treatments-xml/data/3A/E6/0F/3AE60F1A7233C6737C05C1E209B27BBC.xml

<document ID-CLB-Dataset="28264" ID-DOI="http://dx.doi.org/10.3897/phytokeys.110.25846" ID-GBIF-Dataset="44b94249-6f5a-4a6c-8039-c441517a1144" ID-PMC="PMC6224366" ID-Pensoft-Pub="1314-2003-110-23" ID-Pensoft-UUID="FFD8FFA5FFABFC45FF8EFE1DD80CFF9C" ID-PubMed="30416344" ID-Zenodo-Dep="1481316" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-2003-110-23" ModsDocOrigin="PhytoKeys " ModsDocTitle="Begoniayenyeniae (Begoniaceae), a new species from Endau Rompin National Park, Johor, Malaysia" checkinTime="1553125356695" checkinUser="pensoft" docAuthor="Tan, Joanne Pei-Chih, Tam, Sheh May &amp; Kiew, Ruth" docDate="2018" docId="3AE60F1A7233C6737C05C1E209B27BBC" docLanguage="en" docName="PhytoKeys 110: 23-37" docOrigin="PhytoKeys 110" docSource="http://dx.doi.org/10.3897/phytokeys.110.25846" docTitle="Begonia rajah Ridl. ex Rolfe" docType="treatment" docVersion="8" id="FFD8FFA5FFABFC45FF8EFE1DD80CFF9C" lastPageNumber="23" masterDocId="FFD8FFA5FFABFC45FF8EFE1DD80CFF9C" masterDocTitle="Begonia yenyeniae (Begoniaceae), a new species from Endau Rompin National Park, Johor, Malaysia" masterLastPageNumber="37" masterPageNumber="23" pageNumber="23" updateTime="1732940009691" updateUser="ExternalLinkService">
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<mods:title id="67E6BCFDAE3E70A1C42A5C210C484857">Begonia yenyeniae (Begoniaceae), a new species from Endau Rompin National Park, Johor, Malaysia</mods:title>
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<mods:namePart id="DF654E2F46923017A89525AC96049963">Tan, Joanne Pei-Chih</mods:namePart>
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<mods:namePart id="AB7A2B07E2E1EF2CAFF37365456CF659">Tam, Sheh May</mods:namePart>
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<mods:namePart id="8A5678BC26335AAF61BC367015EB2F42">Kiew, Ruth</mods:namePart>
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<mods:title id="074FBEE2D3AAC213EA7C0CFCE306FD25">PhytoKeys</mods:title>
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<mods:date id="69AA1F31629752B4EA83D60F870B049C">2018</mods:date>
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<mods:number id="E8ECBDB5549A8C812077FAA5F60F7631">110</mods:number>
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<mods:classification id="FE570FF515D979226307408B238B14CD">journal article</mods:classification>
<mods:identifier id="150DB7BD356B13A127EF959B8D9423AF" type="DOI">http://dx.doi.org/10.3897/phytokeys.110.25846</mods:identifier>
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<treatment id="3AE60F1A7233C6737C05C1E209B27BBC" ID-GBIF-Taxon="154472608" LSID="urn:lsid:plazi:treatment:3AE60F1A7233C6737C05C1E209B27BBC" httpUri="http://treatment.plazi.org/id/3AE60F1A7233C6737C05C1E209B27BBC" lastPageNumber="23" pageId="0" pageNumber="23" scope_family="Begoniaceae">
<subSubSection id="83FF4E21FFDE943FE1D8427FD0D84767" pageId="0" pageNumber="23" type="nomenclature">
<paragraph id="998E3C26763E78D79C65911217870FEA" pageId="0" pageNumber="23">
<taxonomicName id="52C879E6B00E57F766C797A9801205AE" ID-CoL="7SH5X" LSID="3AE60F1A-7233-C673-7C05-C1E209B27BBC" authority="Ridl. ex Rolfe" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia rajah" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="species" species="rajah">Begonia rajah Ridl. ex Rolfe</taxonomicName>
<figureCitation id="BCDAE0189DA207E39ACF1F00F5441409" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Watercolour painting of Begonia rajah of an original wild-collected plant grown in the Botanic Gardens, Singapore. (Reproduced with permission of the Singapore Botanic Gardens)" figureDoi="10.3897/phytokeys.110.25846.figure1" httpUri="https://binary.pensoft.net/fig/239605" pageId="0" pageNumber="23">Figure 1</figureCitation>
<figureCitation id="FA7DEA06DFB37C1977A424E591F67E73" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Begonia rajah Ridley (cultivated) A Upper leaf surface B under surface C Young blade." figureDoi="10.3897/phytokeys.110.25846.figure3" httpUri="https://binary.pensoft.net/fig/239612" pageId="0" pageNumber="23">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5CEEE858ABA8E721787C7B4568142FDA" pageId="0" pageNumber="23" type="reference_group">
<paragraph id="7A80036E327EDA90588956639BB6C3E9" pageId="0" pageNumber="23">
<normalizedToken id="5C6979FE83CD32A72A8ECD73290AAA28" originalValue="Gardeners’">Gardeners'</normalizedToken>
Chronicle XVI (1894) 213, nom. nud.; Rolfe, Bulletin of Miscellaneous Information (1914) 327; Irmscher, Mitteilungen aus dem Institut 
<normalizedToken id="E6A32B15DD110BCC86D3D6BD5E9DACFC" originalValue="für">fuer</normalizedToken>
allgemeine Botanik in Hamburg 8 (1929) 96; Tebbitt, Begonias: Cultivation, Identification, and Natural History (2005) 198; Kiew, Begonias of Peninsular Malaysia (2005) 216, pro parte.
