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<mods:title id="22700AD18783EB860693577D1BDBC01B">A monographic catalogue on the systematics and phylogeny of the South American iguanian lizard family Liolaemidae (Squamata, Iguania)</mods:title>
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<heading id="E741FC1EFF82612A6CFBED88FE2C8FE8" bold="true" box="[151,483,1687,1713]" fontSize="11" level="1" pageId="50" pageNumber="51" reason="1">
<taxonomicName id="7BB630F1FF82612A6CFBED88FE2C8FE8" ID-CoL="3V6J9" authority="Abdala" authorityName="Abdala" authorityYear="2003" box="[151,483,1687,1713]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="martorii">
<emphasis id="8EC29760FF82612A6CFBED88FE2C8FE8" bold="true" box="[151,483,1687,1713]" pageId="50" pageNumber="51">
<emphasis id="8EC29760FF82612A6CFBED88FE4F8FE8" bold="true" box="[151,384,1687,1713]" italics="true" pageId="50" pageNumber="51">Liolaemus martorii</emphasis>
Abdala
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<treatmentCitation id="3D176D63FF82612A6CFBEDDFFD1D8F8E" author="Vega, L. E. &amp; Bellagamba, P. J." box="[151,722,1728,1751]" page="142" pageId="50" pageNumber="51" year="1994">
<taxonomicName id="7BB630F1FF82612A6CFBEDDFFD1D8F8E" ID-CoL="3V6DB" authority="Vega &amp; Bellagamba 1994: 142" authorityName="Vega &amp; Bellagamba" authorityPageNumber="142" authorityYear="1994" box="[151,722,1728,1751]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="8EC29760FF82612A6CFBEDDFFE4E8F8E" box="[151,385,1728,1751]" italics="true" pageId="50" pageNumber="51">Liolaemus boulengeri</emphasis>
<bibRefCitation id="D8273683FF82612A6DEBEDDFFD1D8F8E" author="Vega &amp; Bellagamba" box="[391,722,1728,1751]" firstAuthor="Vega" page="142" pageId="50" pageNumber="51" pagination="141 - 145" refId="ref73697" refString="Vega, L. E. &amp; Bellagamba, P. J. (1994) Reptiles de la Reserva de Usos Multiples Caleta de los Loros, Rio Negro, Argentina. Cuadernos de Herpetologia, 8, 141 - 145." type="journal article" year="1994">Vega &amp; Bellagamba 1994: 142</bibRefCitation>
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.
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<treatmentCitation id="3D176D63FF82612A6CFBEDFFFDEF8FAE" author="Abdala, C. S." box="[151,544,1760,1783]" page="15" pageId="50" pageNumber="51" year="2003">
<taxonomicName id="7BB630F1FF82612A6CFBEDFFFDEF8FAE" ID-CoL="3V6J9" authority="Abdala 2003: 15" authorityName="Abdala" authorityPageNumber="15" authorityYear="2003" box="[151,544,1760,1783]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="martorii">
<emphasis id="8EC29760FF82612A6CFBEDFFFEA98FAE" box="[151,358,1760,1783]" italics="true" pageId="50" pageNumber="51">Liolaemus martorii</emphasis>
<bibRefCitation id="D8273683FF82612A6D01EDFFFDEF8FAE" author="Abdala" box="[365,544,1760,1783]" firstAuthor="Abdala" page="15" pageId="50" pageNumber="51" pagination="3 - 32" refId="ref50071" refString="Abdala, C. S. (2003) Cuatro nuevas especies del genero Liolaemus (Iguania: Liolaemidae), pertenecientes al grupo boulengeri, de la Patagonia, Argentina. Cuadernos de Herpetologia, 17, 3 - 32." type="journal article" year="2003">Abdala 2003: 15</bibRefCitation>
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(
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locality: Las Grutas, Western San Antonio Department, 
<collectingRegion id="7E728590FF82612A68D3EDFFFA578FAE" box="[1215,1432,1760,1783]" country="Uruguay" name="Rio Negro" pageId="50" pageNumber="51">Río Negro Province</collectingRegion>
, 
<collectingCountry id="C4A10BE2FF82612A6CABEC1FFEFA8E4E" box="[199,309,1792,1815]" name="Argentina" pageId="50" pageNumber="51">Argentina</collectingCountry>
), 2007: 44.
