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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1151.96715" ID-Pensoft-Pub="1313-2970-1151-1" ID-Pensoft-UUID="9C49BF4600E05E54B38F840D28CD0998" ID-ZooBank="4A79044CAE1747D18661BFD317A911E8" ModsDocID="1313-2970-1151-1" checkinTime="1677663983629" checkinUser="pensoft" docAuthor="Boonyanusith, Chaichat &amp; Wongkamhaeng, Koraon" docDate="2023" docId="3F794CB11F5A5A4BA05649DFA5E4C01C" docLanguage="en" docName="ZooKeys 1151: 1-29" docOrigin="ZooKeys 1151" docPubDate="2023-02-28" docSource="http://dx.doi.org/10.3897/zookeys.1151.96715" docTitle="Cletocamptus thailandensis Boonyanusith &amp; Wongkamhaeng 2023, sp. nov." docType="treatment" docUuid="31517B17-C7FE-484F-B96B-69F54CBA879E" docUuidSource="ZooBank" docVersion="2" id="9C49BF4600E05E54B38F840D28CD0998" lastPageNumber="1" masterDocId="9C49BF4600E05E54B38F840D28CD0998" masterDocTitle="A new species of Cletocamptus Schmankevitsch, 1875 (Copepoda, Harpacticoida, Canthocamptidae) from Rayong Province, Eastern Thailand" masterLastPageNumber="29" masterPageNumber="1" pageNumber="1" updateTime="1677664284976" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Cletocamptus Schmankevitsch, 1875 (Copepoda, Harpacticoida, Canthocamptidae) from Rayong Province, Eastern Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Boonyanusith, Chaichat</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1487-2160</mods:nameIdentifier>
<mods:affiliation>Biology Program, Faculty of Science and Technology, Nakhon Ratchasima Rajabhat University, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">chaichat.b@nrru.ac.th</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wongkamhaeng, Koraon</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7671-8869</mods:nameIdentifier>
<mods:affiliation>Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1151</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>29</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1151.96715</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1151.96715</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1151-1</mods:identifier>
<mods:identifier type="ZooBank">4A79044CAE1747D18661BFD317A911E8</mods:identifier>
<mods:identifier type="Pensoft-UUID">9C49BF4600E05E54B38F840D28CD0998</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:31517B17-C7FE-484F-B96B-69F54CBA879E" httpUri="http://treatment.plazi.org/id/3F794CB11F5A5A4BA05649DFA5E4C01C" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/31517B17-C7FE-484F-B96B-69F54CBA879E" authority="Boonyanusith &amp; Wongkamhaeng, 2023" authorityName="Boonyanusith &amp; Wongkamhaeng" authorityYear="2023" class="Copepoda" family="Canthocamptidae" genus="Cletocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cletocamptus thailandensis" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thailandensis" status="sp. nov.">Cletocamptus thailandensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cletocamptus thailandensis sp. nov. female, holotype A habitus, dorsal view B habitus, lateral view (Arrows indicate rows of moderately long spinules) C anal somite and caudal rami, dorsal view D caudal ramus, dorsal view (Arrows indicate cuticular pore and additional row of spinules). Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure2" httpUri="https://binary.pensoft.net/fig/817106" pageId="0" pageNumber="1">Figs 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cletocamptus thailandensis sp. nov. female, holotype A rostrum, ventral view B urosome, ventral view C P 5. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure3" httpUri="https://binary.pensoft.net/fig/817107" pageId="0" pageNumber="1">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cletocamptus thailandensis sp. nov. female, holotype A antennule B antenna C mandible. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure4" httpUri="https://binary.pensoft.net/fig/817108" pageId="0" pageNumber="1">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cletocamptus thailandensis sp. nov. female, holotype A maxillule, frontal view B praecoxal arthrite, caudal view C maxilla, caudal view D maxilliped, caudal view. Scale bar: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure5" httpUri="https://binary.pensoft.net/fig/817109" pageId="0" pageNumber="1">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cletocamptus thailandensis sp. nov. female, holotype A P 1 B P 2. Scale bar: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure6" httpUri="https://binary.pensoft.net/fig/817110" pageId="0" pageNumber="1">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cletocamptus thailandensis sp. nov. female, holotype A P 3 B P 4. Scale bar: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure7" httpUri="https://binary.pensoft.net/fig/817111" pageId="0" pageNumber="1">, 7 (female), 8-11 (male)</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2022-07-19" collectingMethod="hand net" collectorName="C. Boonyanusith" country="Thailand" elevation="24" latitude="12.764711" location="Rayong Province" longLatPrecision="1" longitude="101.79865" specimenCount="1" specimenCount-female="1" stateProvince="Rayong Province" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
<collectingCountry name="Thailand">Thailand</collectingCountry>
<specimenCount type="female">1 ♀</specimenCount>
(adult), 670
<normalizedToken originalValue="μm">μm</normalizedToken>
long;
<collectingRegion country="Thailand" name="Rayong">Rayong Province</collectingRegion>
;
<geoCoordinate degrees="12" direction="north" minutes="45" orientation="latitude" precision="1" seconds="52.96" value="12.764711">12°45'52.96&quot;N</geoCoordinate>
,
<geoCoordinate degrees="101" direction="east" minutes="47" orientation="longitude" precision="1" seconds="55.16" value="101.79865">101°47'55.16&quot;E</geoCoordinate>
,
<elevation metricMagnitude="1" metricUnit="m" metricValue="2.4" unit="m" value="24.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="2.4" unit="m" value="24.0">24 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2022-07-19">19 Jul. 2022</collectingDate>
;
<collectorName>C. Boonyanusith</collectorName>
leg.;
<collectingMethod>hand net</collectingMethod>
; completely dissected and mounted on a slide in glycerol and sealed with nail polish; PSUZC-PK2007-01
</materialsCitation>
.
