small tooth. Median line deeply impressed and complete, extended from anterior to posterior border. Disc with six pairs of distinct sub-parallel lateral carinae; carinae five and six closer together than others and terminated at level of posterior pronotal seta. Marginal border of pronotum thick and continued to peduncle. Two pairs of lateral setae, posterior pair recessed from margin. Prosternum between coxae produced into a sharp ridge.
<emphasis id="EBA1EA95FFC96845FDFD11992AE788A7" bold="true" box="[599,749,403,429]" pageId="5" pageNumber="66">Mesothorax</emphasis>
pedunculate with scutellum dorsaly.
</paragraph>
<paragraph id="D96A3687FFC96845FE9211B129EF8B7E" blockId="5.[264,1323,284,1901]" pageId="5" pageNumber="66">
<emphasis id="EBA1EA95FFC96845FE9211B1298288DF" bold="true" box="[312,392,443,469]" pageId="5" pageNumber="66">Elytra</emphasis>
with eight striae; intervals flat except at extreme base. Intervals three, five and seven each with eleven setigerous punctures on disc; base of interval three with additional setigerous pore. Base of interval three and four with small tubercle. Shoulder with small raised tooth. Lateral channel with umbillical series of punctures interrupted for a short distance at middle.
</paragraph>
<paragraph id="D96A3687FFC96845FE9212882B438B96" blockId="5.[264,1323,284,1901]" box="[312,841,642,668]" pageId="5" pageNumber="66">
<emphasis id="EBA1EA95FFC96845FE921288298D8B96" bold="true" box="[312,391,642,668]" pageId="5" pageNumber="66">Wings</emphasis>
fully developed, venation not studied.
</paragraph>
<paragraph id="D96A3687FFC96845FE9212A32A338AD0" blockId="5.[264,1323,284,1901]" pageId="5" pageNumber="66">
<emphasis id="EBA1EA95FFC96845FE9212A329738BC9" bold="true" box="[312,377,681,707]" pageId="5" pageNumber="66">Legs.</emphasis>
Metacoxae separated slightly by posterior extension of metasternum.
<emphasis id="EBA1EA95FFC96845FEA212DB29848BE1" bold="true" box="[264,398,721,747]" pageId="5" pageNumber="66">Profemora</emphasis>
(
<figureCitation id="41EE2A02FFC96845FE0812D829EA8BE6" box="[418,480,722,748]" captionStart="FIGURES 5 – 7" captionStartId="4.[264,382,1012,1036]" captionTargetBox="[287,1302,616,987]" captionTargetId="figure@4.[264,1323,598,988]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Psammocoryza vanemdeni Hogan, new genus and species. 5, holotype, female; 6 – 7, paratype, male. 5, left fore leg, right lateral aspect; 6, parameres, dorsal aspect; 7, phallus, with endophallus inverted, left lateral aspect. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/174320/files/figure.png" pageId="5" pageNumber="66">Fig.5</figureCitation>
) large and flattened on inner and outer side; basaly with conspicuous row of eight round tubercles.
<emphasis id="EBA1EA95FFC96845FDC312F32AD68A19" bold="true" box="[617,732,761,787]" pageId="5" pageNumber="66">Protibiae</emphasis>
externaly with four spines; the three distal spines long and thin, not widened basaly; proximal spine very short and wide. Apical spurs rounded at tip.
<emphasis id="EBA1EA95FFC96845FE1713422ACE8A68" bold="true" box="[445,708,840,866]" pageId="5" pageNumber="66">Meso and metatibiae</emphasis>
lacking external spines; apical spurs short and thin.
<emphasis id="EBA1EA95FFC96845FEA2137A294F8A80" bold="true" box="[264,325,880,906]" pageId="5" pageNumber="66">Tarsi</emphasis>
thin and elongate, mesotarsi 0.75 times length of mesotibiae. Tarsal claws small, shorter than terminal article. First tarsomere long, first tarsomere of protarsi longer than remaining four combined.
