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<mods:title id="7E798F3949FA2FA543B8965C7140671E">Revision of the genus Eadmuna Schaus, 1928 (Lepidoptera, Mimallonidae) with a description of a new species from French Guiana</mods:title>
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<treatment id="392B2FA1EC19DE5E5850D39A70C52E0D" ID-GBIF-Taxon="152059262" LSID="urn:lsid:plazi:treatment:392B2FA1EC19DE5E5850D39A70C52E0D" httpUri="http://treatment.plazi.org/id/392B2FA1EC19DE5E5850D39A70C52E0D" lastPageId="4" lastPageNumber="55" pageId="2" pageNumber="53">
<subSubSection id="5846356D7C3FDB7A27E60811E5B126F6" pageId="2" pageNumber="53" type="multiple">
<paragraph id="3D8A2D0F1BC87F7B8642DAD58580E919" pageId="2" pageNumber="53">Taxon classification Animalia Lepidoptera Mimallonidae</paragraph>
</subSubSection>
<subSubSection id="0F16926ABEF00584AF1FD15A5B905F4B" pageId="2" pageNumber="53" type="nomenclature">
<paragraph id="48836E01DA9E6D622D977578A5840204" pageId="2" pageNumber="53">
<taxonomicName id="9A9B2868A61A2A4A8A06B413B68E9A60" ID-CoL="927NN" authority="Schaus, 1928" authorityName="Schaus" authorityYear="1928" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">Eadmuna Schaus, 1928</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="CFB4A1F032169D8F4163154A87C2C907" pageId="2" pageNumber="53" type="type species">
<paragraph id="5EBE585EB0C0665763EB14A9D0D80E11" pageId="2" pageNumber="53">Type species.</paragraph>
<paragraph id="02D03CDF073E8E07DDC24B8375A98745" pageId="2" pageNumber="53">
<taxonomicName id="CEE53480C00C2BDDF703CA9EA26CD0ED" class="Insecta" family="Mimallonidae" genus="Cicinnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cicinnus esperans" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="esperans">Cicinnus esperans</taxonomicName>
Schaus, 1905
</paragraph>
</subSubSection>
<subSubSection id="17507C4E2BDB2206F321A61B41F2C91D" pageId="2" pageNumber="53" type="diagnosis">
<paragraph id="86EB40F88B32C82F94E180B447835731" pageId="2" pageNumber="53">Diagnosis.</paragraph>
<paragraph id="DC375DDBF2CB2C1AC7089B96AC45789F" pageId="2" pageNumber="53">
<taxonomicName id="74D53D3DAE07D7F523800B82E90BB008" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
can be recognized by broad wings and silvery-gray ground color accented by varying degrees of brown. The forewing bears a discal cell as a hyaline or sub-hyaline patch bisected by the M2 vein creating two separate windows. The hindwings lack any such hyaline markings. Although this marking is not unique within 
<taxonomicName id="F5B62C55C00B6CB3B7C65D06287CE81D" family="Mimallonidae" lsidName="" pageId="2" pageNumber="53" rank="family">Mimallonidae</taxonomicName>
, this character combined with the following two additional characters: the absence of any straight, continuous, vertical or diagonal postmedial lines and the presence of smooth wing margins; are diagnostic for the genus. Male genitalia are simple with a pointed, teardrop-shaped uncus, broad, ovoid tegumen with a pair of prominent, subtriangular, ridged lobes ventrally.
</paragraph>
</subSubSection>
<subSubSection id="0AC06D424CE203F74920EADDC736B1F8" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53" type="description">
<paragraph id="4B86EEFB77CDDE756CD36A43C9A294FE" pageId="2" pageNumber="53">Description.</paragraph>
<paragraph id="123EF82FB2CA07CB240A428CC655C8C4" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53">
Male. 