</paragraph>
</subSubSection>
<subSubSection id="1ECE790EA9306FCC3267815AA1F6FA4E" pageId="0" pageNumber="23" type="section">
<paragraph id="4403C053D49F41AE677871C376392AF5" pageId="0" pageNumber="23">Section.</paragraph>
<paragraph id="35C62164FF79C9F0385C110E1A89C4AA" pageId="0" pageNumber="23">
<taxonomicName id="1DD12EA1494E8197614E937EE90E8F04" class="Magnoliopsida" family="Begoniaceae" genus="Jackia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Jackia" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="genus">
<emphasis id="940B1A8A60C7D5E2635E6EFECCA6F5D2" italics="true" pageId="0" pageNumber="23">Jackia</emphasis>
</taxonomicName>
M.Hughes
</paragraph>
</subSubSection>
<subSubSection id="5E5310DC007320BA52E38C5DF9EBFB43" pageId="0" pageNumber="23" type="materials_examined">
<paragraph id="8F71F37B1BF32B83D5FBB704D0A58D42" pageId="0" pageNumber="23">Type.</paragraph>
<paragraph id="EE1A27893E7FD39319153C74305C7BD4" pageId="0" pageNumber="23">
Peninsular Malaysia. Terengganu, 1892, 
<emphasis id="F7D503B0E49E4D29B2FBE00895E1F8FA" italics="true" pageId="0" pageNumber="23">Native collector s.n.</emphasis>
(holotype SING ex K!). 
<emphasis id="787A0F2E350B8CD905C3A5A95FBEDA76" bold="true" pageId="0" pageNumber="23">Description.</emphasis>
Dwarf herb with rhizomatous stem. Indumentum of soft hairs, usually pale pink, dense on stipule (1.5-2 mm long), petiole (cream-coloured, 2-4 mm), leaf margin (1-1.5 mm) and veins beneath (2-2.5 mm), peduncle (cream-coloured, 2-2.5 mm). 
<emphasis id="71632A507EDABDDF7E4CC7C564F27751" bold="true" pageId="0" pageNumber="23">Stems</emphasis>
creeping, ca. 9 mm thick, light yellowish-green; stipules lanceolate-oblong or triangular, 15-20 
<normalizedToken id="2BF801FA778182ED2D9FADCF2AA92187" originalValue="×">x</normalizedToken>
6-10 mm, apex attenuate terminating in a hair, keeled from base to apex, margin entire and opaque, brownish-pink to scarlet, persistent. 
<emphasis id="32E08F2E5432DCA66E401F436589336F" bold="true" pageId="0" pageNumber="23">Leaves</emphasis>
tufted, alternate, ca. 5 mm apart; petioles suberect, terete, ca. 5 mm across, 8-25 cm long, light yellowish-pink; blades rigid, succulent, glabrous, broadly ovate with an abruptly acute apex or radical oblique cordate, 7-15 
<normalizedToken id="D171844B7144952FB637A582F33C579B" originalValue="×">x</normalizedToken>
6-15 cm, prominently bullate, margin ciliate, angular with abruptly acute teeth, basal lobes cordate, slightly overlapping when mature, notably dark brownish-purple blotches, polished shining surface, young blades brownish-pink to vivid brownish-red, paler beneath; veins palmate, prominent towards the base and slightly impressed near margin, prominent beneath, lateral veins ca. 1-3 pairs, greenish-yellow when young, yellowish-green when mature. 
<emphasis id="857344FC528AF110A844E6089567E12A" bold="true" pageId="0" pageNumber="23">Inflorescences</emphasis>
axillary, almost erect, many-flowered cyme, 20-25 cm long, peduncles 7-12 cm long, brownish-pink, two main branches ca. 1.3 cm long, pedicels slender, ca. 1 cm; bracts in pairs at node of peduncle, glabrous, broadly ovate, bowl-shaped, subacute, margin laciniate 5-8 
<normalizedToken id="C0C27028D7477F74FA5C0C5F318D9E81" originalValue="×">x</normalizedToken>
7.5-8 mm, light yellowish-pink, persistent; bracteole narrowly ovate, 2-3 mm long, pale yellowish-pink. 
<emphasis id="4246F2608EDC480638E0C67A8D10ED61" bold="true" pageId="0" pageNumber="23">Male flowers</emphasis>
with 4 tepals, margin entire, ca. 1.8 cm across, rose pink; outer tepals 2, obovate-orbicular or elliptic-ovate, 6-8 
<normalizedToken id="994BC43AE45F30EA6084F0F33BF66901" originalValue="×">x</normalizedToken>
5-8 mm, inner 2 tepals obovate or narrow oblong, apex rounded or sometimes retuse and impressed along centre, ca. 5 
<normalizedToken id="D03869D5B859634254DDF15C9782861C" originalValue="×">x</normalizedToken>
3 mm; stamen mass globose, symmetric, 3 
<normalizedToken id="52C28291A018525D6E2EB5503585FDCC" originalValue="×">x</normalizedToken>
2.5 mm; anthers broadly obovoid-oblong, tip rounded, ca. 0.7 mm long, dehiscing through 2 longitudinal slits. 
<emphasis id="AA3774BED0B2B7792C6BCF81F300B58F" bold="true" pageId="0" pageNumber="23">Female flowers</emphasis>
with a pale pink ovary with darker pinkish margin, 7-8 mm, locules 3, placentation axile, 1 placenta per locule, each placenta usually with 2 minute branches at the base, usually 3 tepals rarely 2, outer 2 tepals widely elliptic-ovate, concave at centre, ca. 6-7 mm, pale pink; styles and stigmas 3, 2-2.5 mm, style light greenish-yellow, stigma pale yellow, papillose, spiral band. Capsules ca. 1.2 cm across, locules 3, splitting longitudinally between locules, surface glabrous, dangling on a fine, thread-like pedicel 1 cm long. 
