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<document ID-CLB-Dataset="207920" ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.e84187" ID-GBIF-Dataset="bac14654-3ac7-46b5-a981-8873c1b9e291" ID-Pensoft-Pub="2747-8106-96-179" ID-Pensoft-UUID="FE0861D71BB454999EE6637C0D9B2B0C" ID-ZooBank="35B619086E6548B09A177281C2253391" ModsDocID="2747-8106-96-179" checkinTime="1670952513976" checkinUser="pensoft" docAuthor="Gruendel, Joachim, Keupp, Helmut, Lang, Fritz &amp; Nuetzel, Alexander" docDate="2022" docId="36055E12A8B95738BB927475F5DAE368" docLanguage="en" docName="Zitteliana 96: 179-221" docOrigin="Zitteliana 96" docPubDate="2022-12-12" docSource="http://dx.doi.org/10.3897/zitteliana.96.e84187" docTitle="Ptygmatis ursicina" docType="treatment" docVersion="5" id="FE0861D71BB454999EE6637C0D9B2B0C" lastPageNumber="179" masterDocId="FE0861D71BB454999EE6637C0D9B2B0C" masterDocTitle="Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany)" masterLastPageNumber="221" masterPageNumber="179" pageNumber="179" updateTime="1732645988798" updateUser="ExternalLinkService">
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<mods:title id="A934EB4013CFD73BFF54375997FB726D">Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany)</mods:title>
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<mods:namePart id="C5941D1E24A86E9C707BE35703A4B94F">Gruendel, Joachim</mods:namePart>
<mods:affiliation id="FF5AC911D9FAA719861984663A3C5392">Institut fuer Geowissenschaften, Fachrichtung Palaeontologie, Freie Universitaet Berlin, Malteserstrasse 74 - 100, 12249 Berlin, Germany</mods:affiliation>
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<mods:namePart id="D4F3211137330AED9C2596B913EE8F9C">Keupp, Helmut</mods:namePart>
<mods:affiliation id="2DA59E7DEF90CA8D3A7256E9B086E72D">Institut fuer Geowissenschaften, Fachrichtung Palaeontologie, Freie Universitaet Berlin, Malteserstrasse 74 - 100, 12249 Berlin, Germany</mods:affiliation>
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<mods:namePart id="ECE4065529961747B33AA27F6DEF8433">Lang, Fritz</mods:namePart>
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<mods:namePart id="3EE514C908292E45069ABB458F466A33">Nuetzel, Alexander</mods:namePart>
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<mods:affiliation id="A6AB315256BC1D537A8223254EC15DCD">SNSB-Bayerische Staatssammlung fuer Palaeontologie und Geologie, Richard-Wagner-Str. 10, 80333 Muenchen, Germany &amp; Department of Earth and Environmental Sciences, Paleontology and Geobiology, Ludwig-Maximilians-Universitaet Muenchen, Richard-Wagner-Str. 10, 80333 Muenchen, Germany</mods:affiliation>
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<treatment id="36055E12A8B95738BB927475F5DAE368" ID-GBIF-Taxon="206232340" LSID="urn:lsid:plazi:treatment:36055E12A8B95738BB927475F5DAE368" httpUri="http://treatment.plazi.org/id/36055E12A8B95738BB927475F5DAE368" lastPageNumber="179" pageId="0" pageNumber="179" scope_class="Gastropoda" scope_family="Ptygmatididae" scope_subClass="Heterobranchia" scope_subFamily="Ptygmatidinae" scope_superFamily="Nerineoidea">
<subSubSection id="DD3CD22E47092F0288494DC2D3AEF7BE" pageId="0" pageNumber="179" type="nomenclature">
<paragraph id="BDF272D777F8E583AAE9A19F9892C810" pageId="0" pageNumber="179">
<taxonomicName id="04017E8D2A8599A69DE75659BAE3ECC4" ID-CoL="C88N9" LSID="36055E12-A8B9-5738-BB92-7475F5DAE368" authority="(Thurmann, 1861 in Thurmann and Etallon 1861 - 1864)" baseAuthorityName="Thurmann, 1861 in Thurmann and Etallon 1861 -" baseAuthorityYear="1864" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina">
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Ptygmatis? ursicina (Thurmann, 1861 in Thurmann and
<normalizedToken id="68E88EC035B9916B658D6F9A345B0271" originalValue="Étallon">Etallon</normalizedToken>
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1861-1864)
</taxonomicName>
</paragraph>
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<subSubSection id="F3B250622724C409DBD3C8F875E40093" pageId="0" pageNumber="179" type="description">
<paragraph id="AF7928A3B5240B4874B66A57327A6837" pageId="0" pageNumber="179">