</paragraph>
<paragraph id="D96A3687FFC96845FE9213E22B4F8D70" blockId="5.[264,1323,284,1901]" pageId="5" pageNumber="66">
<emphasis id="EBA1EA95FFC96845FE9213E22A328D08" bold="true" box="[312,568,1000,1026]" pageId="5" pageNumber="66">Abdomen. Pygidium</emphasis>
with numerous very small setae and margin entire in both sexes. Pygidium of male with very fine file-like striae along midline, not visible in female.
<emphasis id="EBA1EA95FFC96845FEA2143D2A0D8D5B" bold="true" box="[264,519,1079,1105]" pageId="5" pageNumber="66">Abdominal sternites</emphasis>
three to six of both sexes each with one pair of paramedian setae; sternite seven of both sexes with 2 pairs of setae.
</paragraph>
<paragraph id="D96A3687FFC96845FE92148D2A2B8DC3" blockId="5.[264,1323,284,1901]" pageId="5" pageNumber="66">
<emphasis id="EBA1EA95FFC96845FE92148D2A1F8DAB" bold="true" box="[312,533,1159,1185]" pageId="5" pageNumber="66">Female genitalia.</emphasis>
Gonocoxite (
<figureCitation id="41EE2A02FFC96845FD69148D2B098DAB" box="[707,771,1159,1185]" captionStart="FIGURES 1 – 4" captionStartId="2.[264,382,1571,1595]" captionTargetBox="[264,1323,414,1531]" captionTargetId="figure@2.[264,1323,406,1547]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Psammocoryza vanemdeni Hogan, new genus and species, holotype, female. 1, habitus, latero-dorsal aspect (total length of specimen 3.64 mm); 2, labium, ventral aspect, showing palpi and ligula; 3, left maxilla, ventral aspect; 4, ovipositor styli, dorsal aspect, Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/174319/files/figure.png" pageId="5" pageNumber="66">Fig.4</figureCitation>
) straight, apex rounded, with row of pores externally; setae absent.
</paragraph>
<paragraph id="D96A3687FFC96845FE9214DD2B058C4A" blockId="5.[264,1323,284,1901]" pageId="5" pageNumber="66">
<emphasis id="EBA1EA95FFC96845FE9214DD29FE8DFB" bold="true" box="[312,500,1239,1265]" pageId="5" pageNumber="66">Male genitalia.</emphasis>
Phallus (
<figureCitation id="41EE2A02FFC96845FDC314DD2AA28DFB" box="[617,680,1239,1265]" captionStart="FIGURES 5 – 7" captionStartId="4.[264,382,1012,1036]" captionTargetBox="[287,1302,616,987]" captionTargetId="figure@4.[264,1323,598,988]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Psammocoryza vanemdeni Hogan, new genus and species. 5, holotype, female; 6 – 7, paratype, male. 5, left fore leg, right lateral aspect; 6, parameres, dorsal aspect; 7, phallus, with endophallus inverted, left lateral aspect. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/174320/files/figure.png" pageId="5" pageNumber="66">Fig.7</figureCitation>
) evenly arcuate; endophallus without enlarged collar setae. Parameres unequal, left paramere larger and wider than right, each bearing a single long apical seta. Genital ring sclerite closed.