<emphasis id="E6F1CEC0C88377C9C3A26AE83CFD175F" bold="true" pageId="0" pageNumber="23">Seeds</emphasis>
not seen by us.
</paragraph>
<caption id="B51C504333C8630C2D889435CF523CB4" ID-Zenodo-Dep="1481328" doi="10.3897/phytokeys.110.25846.figure3" httpUri="https://binary.pensoft.net/fig/239612" pageId="0" pageNumber="23" start="Figure 3" startId="F3">
<paragraph id="7737211CA76D41346163D643C5F8299A" pageId="0" pageNumber="23">
<emphasis id="9F509F0E9C305CA56447CEC501623BAA" bold="true" pageId="0" pageNumber="23">Figure 3.</emphasis>
<taxonomicName id="F482AA8DFD9AF556F21BAF358F6D0610" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia rajah" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="species" species="rajah">
<emphasis id="B757F4B5F6234CC88F74BBB98CAEF1D0" italics="true" pageId="0" pageNumber="23">Begonia rajah</emphasis>
</taxonomicName>
Ridley (cultivated) 
<emphasis id="8C79208738F80FBA44C051152EE5FE4F" bold="true" pageId="0" pageNumber="23">A</emphasis>
Upper leaf surface 
<emphasis id="13AE167AC1230B94390DB6DEAF93C555" bold="true" pageId="0" pageNumber="23">B</emphasis>
under surface 
<emphasis id="19C4E44B3CD900FA120E27DF0ED9DE67" bold="true" pageId="0" pageNumber="23">C</emphasis>
Young blade.
</paragraph>
</caption>
<caption id="7464E6E64C511C35180B4B4093B531C1" ID-Zenodo-Dep="1481330" doi="10.3897/phytokeys.110.25846.figure4" httpUri="https://binary.pensoft.net/fig/239613" pageId="0" pageNumber="23" start="Figure 4" startId="F4">
<paragraph id="E876C6B4B72D9488625011FE848CCBCD" pageId="0" pageNumber="23">
<emphasis id="FF399B78FE77087970B8CF585D0508F1" bold="true" pageId="0" pageNumber="23">Figure 4.</emphasis>
<taxonomicName id="94497AE3D6A633799FEC9A1D7152DDBC" authorityName="Kiew" authorityYear="2005" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia reginula" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="species" species="reginula">
<emphasis id="696727FAF905AE539FAFB24975EBA4FD" italics="true" pageId="0" pageNumber="23">Begonia reginula</emphasis>
</taxonomicName>
Kiew 
<emphasis id="0409C92D8DC8E2DF2507C3402A68E18A" bold="true" pageId="0" pageNumber="23">A</emphasis>
<emphasis id="8065D0228749A281AF45F9A7E374221C" italics="true" pageId="0" pageNumber="23">In situ</emphasis>
: clinging to moss-covered granite boulders 
<emphasis id="1F72CD3226FF6D57A90D0C812D557503" bold="true" pageId="0" pageNumber="23">B</emphasis>
Young blade 
<emphasis id="2A62C732E2B55A00AFC2AD1340E3BC21" bold="true" pageId="0" pageNumber="23">C</emphasis>
Mature leaf of coloured form.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="12F1397910EACB12844073534D9254B5" pageId="0" pageNumber="23" type="distribution">
<paragraph id="B31A5481424BE8FE34715440EB9BCF21" pageId="0" pageNumber="23">Distribution.</paragraph>
<paragraph id="59BE5296016C171C770305D3E819240F" pageId="0" pageNumber="23">Endemic in Peninsular Malaysia, Terengganu (without specific locality). Apparently restricted in its distribution because, despite continuous botanical collecting in Terengganu and elsewhere in Peninsular Malaysia, it has not been re-found since it was first collected in 1892 (Kiew, 2005).</paragraph>
<caption id="D6631C285882042F5F9C3043913279EE" ID-Zenodo-Dep="1481332" doi="10.3897/phytokeys.110.25846.figure5" httpUri="https://binary.pensoft.net/fig/239609" pageId="0" pageNumber="23" start="Figure 5" startId="F5">
<paragraph id="AB04E30B67CCC111D0E1C7F97F33E9CE" pageId="0" pageNumber="23">
<emphasis id="1962489F204F6EC8C52D3048D3D4CD27" bold="true" pageId="0" pageNumber="23">Figure 5.</emphasis>
Part of the Bayesian 50% majority rule consensus tree from the analysis of chloroplast 
<emphasis id="23F2C37AF87EA4D96D35E02012E98D17" italics="true" pageId="0" pageNumber="23">ndhF</emphasis>
- 
<emphasis id="C4B269117F560C0318469DCB90FEE0C2" italics="true" pageId="0" pageNumber="23">rpl32</emphasis>
region depicting relationships amongst 
<taxonomicName id="C73754AA3C341B308963AF9BCF9C2D54" class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. foxworthyi" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="foxworthyi">
<emphasis id="A7803F22EB3AB9BDCCC33C64A523F1D5" italics="true" pageId="0" pageNumber="23">B. foxworthyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="22654F89CCCB55FFAFBA8AC86C8AE026" class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. rajah" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="rajah">
<emphasis id="75C334DD7F3F338CDC82536E41BA5E25" italics="true" pageId="0" pageNumber="23">B. rajah</emphasis>
</taxonomicName>
, 
<taxonomicName id="7F685FD7C7286B0FF5BC2D8A6127A0D6" class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. reginula" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="reginula">
<emphasis id="E14CA26C13D3C00D33B5846CD7D0A472" italics="true" pageId="0" pageNumber="23">B. reginula</emphasis>
</taxonomicName>
and 
<taxonomicName id="AF4D4F906AD9436408083A0029A9298D" class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. yenyeniae" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="yenyeniae">
<emphasis id="1B40C2C604EB0A56A2F20378B84D6F63" italics="true" pageId="0" pageNumber="23">B. yenyeniae</emphasis>
</taxonomicName>
. The locality is indicated at the end of the species name for species with multiple samples. Numbers on the branches indicate Bayesian posterior probability (BPP) support values.
