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<document id="B2ED7A6489B0CE7DB365F4283D1EFE55" ID-CLB-Dataset="26555" ID-DOI="10.11646/zootaxa.4614.1.6" ID-GBIF-Dataset="25e4c643-3a35-4135-9809-58f25fcca05c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3241194" ID-ZooBank="7F5EAB98-0D41-4C8C-AA0B-1663E39EDFB2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1559897275064" checkinUser="plazi" docAuthor="Németh, Tamás" docDate="2019" docId="1E2F8784737DFFF1FF3547F3FB294C9C" docLanguage="en" docName="zootaxa.4614.1.6.pdf" docOrigin="Zootaxa 4614 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megathous tannourinensis Németh 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="140" masterDocId="E216FFFC7370FFFCFFA2476AFFAA4B0B" masterDocTitle="Elateridae (Coleoptera) of Lebanon" masterLastPageNumber="159" masterPageNumber="127" pageNumber="140" updateTime="1698733648462" updateUser="ExternalLinkService">
<mods:mods id="5D837A2FFD50471E42FBEC3574346185" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A0A75BAB35932A93B7B6A6CE912C159D">
<mods:title id="AF6F348D35990A46E9A0A6339EB20D99">Elateridae (Coleoptera) of Lebanon</mods:title>
</mods:titleInfo>
<mods:name id="FCC854A3BC83F51ABA3F7B3263F4D6D4" type="personal">
<mods:role id="77BDDC0FF51EADB0E97527EB4E25FE2A">
<mods:roleTerm id="104F5E8509B03113654BF6002EF0E0A6">Author</mods:roleTerm>
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<mods:namePart id="836FBDEAB3BCC41113FEEEAA69200EFB">Németh, Tamás</mods:namePart>
</mods:name>
<mods:typeOfResource id="D81490F75C9C91A2F6CBB1813626436C">text</mods:typeOfResource>
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<mods:title id="B3F203539138A082AB5880082A16D624">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="43190CC7D822BA57A78ABF1CCF426F74">
<mods:date id="5F5E053BB08CC76AB585DDE0353CC652">2019</mods:date>
<mods:detail id="B0267B2A5033166E3970B47A63A95982" type="pubDate">
<mods:number id="DC61AE5EAA978981B367FBDAA848BE44">2019-06-07</mods:number>
</mods:detail>
<mods:detail id="4B405B76D1F0834B29D4A325C6EA47FB" type="volume">
<mods:number id="F18FA26132007A8045B0CFE8D1EE2999">4614</mods:number>
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<mods:number id="59F0ACF4E2CBF537545A8CB3AFE8FC0C">1</mods:number>
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<mods:classification id="E88E3CEAA325FF7F1C5EC9E958439C40">journal article</mods:classification>
<mods:identifier id="C462833639AC788F4653ED9C8CE9DF60" type="CLB-Dataset">26555</mods:identifier>
<mods:identifier id="CAD3353086F888E2F0397CF4DB6439D7" type="DOI">10.11646/zootaxa.4614.1.6</mods:identifier>
<mods:identifier id="04BA097D14AE8CDAD75F9EF8D4A95872" type="GBIF-Dataset">25e4c643-3a35-4135-9809-58f25fcca05c</mods:identifier>
<mods:identifier id="FCC970A8E0532459CE2BA45B1E583346" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B96B39EBCFD90166707D0BE43894F267" type="Zenodo-Dep">3241194</mods:identifier>
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</mods:mods>
<treatment id="1E2F8784737DFFF1FF3547F3FB294C9C" ID-DOI="http://doi.org/10.5281/zenodo.5935952" ID-GBIF-Taxon="157436076" ID-Zenodo-Dep="5935952" LSID="urn:lsid:plazi:treatment:1E2F8784737DFFF1FF3547F3FB294C9C" httpUri="http://treatment.plazi.org/id/1E2F8784737DFFF1FF3547F3FB294C9C" lastPageNumber="140" pageId="13" pageNumber="140">