</paragraph>
<paragraph id="D96A3687FFC96845FEEC15442A558F0D" blockId="5.[264,1323,284,1901]" pageId="5" pageNumber="66">
<emphasis id="EBA1EA95FFC96845FEEC15442A0F8C62" bold="true" box="[326,517,1358,1384]" pageId="5" pageNumber="66">Measurements.</emphasis>
Head width including eyes
<quantity id="1E2D9B62FFC96845FCC715442BD98C62" box="[877,979,1358,1384]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="5" pageNumber="66" unit="mm" value="0.73">0.73mm</quantity>
. Head length from neck to antero-lateral angle of clypeus
<quantity id="1E2D9B62FFC96845FD23157C2AE58C9A" box="[649,751,1398,1424]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="5" pageNumber="66" unit="mm" value="0.83">0.83mm</quantity>
. Length of pronotum along mid-line
<quantity id="1E2D9B62FFC96845FB68157C2D2D8C9A" box="[1218,1319,1398,1424]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="66" unit="mm" value="0.7">0.70mm</quantity>
. Length of left elytron excluding peduncle
<quantity id="1E2D9B62FFC96845FCB715942B748CB2" box="[797,894,1438,1464]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="5" pageNumber="66" unit="mm" value="1.92">1.92mm</quantity>
(female
<typeStatus id="066E8825FFC96845FC5E15942C558CB2" box="[1012,1119,1438,1464]" pageId="5" pageNumber="66" type="holotype">holotype</typeStatus>
),
<quantity id="1E2D9B62FFC96845FBD315942CD08CB2" box="[1145,1242,1438,1464]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="5" pageNumber="66" unit="mm" value="1.78">1.78mm</quantity>
(male
<typeStatus id="066E8825FFC96845FEA215CC29788CEA" box="[264,370,1478,1504]" pageId="5" pageNumber="66" type="paratype">paratype</typeStatus>
). Total body length from antero-lateral angle of clypeus to apex of elytra, including peduncle,
<quantity id="1E2D9B62FFC96845FE5C15E72A518F0D" box="[502,603,1517,1543]" metricMagnitude="-3" metricUnit="m" metricValue="3.64" pageId="5" pageNumber="66" unit="mm" value="3.64">3.64mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="91CF650CFFC96845FE92161F2C2D8F75" pageId="5" pageNumber="66" type="etymology">
<paragraph id="D96A3687FFC96845FE92161F2B4F8F5D" blockId="5.[264,1323,284,1901]" pageId="5" pageNumber="66">
<emphasis id="EBA1EA95FFC96845FE92161F29CE8F25" bold="true" box="[312,452,1557,1583]" pageId="5" pageNumber="66">Etymology.</emphasis>
This species is named in honour of the distinguished carabidologist and dipterist Professor F. I. van Emden (
<date id="AD6B1047FFC96845FD0516372B338F5D" box="[687,825,1597,1623]" pageId="5" pageNumber="66" value="1898" valueMax="1958">1898–1958</date>
).
</paragraph>
<paragraph id="D96A3687FFC96845FE92166E2C2D8F75" blockId="5.[264,1323,284,1901]" box="[312,1063,1636,1663]" pageId="5" pageNumber="66">
<emphasis id="EBA1EA95FFC96845FE92166E2A8E8F74" bold="true" box="[312,644,1636,1662]" pageId="5" pageNumber="66">Geographical distribution.</emphasis>
Known only from the
<typeStatus id="066E8825FFC96845FC25166F2BC88F75" box="[911,962,1637,1663]" pageId="5" pageNumber="66">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="91CF650CFFC96846FE9216862A888E19" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66" type="description">
<paragraph id="D96A3687FFC96845FE9216862ACA8E67" blockId="5.[264,1323,284,1901]" pageId="5" pageNumber="66">
<emphasis id="EBA1EA95FFC96845FE9216862A038FAC" bold="true" box="[312,521,1676,1702]" pageId="5" pageNumber="66">Natural History.</emphasis>
The
<typeStatus id="066E8825FFC96845FDE716862A758FAC" box="[589,639,1676,1702]" pageId="5" pageNumber="66">type</typeStatus>
locality is coastal and sand grains adhering to the
<typeStatus id="066E8825FFC96845FB5216862D218FAC" box="[1272,1323,1676,1702]" pageId="5" pageNumber="66">type</typeStatus>
specimens suggest
<taxonomicName id="1ED54D04FFC96845FE4616BF2A958FC4" box="[492,671,1717,1742]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFC96845FE4616BF2A958FC4" box="[492,671,1717,1742]" italics="true" pageId="5" pageNumber="66">Psammocoryza</emphasis>
</taxonomicName>
inhabits marine beaches. This hypothesis is supported by the elongate form of the pro-tibial teeth which is similar to those of other psammophilous