<pageBreakToken id="27A604579BE1F0EE4A8B156146C89B46" pageId="3" pageNumber="54" start="start">Head</pageBreakToken>
: Very small, scales on frons swept forward, eyes large comprising roughly two-thirds of head area, bordered posteriorly by darkbrown scales, border of darker scales continues down head reaching beneath labial palpi, labial palpi very small, segments smaller distally, hardly extending beyond frons, basal two segments tufted ventrally, dorsally covered in darkbrown scales greatly contrasting with overall straw coloration of head. Antenna bipectinate, scape and pedicel tufted. Ocelli and putative chaetosemata present. Thorax: Densely covered in long, hair-like scales interspersed with widened, darker, petiolate scales giving a speckled appearance. Legs: Vestiture thick, scales long, especially on femur and tibia, coloration as for thorax, petiolate scales present. Tibial spurs about one fifth length of tibia, thick, triangular in cross section, ridged, ridges finely serrate along ventral length. Forewing dorsum: Forewing length: 16-20 mm, n=40. Triangular, convex outer margins becoming concave near apex in some species, apex accentuated. Silvery gray-brown ground color with extensive speckling due to dark, petiolate scales in similar manner to that of thorax. Discal spot prominent, hyaline or partially covered in translucent scales, with M2 vein covered in dark scales separating hyaline patch into two distinct regions. Postmedial line usually present, though often faint, bulged in costal half, brown, dentate. Overall, scales become smaller and finer distally from wing base. Forewing venter: As for dorsum but usually lighter, postmedial lines generally more pronounced. Hindwing dorsum: Rounded, somewhat accentuated anal angle, essentially bearing same coloration and scale pattern as forewings though postmedial line usually fainter, if present. No hyaline patches present. Spatulate scales denser on inner margin. Hindwing venter: As for dorsum but usually lighter, postmedial lines generally more pronounced, frenulum with single bristle. Wing venation: As for 
<taxonomicName id="8C5DF9442A07FD30BE216593315BB407" class="Insecta" family="Mimallonidae" genus="Cicinnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cicinnus melsheimeri" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="melsheimeri">Cicinnus melsheimeri</taxonomicName>
(Harris, 1841) but R4 + R5 much shorter stalked. Abdomen: Somewhat compressed laterally, short, depth equal to that of thorax, rather triangular due to sudden truncation to slightly upturned tip, coloration a continuation of thoracic color, matching essentially dorsal wing coloration. Genitalia: Simple, uncus abruptly narrowed at base, extended apically. Tegumen broad, ovoid, with prominent, subtriangular, ridged lobes. Anal tube barely discernable, lightly sclerotized, with apex roughly halfway to distal tip of uncus. Valves simple, lightly sclerotized, basal half wider than distal half, sacculus half to one third width of valve at base, extending to half or two-thirds valve length. Juxta ventrally with quadrate lip and with two triangular arm-like spurs, one on either side of phallus. Juxtal spurs reach roughly midway along length of phallus. A small relatively quadrate sclerotized plate present dorsally to juxta/phallus. Vinculum broadly ovoid though flattened on dorsal and ventral margins, somewhat quadrate. Phallus simple, cylindrical, vesica sac-like or elongated with scobinate patch or with multidentate cornutus. Female. Similar to male except for: Head: Eyes greater than two thirds area of head, labial palpi smaller, region of brown scales bordering posterior of eyes thicker, extending to prothorax ventrally. Legs: Small scales nearly completely cover tibial spurs. Forewing dorsum: Forewing length range: 22-24 mm, n=3. Compared to male, forewing much broader overall, postmedial region lighter, more silvery gray than medial area, hyaline discal spot large, prominent. Postmedial line present, more pronounced than for male, brown, dentate, narrowly interrupted by veins, dark wedge where postmedial line meets costa. Antemedial lines present, bilobed, B-shaped. Forewing venter: As for dorsum, but lighter, postmedial line more contrasting. Hindwing dorsum: Broader, hardly accentuated anal angle, essentially bearing same coloration as forewings. Unlike males, entire hindwing, save for postmedial line, concolorous silvery gray, without a brown edge and without darker medial area present in forewings. Dentate postmedial line dark and well pronounced, narrowly interrupted by veins, slightly darker than that of forewing. Hindwing venter: As for dorsum, but lighter, postmedial line more contrasting, frenulum rudimentary with numerous bristles hidden by hindwing scales. Wing venation: As for male but Rs appears to originate closer to middle of cell. Abdomen: Much broader than that of male. Coloration a continuation of thoracic color, though darkening somewhat distally. Two very elongated sclerotized plates present on venter of eighth segment. Genitalia: Papillae anales elongated or stocky, covered in fine setae, apophyses posteriores shorter or same length as apophyses anteriores. Ductus bursae short, ostium opening immediately into corpus bursae. Corpus bursae, round, with or without sclerotized structures reinforcing membrane, elongated appendix bursae.
