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<document id="8A33F0AC4DDB15D55AE8A5D6907F3233" ID-DOI="10.11646/zootaxa.2062.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5320706" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629618014742" checkinUser="felipe" docAuthor="García-López, Alejandra, Morón, Miguel Ángel, Micó, Estefanía &amp; Galante, Eduardo" docDate="2009" docId="2D3DDF298F335D2BFF4597A8FC0600F3" docLanguage="en" docName="zt02062p045.pdf" docOrigin="Zootaxa 2062 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2062.1.3" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Phyllophaga (Phyllophaga) copalensis Garcia-Lopez &amp; Moron, sp. nov." docType="treatment" docVersion="4" lastPageNumber="41" masterDocId="D104A7518F315D2FFFD2932CFFD50256" masterDocTitle="Two new species of Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) from Costa Rica" masterLastPageNumber="45" masterPageNumber="37" pageNumber="39" updateTime="1699084035901" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="018F632B78C0B7F5C87673B47F9BF83B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DC2DC47DC3CA3009BF28B6BC09897B22">
<mods:title id="AF9F73C701332F137F55B92CA58AFCD0">Two new species of Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) from Costa Rica</mods:title>
</mods:titleInfo>
<mods:name id="39E0E55425FCF3F6722F6012C8DA9EC8" type="personal">
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<mods:roleTerm id="DB9A790479FCAE1C2F5D9B3030FF5BA1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4C62D50E243D40B9BEC9096555C8D765">García-López, Alejandra</mods:namePart>
<mods:affiliation id="26F85A105598D810FE00BEFA6C840890">Centro Iberoamericano de la Biodiversidad (CIBIO), Universidad de Alicante, 03080 Alicante, Spain.</mods:affiliation>
</mods:name>
<mods:name id="399C559B48FE6A0A1CBB5D36A3507DAF" type="personal">
<mods:role id="935FD02E08E6428C88A07B17034A0386">
<mods:roleTerm id="E330F4DC013574A222D632987B83A5F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF9FC4008E97294A14B95C9AC7866235">Morón, Miguel Ángel</mods:namePart>
<mods:affiliation id="7CE1D8ACE4EE4E8B07B8BD424AC3645C">Departamento de Entomología, Instituto de Ecología, A. C. Apdo. Postal 63, Xalapa, Veracruz 91000, México.</mods:affiliation>
</mods:name>
<mods:name id="DDD07EE47FA829981FE8FD00273F6DC1" type="personal">
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<mods:roleTerm id="1A5B47EC875B54BCE7933CA02F71B279">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F421935469BA0855043283FEE21F1612">Micó, Estefanía</mods:namePart>
<mods:affiliation id="63446EFB148CD7165FFA296FCDB8AC91">Centro Iberoamericano de la Biodiversidad (CIBIO), Universidad de Alicante, 03080 Alicante, Spain.</mods:affiliation>
</mods:name>
<mods:name id="65FDAC3EFE7C4A491F1D888818919855" type="personal">
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<mods:roleTerm id="16BF66980CAA87C2B7CC03EA00841667">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF20506D4A191E8F05168E37CE4B40B8">Galante, Eduardo</mods:namePart>
<mods:affiliation id="43768ADC9D935EF26DF65ECE4F44F451">Centro Iberoamericano de la Biodiversidad (CIBIO), Universidad de Alicante, 03080 Alicante, Spain.</mods:affiliation>
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<mods:typeOfResource id="3E37549DBCB2FAD798CBE4FCEE419F7F">text</mods:typeOfResource>
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<mods:title id="037E0E1A05DABD59997BB5687327EC7D">Zootaxa</mods:title>
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<mods:part id="221584FE44D4FE90B7589FC0C4F5EF77">
<mods:date id="2F4090191241E73614949CD31E3FDA08">2009</mods:date>
<mods:detail id="F02BECA890362F58B7BC9613ED36A6C2" type="pubDate">
<mods:number id="480DE0356F49DFCB03879FB1439D6FE9">2009-04-03</mods:number>