<taxonomicName id="1ED54D04FFC96845FE63170E2A348E14" box="[457,574,1796,1822]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Scaritinae">Scaritinae</taxonomicName>
such as species of
<taxonomicName id="1ED54D04FFC96845FCB4170E2BAD8E17" box="[798,935,1796,1821]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFC96845FCB4170E2BAD8E17" box="[798,935,1796,1821]" italics="true" pageId="5" pageNumber="66">Halocoryza</emphasis>
</taxonomicName>
,
<taxonomicName id="1ED54D04FFC96845FC1C170E2C458E17" box="[950,1103,1796,1821]" class="Insecta" family="Carabidae" genus="Lophocoryza" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFC96845FC1C170E2C458E17" box="[950,1103,1796,1821]" italics="true" pageId="5" pageNumber="66">Lophocoryza</emphasis>
</taxonomicName>
and
<taxonomicName id="1ED54D04FFC96845FB22170E29978E4C" authority="Dejean. Pale" authorityName="Dejean. Pale" class="Insecta" family="Carabidae" genus="Acanthoscelis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFC96845FB22170E2D218E17" box="[1160,1323,1796,1821]" italics="true" pageId="5" pageNumber="66">Acanthoscelis</emphasis>
Dejean. Pale
</taxonomicName>
testaceous coloration and coarse isodiametric microsculpture are also seen in other coastal
<taxonomicName id="1ED54D04FFC96845FE0E17592AB68E67" authority="(Erwin 2004)" baseAuthorityName="Erwin" baseAuthorityYear="2004" box="[420,700,1875,1901]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="family">
carabidae (
<bibRefCitation id="BD444B76FFC96845FD8917592ABE8E67" author="Erwin" box="[547,692,1875,1901]" pageId="5" pageNumber="68" refString="Erwin, T. L. (2004) The beetle family Carabidae of Costa Rica and Panama: Descriptions of four new genera and six new species with notes on their way of life (Insecta: Coleoptera). Zootaxa, 537, 1 - 18." type="journal article" year="2004">Erwin 2004</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption id="8DAA660FFFCA6846FEA2111129B8885F" ID-Table-UUID="8DAA660FFFCA6846FEA2111129B8885F" httpUri="http://table.plazi.org/id/8DAA660FFFCA6846FEA2111129B8885F" pageId="6" pageNumber="67" targetBox="[264,1309,374,1616]" targetIsTable="true" targetPageId="6">
<paragraph id="D96A3687FFCA6846FEA2111129B8885F" blockId="6.[264,1323,283,342]" pageId="6" pageNumber="67">
<emphasis id="EBA1EA95FFCA6846FEA21111298C8839" bold="true" box="[264,390,283,307]" pageId="6" pageNumber="67">TABLE 1.</emphasis>
Diagnostic characters of the genera
<emphasis id="EBA1EA95FFCA6846FCE611162C548839" box="[844,1118,284,307]" italics="true" pageId="6" pageNumber="67">
<taxonomicName id="1ED54D04FFCA6846FCE611162C298839" box="[844,1059,284,307]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="sensu">
Schizogenius
<taxonomicNameLabel id="F09257EEFFCA6846FC4211162C298839" box="[1000,1059,284,307]" pageId="6" pageNumber="67" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato
</emphasis>
,
<taxonomicName id="1ED54D04FFCA6846FBDC11162CF88839" box="[1142,1266,284,307]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCA6846FBDC11162CF88839" box="[1142,1266,284,307]" italics="true" pageId="6" pageNumber="67">Halocoryza</emphasis>
</taxonomicName>
and
<taxonomicName id="1ED54D04FFCA6846FEA2113529A1885C" box="[264,427,319,342]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCA6846FEA2113529A1885C" box="[264,427,319,342]" italics="true" pageId="6" pageNumber="67">Psammocoryza</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="D96A3687FFCA6846FEBA117C29E78F5A" pageId="6" pageNumber="67">
<table id="ABD5C427FFCA97BFFEA2117C2D178F5A" box="[264,1309,374,1616]" gridcols="4" gridrows="11" pageId="6" pageNumber="67">
<tr id="67E534C5FFCA97BFFEA2117C2D1788BB" box="[264,1309,374,433]" gridrow="0" pageId="6" pageNumber="67">
<th id="24345DB9FFCA97BFFEA2117C2A0D88BB" box="[264,519,374,433]" gridcol="0" gridrow="0" pageId="6" pageNumber="67">Character</th>
<th id="24345DB9FFCA97BFFDB1117C2B1A88BB" box="[539,784,374,433]" gridcol="1" gridrow="0" pageId="6" pageNumber="67">
<taxonomicName id="1ED54D04FFCA6846FDB1117D2A9D8884" box="[539,663,375,398]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCA6846FDB1117D2A9D8884" box="[539,663,375,398]" italics="true" pageId="6" pageNumber="67">Halocoryza</emphasis>
</taxonomicName>
.