</paragraph>
</subSubSection>
<subSubSection id="0C7A1264CB5203993CDF6EA0319A0144" pageId="3" pageNumber="54" type="remarks">
<paragraph id="DA5366B107215155E701934D382CFD36" pageId="3" pageNumber="54">Remarks.</paragraph>
<paragraph id="EABAE0FE995F5DDDC61ECD238650E283" pageId="3" pageNumber="54">
Despite 
<bibRefCitation id="B90307093856A92CD8E547EAD4FACF87" author="Schaus, W" editor="Seitz, A" journalOrPublisher="A. Kernen, Stuttgart" pageId="12" pageNumber="63" pagination="635 - 672" title="Familie Mimallonidae." volumeTitle="Die Gross-Schmetterlinge der Erde. 6. Die amerikanischen Spinner und Schwaermer" year="1928">
<normalizedToken id="44C0F6A557A1B11931F0C73554143994" originalValue="Schaus’">Schaus'</normalizedToken>
(1928)
</bibRefCitation>
comment that 
<taxonomicName id="4368F5F1F6BAF7D1102FCFA55DFA7AC3" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
genitalia are allied to those of 
<taxonomicName id="DFFA33C5430B2FFED93557765773F257" class="Insecta" family="Mimallonidae" genus="Psychocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psychocampa" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Psychocampa</taxonomicName>
(unspecified species), this has not been found when comparing 
<taxonomicName id="9280780FF32748DF6F08B3D3F50487D0" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
genitalia to those of some representative 
<taxonomicName id="CEEA5ECF34F7C7E267BBB68313911431" class="Insecta" family="Mimallonidae" genus="Psychocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psychocampa" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Psychocampa</taxonomicName>
. No 
<taxonomicName id="B157BAC8D065D95B67281D6E3EBB342F" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
genitalia resemble any of the 
<taxonomicName id="F15B42DAC5453981717CD13762159666" class="Insecta" family="Mimallonidae" genus="Psychocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psychocampa" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Psychocampa</taxonomicName>
genitalia figured in 
<bibRefCitation id="424BF45E263C22694ABD78D95743EE98" author="Herbin, D" journalOrPublisher="European Entomologist" pageId="12" pageNumber="63" pagination="1 - 31" title="Descriptions of a new genus and ten new species of Mimallonidae (Lepidoptera: Mimallonoidea)." volume="4" year="2012">Herbin (2012)</bibRefCitation>
, including 
<taxonomicName id="F5958674A4AD88F7562F0DA55FD08D67" class="Insecta" family="Mimallonidae" genus="Psychocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psychocampa kohlii" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="kohlii">Psychocampa kohlii</taxonomicName>
Herbin which greatly resemble the genitalia of the type species 
<taxonomicName id="0BC06A8FA1ACA2F9D1A1BFDC24249FF9" class="Insecta" family="Mimallonidae" genus="Psychocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psychocampa concolor" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="concolor">Psychocampa concolor</taxonomicName>
Grote &amp; Robinson, 1866 (
<bibRefCitation id="23639C2B04CA6197E354E5F1211E1579" author="Herbin, D" journalOrPublisher="European Entomologist" pageId="12" pageNumber="63" pagination="1 - 31" title="Descriptions of a new genus and ten new species of Mimallonidae (Lepidoptera: Mimallonoidea)." volume="4" year="2012">Herbin 2012</bibRefCitation>
).