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<mods:detail id="FB968519620C22C69DB64C8715A26EDE" type="volume">
<mods:number id="53E455736E2FECAB36F430AFD4989514">2062</mods:number>
</mods:detail>
<mods:detail id="1EC4FF88970914BB841C0BE384FFC172" type="issue">
<mods:number id="B9502CD43EB8DDE49C9D570124A94174">1</mods:number>
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<mods:start id="C7FD16A9958439439EA844C3B1F40B24">37</mods:start>
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<mods:url id="2BF0B13859FB509842848238413A15C2">https://biotaxa.org/Zootaxa/article/view/zootaxa.2062.1.3</mods:url>
</mods:location>
<mods:classification id="16A570E17C5CA5FDF2BA753A6C10270F">journal article</mods:classification>
<mods:identifier id="F42D7B0AEF24094355861875281BB40A" type="DOI">10.11646/zootaxa.2062.1.3</mods:identifier>
<mods:identifier id="50797948465A18FB05CB32B538A37B09" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F40F8658DA2177A8E483C3E06F3E72FB" type="Zenodo-Dep">5320706</mods:identifier>
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<treatment id="2D3DDF298F335D2BFF4597A8FC0600F3" ID-DOI="http://doi.org/10.5281/zenodo.5333551" ID-Zenodo-Dep="5333551" LSID="urn:lsid:plazi:treatment:2D3DDF298F335D2BFF4597A8FC0600F3" httpUri="http://treatment.plazi.org/id/2D3DDF298F335D2BFF4597A8FC0600F3" lastPageId="4" lastPageNumber="41" pageId="2" pageNumber="39">
<subSubSection id="ED8E3DB48F335D2DFF4597A8FC2B06C8" box="[151,1022,1156,1182]" pageId="2" pageNumber="39" type="nomenclature">
<paragraph id="A52B6E3F8F335D2DFF4597A8FC2B06C8" blockId="2.[151,1022,1156,1220]" box="[151,1022,1156,1182]" pageId="2" pageNumber="39">
<heading id="FE63D9538F335D2DFF4597A8FC2B06C8" bold="true" box="[151,1022,1156,1182]" fontSize="11" level="1" pageId="2" pageNumber="39" reason="1">
<emphasis id="97E0B22D8F335D2DFF4597A8FC2B06C8" bold="true" box="[151,1022,1156,1182]" pageId="2" pageNumber="39">
<taxonomicName id="629415BC8F335D2DFF4597A8FC4C06C8" authority="Garcia-Lopez &amp; Moron" authorityName="Garcia-Lopez &amp; Moron" box="[151,921,1156,1182]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="copalensis" status="sp. nov." subGenus="Phyllophaga">
<emphasis id="97E0B22D8F335D2DFF4597A8FDBD06C8" bold="true" box="[151,616,1156,1182]" italics="true" pageId="2" pageNumber="39">Phyllophaga (Phyllophaga) copalensis</emphasis>
García-López &amp; Morón
</taxonomicName>
<taxonomicNameLabel id="8CD30F568F335D2DFC7297A8FC2B06C8" box="[928,1022,1156,1182]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ED8E3DB48F335D2DFF459786FB6805F5" pageId="2" pageNumber="39" type="description">
<paragraph id="A52B6E3F8F335D2DFF459786FEE10692" blockId="2.[151,1022,1156,1220]" box="[151,308,1194,1220]" pageId="2" pageNumber="39">
(
<figureCitation id="3DAF72BA8F335D2DFF4D9786FF240692" box="[159,241,1194,1220]" captionStart="FIGURE 1" captionStartId="3.[151,255,1232,1256]" captionText="FIGURE 1. Phyllophaga copalensis sp. nov. dorsal view of holotype." pageId="2" pageNumber="39">Figs. 1</figureCitation>
,
<figureCitation id="3DAF72BA8F335D2DFF2D9786FEF90692" box="[255,300,1194,1220]" captionStart="FIGURES 2–4" captionStartId="3.[151,271,1942,1966]" captionTargetBox="[156,1389,1296,1927]" captionTargetId="figure-58@3.[151,1436,1296,1931]" captionTargetPageId="3" captionText="FIGURES 2–4. Phyllophaga copalensis, male genital capsule. 2—lateral view with aedeagus extended. 3—Apex of parameres, ventral view. 4—Dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320710" httpUri="https://zenodo.org/record/5320710/files/figure.png" pageId="2" pageNumber="39">2–4</figureCitation>
)
</paragraph>
<paragraph id="A52B6E3F8F335D2DFF4597D4FB6805F5" blockId="2.[151,1437,1272,2033]" pageId="2" pageNumber="39">
<emphasis id="97E0B22D8F335D2DFF4597D4FE100744" bold="true" box="[151,453,1272,1298]" pageId="2" pageNumber="39">
Description.