<materialsCitation collectingDate="2022-05-22" collectorName="C. Boonyanusith" country="Thailand" elevation="24" latitude="12.764711" location="Thailand" longLatPrecision="1" longitude="101.79865" specimenCount="1" specimenCount-male="1" stateProvince="Rayong" typeStatus="Allotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Allotype</typeStatus>
</emphasis>
:
<collectingCountry name="Thailand">Thailand</collectingCountry>
<specimenCount type="male">1 ♂</specimenCount>
(adult), 483
<normalizedToken originalValue="μm">μm</normalizedToken>
long;
<collectingDate value="2022-05-22">22 May 2022</collectingDate>
; other collection data as for holotype; PSUZC-PK2007-02
</materialsCitation>
.
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Thailand" location="Thailand" specimenCount="1" specimenCount-female="1" typeStatus="Paratypes">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
<collectingCountry name="Thailand">Thailand</collectingCountry>
<specimenCount type="female">1 ♀</specimenCount>
(adult) and
</materialsCitation>
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="C. Boonyanusith" country="Thailand" elevation="24" latitude="12.764711" location="Rayong Province" longLatPrecision="1" longitude="101.79865" specimenCount="1" specimenCount-male="1" stateProvince="Rayong" typeStatus="paratype">
<specimenCount type="male">1 ♂</specimenCount>
(adult); same data as for holotype; PSUZC-PK2007-03 and PSUZC-PK2007-04, respectively
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Thailand" location="Thailand" specimenCount="1" specimenCount-male="1">
<collectingCountry name="Thailand">Thailand</collectingCountry>
<specimenCount type="male">1 ♂</specimenCount>
(adult)
</materialsCitation>
,
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Thailand" location="Thailand" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1 ♀</specimenCount>
(adult); same data as for holotype; preserved in 70% ethanol; retained in collection of the first author (CB)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Female</emphasis>
: Body fusiform. All abdominal somites with row of moderately long spinules on lateral surface. Anal operculum with two rows of spinules. Caudal ramus ca. 1.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, with three rows of spinules on inner margin. Antennary exopod with three setae. Mandibular palp with two setae on free segment and one seta arising nearby. Praecoxal arthrite of maxillule with moderately strong, lateral seta. P1Endp-1 reaching distal fourth of Exp-2, with inner seta. Armature complement of Exp-3 and Endp-2, from P1-P4: 4.5.6.5 and 3.3.5.2, respectively. Inner apical seta on P3Endp-3 reaching mid length of outer one. P5 with large notch between baseoendopodal lobe and exopodal one, with six marginal setae on the former and five setae on the latter. P6 reduced to small protuberance with one apical seta on peduncle.
<emphasis bold="true" pageId="0" pageNumber="1">Male</emphasis>
: Left and right legs of P5 fused at base, with three marginal setae on baseoendopodal lobe and four setae on exopodal one. P6 reduced to simple unarmed plate.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description of adult female.</paragraph>
<paragraph pageId="0" pageNumber="1">
Total body length, excluding caudal setae, ranging from 630
<normalizedToken originalValue="µm">µm</normalizedToken>
to 680
<normalizedToken originalValue="µm">µm</normalizedToken>
(mean = 655;
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 4). Habitus tapering posteriorly, with maximum width at posterior fourth of cephalothorax (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cletocamptus thailandensis sp. nov. female, holotype A habitus, dorsal view B habitus, lateral view (Arrows indicate rows of moderately long spinules) C anal somite and caudal rami, dorsal view D caudal ramus, dorsal view (Arrows indicate cuticular pore and additional row of spinules). Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure2" httpUri="https://binary.pensoft.net/fig/817106" pageId="0" pageNumber="1">2A, B</figureCitation>
). Rostrum (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cletocamptus thailandensis sp. nov. female, holotype A rostrum, ventral view B urosome, ventral view C P 5. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure3" httpUri="https://binary.pensoft.net/fig/817107" pageId="0" pageNumber="1">3A</figureCitation>
) well developed, distinct, with broad base and rounded tip; dorsal surface with pair of sensilla; ventral surface with arch row of long spinules subdistally. Prosome ca. 1.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as urosome (including caudal rami), comprising cephalothorax and three free pedigerous somites (P2-P4-bearing somites). Cephalothorax ca. 1.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide and ca. 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as length of prosome, furnished with numerous pits, with numerous sensilla and with long spinules along margin dorsally and laterally (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cletocamptus thailandensis sp. nov. female, holotype A habitus, dorsal view B habitus, lateral view (Arrows indicate rows of moderately long spinules) C anal somite and caudal rami, dorsal view D caudal ramus, dorsal view (Arrows indicate cuticular pore and additional row of spinules). Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure2" httpUri="https://binary.pensoft.net/fig/817106" pageId="0" pageNumber="1">2A, B</figureCitation>
). Dorsal and lateral surfaces of free pedigerous somites with numerous transverse rows of spinules, with stronger spinules and subdistal sensilla near posterior margin; posterior margin with long spinules.