</paragraph>
<paragraph id="7163ADCFD2338F0C7AA1DCDC898702B4" pageId="3" pageNumber="54">
<taxonomicName id="2F9E6CB5F21581EA802C9DF3A9D73CAC" genus="Aceclosteria" lsidName="Aceclosteria villaricensis" pageId="3" pageNumber="54" rank="species" species="villaricensis">Aceclosteria villaricensis</taxonomicName>
(Schaus, 1933) was originally described in 
<taxonomicName id="2D66148D3708BEEC2CB74E23DF7C39E3" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
. Currently the genus 
<taxonomicName id="DFEC10E46E5248DFE834D2A4F36CF027" genus="Aceclosteria" lsidName="Aceclosteria" pageId="3" pageNumber="54" rank="genus">Aceclosteria</taxonomicName>
Vuillot, 1893 contains one species, 
<taxonomicName id="49C476718C60DE74184C091BA734A891" genus="Aceclosteria" lsidName="Aceclosteria mus" pageId="3" pageNumber="54" rank="species" species="mus">Aceclosteria mus</taxonomicName>
Vuillot, 1893. Previously Schaus described a female 
<taxonomicName id="62779B451B099BE55F904CD3F2474D58" genus="Aceclosteria" lsidName="Aceclosteria" pageId="3" pageNumber="54" rank="genus">Aceclosteria</taxonomicName>
specimen as 
<taxonomicName id="14E0D814F4087F797B9FC96AD81DA882" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna villaricensis" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="villaricensis">Eadmuna villaricensis</taxonomicName>
due to it being allied with &quot; 
<taxonomicName id="F1E033EBB9CC9920D44A98E75DE2C3EE" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna esperanza" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="esperanza">Eadmuna esperanza</taxonomicName>
,&quot; [sic] (Schaus, 1933) though the two species are quite dissimilar. For instance, 
<taxonomicName id="CDA1BC3219BE7F422AB72A651BEC3E96" genus="Aceclosteria" lsidName="Aceclosteria mus" pageId="3" pageNumber="54" rank="species" species="mus">Aceclosteria mus</taxonomicName>
has a continuous, non-dentate postmedial line. Additionally, in a single male genitalia dissection of 
<taxonomicName id="69FCE0245A63FB93D3421D8437D01EFD" genus="Aceclosteria" lsidName="Aceclosteria mus" pageId="3" pageNumber="54" rank="species" species="mus">Aceclosteria mus</taxonomicName>
from Rio Grande do Sul, Brazil (CUIC genitalia dissection 10-8-14:2), the genitalia were found to be highly complex structurally and asymmetrical, completely unlike 
<taxonomicName id="FBF21C9A8C37FBC87EDC166E241D8FF8" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
. 
<bibRefCitation id="7DAE285BD37C4BA6AAC49F4680F64FEE" author="Becker, VO" editor="Heppner, JB" journalOrPublisher="Association for Tropical Lepidoptera &amp; Scientific Publishers, Gainesville, Florida" pageId="12" pageNumber="63" pagination="17 - 19" title="Atlas of Neotropical Lepidoptera, Checklist. Part 4 B. Drepanoidea, Bombycoidea, Sphingoidea." year="1996">Becker (1996)</bibRefCitation>
synonymized 
<taxonomicName id="D82892453BF59FB821167B4ED7188BE0" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna villaricensis" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="villaricensis">Eadmuna villaricensis</taxonomicName>
with 
<taxonomicName id="B0FA1991BBB76C6A863B31F61EC7D749" genus="Aceclosteria" lsidName="Aceclosteria mus" pageId="3" pageNumber="54" rank="species" species="mus">Aceclosteria mus</taxonomicName>
. An external examination of the holotype of 
<taxonomicName id="CD1B1349D9EAD317E0897CBBC9545A1E" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna villaricensis" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="villaricensis">Eadmuna villaricensis</taxonomicName>
supports 
<normalizedToken id="2022B8C7209A7B826A0E87F3C794B609" originalValue="Becker’s">Becker's</normalizedToken>
synonymy.
