<documentid="0262C8642A1C01935041054FAA44C38F"ID-CLB-Dataset="307155"ID-DOI="10.11646/zootaxa.5562.1.4"ID-GBIF-Dataset="0e124a17-4939-4466-b5dd-33de19d0be92"ID-ISSN="1175-5326"ID-Zenodo-Dep="14595706"ID-ZooBank="91C860B5-34D1-46A6-B10A-5C4A2C4C2F07"IM.bibliography_approvedBy="julia"IM.illustrations_approvedBy="julia"IM.materialsCitations_approvedBy="julia"IM.metadata_approvedBy="felipe"IM.tables_approvedBy="julia"IM.taxonomicNames_approvedBy="felipe"IM.treatments_approvedBy="julia"checkinTime="1735932052183"checkinUser="plazi"docAuthor="Boderau, Mathieu, Nel, André, Maksoud, Sibelle, Abi-Saad, Pierre & Azar, Dany"docDate="2024"docId="2A5265588E42FF83FF632BACFBA5FCA5"docLanguage="en"docName="zootaxa.5562.1.4.pdf"docOrigin="Zootaxa 5562 (1)"docSource="https://doi.org/10.11646/zootaxa.5562.1.4"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Libanobelostoma calineae Boderau, Nel, Maksoud, Abi-Saad & Azar, 2024, sp. nov."docType="treatment"docVersion="7"lastPageNumber="17"masterDocId="D66B1D208E47FF85FFF42D5CFF8FFF9C"masterDocTitle="The first aquatic insect fossil from the Cenomanian Konservat-Lagerstätte of Haqel (Lebanon) fills a gap in the fossil record of giant water bugs (Heteroptera: Belostomatidae)"masterLastPageNumber="22"masterPageNumber="11"pageNumber="16"updateTime="1736168727340"updateUser="julia"zenodo-license-document="CLOSED">
<mods:titleid="8572B65AD2BC9475B733395064D7CE0B">The first aquatic insect fossil from the Cenomanian Konservat-Lagerstätte of Haqel (Lebanon) fills a gap in the fossil record of giant water bugs (Heteroptera: Belostomatidae)</mods:title>
<mods:affiliationid="C9A4CB8CA19D27FF58727FA4F24A964C">Institut de Systématique, Évolution, Biodiversité (UMR 7205) Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, F- 75005 Paris, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, China</mods:affiliation>
<mods:affiliationid="6D25448FDF7947C94E8CE27EC1857046">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, China & Lebanese University, Faculty of Sciences II, Department of Natural Sciences, Fanar, P. O. Box 26110217, Fanar-Matn, Lebanon</mods:affiliation>
<mods:affiliationid="8CCC681C605904AE86711BD77CB6630C">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, China & Lebanese University, Faculty of Sciences II, Department of Natural Sciences, Fanar, P. O. Box 26110217, Fanar-Matn, Lebanon & Institut de Systématique, Évolution, Biodiversité (UMR 7205) Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, F- 75005 Paris, France</mods:affiliation>
<taxonomicNameid="65FBAFCD8E42FF80FF632BACFE5BF897"authorityName="Boderau & Nel & Maksoud & Abi-Saad & Azar"authorityYear="2024"box="[151,468,1776,1803]"class="Insecta"family="Belostomatidae"genus="Libanobelostoma"kingdom="Animalia"order="Hemiptera"pageId="5"pageNumber="16"phylum="Arthropoda"rank="species"species="calineae"status="sp. nov.">
<figureCitationid="3AC0C8CB8E42FF80FF632A48FF77F8B2"box="[151,248,1812,1838]"captionStart-0="FIGURE 2"captionStart-1="FIGURE 3"captionStartId-0="4.[151,251,1943,1967]"captionStartId-1="5.[151,250,1490,1514]"captionTargetBox-0="[165,1422,179,1917]"captionTargetBox-1="[151,1436,179,1464]"captionTargetId-0="figure-41@4.[165,1422,179,1917]"captionTargetId-1="figure-22@5.