<typeStatus id="7A2FD09D8F335D2DFE9697D4FE150744" box="[324,448,1272,1298]" pageId="2" pageNumber="39" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male: Total body length:
<quantity id="626CC3DA8F335D2DFCC497D4FC5F0744" box="[790,906,1272,1298]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="2" pageNumber="39" unit="mm" value="18.6">18.6 mm</quantity>
. Humeral width:
<quantity id="626CC3DA8F335D2DFBBB97D4FB190744" box="[1129,1228,1272,1298]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="2" pageNumber="39" unit="mm" value="8.1">8.1 mm</quantity>
. Clypeus, frons, pronotum and elytra dark brown; head and pronotum shiny with erect, mixed long and short setae on entire surface; mouth parts, sterna, pygidium and legs shiny reddish brown (
<figureCitation id="3DAF72BA8F335D2DFC0E9669FBF70709" box="[988,1058,1349,1375]" captionStart="FIGURE 1" captionStartId="3.[151,255,1232,1256]" captionText="FIGURE 1. Phyllophaga copalensis sp. nov. dorsal view of holotype." pageId="2" pageNumber="39">Fig. 1</figureCitation>
). Clypeus 3.6 times wider than long, surface densely rugopunctate, with many short, erect setae. Frons 2 times wider than long, convex, densely rugopunctate. Frons 4 times wider than dorsal diameter of eye. Eye canthus long and wide, with 11–14 setae. Mentum with anterior border moderately sinuate. Pronotum 2 times wider than long and 2.4 times wider than frons. Pronotal disk with large, shallow, round punctures irregularly separated from one another by less than 1 diameter to 2–3 diameters; lateral borders strongly angled, lateral marginal bead crenulate, with long, slender setae evenly distributed; anterior angles right, clearly prominent; posterior angles obtuse, prominent. Scutellum 1.4 times wider than long. Elytra 2.6 times longer than wider, densely setiferous punctate. Anal plate with transverse row of 6 long setae near the posterior border. Mesotibia with upper apical spur as long as lower spur. Upper apical spur of the metatibia with apex acute, 1.2 times longer than basal metatarsomere, and 1.37 times longer than lower spur. Protarsomeres 1–3 each with subapical small, toothlike prominence. Genital capsule with one lateroapical tooth at each side; apex straight, progressively narrowed, with the ventral border briefly notched (
<figureCitation id="3DAF72BA8F335D2DFCC59439FC460579" box="[791,915,1813,1839]" captionStart="FIGURES 2–4" captionStartId="3.[151,271,1942,1966]" captionTargetBox="[156,1389,1296,1927]" captionTargetId="figure-58@3.[151,1436,1296,1931]" captionTargetPageId="3" captionText="FIGURES 2–4. Phyllophaga copalensis, male genital capsule. 2—lateral view with aedeagus extended. 3—Apex of parameres, ventral view. 4—Dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320710" httpUri="https://zenodo.org/record/5320710/files/figure.png" pageId="2" pageNumber="39">Figs. 2–4</figureCitation>