</paragraph>
<caption doi="10.3897/zookeys.1151.96715.figure2" httpUri="https://binary.pensoft.net/fig/817106" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<taxonomicName authorityName="Boonyanusith &amp; Wongkamhaeng" authorityYear="2023" class="Copepoda" family="Canthocamptidae" genus="Cletocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cletocamptus thailandensis" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis italics="true" pageId="0" pageNumber="1">Cletocamptus thailandensis</emphasis>
</taxonomicName>
sp. nov. female, holotype
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
habitus, lateral view (Arrows indicate rows of moderately long spinules)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
anal somite and caudal rami, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
caudal ramus, dorsal view (Arrows indicate cuticular pore and additional row of spinules). Scale bars: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1151.96715.figure3" httpUri="https://binary.pensoft.net/fig/817107" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName authorityName="Boonyanusith &amp; Wongkamhaeng" authorityYear="2023" class="Copepoda" family="Canthocamptidae" genus="Cletocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cletocamptus thailandensis" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis italics="true" pageId="0" pageNumber="1">Cletocamptus thailandensis</emphasis>
</taxonomicName>
sp. nov. female, holotype
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
rostrum, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
urosome, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
P5. Scale bars: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
Urosome (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cletocamptus thailandensis sp. nov. female, holotype A habitus, dorsal view B habitus, lateral view (Arrows indicate rows of moderately long spinules) C anal somite and caudal rami, dorsal view D caudal ramus, dorsal view (Arrows indicate cuticular pore and additional row of spinules). Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure2" httpUri="https://binary.pensoft.net/fig/817106" pageId="0" pageNumber="1">2A, B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cletocamptus thailandensis sp. nov. female, holotype A rostrum, ventral view B urosome, ventral view C P 5. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure3" httpUri="https://binary.pensoft.net/fig/817107" pageId="0" pageNumber="1">3B</figureCitation>
) comprising fifth pedigerous somite (P5-bearing somite), genital double-somite, and three free abdominal somites; lateral surface of all abdominal somites with row of moderately long spinules (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cletocamptus thailandensis sp. nov. female, holotype A habitus, dorsal view B habitus, lateral view (Arrows indicate rows of moderately long spinules) C anal somite and caudal rami, dorsal view D caudal ramus, dorsal view (Arrows indicate cuticular pore and additional row of spinules). Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure2" httpUri="https://binary.pensoft.net/fig/817106" pageId="0" pageNumber="1">2B</figureCitation>
). Fifth pedigerous somite (first urosomite) with numerous transverse rows of spinules, with spinules and subdistal sensilla near posterior margin; posterior margin with larger spinules dorsally and long spinules laterally. Second and third urosomites fused ventrally forming genital double-somite, ca. 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as wide as long, with subchitinous rib representing former division of genital somite and first abdominal one and dividing genital double-somite into anterior and posterior portions; dorsal and lateral surfaces of both portions with numerous transverse rows of spinules, with spinules along margin of subchitinous rib and with hair-like spinules near posterior margin of posterior portion; ventral surface with row of spinules medially, with ventral pore beside medial row of spinules (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cletocamptus thailandensis sp. nov. female, holotype A rostrum, ventral view B urosome, ventral view C P 5. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure3" httpUri="https://binary.pensoft.net/fig/817107" pageId="0" pageNumber="1">3B</figureCitation>
); posterior margin of double-somite with moderately long spinules. Genital field (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cletocamptus thailandensis sp. nov. female, holotype A rostrum, ventral view B urosome, ventral view C P 5. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure3" httpUri="https://binary.pensoft.net/fig/817107" pageId="0" pageNumber="1">3B</figureCitation>
) with ovipore medially at the middle of genital double-somite. Second and third abdominal somites (fourth and fifth urosomites) with numerous transverse rows of spinules, with hair-like spinules dorsally and sensilla near posterior margin and with moderately longer spinules along posterior margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Anal somite (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cletocamptus thailandensis sp. nov. female, holotype A habitus, dorsal view B habitus, lateral view (Arrows indicate rows of moderately long spinules) C anal somite and caudal rami, dorsal view D caudal ramus, dorsal view (Arrows indicate cuticular pore and additional row of spinules). Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure2" httpUri="https://binary.pensoft.net/fig/817106" pageId="0" pageNumber="1">2C</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cletocamptus thailandensis sp. nov. female, holotype A rostrum, ventral view B urosome, ventral view C P 5. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure3" httpUri="https://binary.pensoft.net/fig/817107" pageId="0" pageNumber="1">3B</figureCitation>
) with pair of dorsal sensilla in front of anal operculum, with numerous paired rows of moderately long spinules on dorsal, lateral, and ventral surfaces as shown, accompanied with two pairs of cuticular pores ventrally. Anal operculum slightly convex, with two rows of spinules.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Caudal rami (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cletocamptus thailandensis sp. nov. female, holotype A habitus, dorsal view B habitus, lateral view (Arrows indicate rows of moderately long spinules) C anal somite and caudal rami, dorsal view D caudal ramus, dorsal view (Arrows indicate cuticular pore and additional row of spinules). Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure2" httpUri="https://binary.pensoft.net/fig/817106" pageId="0" pageNumber="1">2C, D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cletocamptus thailandensis sp. nov. female, holotype A rostrum, ventral view B urosome, ventral view C P 5. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure3" httpUri="https://binary.pensoft.net/fig/817107" pageId="0" pageNumber="1">3B</figureCitation>
) slightly divergent, each ca. 1.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, with seven setae; ornamentation comprised of three transverse rows of spinules on inner margin, row of spinules dorsally near distal margin, oblique row of strong spinules near insertion of seta IV, row of long spinules ventrally near insertion of seta IV and seta V, and cuticular pore near base of seta II. Seta I inserted ventro-laterally near insertion of rami; seta II and seta III inserted closely to each another at anterior third of ramus; seta IV and V, ca. 0.22 and ca. 0.58 of body length, respectively, with fracture plane in seta V; seta VI slender, inserted on inner distal corner; seta VII biarticulate, inserted at mid length of ramus medially.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Antennule (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cletocamptus thailandensis sp. nov. female, holotype A antennule B antenna C mandible. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure4" httpUri="https://binary.pensoft.net/fig/817108" pageId="0" pageNumber="1">4A</figureCitation>
) six-segmented. First segment with two rows of spinules medially. Fourth segment with aesthetasc on peduncle; aesthetasc slightly elongated,&gt; 45% of length of aesthetasc surpassing the tip of antennule. Ultimate segment ca. 4.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, with aesthetasc. Aesthetascs fused to seta at base. Armature formula: 1-[1], 2-[9], 3-[6], 4-[1 + (1 + ae)], 5-[1], 6-[9 + (1 + ae)].