<figureCitation id="6CD8AB303ECB8B3FF437AFC6D36B1B53" captionStart="Plate 12" captionStartId="F12" captionText="Plate 12. (1 - 3) Ptygmatis clio (d'Orbigny, 1852). (1 - 2) SNSB-BSPG 2021 XV 59, Saal (collection Lang); (1) section of fragment, height 21 mm; (2) detail, height 3.7 mm. (3) SNSB-BSPG 2021 XV 60, Saal (collection Lang), lateral view, height 108 mm. (4) Ptygmatis? polyspira (Quenstedt, 1881 - 1884), holotype, collection University Tuebingen (collection Quenstedt); Nattheim, lateral view, height 32 mm (original to Quenstedt 1881 - 1884, pl. 207, fig. 3. (11 - 12) Bactroptyxis teres (Muenster in Goldfuss, 1844), collection Sauerborn, Nattheim area. (5 - 7) Ptygmatis? ursicina (Thurmann in Thurmann &amp; Etallon, 1861). (5 - 6) SNSB-BSPG 2021 XV 30, Saal (collection Lang); (5) last whorl with aperture, height 4 mm; (6) lateral view, height 9.5 mm. (7) SNSB-BSPG 2021 XV 31, Saal (collection Lang), aperture and ornament of base, width 2.5 mm." figureDoi="10.3897/zitteliana.96.e84187.plate12" httpUri="https://binary.pensoft.net/fig/780390" pageId="0" pageNumber="179">Plate 12: figs 5-7</figureCitation>
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</paragraph>
</subSubSection>
<subSubSection id="0563CE9119AEDC04644B55595681B639" pageId="0" pageNumber="179" type="reference_group">
<paragraph id="FA4EB4769346712403FF8C4BA38EA976" pageId="0" pageNumber="179">
<taxonomicName id="F675B7C5B6051021F2E2DC6D0C40425D" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea visurgis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="visurgis">?1844 - Nerinea visurgis</taxonomicName>
<normalizedToken id="173FA23F75DFA56B0422173CB812B579" originalValue="Röm">Roem</normalizedToken>
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. - Goldfuss: 44, pl. 176, fig. 6.
</paragraph>
<paragraph id="D8D520CFD7E08DABAE9A5FBC69C1B2B1" pageId="0" pageNumber="179">
<taxonomicName id="D888C10F3733C381952B5E1FA6C6D4B3" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea visurgis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="visurgis">1852 - Nerinea visurgis</taxonomicName>
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Roemer, 1836 -
<normalizedToken id="28F65BB902837BE0FBF1E2ED118A2D74" originalValue="dOrbigny">d'Orbigny</normalizedToken>
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: 122, pl. 268, figs 5-7.
</paragraph>
<paragraph id="0474D41B81406180A19BD55E9B5F53DC" pageId="0" pageNumber="179">
<taxonomicName id="0808819BB492B02957C61B5928446D12" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina">*1861-1864 - Nerinea ursicina</taxonomicName>
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Th. - Thurmann and
<normalizedToken id="24D4717363C395EDCB3D5015EBAF5EF2" originalValue="Étallon">Etallon</normalizedToken>
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: 103, pl. 8, fig. 50.
</paragraph>
<paragraph id="42C587CCFB1C2DB9FBB48A7F87D9FF13" pageId="0" pageNumber="179">
<taxonomicName id="ADB18B5722199A4701088C914AB1F8F4" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea pseudospeciosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="pseudospeciosa">?1872 - Nerinea pseudospeciosa</taxonomicName>
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P. de Loriol, 1871 - Loriol, Royer and Tombeck: 89, pl. 6, fig. 7.
</paragraph>
<paragraph id="6680E3F47B7DB64CFA8D5B843D23518C" pageId="0" pageNumber="179">
<taxonomicName id="26456D4C03840E403DD9D3E792B919E6" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina">1889 - Nerinea ursicina</taxonomicName>
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Thurmann - Loriol and Koby: 37, pl. 6, figs 1-8.
</paragraph>
<paragraph id="CE50D97EE29DA52D8F2493A5D2F4D5C4" pageId="0" pageNumber="179">
<taxonomicName id="619BA468DBE686C05E6D789763A3A2B4" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea ursicinensis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicinensis">1898 - Nerinea ursicinensis</taxonomicName>
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Thurmann - Cossmann: 37, pl. 3, figs 11, 12.
</paragraph>
<paragraph id="395F3A5634EF6239038116FC124CEED9" pageId="0" pageNumber="179">
<taxonomicName id="12EBF90CA077BCDC9D6308E31F3EC51D" class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea ursicinensis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicinensis">1927 - Nerinea ursicinensis</taxonomicName>
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Thurmann - Maire: 142, pl. 7, figs 15, 16.