<subSubSection id="DE9C6519737DFFF1FF3547F3FDEB4BBF" box="[151,577,153,180]" pageId="13" pageNumber="140" type="nomenclature">
<paragraph id="96393692737DFFF1FF3547F3FDEB4BBF" blockId="13.[151,577,153,215]" box="[151,577,153,180]" pageId="13" pageNumber="140">
<heading id="CD7181FE737DFFF1FF3547F3FDEB4BBF" bold="true" box="[151,577,153,180]" fontSize="11" level="1" pageId="13" pageNumber="140" reason="1">
<emphasis id="A4F2EA80737DFFF1FF3547F3FDEB4BBF" bold="true" box="[151,577,153,180]" pageId="13" pageNumber="140">
<taxonomicName id="51864D11737DFFF1FF3547F3FE774BB8" authority="Németh, 2019" authorityName="Németh" authorityYear="2019" box="[151,477,153,180]" class="Insecta" family="Elateridae" genus="Megathous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="tannourinensis" status="sp. nov.">
<emphasis id="A4F2EA80737DFFF1FF3547F3FE774BB8" bold="true" box="[151,477,153,180]" italics="true" pageId="13" pageNumber="140">Megathous tannourinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFC157FB737DFFF1FE4647F0FDEB4BBF" box="[484,577,154,180]" pageId="13" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE9C6519737DFFF1FF3547D7FE0F4BDC" box="[151,421,189,215]" pageId="13" pageNumber="140" type="description">
<paragraph id="96393692737DFFF1FF3547D7FE0F4BDC" blockId="13.[151,577,153,215]" box="[151,421,189,215]" pageId="13" pageNumber="140">
(
<figureCitation id="0EBD2A17737DFFF1FF3D47D7FE8A4BDC" box="[159,288,189,215]" captionStart="FIGURES 31–36" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[227,1326,185,1911]" captionTargetId="figure@14.[190,1397,174,1918]" captionTargetPageId="14" captionText="FIGURES 31–36. Habitus. Megathous tannourinensis sp. nov., male, holotype, 31—dorsal view, 34—lateral view; M. lizlerwerneri Mertlik, 2005, male, holotype, 32—dorsal view, 35—lateral view; M. menetriesi (Reitter, 1890), male, holotype, 33—dorsal view, 36—lateral view. Scale bars=2.0 mm." httpUri="https://zenodo.org/record/3241202/files/figure.png" pageId="13" pageNumber="140">Figs 31, 34</figureCitation>
,
<figureCitation id="0EBD2A17737DFFF1FE8E47D7FE374BDC" box="[300,413,189,215]" captionStart="FIGURES 37–45" captionStartId="15.[151,264,1326,1351]" captionTargetBox="[151,1434,186,1297]" captionTargetId="figure@15.[151,1436,181,1306]" captionTargetPageId="15" captionText="FIGURES 37–45. Antennomeres. 37—M. tannourinensis sp. nov., male, holotype; 38—M. lizlerwerneri Mertlik, 2005, male, holotype; 39—M. menetriesi (Reitter, 1890), male, holotype. Prosternal process, lateral view. 40—M. tannourinensis sp. nov., male, holotype; 41—M. lizlerwerneri Mertlik, 2005, male, holotype; 42—M. menetriesi (Reitter, 1890), male, holotype.Aedeagus. 43—M. tannourinensis sp. nov., male, holotype. 44—M. lizlerwerneri Mertlik, 2005 male, holotype. 45—M. menetriesi (Reitter, 1890), male, holotype. Scale bars=0.5 mm (Figs 43–45), 1.0 mm (Figs 37–42)." httpUri="https://zenodo.org/record/3241206/files/figure.png" pageId="13" pageNumber="140">37, 40, 43</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DE9C6519737DFFF1FF35466CFAEB4A63" pageId="13" pageNumber="140" type="materials_examined">
<paragraph id="96393692737DFFF1FF35466CFAEB4A63" blockId="13.[151,1437,261,1944]" pageId="13" pageNumber="140">
<emphasis id="A4F2EA80737DFFF1FF35466CFEEF4A2B" bold="true" box="[151,325,261,288]" pageId="13" pageNumber="140">Type material.</emphasis>