</paragraph>
<caption doi="10.3897/zookeys.1151.96715.figure4" httpUri="https://binary.pensoft.net/fig/817108" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<taxonomicName authorityName="Boonyanusith &amp; Wongkamhaeng" authorityYear="2023" class="Copepoda" family="Canthocamptidae" genus="Cletocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cletocamptus thailandensis" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis italics="true" pageId="0" pageNumber="1">Cletocamptus thailandensis</emphasis>
</taxonomicName>
sp. nov. female, holotype
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
antennule
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
antenna
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
mandible. Scale bars: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
Antenna (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cletocamptus thailandensis sp. nov. female, holotype A antennule B antenna C mandible. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure4" httpUri="https://binary.pensoft.net/fig/817108" pageId="0" pageNumber="1">4B</figureCitation>
) biramous, comprising coxa, allobasis, one-segmented Exp, and one-segmented Endp. Coxa short, with arch row of spinules on outer margin. Allobasis with two abexopodal setae. Exp cylindrical, with few spinules subdistally and armed with one lateral seta and two apical setae (one of the apical setae short and slim). Endp club-shaped, with one short seta and two spines medially on inner margin, and five apical setae of which two innermost ones spiniform, two median ones geniculate, outermost one spiniform with two strong spinules half-length of element and bipinnate along distal half; spinule ornamentation comprised of proximal and distal rows of strong spinules, and row of spinules distally.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Mandible (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cletocamptus thailandensis sp. nov. female, holotype A antennule B antenna C mandible. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure4" httpUri="https://binary.pensoft.net/fig/817108" pageId="0" pageNumber="1">4C</figureCitation>
) comprising sclerotized coxal gnathobase and mandibular palp. Coxal gnathobase with two arch rows of spinules as shown; cutting edge with five bicuspidate and multicuspidate teeth, with one pyriform element and pinnate, ventral seta. Mandibular palp one-segmented, with two apical setae, accompanied with one short seta inserted nearby.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Maxillule (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cletocamptus thailandensis sp. nov. female, holotype A maxillule, frontal view B praecoxal arthrite, caudal view C maxilla, caudal view D maxilliped, caudal view. Scale bar: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure5" httpUri="https://binary.pensoft.net/fig/817109" pageId="0" pageNumber="1">5A, B</figureCitation>
) comprising robust praecoxa, coxa, and basal complex, the latter composed of basis with Endp and Exp completely incorporated to the latter. Praecoxal arthrite with row of spinules on caudal surface and on lateral margin, armed with surface seta on frontal surface and nine distal elements: seven of which strong, curved spines, subdistal seta slim and pinnate, lateral seta moderately strong and pinnate. Coxa with cylindrical endite bearing two smooth apical setae, and with few spinules. Basis seemingly with three apical setae of which middle robust and pinnate. Endp and Exp completely incorporated to basis, seemingly represented by three setae each.
</paragraph>
<caption doi="10.3897/zookeys.1151.96715.figure5" httpUri="https://binary.pensoft.net/fig/817109" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
<taxonomicName authorityName="Boonyanusith &amp; Wongkamhaeng" authorityYear="2023" class="Copepoda" family="Canthocamptidae" genus="Cletocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cletocamptus thailandensis" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis italics="true" pageId="0" pageNumber="1">Cletocamptus thailandensis</emphasis>
</taxonomicName>
sp. nov. female, holotype
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
maxillule, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
praecoxal arthrite, caudal view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
maxilla, caudal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
maxilliped, caudal view. Scale bar: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
Maxilla (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cletocamptus thailandensis sp. nov. female, holotype A maxillule, frontal view B praecoxal arthrite, caudal view C maxilla, caudal view D maxilliped, caudal view. Scale bar: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure5" httpUri="https://binary.pensoft.net/fig/817109" pageId="0" pageNumber="1">5C</figureCitation>
) composed of syncoxa and allobasis, and Endp fully incorporated to the latter. Syncoxa with two endites, each endite armed with three elements apically, one of which slender. Allobasis drawn out into claw, with one seta basally on caudal surface. Endp completely incorporated to allobasis, represented by three smooth setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Maxilliped (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cletocamptus thailandensis sp. nov. female, holotype A maxillule, frontal view B praecoxal arthrite, caudal view C maxilla, caudal view D maxilliped, caudal view. Scale bar: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure5" httpUri="https://binary.pensoft.net/fig/817109" pageId="0" pageNumber="1">5D</figureCitation>
) subchelate, three-segmented, comprising syncoxa, basis, and Endp. Syncoxa with curved row of spinules on caudal surface, with one pinnate seta on inner distal corner. Basis with longitudinal row of spinules on frontal and caudal surfaces, and two transverse rows of spinules on outer margin. Endp drawn out into strong claw and armed with one minute seta near base.