[151,1436,179,1464]"captionTargetPageId-0="4"captionTargetPageId-1="5"captionText-0="FIGURE 2. Libanobelostoma calineae gen et sp. nov., holotype I-26247/2A (part) and B (counterpart), general habitus. A–B, Dorsum under natural light (A) and under alcohol (B). C–D, Venter under natural light (C) and under alcohol (D). Scale bars = 1 cm."captionText-1="FIGURE 3. Libanobelostoma calineae gen et sp. nov., holotype I-26247/2A (part) and B (counterpart). A, Ventral view of head. B, Reconstruction of rostrum. C, Dorsal view of head and prothoracic legs. D, Reconstruction of right raptorial fore leg. E, Abdominal sternum. F, Hind leg in dorsal view. Abbreviations: ant = antennae; as = air straps (siphon); clw = claw; ffm = forefemur; flp = finger-like projections; ftb = foretibia; fts = foretarsus; hfm = hind femur; htb = hind tibia; hts = hind tarsus; lbr1 = first labial segment; lbr2 = second labial segment; lbr3 = third labial segment; rfl = right foreleg; set = setae. IV–VII: abdominal sterna IV–VII. Scale bars: A–D = 1 mm; E–F = 2 mm."figureDoi-0="http://doi.org/10.5281/zenodo.14595712"figureDoi-1="http://doi.org/10.5281/zenodo.14595716"httpUri-0="https://zenodo.org/record/14595712/files/figure.png"httpUri-1="https://zenodo.org/record/14595716/files/figure.png"pageId="5"pageNumber="16">Figs 2–3</figureCitation>
<materialsCitationid="1293DE138E42FF80FEA42ADCFDBAF822"collectionCode="I"pageId="5"pageNumber="16"specimenCode="I-26247/2A and B"specimenCount="1"typeStatus="holotype">
<figureCitationid="3AC0C8CB8E41FF83FD362C5FFCCDFE81"box="[706,834,259,285]"captionStart="FIGURE 2"captionStartId="4.[151,251,1943,1967]"captionTargetBox="[165,1422,179,1917]"captionTargetId="figure-41@4.[165,1422,179,1917]"captionTargetPageId="4"captionText="FIGURE 2. Libanobelostoma calineae gen et sp. nov., holotype I-26247/2A (part) and B (counterpart), general habitus. A–B, Dorsum under natural light (A) and under alcohol (B). C–D, Venter under natural light (C) and under alcohol (D). Scale bars = 1 cm."figureDoi="http://doi.org/10.5281/zenodo.14595712"httpUri="https://zenodo.org/record/14595712/files/figure.png"pageId="6"pageNumber="17">Fig. 2A–B</figureCitation>
<figureCitationid="3AC0C8CB8E41FF83FBE62C17FBE2FEF9"box="[1042,1133,331,357]"captionStart="FIGURE 1"captionStartId="3.[151,250,1829,1853]"captionTargetBox="[183,1403,235,1803]"captionTargetId="figure-59@3.[183,1403,234,1803]"captionTargetPageId="3"captionText="FIGURE 1. Mid-Cretaceous fossil site Konservat-Lagerstätte of Haqel, Lebanon. A, Geological map of outcrop (modified from Dubertret, 1945; Dubertret & Wetzel, 1945). J6 = uppermost Jurassic; C2a = lower Barremian ‘Grès du Liban’ sandstone; C2b = Barremian clay and oolitic deposition of upper part of ‘Grès du Liban’ and oolitic deposition of lower part of Jezzinian; C3 = micritic part of Jezzinian (uppermost Barremian-lowermost Aptian); C4 = Albian; C5 = Cenomanian; Q = Quaternary scree; βJ6 = Kimmeridgian volcanic deposition; βC3 = Jezzinian volcanic deposition. Thick lines represent faults. B, General view of outcrop. C–D, Details of stratigraphic layers of outcrop."figureDoi="http://doi.org/10.5281/zenodo.14595710"httpUri="https://zenodo.org/record/14595710/files/figure.png"pageId="6"pageNumber="17">Fig. 1A</figureCitation>