). Aedeagus long, with dorsal spatula-like sclerotized support, with distal tuft of 8 long bristles; ventroapical border of the sclerotized support downturned, widely rounded; preapical area of inner sac with patches of golden setae and strongly sclerotized spines. Length of genital capsule from apex of parameres to border of phallobase:
<quantity id="626CC3DA8F335D2DFB8994A5FB6C05F5" box="[1115,1209,1929,1955]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="39" unit="mm" value="4.5">4.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="ED8E3DB48F335D2BFF14949CFC9003C1" lastPageId="4" lastPageNumber="41" pageId="2" pageNumber="39" type="materials_examined">
<paragraph id="A52B6E3F8F335D2DFF14949CFC0505A7" blockId="2.[151,1437,1272,2033]" pageId="2" pageNumber="39">
<emphasis id="97E0B22D8F335D2DFF14949CFE6B059C" bold="true" box="[198,446,1968,1994]" pageId="2" pageNumber="39">Material examined.</emphasis>
(
<materialsCitation id="15FC64628F335D2DFE06949CFC0505A7" collectingDate="2007-06-15" collectionCode="INBio" collectorName="Selva. Alb. El Copal &amp; A. Garcia &amp; M. Moraga" country="Costa Rica" elevation="1000" location="Pejibaye" municipality="Jimenez" pageId="2" pageNumber="39" specimenCount="1" stateProvince="Cartago" typeStatus="holotype">
<specimenCount id="B392A5B68F335D2DFE06949CFDB5059C" box="[468,608,1968,1994]" pageId="2" pageNumber="39" type="generic">1 specimen</specimenCount>
).
<typeStatus id="7A2FD09D8F335D2DFDA5949CFD31059C" box="[631,740,1968,1994]" pageId="2" pageNumber="39" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="DD832EAF8F335D2DFCEB949CFC39059C" box="[825,1004,1968,1994]" name="Costa Rica" pageId="2" pageNumber="39">COSTA RICA</collectingCountry>
:
<collectingRegion id="6750A0DD8F335D2DFC29949CFB8E059C" box="[1019,1115,1968,1994]" country="Costa Rica" name="Cartago" pageId="2" pageNumber="39">Cartago</collectingRegion>
,
<collectingMunicipality id="454FF4458F335D2DFBB8949CFB18059C" box="[1130,1229,1968,1994]" pageId="2" pageNumber="39">Jiménez</collectingMunicipality>
,
<location id="A04B38E48F335D2DFB0F949CFA90059C" LSID="urn:lsid:plazi:treatment:2D3DDF298F335D2BFF4597A8FC0600F3:A04B38E48F335D2DFB0F949CFA90059C" box="[1245,1349,1968,1994]" country="Costa Rica" municipality="Jimenez" name="Pejibaye" pageId="2" pageNumber="39" stateProvince="Cartago">Pejibaye</location>
,
<collectorName id="08610BE98F335D2DFA86949CFEE905A7" pageId="2" pageNumber="39">Selva. Alb. El Copal</collectorName>
,
<quantity id="626CC3DA8F335D2DFE9594FBFE7705A7" box="[327,418,2007,2033]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="39" unit="m" value="1000.0">
<elevation id="2EB9890C8F335D2DFE9594FBFE7705A7" box="[327,418,2007,2033]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="39" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="D12A48FF8F335D2DFE7C94FBFD8005A7" box="[430,597,2007,2033]" pageId="2" pageNumber="39" value="2007-06-15">
<collectingDate id="C16EB1178F335D2DFE7C94FBFD8005A7" box="[430,597,2007,2033]" pageId="2" pageNumber="39" value="2007-06-15">June 15, 2007</collectingDate>
</date>
.