</paragraph>
<paragraph pageId="0" pageNumber="1">P1-P4 comprised of intercoxal sclerite, praecoxa, coxa, basis, and two rami.</paragraph>
<paragraph pageId="0" pageNumber="1">
P1 (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cletocamptus thailandensis sp. nov. female, holotype A P 1 B P 2. Scale bar: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure6" httpUri="https://binary.pensoft.net/fig/817110" pageId="0" pageNumber="1">6A</figureCitation>
). Intercoxal sclerite as shown, with two rows of minute spinules on each side. Praecoxa triangular, with spinules along distal margin and on outer distal corner. Coxa rectangular, with one row of spinules medially, with two oblique rows of spinules and with strong spinules on outer distal corner. Basis with integumental pore on frontal surface, with oblique row of spinules and three rows of stronger spinules of which innermost at base of inner spine, median between insertion of rami, outermost at base of Exp; armament comprising outer and inner spines, inner spine reaching distal third of Endp-1. Exp three-segmented, all segments with spinule row on outer margin and outer distal corner; Exp-1 with outer spine; Exp-2 with outer spine and inner seta; Exp-3 with four elements: innermost and inner apical ones slender. Endp two-segmented, reaching tip of Exp; Endp-1 reaching distal fourth of Exp-2, with inner seta; Endp-2 slightly longer than Endp-1, with three elements of which outer apical spiniform.
</paragraph>
<caption doi="10.3897/zookeys.1151.96715.figure6" httpUri="https://binary.pensoft.net/fig/817110" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
<taxonomicName authorityName="Boonyanusith &amp; Wongkamhaeng" authorityYear="2023" class="Copepoda" family="Canthocamptidae" genus="Cletocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cletocamptus thailandensis" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis italics="true" pageId="0" pageNumber="1">Cletocamptus thailandensis</emphasis>
</taxonomicName>
sp. nov. female, holotype
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
P1
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
P2. Scale bar: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
P2 (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cletocamptus thailandensis sp. nov. female, holotype A P 1 B P 2. Scale bar: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure6" httpUri="https://binary.pensoft.net/fig/817110" pageId="0" pageNumber="1">6B</figureCitation>
). Intercoxal sclerite as shown, bare. Praecoxa with spinules along distal margin. Coxa with row of spinules medially and strong spinules along outer margin. Basis as that of P1 but lacking inner spine and inner distal corner; spinule row between rami with smaller spinules relative to those of P1; with medial pore proximally. Exp three-segmented, with spinule ornamentation as that of P1, additionally with row of spinule on inner distal corner of Exp-1 and Exp-2; Exp-1 with outer spine; Exp-2 with outer spine and inner seta; Exp-3 with five elements (two outer spines, two distal and one inner seta). Endp two-segmented, reaching middle of Exp-2; Endp-1 small, wider than long, with inner spinules; Endp-2 ca. 3.0
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, with spinule ornamentation on outer and inner margins, armed with three elements of which outer spiniform and slender.
</paragraph>
<paragraph pageId="0" pageNumber="1">
P3 (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cletocamptus thailandensis sp. nov. female, holotype A P 3 B P 4. Scale bar: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure7" httpUri="https://binary.pensoft.net/fig/817111" pageId="0" pageNumber="1">7A</figureCitation>
). Intercoxal sclerite, praecoxa, coxa, and basis as those of P2. Exp three-segmented; Exp-1 and Exp-2 as those of P2; Exp-3 with six elements (two outer spines, two distal and two inner seta). Endp two-segmented; Endp-1 small, wider than long, with inner spinules; Endp-2 ca. 3.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, armed with five elements of which outer spiniform and slender, inner apical one reaching middle of outer apical one, two inner ones subequal in length.
</paragraph>
<caption doi="10.3897/zookeys.1151.96715.figure7" httpUri="https://binary.pensoft.net/fig/817111" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
<taxonomicName authorityName="Boonyanusith &amp; Wongkamhaeng" authorityYear="2023" class="Copepoda" family="Canthocamptidae" genus="Cletocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cletocamptus thailandensis" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis italics="true" pageId="0" pageNumber="1">Cletocamptus thailandensis</emphasis>
</taxonomicName>
sp. nov. female, holotype
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
P3
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
P4. Scale bar: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
P4 (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cletocamptus thailandensis sp. nov. female, holotype A P 3 B P 4. Scale bar: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure7" httpUri="https://binary.pensoft.net/fig/817111" pageId="0" pageNumber="1">7B</figureCitation>
). Intercoxal sclerite, praecoxa, coxa, and basis as those of P2. Exp three-segmented; Exp-1 and Exp-2 as those of P2; Exp-3 with five elements (two outer spines, two distal and one inner seta). Endp two-segmented; Endp-1 small, wider than long, with inner spinules; Endp-2 ca. 2.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, armed with two apical setae of which outer ca. 2.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as inner one.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Armature formula of P1-P4 as in Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Armature formula of P 1 - P 4 of Cletocamptus thailandensis sp. nov. Arabic numerals indicate number of setae; Roman numerals indicate number of spines." httpUri="http://table.plazi.org/id/66CE5D5C41F540FD902E6A8A38B73167" pageId="0" pageNumber="1" tableUuid="66CE5D5C41F540FD902E6A8A38B73167">1</tableCitation>
.