<materialsCitation id="26EE3CCF737DFFF1FEE9466FFDD54A63" ID-GBIF-Occurrence="2248479082" collectingDate="2017-05-01" collectingDateMax="2017-06-19" collectingDateMin="2017-05-01" collectionCode="HNHM" collectorName="A. Kotan &amp; P. Nemes &amp; T. Nemeth" country="Lebanon" county="Tannourine Cedars Nat. Reserve" elevation="1750" latitude="34.209446" location="Northern" longLatPrecision="19" longitude="35.92917" municipality="Tannourine" pageId="13" pageNumber="140" specimenCount="1" typeStatus="holotype">
<typeStatus id="493D8830737DFFF1FEE9466FFE1C4A14" box="[331,438,261,287]" pageId="13" pageNumber="140" type="holotype">Holotype</typeStatus>
, male. “
<collectingCountry id="EE917602737DFFF1FDAC466FFD304A14" box="[526,666,261,287]" name="Lebanon" pageId="13" pageNumber="140">LEBANON</collectingCountry>
,
<location id="93596049737DFFF1FD07466FFCA04A14" LSID="urn:lsid:plazi:treatment:1E2F8784737DFFF1FF3547F3FB294C9C:93596049737DFFF1FD07466FFCA04A14" box="[677,778,261,287]" country="Lebanon" county="Tannourine Cedars Nat. Reserve" latitude="34.209446" longLatPrecision="19" longitude="35.92917" municipality="Tannourine" name="Northern" pageId="13" pageNumber="140">Northern</location>
gov.,
<collectingMunicipality id="765DACE8737DFFF1FCEF466FFC664A14" box="[845,972,261,287]" pageId="13" pageNumber="140">Tannourine</collectingMunicipality>
env.,
<locationDeviation id="878B2FB0737DFFF1FBAC466FFB154A14" box="[1038,1215,261,288]" location="Harissa" pageId="13" pageNumber="140">
<quantity id="517E9B77737DFFF1FBAC466FFBEC4A14" box="[1038,1094,261,287]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="140" unit="km" value="2.0">2 km</quantity>
N Harissa
</locationDeviation>
,
<collectingCounty id="7F584E1E737DFFF1FB6B466FFE864A4F" pageId="13" pageNumber="140">Tannourine Cedars Nat. Reserve</collectingCounty>
, reared from hollow
<taxonomicName id="51864D11737DFFF1FDBE4640FDD04A4F" box="[540,634,298,324]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="140" phylum="Tracheophyta" rank="genus">
<emphasis id="A4F2EA80737DFFF1FDBE4640FDD04A4F" box="[540,634,298,324]" italics="true" pageId="13" pageNumber="140">Quercus</emphasis>
</taxonomicName>
,
<geoCoordinate id="F3B25055737DFFF1FD2B4643FCBE4A48" box="[649,788,297,323]" degrees="34" direction="north" minutes="12" orientation="latitude" pageId="13" pageNumber="140" precision="15" seconds="34" value="34.209446">34°12’34”N</geoCoordinate>
,
<geoCoordinate id="F3B25055737DFFF1FC804643FC004A4F" box="[802,938,297,324]" degrees="35" direction="east" minutes="55" orientation="longitude" pageId="13" pageNumber="140" precision="15" seconds="45" value="35.92917">35°55’45”E</geoCoordinate>
,
<elevation id="1DABD1A1737DFFF1FC154643FBA54A4F" box="[951,1039,297,324]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="13" pageNumber="140" unit="m" value="1750.0">
<quantity id="517E9B77737DFFF1FC154643FBA54A4F" box="[951,1039,297,324]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="13" pageNumber="140" unit="m" value="1750.0">1750 m</quantity>
</elevation>
,
<collectingDate id="F27CE9BA737DFFF1FBBE4643FB7E4A48" box="[1052,1236,297,323]" pageId="13" pageNumber="140" value="2017-05-01" valueMax="2017-06-19" valueMin="2017-05-01">
<date id="E2381052737DFFF1FBBE4643FB7E4A48" box="[1052,1236,297,323]" pageId="13" pageNumber="140" value="2017-05-01" valueMax="2017-06-19" valueMin="2017-05-01">1.V.-19.VI.2017</date>
</collectingDate>
, leg.