</th>
<emphasis id="EBA1EA95FFCA6846FC8E117D2CDB8884" italics="true" pageId="6" pageNumber="67">
<th id="24345DB9FFCA97BFFC8E117C2C1388BB" box="[804,1049,374,433]" gridcol="2" gridrow="0" pageId="6" pageNumber="67">
<emphasis id="EBA1EA95FFCA6846FC8E117D2B5F88BB" italics="true" pageId="6" pageNumber="67">
<taxonomicName id="1ED54D04FFCA6846FC8E117D2BFB8884" box="[804,1009,375,398]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="sensu">
Schizogenius
<taxonomicNameLabel id="F09257EEFFCA6846FC1C117D2BFB8884" box="[950,1009,375,398]" pageId="6" pageNumber="67" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato.
</emphasis>
</th>
<th id="24345DB9FFCA97BFFB84117C2D1788BB" box="[1070,1309,374,433]" gridcol="3" gridrow="0" pageId="6" pageNumber="67">
<taxonomicName id="1ED54D04FFCA6846FB84117D2CDB8884" box="[1070,1233,375,398]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="EBA1EA95FFCA6846FB84117D2CDB8884" box="[1070,1233,375,398]" italics="true" pageId="6" pageNumber="67">Psammocoryza</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F09257EEFFCA6846FB7D117C2C7A88BA" pageId="6" pageNumber="67" rank="genus">new genus</taxonomicNameLabel>
.
</th>
</emphasis>
</tr>
<tr id="67E534C5FFCA97BFFEA211CE2D178B2A" box="[264,1309,452,544]" gridrow="1" pageId="6" pageNumber="67">
<th id="24345DB9FFCA97BFFEA211CE2A0D8B2A" box="[264,519,452,544]" gridcol="0" gridrow="1" pageId="6" pageNumber="67">Clypeal suture.</th>
<td id="24345DB9FFCA97BFFDB111CE2B1A8B2A" box="[539,784,452,544]" gridcol="1" gridrow="1" pageId="6" pageNumber="67">Obsolete.</td>
<td id="24345DB9FFCA97BFFC8E11CE2C138B2A" box="[804,1049,452,544]" gridcol="2" gridrow="1" pageId="6" pageNumber="67">Feebly to sharply impressed.</td>
<td id="24345DB9FFCA97BFFB8411CE2D178B2A" box="[1070,1309,452,544]" gridcol="3" gridrow="1" pageId="6" pageNumber="67">Sharply impressed laterally, obsolete medially.</td>
</tr>
<tr id="67E534C5FFCA97BFFEA2123E2D178B67" box="[264,1309,564,621]" gridrow="2" pageId="6" pageNumber="67">
<th id="24345DB9FFCA97BFFEA2123E2A0D8B67" box="[264,519,564,621]" gridcol="0" gridrow="2" pageId="6" pageNumber="67">Setation of lacinia.</th>
<td id="24345DB9FFCA97BFFDB1123E2B1A8B67" box="[539,784,564,621]" gridcol="1" gridrow="2" pageId="6" pageNumber="67">Setose along inner margin only.</td>
<td id="24345DB9FFCA97BFFC8E123E2C138B67" box="[804,1049,564,621]" gridcol="2" gridrow="2" pageId="6" pageNumber="67">Setose along inner and outer margin.</td>
<td id="24345DB9FFCA97BFFB84123E2D178B67" box="[1070,1309,564,621]" gridcol="3" gridrow="2" pageId="6" pageNumber="67">Setose along inner and outer margin.</td>
</tr>
<tr id="67E534C5FFCA97BFFEA2128B2D178A0A" box="[264,1309,641,768]" gridrow="3" pageId="6" pageNumber="67">
<th id="24345DB9FFCA97BFFEA2128B2A0D8A0A" box="[264,519,641,768]" gridcol="0" gridrow="3" pageId="6" pageNumber="67">Gula.</th>
<td id="24345DB9FFCA97BFFDB1128B2B1A8A0A" box="[539,784,641,768]" gridcol="1" gridrow="3" pageId="6" pageNumber="67">Broad, more than 0.4 times maximum width of mentum at narrowest part.</td>