</paragraph>
<caption ID-Table-UUID="66CE5D5C41F540FD902E6A8A38B73167" httpUri="http://table.plazi.org/id/66CE5D5C41F540FD902E6A8A38B73167" pageId="0" pageNumber="1" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Table 1.</emphasis>
Armature formula of P1-P4 of
<taxonomicName authorityName="Boonyanusith &amp; Wongkamhaeng" authorityYear="2023" class="Copepoda" family="Canthocamptidae" genus="Cletocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cletocamptus thailandensis" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis italics="true" pageId="0" pageNumber="1">Cletocamptus thailandensis</emphasis>
</taxonomicName>
sp. nov. Arabic numerals indicate number of setae; Roman numerals indicate number of spines.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">Legs</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">Basis</th>
<th colspan="3" pageId="0" pageNumber="1" rowspan="1">Exopod</th>
<th colspan="3" pageId="0" pageNumber="1" rowspan="1">Endopod</th>
</tr>
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">1</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">2</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">3</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">1</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">2</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">3</th>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">P1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-I</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-I1-1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0-I1-1</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">P2</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">II-2-1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1-1</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">P3</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">II-2-2</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-2-2</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">P3 (male)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">II-2-2</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0-2-0</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">P4</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">II-2-1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0-0</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0-2-0</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="1">
P5 (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cletocamptus thailandensis sp. nov. female, holotype A rostrum, ventral view B urosome, ventral view C P 5. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure3" httpUri="https://binary.pensoft.net/fig/817107" pageId="0" pageNumber="1">3C</figureCitation>
). Baseoendopod and Exp completely fused basally, with large notch between them; left and right legs completely separated. Baseoendopodal lobe ca. 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as length of exopodal one, with one outer, two apical and three inner setae. Exopodal lobe with five elements accompanied with slender, outer seta of basis; relative length and characteristics of setae on both baseoendopodal and exopodal lobes as shown.
</paragraph>
<paragraph pageId="0" pageNumber="1">
P6 (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cletocamptus thailandensis sp. nov. female, holotype A rostrum, ventral view B urosome, ventral view C P 5. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure3" httpUri="https://binary.pensoft.net/fig/817107" pageId="0" pageNumber="1">3B</figureCitation>
) reduced to minute prominence, forming simple plate near anterior margin of genital double-somite, with row of minute spinules and one short seta on each side.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description of adult male.</paragraph>
<paragraph pageId="0" pageNumber="1">
Body smaller than in female. Total body length, excluding caudal setae, ranging from 483
<normalizedToken originalValue="µm">µm</normalizedToken>
to 505
<normalizedToken originalValue="µm">µm</normalizedToken>
(mean = 493;
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 4). Prosome ca. 1.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as urosome (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cletocamptus thailandensis sp. nov. male, allotype A habitus, dorsal view B habitus, lateral view C rostrum, frontal view D anal somite and caudal rami, dorsal view. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure8" httpUri="https://binary.pensoft.net/fig/817112" pageId="0" pageNumber="1">8A, B</figureCitation>
). Cephalothorax slightly longer than wide, ca. 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as length of prosome. Rostrum (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cletocamptus thailandensis sp. nov. male, allotype A habitus, dorsal view B habitus, lateral view C rostrum, frontal view D anal somite and caudal rami, dorsal view. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure8" httpUri="https://binary.pensoft.net/fig/817112" pageId="0" pageNumber="1">8C</figureCitation>
) well developed, distinct, narrower than that of female, with rounded tip; dorsal surface with pair of sensilla, ventral surface with arch row of long spinules subdistally. Ornamentation of cephalothorax and free prosomite as those of female.
</paragraph>
<caption doi="10.3897/zookeys.1151.96715.figure8" httpUri="https://binary.pensoft.net/fig/817112" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
<taxonomicName authorityName="Boonyanusith &amp; Wongkamhaeng" authorityYear="2023" class="Copepoda" family="Canthocamptidae" genus="Cletocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cletocamptus thailandensis" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis italics="true" pageId="0" pageNumber="1">Cletocamptus thailandensis</emphasis>
</taxonomicName>
sp. nov. male, allotype
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
habitus, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
rostrum, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
anal somite and caudal rami, dorsal view. Scale bars: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
Urosome (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cletocamptus thailandensis sp. nov. male, allotype A habitus, dorsal view B habitus, lateral view C rostrum, frontal view D anal somite and caudal rami, dorsal view. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure8" httpUri="https://binary.pensoft.net/fig/817112" pageId="0" pageNumber="1">8A, B</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cletocamptus thailandensis sp. nov. male, allotype (A, B); paratype (C) A urosome, ventral view B antennule C segments 4 - 6 of antennule. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure9" httpUri="https://binary.pensoft.net/fig/817113" pageId="0" pageNumber="1">9A</figureCitation>
) six-segmented, comprising fifth pedigerous somite, genital somite and four free abdominal somites; lateral surface of all abdominal somites with row of moderately long spinules. Dorsal and lateral surfaces of fifth pedigerous somite with numerous transverse rows of spinules, with spinules and subdistal sensilla near posterior margin; posterior margin with long spinules. Genital somite with numerous transverse rows of spinules, with spinules and subdistal sensilla near posterior margin; posterior margin with larger spinules dorsally and long spinules laterally. Genital opening positioned mid-ventrally near anterior margin of genital somite (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cletocamptus thailandensis sp. nov. male, allotype (A, B); paratype (C) A urosome, ventral view B antennule C segments 4 - 6 of antennule. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure9" httpUri="https://binary.pensoft.net/fig/817113" pageId="0" pageNumber="1">9A</figureCitation>
). First to third abdominal somites as those of female, but lacking hair-like spinules near posterior margin; spinule ornamentation lesser developed in comparison to those of female, each somite with three medial rows of spinules ventrally. Anal somite (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cletocamptus thailandensis sp. nov. male, allotype A habitus, dorsal view B habitus, lateral view C rostrum, frontal view D anal somite and caudal rami, dorsal view. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure8" httpUri="https://binary.pensoft.net/fig/817112" pageId="0" pageNumber="1">8D</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cletocamptus thailandensis sp. nov. male, allotype (A, B); paratype (C) A urosome, ventral view B antennule C segments 4 - 6 of antennule. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure9" httpUri="https://binary.pensoft.net/fig/817113" pageId="0" pageNumber="1">9A</figureCitation>
) as that of female.