</paragraph>
<paragraph id="DDD074E21D147B27D36F558298C0263B" pageId="3" pageNumber="54">
One or two other undescribed species from Costa Rica are currently considered to belong to 
<taxonomicName id="99942B8A68C7F7D9D689CB312C08A433" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
by Daniel Herbin (pers. comm.). These golden-colored species, superficially somewhat similar to 
<taxonomicName id="5BBACF86DB70FE1BC3EEE60538EDBA76" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna guianensis" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="guianensis">Eadmuna guianensis</taxonomicName>
, new species and 
<taxonomicName id="913650CAC559046D7502FC4DAEFA511C" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna esperans" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="esperans">Eadmuna esperans</taxonomicName>
, have broad wings, dentate postmedial lines, and bisected forewing hyaline areas. However, the genitalia are very distinct (MGCL dissection number 9-24-14:1). In one of these undescribed species, the uncus is not truncate and is rather triangular and flattened apically, the juxta has two extremely long, curved tusk-like projections, pointed outwards above the phallus. Finally, somewhat triangular tegumen lobes are present, as in 
<taxonomicName id="B952FC1FB994F26737259A59F8F648E8" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
, but are significantly elongated and without numerous ridges as in 
<taxonomicName id="F42E1862FA40E35A21B736D37F9E3BE6" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
. Thus, these species from Costa Rica cannot be considered 
<taxonomicName id="23ED908C2C5CB03EB115A2C7DB244C1C" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
.
</paragraph>
<paragraph id="42DC29721A6876208E99CE02E61173C8" pageId="3" pageNumber="54">
The geographic distribution of the species 
<taxonomicName id="813401FFB5B707BFD4EDB856B7345089" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna esperans" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="esperans">Eadmuna esperans</taxonomicName>
and 
<taxonomicName id="5F582FFBA908EAC460D7DAF64C555383" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna paloa" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="paloa">Eadmuna paloa</taxonomicName>
, and possibly 
<taxonomicName id="F91A5B27BE32CE5F70B099686BE00C92" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna pulverula" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="pulverula">Eadmuna pulverula</taxonomicName>
, very clearly follows the Atlantic coastal rainforest of Brazil (see Figure 18) (IBGE). This biome is of particular conservation interest due to a massive loss of habitat, such that it has been estimated that only approximately 11% of the Brazilian Atlantic forest remains (
<bibRefCitation id="3FE55D82EBC341BD5D698C83D4015291" author="Ribeiro, MC" journalOrPublisher="Biological Conservation" pageId="12" pageNumber="63" pagination="1141 - 1153" title="The Brazilian Atlantic Forest: How much is left, and how is the remaining forest distributed? Implications for conservation." url="10.1016/j.biocon.2009.02.021" volume="142" year="2009">Ribeiro et al. 2009</bibRefCitation>
). The association of these two or three species with this biome, along with the almost complete lack of recent material of these species in any of the visited collections, presents further justification for the conservation of this area of high species richness (
<bibRefCitation id="96667CB760E529C2BB3780941D5C5674" author="Ribeiro, MC" journalOrPublisher="Biological Conservation" pageId="12" pageNumber="63" pagination="1141 - 1153" title="The Brazilian Atlantic Forest: How much is left, and how is the remaining forest distributed? Implications for conservation." url="10.1016/j.biocon.2009.02.021" volume="142" year="2009">Ribeiro et al. 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F661F970F308F9CE0392A23B81E1E906" pageId="4" pageNumber="55" type="key to species of eadmuna">
<paragraph id="DC0E97405DCA91AF245E14D0307D1DDF" pageId="4" pageNumber="55">
<pageBreakToken id="964987DF617CCAC8395B1504B0F841DA" pageId="4" pageNumber="55" start="start">Key</pageBreakToken>
to species of 
<taxonomicName id="7B8F2B6525CF099122DD126DCA07D07C" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
*
</paragraph>
<paragraph id="9F4D4EDB310874F13F7B223135C83593" pageId="4" pageNumber="55">