</paragraph>
<paragraph id="7F8804B3B50CB8B8FBC2D5CCE81BC86F" pageId="0" pageNumber="179">
<taxonomicName id="5A766C03788B0A50B992327308C20EBA" class="Gastropoda" family="Ptygmatididae" genus="Cossmannea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cossmannea (Eunerinea) ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina" subGenus="Eunerinea">1997 - Cossmannea (Eunerinea) ursicina</taxonomicName>
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(Thurmann, 1861) - Fischer and Weber: 40, pl. 9, fig. 2.
</paragraph>
</subSubSection>
<subSubSection id="8BE94F77A1E96A1D1A0DBFA0C31BFC02" pageId="0" pageNumber="179" type="materials_examined">
<paragraph id="739767AEFB135A38E61DDADD12C9CA6C" pageId="0" pageNumber="179">Material.</paragraph>
<paragraph id="F2520C4F58370447317A35D7F4BFE503" pageId="0" pageNumber="179">Nineteen juvenile specimens from Saal (collection Lang), of which two are illustrated (SNSB-BSPG 2021 XV 30, 31).</paragraph>
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</subSubSection>
<subSubSection id="E599E048C838FE98EAFEE7E2751482A9" pageId="0" pageNumber="179" type="description">
<paragraph id="A6D72BFBFF3D3D1AE998DECFFE00BB57" pageId="0" pageNumber="179">Description.</paragraph>
<paragraph id="7817CAF7E5EBB01B6B146E33EC167894" pageId="0" pageNumber="179">The largest specimen from Saal is 9 mm high. The shell is slender to very slender. The whorls are regularly increasing in width. The whorl face is concave, with a strong, sometimes ramp-like bulge. The bulge is formed at the suture by both whorls. The suture is situated somewhat above the middle of the bulge. The whorl face between the bulges is initially ornamented with one spiral cord, later whorls with 4-5 spiral cords: two of these spiral cords may be stronger than the others. The bulges and stronger spiral cords are possibly nodular (unclear due to preservation). The base is flat, with an almost rectangular transition to the whorl face. The transition has a strong, protruding bulge that is largely covered by the following whorls of the spire. The base is densely covered with weak spiral cords. The aperture has an approximately rectangular outline and distinct oblique siphonal canal. The aperture has a single parietal plait, two columellar plaits, and a palatal plait.</paragraph>
<caption id="67B1177005CFDA73F9FA9A7243E23FA8" doi="10.3897/zitteliana.96.e84187.plate12" httpUri="https://binary.pensoft.net/fig/780390" pageId="0" pageNumber="179" start="Plate 12" startId="F12">
<paragraph id="E52C2EF9DD6FFACFCA9AC9D373DC0719" pageId="0" pageNumber="179">
<emphasis id="3F919ED449414355E00BC27DD7B32D7A" bold="true" pageId="0" pageNumber="179">Plate 12.</emphasis>
<emphasis id="3CFE4977CC76ED23FDA4D65EE134D0C6" bold="true" pageId="0" pageNumber="179">(1-3)</emphasis>
<taxonomicName id="AD1797BE8542FA10869896F57A0F9477" baseAuthorityName="d'Orbigny" baseAuthorityYear="1852" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis clio" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="clio">
<emphasis id="2C03D3BAA9250442A203C27487521C3A" italics="true" pageId="0" pageNumber="179">Ptygmatis clio</emphasis>
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</taxonomicName>
(
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, 1852).
<emphasis id="01A4A538B95DA140ED8A743156D6BCDC" bold="true" pageId="0" pageNumber="179">(1-2)</emphasis>
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SNSB-BSPG 2021 XV 59, Saal (collection Lang);
<emphasis id="6A3A94A7447A8AB49D6DCFB0A4FA543C" bold="true" pageId="0" pageNumber="179">(1)</emphasis>
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section of fragment, height 21 mm;
<emphasis id="6DBD4BF68F8C557D63FFCCB473762712" bold="true" pageId="0" pageNumber="179">(2)</emphasis>
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detail, height 3.7 mm.
<emphasis id="6571FAE78B0F0F8968D93E5CF42F7577" bold="true" pageId="0" pageNumber="179">(3)</emphasis>
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SNSB-BSPG 2021 XV 60, Saal (collection Lang), lateral view, height 108 mm.