<td id="24345DB9FFCA97BFFC8E128B2C138A0A" box="[804,1049,641,768]" gridcol="2" gridrow="3" pageId="6" pageNumber="67">Narrow, not more than 0.3 times maximum width of mentum at narrowest part.</td>
<td id="24345DB9FFCA97BFFB84128B2D178A0A" box="[1070,1309,641,768]" gridcol="3" gridrow="3" pageId="6" pageNumber="67">Narrow, about 0.2 times maximun width of mentum at narrowest part.</td>
</tr>
<tr id="67E534C5FFCA97BFFEA2131E2D178A7A" box="[264,1309,788,880]" gridrow="4" pageId="6" pageNumber="67">
<th id="24345DB9FFCA97BFFEA2131E2A0D8A7A" box="[264,519,788,880]" gridcol="0" gridrow="4" pageId="6" pageNumber="67">Median sulcus of frons. (area between first carinae)</th>
<td id="24345DB9FFCA97BFFDB1131E2B1A8A7A" box="[539,784,788,880]" gridcol="1" gridrow="4" pageId="6" pageNumber="67">Median sulcus almost equal in width to remaining sulci.</td>
<td id="24345DB9FFCA97BFFC8E131E2C138A7A" box="[804,1049,788,880]" gridcol="2" gridrow="4" pageId="6" pageNumber="67">Median sulcus wider than first paramedian sulci.</td>
<td id="24345DB9FFCA97BFFB84131E2D178A7A" box="[1070,1309,788,880]" gridcol="3" gridrow="4" pageId="6" pageNumber="67">Median sulcus narower than first paramedian sulci.</td>
</tr>
<tr id="67E534C5FFCA97BFFEA2138E2D178AEA" box="[264,1309,900,992]" gridrow="5" pageId="6" pageNumber="67">
<th id="24345DB9FFCA97BFFEA2138E2A0D8AEA" box="[264,519,900,992]" gridcol="0" gridrow="5" pageId="6" pageNumber="67">Second article of antenna. (pedicel)</th>
<td id="24345DB9FFCA97BFFDB1138E2B1A8AEA" box="[539,784,900,992]" gridcol="1" gridrow="5" pageId="6" pageNumber="67">Plurisetose.</td>
<td id="24345DB9FFCA97BFFC8E138E2C138AEA" box="[804,1049,900,992]" gridcol="2" gridrow="5" pageId="6" pageNumber="67">Uni- or bi- setose, exceptionally plurisetose.</td>
<td id="24345DB9FFCA97BFFB84138E2D178AEA" box="[1070,1309,900,992]" gridcol="3" gridrow="5" pageId="6" pageNumber="67">Unisetose.</td>
</tr>
<tr id="67E534C5FFCA97BFFEA213FE2D178D27" box="[264,1309,1012,1069]" gridrow="6" pageId="6" pageNumber="67">
<th id="24345DB9FFCA97BFFEA213FE2A0D8D27" box="[264,519,1012,1069]" gridcol="0" gridrow="6" pageId="6" pageNumber="67">Profemur.</th>
<td id="24345DB9FFCA97BFFDB113FE2B1A8D27" box="[539,784,1012,1069]" gridcol="1" gridrow="6" pageId="6" pageNumber="67">Without tubercles.</td>
<td id="24345DB9FFCA97BFFC8E13FE2C138D27" box="[804,1049,1012,1069]" gridcol="2" gridrow="6" pageId="6" pageNumber="67">Without tubercles.</td>
<td id="24345DB9FFCA97BFFB8413FE2D178D27" box="[1070,1309,1012,1069]" gridcol="3" gridrow="6" pageId="6" pageNumber="67">Base with row of eight tubercles.</td>
</tr>
<tr id="67E534C5FFCA97BFFEA2144B2D178C0F" box="[264,1309,1089,1285]" gridrow="7" pageId="6" pageNumber="67">
<th id="24345DB9FFCA97BFFEA2144B2A0D8C0F" box="[264,519,1089,1285]" gridcol="0" gridrow="7" pageId="6" pageNumber="67">Macrosculpture of pronotum.</th>