</paragraph>
</caption>
<caption id="3872EDFE9849FFA654ED556A72FF705D" ID-Table-UUID="3872EDFE9849FFA654ED556A72FF705D" httpUri="http://table.plazi.org/id/3872EDFE9849FFA654ED556A72FF705D" pageId="0" pageNumber="23" start="Table 1" startId="T1">
<paragraph id="F2A1ABAEB37F2CED39F8044D35D51303" pageId="0" pageNumber="23">
<emphasis id="9DACAF9FA2A289B1CCA3B22FE0A0DFD0" bold="true" pageId="0" pageNumber="23">Table 1.</emphasis>
<taxonomicName id="3C211EDA113A482E96A6756CCFAA02F4" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia rajah" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="species" species="rajah">
<emphasis id="08902ACBC38F6AD73C465626B1C2790A" italics="true" pageId="0" pageNumber="23">Begonia rajah</emphasis>
</taxonomicName>
, 
<taxonomicName id="30CA4766BF51019B3A89FE1766709ADD" class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. yenyeniae" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="yenyeniae">
<emphasis id="518916807570303EB4E4D85E20FA5DC4" italics="true" pageId="0" pageNumber="23">B. yenyeniae</emphasis>
</taxonomicName>
, 
<taxonomicName id="354ADC6520F51459B590E24A396D51F8" class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. reginula" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="reginula">
<emphasis id="4A2A8995D9BEE613865C0ACA1115963E" italics="true" pageId="0" pageNumber="23">B. reginula</emphasis>
</taxonomicName>
and 
<taxonomicName id="ECD5A988042CBDDE780A17AB0CB1FAB2" class="Monocotyledoneae" family="Zingiberaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="B. foxworthyi" order="Scitamineae" pageId="0" pageNumber="23" phylum="Angiospermae" rank="species" species="foxworthyi">
<emphasis id="E22E16D48E6E31B2A8F5271C5B09B422" italics="true" pageId="0" pageNumber="23">B. foxworthyi</emphasis>
</taxonomicName>
compared.
</paragraph>
</caption>
<paragraph id="89F25F2DB7245E6F54BC9C40190ACB4A" pageId="0" pageNumber="23">
<table id="4A65420CE893FEF0101139D863106CE1" pageId="0" pageNumber="23">
<tr id="34158FD534F1876806206B43EC0568B2" pageId="0" pageNumber="23">
<th id="AB3E66FEBD982B5A0A1CDF5FFB5AA812" colspan="1" pageId="0" pageNumber="23" rowspan="1">Character</th>
<th id="4F5B22A1B7A62C52AF006F6D7B0756E8" colspan="1" pageId="0" pageNumber="23" rowspan="1">
<taxonomicName id="17F88C7BBDA44A1CF449584B6ADC19AC" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia rajah" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="species" species="rajah">
<emphasis id="F08589616A69CE1D27DA34B11D972335" italics="true" pageId="0" pageNumber="23">Begonia rajah</emphasis>
</taxonomicName>
</th>
<th id="8CD4D9C1B1623F76CB7974EA29E530D6" colspan="1" pageId="0" pageNumber="23" rowspan="1">
<taxonomicName id="B6E066736BF680DFAD808F224B59D950" authorityName="J. P. C. Tan" authorityYear="2018" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia yenyeniae" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="species" species="yenyeniae">
<emphasis id="5A5FAAB09308758E735ABE00C1FD3384" italics="true" pageId="0" pageNumber="23">Begonia yenyeniae</emphasis>
</taxonomicName>
sp. nov.
</th>
<th id="50B35BF59B4FA736910F7DE245F30CA0" colspan="1" pageId="0" pageNumber="23" rowspan="1">
<taxonomicName id="D1653E50B324271961156A66A7F7097C" authorityName="Kiew" authorityYear="2005" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia reginula" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="species" species="reginula">
<emphasis id="40110CA1FC3B4AE6EBA6A0B249DD9C16" italics="true" pageId="0" pageNumber="23">Begonia reginula</emphasis>
</taxonomicName>
</th>
<th id="58AD1DE35DDD3B95CD5AF4E52F8C7244" colspan="1" pageId="0" pageNumber="23" rowspan="1">
<taxonomicName id="B044129A86047E27422E97B6A1906075" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Begonia foxworthyi" order="Cucurbitales" pageId="0" pageNumber="23" phylum="Tracheophyta" rank="species" species="foxworthyi">
<emphasis id="B108732590557A34C59B363695200FD0" italics="true" pageId="0" pageNumber="23">Begonia foxworthyi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="263F320C1ACCF974E35930EF02C47232" pageId="0" pageNumber="23">
<td id="03863B0603370D61CB506E26E6F3C840" colspan="1" pageId="0" pageNumber="23" rowspan="1">Leaf: texture</td>