<emphasis id="A0CDAC6FD80B0FFC66AA4A6104B62C9C" bold="true" pageId="0" pageNumber="179">(4)</emphasis>
<taxonomicName id="97196B2B23F8B31B99FBCB85972AC2EB" baseAuthorityName="Quenstedt" baseAuthorityYear="1884" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis polyspira" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="polyspira">Ptygmatis? polyspira</taxonomicName>
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(Quenstedt, 1881-1884), holotype, collection University
<normalizedToken id="E17C9BEF15DF6DD78EA29CFBFD3EA581" originalValue="Tübingen">Tuebingen</normalizedToken>
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(collection Quenstedt); Nattheim, lateral view, height 32 mm (original to
<bibRefCitation id="B3EF9A36B48B47B0CC02826E17A71A9F" author="Quenstedt, FA" journalOrPublisher="Fues, Leipzig" pageId="0" pageNumber="179" refId="B82" refString="Quenstedt, FA, 1881-1884. Petrefaktenkunde Deutschlands. Erste Abteilung, Band 7: Gasteropoden. Fues, Leipzig" title="Petrefaktenkunde Deutschlands. Erste Abteilung, Band 7: Gasteropoden." year="1881 - 1884">Quenstedt 1881-1884</bibRefCitation>
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, pl. 207, fig. 3. (11-12)
<taxonomicName id="F2E96524DC0DEBEBEBBB775AA5FCC025" baseAuthorityName="Muenster in Goldfuss" baseAuthorityYear="1844" class="Gastropoda" family="Ptygmatididae" genus="Bactroptyxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bactroptyxis teres" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="teres">
<emphasis id="2894F933E5D1A21173E113C5B3257CDE" italics="true" pageId="0" pageNumber="179">Bactroptyxis teres</emphasis>
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</taxonomicName>
(
<normalizedToken id="98DF2B147BFFD84C80E0AFF0E52F1E4E" originalValue="Münster">Muenster</normalizedToken>
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in Goldfuss, 1844), collection Sauerborn, Nattheim area.
<emphasis id="DBA021A8E246505A792C382B2457DC7E" bold="true" pageId="0" pageNumber="179">(5-7)</emphasis>
<taxonomicName id="A17B0FBC958982F9E5F1B90A6CD8AFBD" baseAuthorityName="Thurmann, 1861 in Thurmann and Etallon 1861 -" baseAuthorityYear="1864" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina">Ptygmatis? ursicina</taxonomicName>
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(Thurmann in Thurmann &amp;
<normalizedToken id="2FB1F924F840BD5271553D0564028503" originalValue="Étallon">Etallon</normalizedToken>
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, 1861).
<emphasis id="AA63B134DD74B7192E4119A579C8B349" bold="true" pageId="0" pageNumber="179">(5-6)</emphasis>
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SNSB-BSPG 2021 XV 30, Saal (collection Lang);
<emphasis id="8C5D903972C91CD40EA86DA0FE8EA902" bold="true" pageId="0" pageNumber="179">(5)</emphasis>
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last whorl with aperture, height 4 mm;
<emphasis id="75EA48B975E2EBC58ADEECB9373714B3" bold="true" pageId="0" pageNumber="179">(6)</emphasis>
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lateral view, height 9.5 mm.
<emphasis id="B8A098D57D4687C0F798903F3A567751" bold="true" pageId="0" pageNumber="179">(7)</emphasis>
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SNSB-BSPG 2021 XV 31, Saal (collection Lang), aperture and ornament of base, width 2.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C1157329CB039108458C84765CB02CB6" pageId="0" pageNumber="179" type="remarks">
<paragraph id="C4E396A39A22D8F1B86B72F90CDCBB85" pageId="0" pageNumber="179">Remarks.</paragraph>
<paragraph id="B4C2E31271A4E96E35EA33AF62A78286" pageId="0" pageNumber="179">
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The identity of the present juvenile shells with much larger growing taxa is uncertain (see synonymy list). Thurmann in
<bibRefCitation id="4C68D3D767C02CB4B71A41F1FE8E31D5" author="Thurmann, J" journalOrPublisher="Memoires de la Societe du museum d'histoire naturelle de Strasbourg" pageId="0" pageNumber="179" refId="B100" refString="Thurmann, J, Etallon, A, 1861-1864. Lethea bruntrutana ou etudes paleontologiques et stratigraphiques sur le Jura bernois et en particulier les environs de Porrentruy. Neue Denkschrift der allgemeinen Schweizerischen Gesellschaft fuer die Naturwissenschaften 18-20: 1-500." title="Lethea bruntrutana ou etudes paleontologiques et stratigraphiques sur le Jura bernois et en particulier les environs de Porrentruy. Neue Denkschrift der allgemeinen Schweizerischen Gesellschaft fuer die Naturwissenschaften 18 - 20: 1 - 500." year="1861 - 1864">
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Thurmann and
<normalizedToken id="FB1DDD6C41F430F1698A2E75233B2CEF" originalValue="Étallon">Etallon</normalizedToken>
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(1861-1864)
</bibRefCitation>
reported that
<taxonomicName id="A32433987EFFAC37D577D5B3699ABED0" class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina">
<emphasis id="E51E94411D5C766FA59AC18E46E93668" italics="true" pageId="0" pageNumber="179">Nerinea ursicina</emphasis>
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</taxonomicName>
has distinct knobs on the bulges and partly also on the spiral cords between the bulges. Such a knobby ornament could not be substantiated for the present material, due perhaps to preservation.