<collectorName id="3B735344737DFFF1FAB34640FAD64A48" box="[1297,1404,297,324]" pageId="13" pageNumber="140">A. Kotán</collectorName>
,
<collectorName id="3B735344737DFFF1FA2B4640FF4F4A63" pageId="13" pageNumber="140">P. Nemes</collectorName>
&amp;
<collectorName id="3B735344737DFFF1FEAA4624FED44A6C" box="[264,382,333,360]" pageId="13" pageNumber="140">T. Németh</collectorName>
(No. 30)” (1,
<collectionCode id="F097AE57737DFFF1FDBF4624FDD24A63" box="[541,632,334,360]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="13" pageNumber="140">HNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="26EE3CCF737DFFF1FD2B4624FA974A63" ID-GBIF-Occurrence="2248479066" box="[649,1341,333,360]" collectingDate="2017-05-01" collectingDateMax="2017-06-19" collectingDateMin="2017-05-01" collectionCode="HNHM" collectorName="A. Kotan &amp; P. Nemes &amp; T. Nemeth" country="Lebanon" county="Tannourine Cedars Nat. Reserve" elevation="1750" latitude="34.209446" location="Northern" longLatPrecision="19" longitude="35.92917" municipality="Tannourine" pageId="13" pageNumber="140" specimenCount="1" typeStatus="paratype">
<typeStatus id="493D8830737DFFF1FD2B4624FD444A63" box="[649,750,334,360]" pageId="13" pageNumber="140" type="paratype">Paratype</typeStatus>
, male, the same data as for the holotype (1,
<collectionCode id="F097AE57737DFFF1FB794624FA9C4A63" box="[1243,1334,334,360]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="13" pageNumber="140">HNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DE9C6519737DFFF1FF65461BFC0F4854" pageId="13" pageNumber="140" type="diagnosis">
<paragraph id="96393692737DFFF1FF65461BFC684AA4" blockId="13.[151,1437,261,1944]" pageId="13" pageNumber="140">
<emphasis id="A4F2EA80737DFFF1FF65461BFEE84A87" bold="true" box="[199,322,369,396]" pageId="13" pageNumber="140">Diagnosis.</emphasis>
<taxonomicName id="51864D11737DFFF1FEE8461BFDDF4A80" authority="Németh, 2019" authorityName="Németh" authorityYear="2019" box="[330,629,369,395]" class="Insecta" family="Elateridae" genus="Megathous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="tannourinensis" status="sp. nov.">
<emphasis id="A4F2EA80737DFFF1FEE8461BFDDF4A80" box="[330,629,369,395]" italics="true" pageId="13" pageNumber="140">Megathous tannourinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFC157FB737DFFF1FDDC4618FD704A87" box="[638,730,370,396]" pageId="13" pageNumber="140" rank="species">
<emphasis id="A4F2EA80737DFFF1FDDC4618FD704A87" bold="true" box="[638,730,370,396]" pageId="13" pageNumber="140">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from the related species of the region by its pronotum shape, body colorisation, shape of prosternal process and male genitalia.