<td id="A42A725D618F14FB822C0536E3F76518" colspan="1" pageId="0" pageNumber="23" rowspan="2">thickly succulent polished shining prominent bullate</td>
<td id="24B9D682CB04AD9721423E4816418218" colspan="1" pageId="0" pageNumber="23" rowspan="1">thinly succulent</td>
<td id="DEA0C78B8C5C589F4733AEF308EAAD43" colspan="1" pageId="0" pageNumber="23" rowspan="2">thinly succulent slightly raised in between veins</td>
<td id="DDC768594C9437A58FCDF7F3F8EB4A42" colspan="1" pageId="0" pageNumber="23" rowspan="1">thinly succulent</td>
</tr>
<tr id="7D0A74C6D5761193835BBF1447177CAE" pageId="0" pageNumber="23">
<td id="9B3848713EC09726894380784C896A72" colspan="1" pageId="0" pageNumber="23" rowspan="1">Surface</td>
<td id="90B3EB09F8AFB81B44133FC6C96F1456" colspan="1" pageId="0" pageNumber="23" rowspan="1">moderately raised in between veins</td>
<td id="4BE5F82A3AFBDDBE9F5DF6F47268F9E7" colspan="1" pageId="0" pageNumber="23" rowspan="1">glossy, veins slightly impressed</td>
</tr>
<tr id="C10D222A2D950ABDC20ADD9EB4EC5D5D" pageId="0" pageNumber="23">
<td id="2830E073B8C4336A0A01492FA76A433F" colspan="1" pageId="0" pageNumber="23" rowspan="3">Leaf shape</td>
<td id="5A1E933A4A639CE63047F3573E1338CD" colspan="1" pageId="0" pageNumber="23" rowspan="1">subrotund</td>
<td id="FBE307B9065CB4BFB6F5E97C497A837E" colspan="1" pageId="0" pageNumber="23" rowspan="1">orbicular-reniform</td>
<td id="0E0FD13DC8B79E0DCCAE452A788B9B1E" colspan="1" pageId="0" pageNumber="23" rowspan="1">broadly ovate</td>
<td id="E3B05CA717BDA420E37012BD937AB749" colspan="1" pageId="0" pageNumber="23" rowspan="1">almost rotund apex short- pointed</td>
</tr>
<tr id="7AE3FD36A513DE3852E478B369ACC867" pageId="0" pageNumber="23">
<td id="AF5A72BA44D692E88092477478BB6B51" colspan="1" pageId="0" pageNumber="23" rowspan="1">apex abruptly acute</td>
<td id="C54992845211550A3297966F27184649" colspan="1" pageId="0" pageNumber="23" rowspan="1">apex rounded</td>
<td id="597DCDAC500E6786E3BFF8E788F3C4F8" colspan="1" pageId="0" pageNumber="23" rowspan="1">apex abruptly attenuate angular</td>
<td id="D9A14894386FB0322A2A21C80A585889" colspan="1" pageId="0" pageNumber="23" rowspan="1">not angular</td>
</tr>
<tr id="45FB18AE4E06659283DB835B05BF3827" pageId="0" pageNumber="23">
<td id="241306ED7CD2AE5926CB94BC42781D63" colspan="1" pageId="0" pageNumber="23" rowspan="1">angular</td>
<td id="3813E67B47A49CCDC473EC6192A29F58" colspan="1" pageId="0" pageNumber="23" rowspan="1">scarcely angular</td>
<td id="D0DA2885044F4B848AE24C164CAE9E8B" colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
<td id="3110D73DAF5D641E0FC1103B288BAAD5" colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
</tr>
<tr id="E5A88B71F0BB03E43BEF31351D016BE7" pageId="0" pageNumber="23">
<td id="A8AA414302C8C57DC0942B4C35DEDC28" colspan="1" pageId="0" pageNumber="23" rowspan="1">Tepal no. in male flower</td>
<td id="C53329EB0DBEB21C18C4AE71FFF294FC" colspan="1" pageId="0" pageNumber="23" rowspan="1">4</td>
<td id="CFFEB992316FFFD69D24ECA7140382C4" colspan="1" pageId="0" pageNumber="23" rowspan="1">4</td>
<td id="EDE639A6E7ABCE03B926BB0A7E2F160E" colspan="1" pageId="0" pageNumber="23" rowspan="1">2</td>
<td id="D0351941A1ED83DFE3C27EA93B756EDD" colspan="1" pageId="0" pageNumber="23" rowspan="1">2</td>
</tr>
<tr id="04B50E78851510CB97D40BB4CB450DF3" pageId="0" pageNumber="23">
<td id="35C8DCB2EE4510DE2A7332AFDF5D4F7B" colspan="1" pageId="0" pageNumber="23" rowspan="1">Tepal no. in female flower</td>
<td id="7D123D5CFC34C3AB07F1B4B18857519C" colspan="1" pageId="0" pageNumber="23" rowspan="1">3</td>
<td id="F968F999B088D66B9263BCD9D9A3F0B4" colspan="1" pageId="0" pageNumber="23" rowspan="1">3</td>
<td id="6F0FBCC25D053DF296CCCA81E3D83252" colspan="1" pageId="0" pageNumber="23" rowspan="1">2</td>
<td id="D78F88D8F8B3C01C104F2CE21093CF2B" colspan="1" pageId="0" pageNumber="23" rowspan="1">2</td>
</tr>
<tr id="6C98FF38A92F5794BB16CE255F518BA6" pageId="0" pageNumber="23">
<td id="E7B3E13344DD502ECABD1114BEFDB5B5" colspan="1" pageId="0" pageNumber="23" rowspan="1">Outer tepal size (mm)</td>
<td id="66DB4563D2AFD52BF274F1F7C41E6E15" colspan="1" pageId="0" pageNumber="23" rowspan="1">
6-8 
<normalizedToken id="1633F7893A8F1A8766C087CC64E624A1" originalValue="×">x</normalizedToken>
5-8
</td>