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<paragraph id="BC094B72FF82612B6CFBEC58FDCA880B" blockId="50.[151,1436,1863,2010]" lastBlockId="51.[151,1436,152,338]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
<emphasis id="8EC29760FF82612A6CFBEC58FE898E38" bold="true" box="[151,326,1863,1889]" pageId="50" pageNumber="51">Observations:</emphasis>
The taxonomic status of this species should be reevaluated on the basis of comparative evidence between its 
<typeStatus id="630DF5D0FF82612A6D15EC6FFE648ED3" box="[377,427,1904,1930]" pageId="50" pageNumber="51">type</typeStatus>
series and specimens of the Patagonian species 
<taxonomicName id="7BB630F1FF82612A6F91EC6FFB2B8ED0" authorityName="Muller &amp; Hellmich" authorityYear="1938" box="[1021,1252,1904,1929]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="goetschi">
<emphasis id="8EC29760FF82612A6F91EC6FFB2B8ED0" box="[1021,1252,1904,1929]" italics="true" pageId="50" pageNumber="51">Liolaemus goetschi</emphasis>
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. When 
<bibRefCitation id="D8273683FF82612A692BEC6FFF2B8EEB" author="Abdala" firstAuthor="Abdala" pageId="50" pageNumber="51" pagination="3 - 32" refId="ref50071" refString="Abdala, C. S. (2003) Cuatro nuevas especies del genero Liolaemus (Iguania: Liolaemidae), pertenecientes al grupo boulengeri, de la Patagonia, Argentina. Cuadernos de Herpetologia, 17, 3 - 32." type="journal article" year="2003">Abdala (2003)</bibRefCitation>
proposed 
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<emphasis id="8EC29760FF82612A6D33EC87FE2F8EE8" box="[351,480,1944,1969]" italics="true" pageId="50" pageNumber="51">L. martorii</emphasis>
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as a new species, the taxon 
<taxonomicName id="7BB630F1FF82612A6F47EC87FC638EE8" authorityName="Muller &amp; Hellmich" authorityYear="1938" box="[811,940,1944,1969]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="goetschi">
<emphasis id="8EC29760FF82612A6F47EC87FC638EE8" box="[811,940,1944,1969]" italics="true" pageId="50" pageNumber="51">L. goetschi</emphasis>
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was still considered a synonym of 
<taxonomicName id="7BB630F1FF82612A6925EC87FF128E80" authorityName="Etheridge" authorityYear="1993" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="melanops">
<emphasis id="8EC29760FF82612A6925EC87FF128E80" italics="true" pageId="50" pageNumber="51">L. melanops</emphasis>
</taxonomicName>
(
<emphasis id="8EC29760FF82612A6C82ECDFFED78E80" box="[238,280,1984,2009]" italics="true" pageId="50" pageNumber="51">e.g.</emphasis>
<bibRefCitation id="D8273683FF82612A6D4DECDFFE5F8E83" author="Cei" box="[289,400,1984,2010]" firstAuthor="Cei" pageId="50" pageNumber="51" refId="ref54786" refString="Cei, J. M. (1986) Reptiles del centro, centro-oeste y sur de la Argentina. Herpetofauna de las zonas aridas y semiaridas. Museo Regionale di Scienze Naturali di Torino, Monografie 4, 528 pp." type="book" year="1986">Cei 1986</bibRefCitation>
). Nevertheless, 