</paragraph>
</subSubSection>
<subSubSection id="764CBA0824D69395ADE8C3C10E1BB609" pageId="0" pageNumber="179" type="relationships">
<paragraph id="E0FEE84BC4389A6799E90885541F2A7F" pageId="0" pageNumber="179">Relationships.</paragraph>
<paragraph id="17768CA8DCFCC3B26C699CFCEB0D0C65" pageId="0" pageNumber="179">
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Differences to
<taxonomicName id="20FC892B8498A4CD11EC0046FAD04452" baseAuthorityName="Quenstedt" baseAuthorityYear="1884" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis polyspira" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="polyspira">Ptygmatis? polyspira</taxonomicName>
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(Quenstedt, 1881-1884) are herein discussed in the treatment of this taxon.
<taxonomicName id="C49E46AF64D34885F9612BBD83C34E0D" class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea subscalaris" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="subscalaris">
<emphasis id="8456523C22487BBCB0E331C6C1DD0830" italics="true" pageId="0" pageNumber="179">Nerinea subscalaris</emphasis>
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</taxonomicName>
<normalizedToken id="3FA7BB71D8EE8A157F7EE4697006810E" originalValue="Münster">Muenster</normalizedToken>
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in Goldfuss (1844) has higher whorls in relation to whorl width and it has more oblique sutures.
<taxonomicName id="F3E716F6F24507E20A217A67428B9E2B" class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea lafayettensis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="lafayettensis">Nerinea? lafayettensis</taxonomicName>
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Imlay, 1945 has, among other differences, only two spiral cords between the bulges.
<taxonomicName id="90EA9E349EFF45EACEA7D5E2F89EBDCB" class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea speciosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="speciosa">
<emphasis id="38853403CC8F55F6D17BCD3C94BFFA38" italics="true" pageId="0" pageNumber="179">Nerinea speciosa</emphasis>
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</taxonomicName>
Voltz sensu
<bibRefCitation id="EE1ABC4F4C70440340BFC38EFD06EC20" author="Maire, V" journalOrPublisher="Bulletin de la Societe grayloise d'emulation Lettress, Sciences, Beaux-arts" pageId="0" pageNumber="179" refId="B61" refString="Maire, V, 1927. Etudes geologiques et paleontologiques sur l'arrondissement de Gray. Les Gastropodes du Jurassique superieur graylois (fin): 85-173." title="Etudes geologiques et paleontologiques sur l'arrondissement de Gray. Les Gastropodes du Jurassique superieur graylois (fin): 85 - 173." year="1927">Maire (1927)</bibRefCitation>
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has a broader shell with lower whorls and its bulges are not as much protruding.
<taxonomicName id="EF8BF01544B85193DA865B23E881806B" class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea mariae" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="mariae">
<emphasis id="53805237BA8C1E6504E447D06A44CE2C" italics="true" pageId="0" pageNumber="179">Nerinea mariae</emphasis>
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</taxonomicName>
<normalizedToken id="F37B1D9CBC1BD671AB025E202BB1A714" originalValue="dOrbigny">d'Orbigny</normalizedToken>
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sensu
<bibRefCitation id="D6B394B290D5A044EA00F0926B295F3A" author="Cossmann, M" journalOrPublisher="The author and Comptoir Geologique, Paris" pageId="0" pageNumber="179" refId="B12" refString="Cossmann, M, 1898. Contribution a la Paleontologie francaise des terrains jurassiques: Nerinees. Memoires de la Societe gelogique de France, ser. Paleontologie (2) 8: 1-179." title="Contribution a la Paleontologie francaise des terrains jurassiques: Nerinees. Memoires de la Societe gelogique de France, ser. Paleontologie (2) 8: 1 - 179." year="1898">Cossmann (1898)</bibRefCitation>
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is slenderer and has higher whorls.
</paragraph>
</subSubSection>
</treatment>
</document>