<td id="A418FFD114A3AE20CF2C704247D85A38" colspan="1" pageId="0" pageNumber="23" rowspan="1">
5-7 
<normalizedToken id="523A6BA9E254E68ED35A870BF2451CC5" originalValue="×">x</normalizedToken>
6
</td>
<td id="F4CDA44EFA481266AE7C84181037FB61" colspan="1" pageId="0" pageNumber="23" rowspan="1">
6-10 
<normalizedToken id="1A549DF6E746621B1D6880AE8E3EA249" originalValue="×">x</normalizedToken>
9-11
</td>
<td id="D34D5B9570D5CE399F1B87E47E6C2EC0" colspan="1" pageId="0" pageNumber="23" rowspan="1">
4-6 
<normalizedToken id="DBE54B66236B41CCA81FDD610A7B2EB8" originalValue="×">x</normalizedToken>
4-6
</td>
</tr>
<tr id="8BDC7D0E66B26FF4B20E0B6E63FD499C" pageId="0" pageNumber="23">
<td id="044A804F006FD71155F67B3E8C03DBC3" colspan="1" pageId="0" pageNumber="23" rowspan="1">Outer tepal: shape</td>
<td id="7BB562B8ED9549DEC014E4363A0961E8" colspan="1" pageId="0" pageNumber="23" rowspan="1">widely ovate or cordate</td>
<td id="8E2F0EB983CD2A0468EE609FDC18189A" colspan="1" pageId="0" pageNumber="23" rowspan="1">rotund</td>
<td id="BE43ABABC73F7B57BCAE69307201AFF0" colspan="1" pageId="0" pageNumber="23" rowspan="1">widely ovate or rotund</td>
<td id="44AFFE4CCF2CB39741AD59B2B809D733" colspan="1" pageId="0" pageNumber="23" rowspan="1">broadly ovate to rotund rounded</td>
</tr>
<tr id="ECBF2B330626B4FEA1B4F656F76B7709" pageId="0" pageNumber="23">
<td id="52173DEA9D6868DC34F1A3E27B5AF7F1" colspan="1" pageId="0" pageNumber="23" rowspan="1">apex</td>
<td id="EAF0D0742B2A2A96E7A652320D8E2214" colspan="1" pageId="0" pageNumber="23" rowspan="1">obtuse</td>
<td id="E7FD105CD40116C0960DCDD8896AFAC1" colspan="1" pageId="0" pageNumber="23" rowspan="1">rounded</td>
<td id="198A27B88F9A5D3343FE4ABC894E830B" colspan="1" pageId="0" pageNumber="23" rowspan="1">rounded or acute subcordate</td>
<td id="B78FC19F1164E850E8497F076DB56269" colspan="1" pageId="0" pageNumber="23" rowspan="1">subcordate</td>
</tr>
<tr id="20B3FB079DFE06D5DDE192D382125749" pageId="0" pageNumber="23">
<td id="6290920E9E7648F16F2D7E4BCFA2BF00" colspan="1" pageId="0" pageNumber="23" rowspan="1">base</td>
<td id="A3A98F1712AB6C805A3D2D4B42A9A57D" colspan="1" pageId="0" pageNumber="23" rowspan="1">subcordate</td>
<td id="55E9F4541C6807939EA8E4A422C194A7" colspan="1" pageId="0" pageNumber="23" rowspan="1">rounded</td>
<td id="6E90398BBD0D9BC06046F28D8336CA5C" colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
<td id="BEF07B3F87C7DFCC2C22A8FB3CA09369" colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
</tr>
<tr id="EF614D8E7AC558A87750FDC8352448FC" pageId="0" pageNumber="23">
<td id="8C43146069DACDEFC16EF202391DD55A" colspan="1" pageId="0" pageNumber="23" rowspan="1">Stipule: shape</td>
<td id="0AFF9BBF6580C13D5F1C0EBA6A13B3ED" colspan="1" pageId="0" pageNumber="23" rowspan="1">lanceolate-oblong or triangular</td>
<td id="AB546C7B5137D31992A1DB28563617E3" colspan="1" pageId="0" pageNumber="23" rowspan="1">narrowly triangular,</td>
<td id="9F6E02686C97EB5A437F2D0D1ED243FE" colspan="1" pageId="0" pageNumber="23" rowspan="1">narrowly triangular,</td>
<td id="F38C9AD6EAAC0E2957CC9999B0672545" colspan="1" pageId="0" pageNumber="23" rowspan="1">narrowly triangular</td>
</tr>
<tr id="442FCB1BD36F020C6C6D1A45F35EF1BB" pageId="0" pageNumber="23">
<td id="70207051535F4EB46B9F2E5E0887E34F" colspan="1" pageId="0" pageNumber="23" rowspan="1">size (mm)</td>
<td id="19C2D2924DA4A7584B6A5F4518DC269E" colspan="1" pageId="0" pageNumber="23" rowspan="1">
15-20 
<normalizedToken id="1817A3636A88263E3039B8D318E50491" originalValue="×">x</normalizedToken>
10-12
</td>
<td id="F980075B7D3686EB0FE61287FC29BC49" colspan="1" pageId="0" pageNumber="23" rowspan="1">
9-12 
<normalizedToken id="D5675ABC8EEE13271644A5B7EB9213CD" originalValue="×">x</normalizedToken>
3-4
</td>
<td id="B948B7EAB1EE9F7B07630266FEA05AEF" colspan="1" pageId="0" pageNumber="23" rowspan="1">
5-7 
<normalizedToken id="CCF7A49AA9358891055A2EF0F06230B3" originalValue="×">x</normalizedToken>
2-3
</td>
<td id="4F6C71B1BE84C17318F58A6E597F5710" colspan="1" pageId="0" pageNumber="23" rowspan="1">
7-12 
<normalizedToken id="6582ED117F8C545A2223D1267C12ABC6" originalValue="×">x</normalizedToken>
2-5
</td>
</tr>
<tr id="954C90367CD50AA1E7E3332687363563" pageId="0" pageNumber="23">