<bibRefCitation id="D8273683FF82612A6E20ECDFFC858E83" author="Cei &amp; Scolaro" box="[588,842,1984,2010]" firstAuthor="Cei" pageId="50" pageNumber="51" pagination="107 - 112" refId="ref72364" refString="Scolaro, J. A. &amp; Cei, J. M. (2003) Una excepcional nueva especie de Phymaturus de la precordillera de Chubut, Argentina (Liolaemidae, Iguania, Lacertilia, Reptilia). Facena, 19, 107 - 112." type="journal article" year="2003">Cei &amp; Scolaro (2003)</bibRefCitation>
suggested that 
<taxonomicName id="7BB630F1FF82612A686FECDFFB4B8E80" authorityName="Muller &amp; Hellmich" authorityYear="1938" box="[1027,1156,1984,2009]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="goetschi">
<emphasis id="8EC29760FF82612A686FECDFFB4B8E80" box="[1027,1156,1984,2009]" italics="true" pageId="50" pageNumber="51">L. goetschi</emphasis>
</taxonomicName>
and 
<taxonomicName id="7BB630F1FF82612A68D2ECDFFA818E80" authorityName="Etheridge" authorityYear="1993" box="[1214,1358,1984,2009]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="melanops">
<emphasis id="8EC29760FF82612A68D2ECDFFA818E80" box="[1214,1358,1984,2009]" italics="true" pageId="50" pageNumber="51">L. melanops</emphasis>
</taxonomicName>
represent two different species. This might explain why 
<bibRefCitation id="D8273683FF83612B6E90EB87FC6889EB" author="Abdala" box="[764,935,152,178]" firstAuthor="Abdala" pageId="51" pageNumber="52" pagination="3 - 32" refId="ref50071" refString="Abdala, C. S. (2003) Cuatro nuevas especies del genero Liolaemus (Iguania: Liolaemidae), pertenecientes al grupo boulengeri, de la Patagonia, Argentina. Cuadernos de Herpetologia, 17, 3 - 32." type="journal article" year="2003">Abdala (2003)</bibRefCitation>
did not provide comparisons between the populations identified as 
<taxonomicName id="7BB630F1FF83612B6DD1EBDFFDF18980" authorityName="Muller &amp; Hellmich" authorityYear="1938" box="[445,574,192,217]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="goetschi">
<emphasis id="8EC29760FF83612B6DD1EBDFFDF18980" box="[445,574,192,217]" italics="true" pageId="51" pageNumber="52">L. goetschi</emphasis>
</taxonomicName>
and 
<taxonomicName id="7BB630F1FF83612B6E1AEBDFFD398980" authorityName="Abdala" authorityYear="2003" box="[630,758,192,217]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="martorii">
<emphasis id="8EC29760FF83612B6E1AEBDFFD398980" box="[630,758,192,217]" italics="true" pageId="51" pageNumber="52">L. martorii</emphasis>
</taxonomicName>
. Morphological and field observations between these two taxa carried out by one of us (JAS) suggest that they may be conspecific. Distribution, morphology and coloration exhibit strong overlap between 
<taxonomicName id="7BB630F1FF83612B6E37EA0FFD118870" authorityName="Muller &amp; Hellmich" authorityYear="1938" box="[603,734,272,297]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="goetschi">
<emphasis id="8EC29760FF83612B6E37EA0FFD118870" box="[603,734,272,297]" italics="true" pageId="51" pageNumber="52">L. goetschi</emphasis>
</taxonomicName>
and 
<taxonomicName id="7BB630F1FF83612B6F72EA0FFC6D8870" authorityName="Abdala" authorityYear="2003" box="[798,930,272,297]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="martorii">
<emphasis id="8EC29760FF83612B6F72EA0FFC6D8870" box="[798,930,272,297]" italics="true" pageId="51" pageNumber="52">L. martorii</emphasis>
</taxonomicName>
. The problem should be reassessed using appropriate statistical analyses.
</paragraph>
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