<figureCitationid="3AC0C8CB8E41FF83FB122C33FAEDFE15"box="[1254,1378,367,393]"captionStart="FIGURE 1"captionStartId="3.[151,250,1829,1853]"captionTargetBox="[183,1403,235,1803]"captionTargetId="figure-59@3.[183,1403,234,1803]"captionTargetPageId="3"captionText="FIGURE 1. Mid-Cretaceous fossil site Konservat-Lagerstätte of Haqel, Lebanon. A, Geological map of outcrop (modified from Dubertret, 1945; Dubertret & Wetzel, 1945). J6 = uppermost Jurassic; C2a = lower Barremian ‘Grès du Liban’ sandstone; C2b = Barremian clay and oolitic deposition of upper part of ‘Grès du Liban’ and oolitic deposition of lower part of Jezzinian; C3 = micritic part of Jezzinian (uppermost Barremian-lowermost Aptian); C4 = Albian; C5 = Cenomanian; Q = Quaternary scree; βJ6 = Kimmeridgian volcanic deposition; βC3 = Jezzinian volcanic deposition. Thick lines represent faults. B, General view of outcrop. C–D, Details of stratigraphic layers of outcrop."figureDoi="http://doi.org/10.5281/zenodo.14595710"httpUri="https://zenodo.org/record/14595710/files/figure.png"pageId="6"pageNumber="17">Fig. 1C–D</figureCitation>
<figureCitationid="3AC0C8CB8E41FF83FA842C33FF52FE31"captionStart="FIGURE 2"captionStartId="4.[151,251,1943,1967]"captionTargetBox="[165,1422,179,1917]"captionTargetId="figure-41@4.[165,1422,179,1917]"captionTargetPageId="4"captionText="FIGURE 2. Libanobelostoma calineae gen et sp. nov., holotype I-26247/2A (part) and B (counterpart), general habitus. A–B, Dorsum under natural light (A) and under alcohol (B). C–D, Venter under natural light (C) and under alcohol (D). Scale bars = 1 cm."figureDoi="http://doi.org/10.5281/zenodo.14595712"httpUri="https://zenodo.org/record/14595712/files/figure.png"pageId="6"pageNumber="17">Fig. 2A–B</figureCitation>
<figureCitationid="3AC0C8CB8E41FF83FF6B2FD3FE93FD35"box="[159,284,655,681]"captionStart="FIGURE 2"captionStartId="4.[151,251,1943,1967]"captionTargetBox="[165,1422,179,1917]"captionTargetId="figure-41@4.[165,1422,179,1917]"captionTargetPageId="4"captionText="FIGURE 2. Libanobelostoma calineae gen et sp. nov., holotype I-26247/2A (part) and B (counterpart), general habitus. A–B, Dorsum under natural light (A) and under alcohol (B). C–D, Venter under natural light (C) and under alcohol (D). Scale bars = 1 cm."figureDoi="http://doi.org/10.5281/zenodo.14595712"httpUri="https://zenodo.org/record/14595712/files/figure.png"pageId="6"pageNumber="17">Fig. 2C–D</figureCitation>
), metacoxa conical, firmly united with metapleuron, femur slightly enlarged at base and narrower at apex, tibia apparently flattened, bearing few poorly preserved setae (
<figureCitationid="3AC0C8CB8E41FF83FB9E2F8BFB4EFD6D"box="[1130,1217,727,753]"captionStart="FIGURE 2"captionStartId="4.[151,251,1943,1967]"captionTargetBox="[165,1422,179,1917]"captionTargetId="figure-41@4.[165,1422,179,1917]"captionTargetPageId="4"captionText="FIGURE 2. Libanobelostoma calineae gen et sp. nov., holotype I-26247/2A (part) and B (counterpart), general habitus. A–B, Dorsum under natural light (A) and under alcohol (B). C–D, Venter under natural light (C) and under alcohol (D). Scale bars = 1 cm."figureDoi="http://doi.org/10.5281/zenodo.14595712"httpUri="https://zenodo.org/record/14595712/files/figure.png"pageId="6"pageNumber="17">Fig. 2F</figureCitation>
<figureCitationid="3AC0C8CB8E41FF83FEA42E43FE2CFCA5"box="[336,419,799,825]"captionStart="FIGURE 2"captionStartId="4.[151,251,1943,1967]"captionTargetBox="[165,1422,179,1917]"captionTargetId="figure-41@4.[165,1422,179,1917]"captionTargetPageId="4"captionText="FIGURE 2. Libanobelostoma calineae gen et sp. nov., holotype I-26247/2A (part) and B (counterpart), general habitus. A–B, Dorsum under natural light (A) and under alcohol (B). C–D, Venter under natural light (C) and under alcohol (D). Scale bars = 1 cm."figureDoi="http://doi.org/10.5281/zenodo.14595712"httpUri="https://zenodo.org/record/14595712/files/figure.png"pageId="6"pageNumber="17">Fig. 2E</figureCitation>