</paragraph>
<caption doi="10.3897/zookeys.1151.96715.figure9" httpUri="https://binary.pensoft.net/fig/817113" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
<taxonomicName authorityName="Boonyanusith &amp; Wongkamhaeng" authorityYear="2023" class="Copepoda" family="Canthocamptidae" genus="Cletocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cletocamptus thailandensis" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis italics="true" pageId="0" pageNumber="1">Cletocamptus thailandensis</emphasis>
</taxonomicName>
sp. nov. male, allotype (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
); paratype (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
urosome, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
antennule
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
segments 4-6 of antennule. Scale bars: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
Caudal rami (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cletocamptus thailandensis sp. nov. male, allotype A habitus, dorsal view B habitus, lateral view C rostrum, frontal view D anal somite and caudal rami, dorsal view. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure8" httpUri="https://binary.pensoft.net/fig/817112" pageId="0" pageNumber="1">8D</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cletocamptus thailandensis sp. nov. male, allotype (A, B); paratype (C) A urosome, ventral view B antennule C segments 4 - 6 of antennule. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure9" httpUri="https://binary.pensoft.net/fig/817113" pageId="0" pageNumber="1">9A</figureCitation>
) as that of female, but inner margin with only two transverse rows of spinules. Seta IV and V, ca. 0.26
<normalizedToken originalValue="×">x</normalizedToken>
and ca. 0.59
<normalizedToken originalValue="×">x</normalizedToken>
body length, respectively. Seta VI relatively shorter than that of female.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Antennule (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cletocamptus thailandensis sp. nov. male, allotype (A, B); paratype (C) A urosome, ventral view B antennule C segments 4 - 6 of antennule. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure9" httpUri="https://binary.pensoft.net/fig/817113" pageId="0" pageNumber="1">9B, C</figureCitation>
) subchirocerate, six-segmented. First segment with two rows of spinules medially. Fourth segment bulbous, with aesthetasc on peduncle. Ultimate segment with two claw-shaped extensions subdistally. Aesthetascs to seta at base. Armature formula: 1-[1], 2-[9], 3-[6], 4-[7 + (1 + ae)], 5-[1], 6-[6 + (1+ae)].
</paragraph>
<paragraph pageId="0" pageNumber="1">Antenna, mandible, maxillule, maxilla, and maxilliped as those of female.</paragraph>
<paragraph pageId="0" pageNumber="1">
P1 (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cletocamptus thailandensis sp. nov. male, allotype A P 1 B P 2 C P 5. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure10" httpUri="https://binary.pensoft.net/fig/817114" pageId="0" pageNumber="1">10A</figureCitation>
) as that of female, except for the projection on inner distal corner of basis.
</paragraph>
<caption doi="10.3897/zookeys.1151.96715.figure10" httpUri="https://binary.pensoft.net/fig/817114" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
<taxonomicName authorityName="Boonyanusith &amp; Wongkamhaeng" authorityYear="2023" class="Copepoda" family="Canthocamptidae" genus="Cletocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cletocamptus thailandensis" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis italics="true" pageId="0" pageNumber="1">Cletocamptus thailandensis</emphasis>
</taxonomicName>
sp. nov. male, allotype
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
P1
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
P2
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
P5. Scale bars: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
P2 (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cletocamptus thailandensis sp. nov. male, allotype A P 1 B P 2 C P 5. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure10" httpUri="https://binary.pensoft.net/fig/817114" pageId="0" pageNumber="1">10B</figureCitation>
) as that of female, except for sexual dimorphic (slightly shorter and stronger) inner seta on Endp-2.
</paragraph>
<paragraph pageId="0" pageNumber="1">
P3 (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Cletocamptus thailandensis sp. nov. male, allotype A P 3 B P 4. Scale bar: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure11" httpUri="https://binary.pensoft.net/fig/817115" pageId="0" pageNumber="1">11A</figureCitation>
) as that of female in intercoxal sclerite, praecoxa, coxa, basis, and Exp. Endp three-segmented; Endp-1 short, wider than long; Endp-2 drawn out into apophysis, with row of spinules on outer margin and on frontal surface near base of Endp-3, apophysis surpassing tip of Endp-3; Endp-3 with two apical setae.
</paragraph>
<caption doi="10.3897/zookeys.1151.96715.figure11" httpUri="https://binary.pensoft.net/fig/817115" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
<taxonomicName authorityName="Boonyanusith &amp; Wongkamhaeng" authorityYear="2023" class="Copepoda" family="Canthocamptidae" genus="Cletocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cletocamptus thailandensis" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis italics="true" pageId="0" pageNumber="1">Cletocamptus thailandensis</emphasis>
</taxonomicName>
sp. nov. male, allotype
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
P3
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
P4. Scale bar: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
P4 (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Cletocamptus thailandensis sp. nov. male, allotype A P 3 B P 4. Scale bar: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure11" httpUri="https://binary.pensoft.net/fig/817115" pageId="0" pageNumber="1">11B</figureCitation>
) as that of female.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Left and right legs of P5 (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cletocamptus thailandensis sp. nov. male, allotype A P 1 B P 2 C P 5. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure10" httpUri="https://binary.pensoft.net/fig/817114" pageId="0" pageNumber="1">10C</figureCitation>
) fused medially at base, with medial notch indicating former separation between them; baseoendopod and Exp fused basally, with large notch indicating division between baseoendopodal and exopodal lobes. Baseoendopodal lobe with three pinnate spiniform setae apically, setae subequal in length; exopodal lobe with spinules at base of outermost apical seta, armed with four elements of which three apical ones pinnate and spiniform, outermost smooth and slender. Smooth soft seta on outer margin of basis, accompanied with row of spinules at base.