<collectorName id="08610BE98F335D2DFDB394FBFD0005A7" box="[609,725,2007,2033]" pageId="2" pageNumber="39">A. García</collectorName>
,
<collectorName id="08610BE98F335D2DFD3394FBFCB005A7" box="[737,869,2007,2033]" pageId="2" pageNumber="39">M. Moraga</collectorName>
(
<collectionCode id="C385F6FA8F335D2DFCA794FBFC1105A7" box="[885,964,2007,2033]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="2" pageNumber="39">INBio</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="F1EB3EB78F325D2CFF4597FCFC5006BE" box="[151,901,1232,1256]" pageId="3" pageNumber="40" startId="3.[151,255,1232,1256]">
<paragraph id="A52B6E3F8F325D2CFF4597FCFC5006BE" blockId="3.[151,901,1232,1256]" box="[151,901,1232,1256]" pageId="3" pageNumber="40">
<emphasis id="97E0B22D8F325D2CFF4597FCFECC06BE" bold="true" box="[151,281,1232,1256]" pageId="3" pageNumber="40">FIGURE 1.</emphasis>
<taxonomicName id="629415BC8F325D2CFEF297FDFDCA06BE" box="[288,543,1233,1256]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="copalensis" status="sp. nov.">
<emphasis id="97E0B22D8F325D2CFEF297FDFDCA06BE" box="[288,543,1233,1256]" italics="true" pageId="3" pageNumber="40">Phyllophaga copalensis</emphasis>
</taxonomicName>
<emphasis id="97E0B22D8F325D2CFDF497FCFDAE06BE" bold="true" box="[550,635,1232,1256]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="8CD30F568F325D2CFDF497FCFDAE06BE" box="[550,635,1232,1256]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
dorsal view of holotype.
</paragraph>
</caption>
<caption id="F1EB3EB78F325D2CFF4594BAFCD00586" ID-DOI="http://doi.org/10.5281/zenodo.5320710" ID-Zenodo-Dep="5320710" httpUri="https://zenodo.org/record/5320710/files/figure.png" pageId="3" pageNumber="40" startId="3.[151,271,1942,1966]" targetBox="[156,1389,1296,1927]" targetPageId="3">
<paragraph id="A52B6E3F8F325D2CFF4594BAFCD00586" blockId="3.[151,1436,1942,2000]" pageId="3" pageNumber="40">
<emphasis id="97E0B22D8F325D2CFF4594BAFE9305F8" bold="true" box="[151,326,1942,1966]" pageId="3" pageNumber="40">FIGURES 2–4.</emphasis>
<taxonomicName id="629415BC8F325D2CFE9C94BBFD8005F8" box="[334,597,1943,1966]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="copalensis">
<emphasis id="97E0B22D8F325D2CFE9C94BBFD8005F8" box="[334,597,1943,1966]" italics="true" pageId="3" pageNumber="40">Phyllophaga copalensis</emphasis>
</taxonomicName>
, male genital capsule. 2—lateral view with aedeagus extended. 3—Apex of parameres, ventral view. 4—Dorsal view. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="A52B6E3F8F355D2BFF1D93B2FED00352" blockId="4.[151,1437,158,677]" pageId="4" pageNumber="41">
<materialsCitation id="15FC64628F355D2BFF1D93B2FED40352" collectorName="Reserva Biologica El Copal" country="Costa Rica" latitude="9.786112" location="Area de Conservacion La Amistad-Pacifico" longLatPrecision="21" longitude="-83.75166" municipality="Parque Nacional Tapanti-Macizo de la Muerte. Province" pageId="4" pageNumber="41" specimenCount="1" stateProvince="Cartago" typeStatus="holotype">
<emphasis id="97E0B22D8F355D2BFF1D93B2FEAE02EE" bold="true" box="[207,379,158,184]" pageId="4" pageNumber="41">
<typeStatus id="7A2FD09D8F355D2BFF1D93B2FED802EE" box="[207,269,158,184]" pageId="4" pageNumber="41">Type</typeStatus>
locality.