<td id="24345DB9FFCA97BFFDB1144B2B1A8C0F" box="[539,784,1089,1285]" gridcol="1" gridrow="7" pageId="6" pageNumber="67">One pair of paramedian longitudinal sulci. Paralateral sulci at least weakly developed.</td>
<td id="24345DB9FFCA97BFFC8E144B2C138C0F" box="[804,1049,1089,1285]" gridcol="2" gridrow="7" pageId="6" pageNumber="67">One pair of paramedian longitudinal sulci. Paralateral sulci present in some species.</td>
<td id="24345DB9FFCA97BFFB84144B2D178C0F" box="[1070,1309,1089,1285]" gridcol="3" gridrow="7" pageId="6" pageNumber="67">Multiple raised carinae, no longitudinal sulci.</td>
</tr>
<tr id="67E534C5FFCA97BFFEA215132D178C59" box="[264,1309,1305,1363]" gridrow="8" pageId="6" pageNumber="67">
<th id="24345DB9FFCA97BFFEA215132A0D8C59" box="[264,519,1305,1363]" gridcol="0" gridrow="8" pageId="6" pageNumber="67">Female genitalia.</th>
<td id="24345DB9FFCA97BFFDB115132B1A8C59" box="[539,784,1305,1363]" gridcol="1" gridrow="8" pageId="6" pageNumber="67">Gonocoxite with single seta at middle.</td>
<td id="24345DB9FFCA97BFFC8E15132C138C59" box="[804,1049,1305,1363]" gridcol="2" gridrow="8" pageId="6" pageNumber="67">Gonocoxite plurisetose at middle.</td>
<td id="24345DB9FFCA97BFFB8415132D178C59" box="[1070,1309,1305,1363]" gridcol="3" gridrow="8" pageId="6" pageNumber="67">Gonocoxite without setae.</td>
</tr>
<tr id="67E534C5FFCA97BFFEA2156C2D178CEF" box="[264,1309,1382,1509]" gridrow="9" pageId="6" pageNumber="67">
<th id="24345DB9FFCA97BFFEA2156C2A0D8CEF" box="[264,519,1382,1509]" gridcol="0" gridrow="9" pageId="6" pageNumber="67">Habitat and substrate preference.</th>
<td id="24345DB9FFCA97BFFDB1156C2B1A8CEF" box="[539,784,1382,1509]" gridcol="1" gridrow="9" pageId="6" pageNumber="67">Coastal, on sandy beaches.</td>
<td id="24345DB9FFCA97BFFC8E156C2C138CEF" box="[804,1049,1382,1509]" gridcol="2" gridrow="9" pageId="6" pageNumber="67">Inland, at margins of flowing water. Usually in river gravels, occasionally in sand.</td>
<td id="24345DB9FFCA97BFFB84156C2D178CEF" box="[1070,1309,1382,1509]" gridcol="3" gridrow="9" pageId="6" pageNumber="67">Coastal, on sandy beaches.</td>
</tr>
<tr id="67E534C5FFCA97BFFEA2163C2D178F5A" box="[264,1309,1590,1616]" gridrow="10" pageId="6" pageNumber="67" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="24345DB9FFCA97BFFEA2163C2A0D8F5A" box="[264,519,1590,1616]" gridcol="0" gridrow="10" pageId="6" pageNumber="67">Acknowledgments</th>
</tr>
</table>
</paragraph>
<paragraph id="D96A3687FFCA6846FEA2168B2A888E19" blockId="6.[264,1323,1665,1811]" pageId="6" pageNumber="67">
The author thanks Dr Roger Booth and Max Barclay (Natural History Museum, London) for arranging the loan of specimens, Menaka Rambukwella (Oxford University Museum) for discussions on digital imaging and Darren
<collectingCountry id="A1C27617FFCA6846FCE316DB2B878FE1" box="[841,909,1745,1771]" name="Isle of Man" pageId="6" pageNumber="67">Mann</collectingCountry>
(Oxford University Museum) for advice on dissection techniques.
</paragraph>
</subSubSection>
</treatment>
</document>
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