</paragraph>
<paragraph pageId="0" pageNumber="1">
P6 (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cletocamptus thailandensis sp. nov. male, allotype (A, B); paratype (C) A urosome, ventral view B antennule C segments 4 - 6 of antennule. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1151.96715.figure9" httpUri="https://binary.pensoft.net/fig/817113" pageId="0" pageNumber="1">9A</figureCitation>
) reduced to simple plate, without armature and ornamentation. Posterior margin smooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The species name is a noun proposed to reflect the name of the country,
<normalizedToken originalValue="Thailand">'Thailand'</normalizedToken>
, where the new species was encountered. It is in the nominative singular, gender masculine.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
The new species was collected from a water body at the base of an isolated limestone mountain (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographical location and details of sampling site A map of Thailand and location of sampling site in Rayong province (indicated by a triangular) B details of the mountain in which, the specimens were collected C details of the type locality." figureDoi="10.3897/zookeys.1151.96715.figure1" httpUri="https://binary.pensoft.net/fig/817105" pageId="0" pageNumber="1">1A, B</figureCitation>
) that is the distalmost part of the Cha Moon-Cha Mao Mountain Range in the Gong Din Subdistrict, Klaeng District, Rayong Province, Eastern Thailand. The coordinates of the type locality are
<geoCoordinate degrees="12" direction="north" minutes="45" orientation="latitude" precision="1" seconds="52.96" value="12.764711">12°45'52.96&quot;N</geoCoordinate>
,
<geoCoordinate degrees="101" direction="east" minutes="47" orientation="longitude" precision="1" seconds="55.16" value="101.79865">101°47'55.16&quot;E</geoCoordinate>
. The mountain is located beside a monastery known in Thai as 'Wat Tham Rakang
<normalizedToken originalValue="Thong">Thong'</normalizedToken>
(Tham Rakang Thong Monastery). The mountain and the water body are surrounded by a manmade concrete road (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographical location and details of sampling site A map of Thailand and location of sampling site in Rayong province (indicated by a triangular) B details of the mountain in which, the specimens were collected C details of the type locality." figureDoi="10.3897/zookeys.1151.96715.figure1" httpUri="https://binary.pensoft.net/fig/817105" pageId="0" pageNumber="1">1C</figureCitation>
), which separates the mountain and the monastery from the mangrove forests. The water is ca. 20 cm in depth and transparent with a brown color. The bottom of the water body is covered by leaf litter and filamentous algae. On the first sampling occasion, the temperature was 27.4 °C, the pH was 5.48, and the dissolved oxygen level was 5.5 mg L-1. The water electroconductivity and total dissolved solids were higher than 3,999
<normalizedToken originalValue="µS">µS</normalizedToken>
cm-1 and 2,000 ppm, respectively. On the second occasion, the temperature was 30.0 °C, the pH was 6.53, the electroconductivity was 2,158
<normalizedToken originalValue="µS">µS</normalizedToken>
cm-1, and the total dissolved solids were 1,080 ppm. The new species was collected with other zooplankton species, such as
<taxonomicName authorityName="Mrazek" authorityYear="1893" class="Hexanauplia" family="Phyllognathopodidae" genus="Phyllognathopus" kingdom="Animalia" lsidName="Phyllognathopus paludosus" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="paludosus">
<emphasis italics="true" pageId="0" pageNumber="1">Phyllognathopus paludosus</emphasis>
</taxonomicName>
(Mrazek, 1893),
<taxonomicName authorityName="Onabamiro" authorityYear="1957" class="Hexanauplia" family="Cyclopidae" genus="Mesocyclops" kingdom="Animalia" lsidName="Mesocyclops ogunnus" order="Cyclopoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ogunnus">
<emphasis italics="true" pageId="0" pageNumber="1">Mesocyclops ogunnus</emphasis>
</taxonomicName>
Onabamiro, 1957 (the most dominant),
<taxonomicName authorityName="Lindberg" authorityYear="1954" class="Hexanauplia" family="Cyclopidae" genus="Apocyclops" kingdom="Animalia" lsidName="Apocyclops borneoensis" order="Cyclopoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis italics="true" pageId="0" pageNumber="1">Apocyclops borneoensis</emphasis>
</taxonomicName>
Lindberg, 1954,
<taxonomicName authorityName="Kiefer" authorityYear="1927" class="Hexanauplia" family="Cyclopidae" genus="Tropocyclops" kingdom="Animalia" lsidName="Tropocyclops" order="Cyclopoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Tropocyclops</emphasis>
</taxonomicName>
sp.,
<taxonomicName authorityName="Boeck" authorityYear="1865" class="Hexanauplia" family="Ameiridae" genus="Nitokra" kingdom="Animalia" lsidName="Nitokra" order="Harpacticoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nitokra</emphasis>
</taxonomicName>
sp., the rotifer of the genus
<taxonomicName class="Eurotatoria" family="Testudinellidae" genus="Testudinella" kingdom="Animalia" lsidName="Testudinella" order="Flosculariaceae" pageId="0" pageNumber="1" phylum="Rotifera" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Testudinella</emphasis>
</taxonomicName>
Bory de St. Vincent, 1822, the cladoceran of the genus
<taxonomicName authorityName="Smirnov" authorityYear="1989" class="Insecta" genus="Leberis" kingdom="Animalia" lsidName="Leberis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Leberis</emphasis>
</taxonomicName>
Smirnov, 1989, and ostracods. Aquatic insects belonging to the order
<taxonomicName authorityName="Smirnov" authorityYear="1989" class="Insecta" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="order">Ephemeroptera</taxonomicName>
and the family
<taxonomicName class="Insecta" family="Chironomidae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
were also observed.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">The species has been known from the type locality only.</paragraph>
</subSubSection>
</treatment>
</document>