</emphasis>
<collectorName id="08610BE98F355D2BFE5A938CFD0302E1" box="[392,726,159,183]" pageId="4" pageNumber="41">Reserva Biológica El Copal</collectorName>
:
<location id="A04B38E48F355D2BFD3593B3FB3202E1" LSID="urn:lsid:plazi:treatment:2D3DDF298F335D2BFF4597A8FC0600F3:A04B38E48F355D2BFD3593B3FB3202E1" box="[743,1255,159,183]" country="Costa Rica" latitude="9.786112" longLatPrecision="21" longitude="-83.75166" municipality="Parque Nacional Tapanti-Macizo de la Muerte. Province" name="Area de Conservacion La Amistad-Pacifico" pageId="4" pageNumber="41" stateProvince="Cartago">Área de Conservación La Amistad-Pacífico</location>
, adjoining the
<collectingMunicipality id="454FF4458F355D2BFF4593E8FCFA0288" box="[151,815,196,222]" pageId="4" pageNumber="41">Parque Nacional Tapantí-Macizo de la Muerte. Province</collectingMunicipality>
of
<collectingRegion id="6750A0DD8F355D2BFC8493E8FC620288" box="[854,951,196,222]" country="Costa Rica" name="Cartago" pageId="4" pageNumber="41">Cartago</collectingRegion>
.
<collectingCountry id="DD832EAF8F355D2BFC1093E8FB970288" box="[962,1090,196,222]" name="Costa Rica" pageId="4" pageNumber="41">Costa Rica</collectingCountry>
(
<geoCoordinate id="C0A008F88F355D2BFB8093E8FB3F0288" box="[1106,1258,196,222]" degrees="9" direction="north" minutes="47" orientation="latitude" pageId="4" pageNumber="41" precision="15" seconds="10" value="9.786112">9º 47’ 10” N</geoCoordinate>
,
<geoCoordinate id="C0A008F88F355D2BFB2493E8FA460288" box="[1270,1427,196,222]" degrees="83" direction="west" minutes="45" orientation="longitude" pageId="4" pageNumber="41" precision="15" seconds="6" value="-83.75166">83º 45’ 6” W</geoCoordinate>
) (
<figureCitation id="3DAF72BA8F355D2BFF4D93C6FF230352" box="[159,246,234,260]" captionStart="FIGURE 10" captionStartId="6.[151,255,1891,1915]" captionTargetBox="[192,1391,778,1844]" captionTargetId="figure-273@6.[151,1436,743,1898]" captionTargetPageId="6" captionText="FIGURE 10. Distribution of Phyllophaga larai, P. copalensis, P. guapiles and P. guapiloides in Costa Rica. Life Zones map provided by GIS Unit of Inbio. Distribution of P. guapiles and P. guapiloides taken from ATTA database (InBio)." figureDoi="http://doi.org/10.5281/zenodo.5320714" httpUri="https://zenodo.org/record/5320714/files/figure.png" pageId="4" pageNumber="41">Fig. 10</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A52B6E3F8F355D2BFF149224FC9003C1" blockId="4.[151,1437,158,677]" pageId="4" pageNumber="41">
<emphasis id="97E0B22D8F355D2BFF149224FE520374" bold="true" box="[198,391,264,290]" pageId="4" pageNumber="41">Biological data.</emphasis>
Phenology: June. Other species of
<taxonomicName id="629415BC8F355D2BFCF19225FC6C0374" authorityName="Harris" authorityYear="1826" box="[803,953,265,290]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="97E0B22D8F355D2BFCF19225FC6C0374" box="[803,953,265,290]" italics="true" pageId="4" pageNumber="41">Phyllophaga</emphasis>
</taxonomicName>
flying at the same time and place were
<taxonomicName id="629415BC8F355D2BFA549225FE89031C" authority="(Moser)" baseAuthorityName="Moser" baseAuthorityYear="1918" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="densata">
<emphasis id="97E0B22D8F355D2BFA549225FF26031F" italics="true" pageId="4" pageNumber="41">P. densata</emphasis>
(Moser)
</taxonomicName>
,
<taxonomicName id="629415BC8F355D2BFEBB921CFD2F031C" authority="(Moron &amp; Solis)" baseAuthorityName="Moron &amp; Solis" box="[361,762,304,330]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="guanacasteca">