<table id="55C4A0D3B201885C4192BAA42758E120" pageId="4" pageNumber="55">
<tr id="26E543714B3486C53B1FA67F2D463B79" pageId="4" pageNumber="55">
<td id="D3871711765F3F53D85DEB3E298301CF" colspan="1" pageId="4" pageNumber="55" rowspan="1">1-6</td>
<td id="022092703CB0B2A56E9DEDA8BB0E1B76" colspan="1" pageId="4" pageNumber="55" rowspan="1">2</td>
</tr>
<tr id="48A9B748F6DA2EDC2F76BD573A74233A" pageId="4" pageNumber="55">
<td id="1965723F2B3B852125C4016FFC1EAA3A" colspan="1" pageId="4" pageNumber="55" rowspan="1">78</td>
<td id="2FE7A676A11552E5FD6E7E6F6FE0161B" colspan="1" pageId="4" pageNumber="55" rowspan="1">4</td>
</tr>
<tr id="90CF6CE0FDC1683B8057F658950DAAC7" pageId="4" pageNumber="55">
<td id="313E247850D3B8487D8EA02BC7EECCCC" colspan="1" pageId="4" pageNumber="55" rowspan="1">5614</td>
<td id="9AF33043343680C50FAAB3B42DBDC93E" colspan="1" pageId="4" pageNumber="55" rowspan="1">
<taxonomicName id="222C8029653795937759C6AF4407B03F" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna paloa" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="paloa">Eadmuna paloa</taxonomicName>
(part)
</td>
</tr>
<tr id="1ED465E27E7F6A8549FAB3AF5672FC46" pageId="4" pageNumber="55">
<td id="57EEBF1494E1B0683A477E9AFD519F8F" colspan="1" pageId="4" pageNumber="55" rowspan="1">1-412</td>
<td id="3AAD3EDAF3B38B567D5A3561383FC5A8" colspan="1" pageId="4" pageNumber="55" rowspan="1">3</td>
</tr>
<tr id="7A4A009CE60A11CFA7E0159791A073D2" pageId="4" pageNumber="55">
<td id="61119DBFE86A37C38851AFE2DCB13808" colspan="1" pageId="4" pageNumber="55" rowspan="1">1213</td>
<td id="87DE21263D768ECB172263BD948C6C1D" colspan="1" pageId="4" pageNumber="55" rowspan="1">
<taxonomicName id="9C440A77488DBF70182991904BA7C074" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna esperans" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="esperans">Eadmuna esperans</taxonomicName>
</td>
</tr>
<tr id="7B01529B04E81DC2ACD0A9D66937BE07" pageId="4" pageNumber="55">
<td id="DF7A5D8B2E3BEA803932559524421D34" colspan="1" pageId="4" pageNumber="55" rowspan="1">3412</td>
<td id="3F9231CA24680A38910125484CBF0FE0" colspan="1" pageId="4" pageNumber="55" rowspan="1">
<taxonomicName id="14368D5301E14486D32A0323B5C23CE0" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna guianensis" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="guianensis">Eadmuna guianensis</taxonomicName>
sp. n.
</td>
</tr>
<tr id="6B90B5F04EB428E448713E420CF8E768" pageId="4" pageNumber="55">
<td id="AC599AAB833FAED19E4DCD1CCF69D56C" colspan="1" pageId="4" pageNumber="55" rowspan="1">71516</td>
<td id="222FFF86725C450EEBC2EA59A9E35DF1" colspan="1" pageId="4" pageNumber="55" rowspan="1">
<taxonomicName id="75C5414F6570B55F796984DC0C9C5A32" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna paloa" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="paloa">Eadmuna paloa</taxonomicName>
(part)
</td>
</tr>
<tr id="973BA01E2638F8C0D413287645A4BD81" pageId="4" pageNumber="55">
<td id="0E7998DB4FCB312C3496D7FD60AB4258" colspan="1" pageId="4" pageNumber="55" rowspan="1">817</td>
<td id="003CF26D231D3B4899270AF0F8196017" colspan="1" pageId="4" pageNumber="55" rowspan="1">
<taxonomicName id="9D48CFD4E2441AEED278F2A623B46F3F" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna pulverula" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="pulverula">Eadmuna pulverula</taxonomicName>
</td>
</tr>
</table>
</paragraph>
<paragraph id="5901F9520DC606C4C586057A01EC8C6F" pageId="4" pageNumber="55">
* Note: the male of 
<taxonomicName id="5BBC1A5C9B609E974A7D49AA6FC0997C" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna pulverula" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="pulverula">Eadmuna pulverula</taxonomicName>
and the females of 
<taxonomicName id="5B500476CD4BBD45C343A7FD78520090" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna esperans" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="esperans">Eadmuna esperans</taxonomicName>
and 
<taxonomicName id="5553A1642EC8CEF565E2C8AFB1784A48" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna guianensis" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="guianensis">Eadmuna guianensis</taxonomicName>
are unknown
</paragraph>
<caption id="2DC32468B7CF94AB7CB01BC080F153FF" pageId="4" pageNumber="55">
<paragraph id="18DA051D38C933A7CFD928DCCFF0C9FB" pageId="4" pageNumber="55">
Figures 1-8. 