<td id="24AFA0FFE5FE803280F38D0D77CAD36D" colspan="1" pageId="0" pageNumber="23" rowspan="1">colour</td>
<td id="5CAEDB5F9C20B20840B07862CE670162" colspan="1" pageId="0" pageNumber="23" rowspan="1">Brownish-pink to scarlet</td>
<td id="CD80E7446BE7A913A4046C68BD41FF96" colspan="1" pageId="0" pageNumber="23" rowspan="1">light yellowish-green sometimes with a faint pink tinge</td>
<td id="30B4B75A3C0D2D9EBED12E2EC951910C" colspan="1" pageId="0" pageNumber="23" rowspan="1">pale greenish yellow or reddish yellow</td>
<td id="817FAC1A999165B62663AC08BB863BED" colspan="1" pageId="0" pageNumber="23" rowspan="1">red</td>
</tr>
<tr id="BF3F8EB48F4BE70F37EC603C4974DFEB" pageId="0" pageNumber="23">
<td id="625D33C7C0159B1FC553B2938B9B01E9" colspan="1" pageId="0" pageNumber="23" rowspan="1">Keel on stipules</td>
<td id="9221BF71CAB93F4A5D144DD72BCD0B5B" colspan="1" pageId="0" pageNumber="23" rowspan="1">from base to apex</td>
<td id="B8299526D754DE5193FD436A525A433E" colspan="1" pageId="0" pageNumber="23" rowspan="1">from base to apex</td>
<td id="249BF740CDE85B74F1F62475BCF275A7" colspan="1" pageId="0" pageNumber="23" rowspan="1">absent</td>
<td id="143A3B641E3D0ADE6531ED6A16761630" colspan="1" pageId="0" pageNumber="23" rowspan="1">from base to apex</td>
</tr>
<tr id="8FFB41FF31615E5A6967BDD79B2C28DA" pageId="0" pageNumber="23">
<td id="4BC290939554E06F6304DBB550F9641E" colspan="1" pageId="0" pageNumber="23" rowspan="1">Bract: shape</td>
<td id="E0610797B3B3134C085C6EC49FA4CE43" colspan="1" pageId="0" pageNumber="23" rowspan="1">broadly ovate, bowl-shaped</td>
<td id="7B06AE0D6C7F4C39155BB596E14B8F40" colspan="1" pageId="0" pageNumber="23" rowspan="1">elliptic-ovate or obovate, curved or flat</td>
<td id="62A9762B3E3D4F6EF04133559781CFAC" colspan="1" pageId="0" pageNumber="23" rowspan="1">obovate, curved or flat</td>
<td id="82362FABBD18437258567A764DBC4499" colspan="1" pageId="0" pageNumber="23" rowspan="1">obovate, flat</td>
</tr>
<tr id="7C74159CFAC48E85B1B7C5D2D61A7748" pageId="0" pageNumber="23">
<td id="ACED86D5294053A8039B86E57992EF1F" colspan="1" pageId="0" pageNumber="23" rowspan="1">size (mm)</td>
<td id="DF5A815D32C6903048B56D27E9232351" colspan="1" pageId="0" pageNumber="23" rowspan="1">
5-8 
<normalizedToken id="43E04A37296E0D6AC84D953A3CAFA8D1" originalValue="×">x</normalizedToken>
7.5-8
</td>
<td id="75516A6239774E2043902FBB39988114" colspan="1" pageId="0" pageNumber="23" rowspan="1">
2-3 
<normalizedToken id="775F99C514C22D4BBECF495797A38A59" originalValue="×">x</normalizedToken>
1.5-2
</td>
<td id="283C85A0F199FA96A30E8FEFF7E1E261" colspan="1" pageId="0" pageNumber="23" rowspan="1">
3-4 
<normalizedToken id="0E7E2AB261C5E803B5A59391ABBD6611" originalValue="×">x</normalizedToken>
1-3
</td>
<td id="93D8404B917AB318E56BA04770358895" colspan="1" pageId="0" pageNumber="23" rowspan="1">
4-6 
<normalizedToken id="2EC0A34E234D3561D1DDBE5D8BFDBDFE" originalValue="×">x</normalizedToken>
2-4
</td>
</tr>
<tr id="3BBF1CD6A245478A1259590B96620D74" pageId="0" pageNumber="23">
<td id="CCB99E375563C20B1731E8949A09C45B" colspan="1" pageId="0" pageNumber="23" rowspan="1">Hair density:</td>
<td id="F0E60A7E222C81224A3DFAD315B7B4AA" colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
<td id="2BF03DD5580F53DC5C9568C6F404ADB3" colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
<td id="DAD3104E016B92C6CC235DDD9A36F3B3" colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
<td id="A0F8020CEBC3E3FDB9B90685A40DACA6" colspan="1" isEmpty="true" pageId="0" pageNumber="23" rowspan="1">-</td>
</tr>
<tr id="42AAFAAEEA0733A8E14E485180BB82F6" pageId="0" pageNumber="23">
<td id="2B1DEA056E1C6D8E54D8D9818D24EB5A" colspan="1" pageId="0" pageNumber="23" rowspan="1">petioles</td>
<td id="A29A018766DDAFBE844F4AD8DBF11A68" colspan="1" pageId="0" pageNumber="23" rowspan="1">hairy</td>
<td id="07790EFFE83E01600FCE3C06B80EFDA4" colspan="1" pageId="0" pageNumber="23" rowspan="1">sparsely hairy, more hairy near leaf base</td>
<td id="33093A6683511427DAE50018129E7B1A" colspan="1" pageId="0" pageNumber="23" rowspan="1">sparsely hairy</td>
<td id="918E09FB5C493A01DA896CA77982D1CB" colspan="1" pageId="0" pageNumber="23" rowspan="1">densely hairy</td>
</tr>
<tr id="5AE418EFFCC98404213E6BCAE7F33477" pageId="0" pageNumber="23">