<emphasis id="97E0B22D8F355D2BFEBB921CFDFC031F" box="[361,553,304,329]" italics="true" pageId="4" pageNumber="41">P. guanacasteca</emphasis>
(Morón &amp; Solís)
</taxonomicName>
,
<taxonomicName id="629415BC8F355D2BFCD5921CFC21031C" authority="(Saylor)" baseAuthorityName="Saylor" box="[775,1012,304,330]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="guapiles">
<emphasis id="97E0B22D8F355D2BFCD5921CFC5C031F" box="[775,905,304,329]" italics="true" pageId="4" pageNumber="41">P. guapiles</emphasis>
(Saylor)
</taxonomicName>
,
<taxonomicName id="629415BC8F355D2BFBD3921CFAAF031C" authority="(Moron &amp; Solis)" baseAuthorityName="Moron &amp; Solis" box="[1025,1402,304,330]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="guapiloides">
<emphasis id="97E0B22D8F355D2BFBD3921CFB7C031F" box="[1025,1193,304,329]" italics="true" pageId="4" pageNumber="41">P. guapiloides</emphasis>
(Morón &amp; Solís)
</taxonomicName>
,
<taxonomicName id="629415BC8F355D2BFA54921CFEA00326" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1888" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="lissopyge">
<emphasis id="97E0B22D8F355D2BFA54921CFED80326" italics="true" pageId="4" pageNumber="41">P. lissopyge</emphasis>
(Bates)
</taxonomicName>
,
<taxonomicName id="629415BC8F355D2BFE57927BFD2C0326" authority="(Moron &amp; Solis)" baseAuthorityName="Moron &amp; Solis" box="[389,761,342,368]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="lorencita">
<emphasis id="97E0B22D8F355D2BFE57927BFDC20326" box="[389,535,343,368]" italics="true" pageId="4" pageNumber="41">P. lorencita</emphasis>
(Morón &amp; Solís)
</taxonomicName>
,
<taxonomicName id="629415BC8F355D2BFCDB927BFC2C0326" authority="(Moser)" baseAuthorityName="Moser" baseAuthorityYear="1918" box="[777,1017,342,368]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="orosina">
<emphasis id="97E0B22D8F355D2BFCDB927BFC5C0326" box="[777,905,343,368]" italics="true" pageId="4" pageNumber="41">P. orosina</emphasis>
(Moser)
</taxonomicName>
,
<taxonomicName id="629415BC8F355D2BFBD9927BFB2E0326" authority="(Bates)" baseAuthorityName="Bates" box="[1035,1275,342,368]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="prolixus">
<emphasis id="97E0B22D8F355D2BFBD9927BFB460326" box="[1035,1171,343,368]" italics="true" pageId="4" pageNumber="41">P. prolixus</emphasis>
(Bates)
</taxonomicName>
,
<taxonomicName id="629415BC8F355D2BFAD9927BFEF303C1" authority="(Blanchard)" baseAuthorityName="Blanchard" baseAuthorityYear="1850" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="pruinosa">
<emphasis id="97E0B22D8F355D2BFAD9927BFA490326" box="[1291,1436,343,368]" italics="true" pageId="4" pageNumber="41">P. pruinosa</emphasis>
(Blanchard)
</taxonomicName>
,
<taxonomicName id="629415BC8F355D2BFEE39251FD8703C1" authority="(Blanchard)" baseAuthorityName="Blanchard" box="[305,594,381,407]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="setidorsis">
<emphasis id="97E0B22D8F355D2BFEE39251FE6B03C0" box="[305,446,381,406]" italics="true" pageId="4" pageNumber="41">P. setidorsis</emphasis>
(Blanchard)
</taxonomicName>
and
<taxonomicName id="629415BC8F355D2BFD599251FD3103C0" authorityName="Garcia-Lopez &amp; Moron" authorityYear="2009" box="[651,740,381,406]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="larai" status="sp. nov.">
<emphasis id="97E0B22D8F355D2BFD599251FD3103C0" box="[651,740,381,406]" italics="true" pageId="4" pageNumber="41">P.larai</emphasis>