<taxonomicName id="A31B7F5CA09E643CC3A01859CD034266" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
adults. 1 
<taxonomicName id="04CFBC0681BF471EBDBCA7E117BA8F7F" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna esperans" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="esperans">Eadmuna esperans</taxonomicName>
holotype male, 
<normalizedToken id="8C943EF4128CE71E6F090022E5E46B28" originalValue="Espírito">Espirito</normalizedToken>
Santo, Brazil (image inverted laterally) [USNM] 2 
<taxonomicName id="32541A573B4875BFE14F61AED4C829E3" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna esperans" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="esperans">Eadmuna esperans</taxonomicName>
male, 
<normalizedToken id="195FD51DD6301444F5D187C2A6716D2B" originalValue="Jaraguá">Jaragua</normalizedToken>
[do Sul], Santa Catarina, Brazil [CUIC] 3 
<taxonomicName id="F961A104AF2925640D0E480496BE8580" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna guianensis" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="guianensis">Eadmuna guianensis</taxonomicName>
holotype male, Mana River, French Guiana [CMNH] 4 
<taxonomicName id="CEE1718B52880DD8C22921B7371916B2" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna guianensis" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="guianensis">Eadmuna guianensis</taxonomicName>
paratype male, Mana River, French Guiana [CMNH] 5 
<taxonomicName id="0EDCC7B144A65748BDB791357DD6D2DC" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna paloa" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="paloa">Eadmuna paloa</taxonomicName>
holotype male, 
<normalizedToken id="673555C59FFE2DD055C57CD0B27CD3C7" originalValue="São">Sao</normalizedToken>
Paulo, Brazil (image inverted laterally) [USNM] 6 
<taxonomicName id="E12F21FD7D641E9ADF026C341C69D9E8" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna paloa" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="paloa">Eadmuna paloa</taxonomicName>
male, Nova Bremen, Santa Catarina, Brazil (image inverted laterally) [CUIC] 7 
<taxonomicName id="5FE417E042A84EBE6DE4439E760C9F2E" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna paloa" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="paloa">Eadmuna paloa</taxonomicName>
female, Rio Vermelho, Santa Catarina, Brazil (image inverted laterally) [AMNH] 8 
<taxonomicName id="660B1C92BB8CFDF608383AA8EC5DA0DD" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna pulverula" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="pulverula">Eadmuna pulverula</taxonomicName>
holotype female, 
<normalizedToken id="F8C686F5CBEA68FC2A61EED500C0EBCA" originalValue="São">Sao</normalizedToken>
Paulo, Brazil [USNM]. Scale bar = 1 cm.
</paragraph>
</caption>
<caption id="AE3DADA1B865BE0ABD618C27A758FCCE" pageId="4" pageNumber="55">
<paragraph id="7F39DD76AEC015065CE0B20FB1091578" pageId="4" pageNumber="55">
Figures 9-14. 
<taxonomicName id="AF2E3911C5C80C7EF745721CDCBD3600" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
male genitalia, valves with phallus and juxta removed, and separated phallus and juxta. 9 
<taxonomicName id="952A2CE6EA67129283284DAA66F8229E" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna guianensis" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="guianensis">Eadmuna guianensis</taxonomicName>
, holotype, Mana River, French Guiana [St Laurent diss.: 9-14-14:3] 10 
<taxonomicName id="6B50095A1315D7B4D8857D35F9F83984" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna esperans" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="esperans">Eadmuna esperans</taxonomicName>
, Est. Biol. Boraceia, 850 m, near Salesopolis, 
<normalizedToken id="91746A5FEB87BDB6767C05384010FBBC" originalValue="São">Sao</normalizedToken>
Paulo, Brazil [St Laurent diss.: 9-14-14:6] 11 
<taxonomicName id="D0D226714AB11AD21DEC22EF0177C917" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna paloa" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="paloa">Eadmuna paloa</taxonomicName>
, Nova Bremen, Santa Catarina, Brazil [St Laurent diss.: 9-14-14:8] 12 
<taxonomicName id="50DB940C299AA8ECEEEBE6802C2E7307" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna guianensis" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="guianensis">Eadmuna guianensis</taxonomicName>
, holotype, Mana River, French Guiana [St Laurent diss.: 9-14-14:3] 13 
<taxonomicName id="36CDA8A408726D9ABAEE2E0B01D8905A" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna esperans" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="esperans">Eadmuna esperans</taxonomicName>
, Est. Biol. Boraceia, 850 m, near Salesopolis, 
<normalizedToken id="62E8CAD284B8AE6BB97147B208CAF491" originalValue="São">Sao</normalizedToken>
Paulo, Brazil [St Laurent diss.: 9-14-14:6] 14 
<taxonomicName id="990E664AA5B96CF38396B47127357279" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna paloa" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="paloa">Eadmuna paloa</taxonomicName>
, Nova Bremen, Santa Catarina, Brazil [St Laurent diss.: 9-14-14:8]. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="916B5965BE369E7067289BC59B6D2FAB" pageId="4" pageNumber="55">
<paragraph id="F425FD0BE1D08ADA413FCDB74F3F153B" pageId="4" pageNumber="55">
Figures 15-17. 