<td id="C78C5E429912F2A9BF5C2DFABED99F88" colspan="1" pageId="0" pageNumber="23" rowspan="1">stipules</td>
<td id="14133AAFFEE3BACD279A866576DB7A39" colspan="1" pageId="0" pageNumber="23" rowspan="1">densely hairy</td>
<td id="A334C5B7AD1DE627EA34116D2A46CA2E" colspan="1" pageId="0" pageNumber="23" rowspan="1">sparsely hairy</td>
<td id="9511FB33A378010DC3584252E959A013" colspan="1" pageId="0" pageNumber="23" rowspan="1">hairy</td>
<td id="B8667BFD51E21AC02BB75A2AE78DEB10" colspan="1" pageId="0" pageNumber="23" rowspan="1">densely hairy</td>
</tr>
<tr id="D0A68D83AF5A18A0D45F97C4F4E0100C" pageId="0" pageNumber="23">
<td id="CA4AE578E84E3FFBB93FA50F62447919" colspan="1" pageId="0" pageNumber="23" rowspan="1">peduncles</td>
<td id="9CCD97DF44DFE559BFF2641764B3D08D" colspan="1" pageId="0" pageNumber="23" rowspan="1">hairy</td>
<td id="48262F1DDF8D636D2ABFC2BF468FC290" colspan="1" pageId="0" pageNumber="23" rowspan="1">sparsely covered, hairs inconspicuous</td>
<td id="91B18EE38528D72581E229C83B48EF2B" colspan="1" pageId="0" pageNumber="23" rowspan="1">without hairs</td>
<td id="E2D3C4149F15A16DF78B70C2756D2E1F" colspan="1" pageId="0" pageNumber="23" rowspan="1">sparsely hairy</td>
</tr>
<tr id="F1D94CF259325A9EB52BFE0333B26519" pageId="0" pageNumber="23">
<td id="ABD5B6364CF9B6C2F300DEBCD57CF7FD" colspan="1" pageId="0" pageNumber="23" rowspan="1">Ovary colour</td>
<td id="2C74A2F6DBA7D50F34ED30C2AC084A5A" colspan="1" pageId="0" pageNumber="23" rowspan="1">pink or pale pink</td>
<td id="6F1D7655097E98488A162CB42946E058" colspan="1" pageId="0" pageNumber="23" rowspan="1">light yellowish-green</td>
<td id="CAC9DF7C1C76F0F336D07C7F41DC6C9C" colspan="1" pageId="0" pageNumber="23" rowspan="1">light yellowish-green</td>
<td id="3868795D78EE7582C2D34F7754187FF3" colspan="1" pageId="0" pageNumber="23" rowspan="1">reddish or dark pink</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="32F0C37BDFE0927D64726217CD63F1C1" pageId="0" pageNumber="23" type="etymology">
<paragraph id="484E1723F138D4875000E9BF1B55F727" pageId="0" pageNumber="23">Etymology.</paragraph>
<paragraph id="C0E89E91159EB8E3A574B4A4EF3BF04B" pageId="0" pageNumber="23">
From a Sanskrit word 
<emphasis id="64C2B0FBFCDD6A8CC085DB148A44F1C5" italics="true" pageId="0" pageNumber="23">raja</emphasis>
= king, presumably referring to it as the most beautiful begonia species.
</paragraph>
</subSubSection>
<subSubSection id="8D3786BAD669A39CEE495C01DEABF4B0" pageId="0" pageNumber="23" type="conservation status">
<paragraph id="2B86E64E16AF37455C88874CBE48779F" pageId="0" pageNumber="23">Conservation status.</paragraph>
<paragraph id="F579B94E4249AEDA561423A4E12F8173" pageId="0" pageNumber="23">
Extinct in the Wild (EW). However, it survives in cultivation since its first introduction into cultivation in 1894 (
<bibRefCitation id="1868359F80E2ABCBB14C9EBB4AF52FB1" author="Anonymous" pageId="0" pageNumber="23" refId="B2" refString="Anonymous, 1894b. New Plants and Flowers. Gardeners' Magazine, 485." title="New Plants and Flowers. Gardeners' Magazine, 485." year="1894 b">Anonymous 1894b</bibRefCitation>
, 
<bibRefCitation id="95859D851A2AED49A006EA34A13C90C1" author="Anonymous" pageId="0" pageNumber="23" refId="B3" refString="Anonymous, 1895. New Garden Plants of the Year 1894. Bulletin of Miscellaneous Information 2 (Appendix): 34." title="New Garden Plants of the Year 1894. Bulletin of Miscellaneous Information 2 (Appendix): 34." year="1895">1895</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E6E4F51D780EA5BD85D44C6BE9DC3065" pageId="0" pageNumber="23" type="ecology">
<paragraph id="264D3FC3740A637736A843C1629DE9DF" pageId="0" pageNumber="23">Ecology.</paragraph>
<paragraph id="D2E12A73DC91C1915AC2014BFF88D4D0" pageId="0" pageNumber="23">
Its natural habitat is not known but judging from its growth requirements in cultivation, it probably grew in cool and shaded conditions. 
<emphasis id="63A31799ED28BA9411A22DB4423038FC" bold="true" pageId="0" pageNumber="23">Other specimens examined. Peninsular Malaysia.</emphasis>
Herbarium specimens made from cultivated plants in Royal Botanic Gardens, Kew Anonymous s.n. 1903 (K!); Sander F. et al., s.n. 1913 (K!); Anonymous s.n. August 1913 (K!).
</paragraph>
</subSubSection>
</treatment>
</document>