</taxonomicName>
<emphasis id="97E0B22D8F355D2BFD3A9251FC9003C1" bold="true" box="[744,837,381,407]" pageId="4" pageNumber="41">
<taxonomicNameLabel id="8CD30F568F355D2BFD3A9251FC9003C1" box="[744,837,381,407]" pageId="4" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="ED8E3DB48F355D2BFF14928FFA43000E" pageId="4" pageNumber="41" type="discussion">
<paragraph id="A52B6E3F8F355D2BFF14928FFA43000E" blockId="4.[151,1437,158,677]" pageId="4" pageNumber="41">
<emphasis id="97E0B22D8F355D2BFF14928FFEEA03EB" bold="true" box="[198,319,419,445]" pageId="4" pageNumber="41">Remarks.</emphasis>
<taxonomicName id="629415BC8F355D2BFE959288FDB403EB" box="[327,609,420,445]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="copalensis">
<emphasis id="97E0B22D8F355D2BFE959288FDB403EB" box="[327,609,420,445]" italics="true" pageId="4" pageNumber="41">Phyllophaga copalensis</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="629415BC8F355D2BFC849288FC7803EB" authorityName="Garcia-Lopez &amp; Moron" authorityYear="2009" box="[854,941,420,445]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="larai">
<emphasis id="97E0B22D8F355D2BFC849288FC7803EB" box="[854,941,420,445]" italics="true" pageId="4" pageNumber="41">P. larai</emphasis>
</taxonomicName>
but the small body size, dark color, shiny pronotum and elytra, shallow large punctures on the pronotum, long elytral setae, and lateroventral border of parameres with rounded tooth on each side will help to separate these species. It is also related to
<taxonomicName id="629415BC8F355D2BFAC992DDFF360064" authority="Saylor" authorityName="Saylor. Moron and Solis" authorityYear="2001" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="guapiles">
<emphasis id="97E0B22D8F355D2BFAC992DDFA49005C" box="[1307,1436,497,522]" italics="true" pageId="4" pageNumber="41">P. guapiles</emphasis>
Saylor
</taxonomicName>
from the same area in
<collectingCountry id="DD832EAF8F355D2BFE209134FDA10064" box="[498,628,536,562]" name="Costa Rica" pageId="4" pageNumber="41">Costa Rica</collectingCountry>
, but the uniform and dense vestiture of the pronotum and elytra, and the ventroapical border of the sclerotized support of the aedeagus are clearly different
<bibRefCitation id="C10513CE8F355D2BFB5B9112FA47000E" author="Moron &amp; Solis" box="[1161,1426,574,600]" firstAuthor="Moron" pageId="4" pageNumber="41" pagination="11 - 29" refId="ref4546" refString="Moron, M. A. &amp; Solis, A. (2001) Seven new species of Phyllophaga (s. str.) Harris from Costa Rica (Coleoptera: Melolonthidae: Melolonthinae). The Coleopterists Bulletin, 55 (1), 11 - 29." type="journal article" year="2001">Morón &amp; Solís (2001)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ED8E3DB48F355D2BFF149149FC0600F3" pageId="4" pageNumber="41" type="etymology">
<paragraph id="A52B6E3F8F355D2BFF149149FC0600F3" blockId="4.[151,1437,158,677]" pageId="4" pageNumber="41">
<emphasis id="97E0B22D8F355D2BFF149149FE860029" bold="true" box="[198,339,613,639]" pageId="4" pageNumber="41">Etymology.</emphasis>
Derived from the locality name El Copal, where the
<typeStatus id="7A2FD09D8F355D2BFC0F9149FB910029" box="[989,1092,613,639]" pageId="4" pageNumber="41" type="holotype">holotype</typeStatus>
was collected. This name is dedicated with gratitude to the people who run the Biological Reserve.
</paragraph>
</subSubSection>
</treatment>
</document>
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