<taxonomicName id="16A280D5086942DD58EB4DA0FA231A7A" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
female genitalia. 15 
<taxonomicName id="53D05E50E18C92CD67636014562F53EE" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna paloa" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="paloa">Eadmuna paloa</taxonomicName>
, ventral, Rio Vermelho, Santa Catarina, Brazil [St Laurent diss.: 10-11-14:3] 16 
<taxonomicName id="61475284C64A424D765FBA5B944F5140" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna paloa" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="paloa">Eadmuna paloa</taxonomicName>
, lateral, Rio Vermelho, Santa Catarina, Brazil [St Laurent diss.: 11-1-14:8] 17 Damaged 
<taxonomicName id="41261EF6D06663B0241651EDCE86D455" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna pulverula" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="pulverula">Eadmuna pulverula</taxonomicName>
, holotype, ventral, 
<normalizedToken id="17F0B1DF892CCEE8D71154415863874E" originalValue="São">Sao</normalizedToken>
Paulo, Brazil [St Laurent diss.: 11-1-14:8]. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="6D86108E42DED8973CB4EF88772BC820" pageId="4" pageNumber="55">
<paragraph id="F03E770EEF910D568786E53868C43120" pageId="4" pageNumber="55">
Figure 18. Distribution of 
<taxonomicName id="D3C82ABD675E00F4B5F7DEF8FA423AB3" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">Eadmuna</taxonomicName>
. 
<taxonomicName id="6FB01EB75A593883EE0C3CCF617F578F" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna esperans" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="esperans">Eadmuna esperans</taxonomicName>
(red circles), 
<taxonomicName id="D4BB2263919D5023B68186DEEA9166AF" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna guianensis" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="guianensis">Eadmuna guianensis</taxonomicName>
(purple circles), 
<taxonomicName id="C807911B529AE1898F2723C489372400" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna paloa" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="paloa">Eadmuna paloa</taxonomicName>
(blue circles), 
<taxonomicName id="4FDE74F5E4D9DAB61AC83C13DCF7B7B3" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna pulverula" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="pulverula">Eadmuna pulverula</taxonomicName>
(green star). Notes: red/blue circle represents the locality where both 
<taxonomicName id="46774FA0634A93C90A8DDE54772D6D35" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna esperans" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="esperans">Eadmuna esperans</taxonomicName>
and 
<taxonomicName id="9DDBB09598305959A3EE38FF4BE19F2F" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna paloa" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="paloa">Eadmuna paloa</taxonomicName>
have been collected, 
<taxonomicName id="D852240B42E6A6D2F9B70FA472FDDDF5" class="Insecta" family="Mimallonidae" genus="Eadmuna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadmuna pulverula" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="pulverula">Eadmuna pulverula</taxonomicName>
is represented by a star placed near the center of the Brazilian state of 
<normalizedToken id="194060D8FA9DA99DEA98DFF10BAFA563" originalValue="São">Sao</normalizedToken>
Paulo because the type locality is 
<normalizedToken id="62A16F7BA5E42DCAF5DFE2B3B58578CA" originalValue="“São">&quot;Sao</normalizedToken>
Paulo,&quot; without further information regarding specific locality.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>