</paragraph>
<paragraph id="96393692737DFFF1FF6546D3FC0F4854" blockId="13.[151,1437,261,1944]" pageId="13" pageNumber="140">
<emphasis id="A4F2EA80737DFFF1FF6546D3FDE24ADF" bold="true" box="[199,584,441,468]" pageId="13" pageNumber="140">Comparison with other species.</emphasis>
<taxonomicName id="51864D11737DFFF1FDF146D3FC2A4AD8" authority="Németh, 2019" authorityName="Németh" authorityYear="2019" box="[595,896,441,467]" class="Insecta" family="Elateridae" genus="Megathous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="tannourinensis" status="sp. nov.">
<emphasis id="A4F2EA80737DFFF1FDF146D3FC2A4AD8" box="[595,896,441,467]" italics="true" pageId="13" pageNumber="140">Megathous tannourinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFC157FB737DFFF1FC2946D0FC424ADF" box="[907,1000,442,468]" pageId="13" pageNumber="140" rank="species">
<emphasis id="A4F2EA80737DFFF1FC2946D0FC424ADF" bold="true" box="[907,1000,442,468]" pageId="13" pageNumber="140">sp. nov.</emphasis>
</taxonomicNameLabel>
shares body size and coloration with
<taxonomicName id="51864D11737DFFF1FF3546B7FDE74AF3" authority="Mertlik, 2005" authorityName="Mertlik" authorityYear="2005" box="[151,589,477,504]" class="Insecta" family="Elateridae" genus="Megathous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="lizlerwerneri">
<emphasis id="A4F2EA80737DFFF1FF3546B7FE064AFC" box="[151,428,477,503]" italics="true" pageId="13" pageNumber="140">Megathous lizlerwerneri</emphasis>
<bibRefCitation id="F2174B63737DFFF1FE1146B7FDE74AF3" author="Mertlik, J." box="[435,589,477,504]" pageId="13" pageNumber="140" pagination="167 - 173" refId="ref21270" refString="Mertlik, J. (2005) Description of three new species of family Elateridae (Coleoptera) from Syria and Turkey. Folia Heyrovskyana, 12, 167 - 173" type="journal article" year="2005">Mertlik, 2005</bibRefCitation>
</taxonomicName>
(
<figureCitation id="0EBD2A17737DFFF1FDFE46B7FD774AF3" box="[604,733,477,504]" captionStart="FIGURES 31–36" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[227,1326,185,1911]" captionTargetId="figure@14.[190,1397,174,1918]" captionTargetPageId="14" captionText="FIGURES 31–36. Habitus. Megathous tannourinensis sp. nov., male, holotype, 31—dorsal view, 34—lateral view; M. lizlerwerneri Mertlik, 2005, male, holotype, 32—dorsal view, 35—lateral view; M. menetriesi (Reitter, 1890), male, holotype, 33—dorsal view, 36—lateral view. Scale bars=2.0 mm." httpUri="https://zenodo.org/record/3241202/files/figure.png" pageId="13" pageNumber="140">Figs 32, 35</figureCitation>
,
<figureCitation id="0EBD2A17737DFFF1FD4B46B7FCF04AFC" box="[745,858,477,503]" captionStart="FIGURES 37–45" captionStartId="15.[151,264,1326,1351]" captionTargetBox="[151,1434,186,1297]" captionTargetId="figure@15.[151,1436,181,1306]" captionTargetPageId="15" captionText="FIGURES 37–45. Antennomeres. 37—M. tannourinensis sp. nov., male, holotype; 38—M. lizlerwerneri Mertlik, 2005, male, holotype; 39—M. menetriesi (Reitter, 1890), male, holotype. Prosternal process, lateral view. 40—M. tannourinensis sp. nov., male, holotype; 41—M. lizlerwerneri Mertlik, 2005, male, holotype; 42—M. menetriesi (Reitter, 1890), male, holotype.Aedeagus. 43—M. tannourinensis sp. nov., male, holotype. 44—M. lizlerwerneri Mertlik, 2005 male, holotype. 45—M. menetriesi (Reitter, 1890), male, holotype. Scale bars=0.5 mm (Figs 43–45), 1.0 mm (Figs 37–42)." httpUri="https://zenodo.org/record/3241206/files/figure.png" pageId="13" pageNumber="140">38, 41, 44</figureCitation>
) from
<collectingCountry id="EE917602737DFFF1FC0546B7FC4F4AFC" box="[935,997,477,503]" name="Syria" pageId="13" pageNumber="140">Syria</collectingCountry>
, but the new species differs in pronotal shape, antennal length, prosternal process and shape of parameres.
<taxonomicName id="51864D11737DFFF1FC384568FBF84910" authorityName="Mertlik" authorityYear="2005" box="[922,1106,513,539]" class="Insecta" family="Elateridae" genus="Megathous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="lizlerwerneri">
<emphasis id="A4F2EA80737DFFF1FC384568FBF84910" box="[922,1106,513,539]" italics="true" pageId="13" pageNumber="140">M. lizlerwerneri</emphasis>
</taxonomicName>
has flattened pronotum with elongated and strongly divergent posterior angles, its antennae not reaching the posterior angles (
<figureCitation id="0EBD2A17737DFFF1FB46454FFAC2494B" box="[1252,1384,549,576]" captionStart="FIGURES 31–36" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[227,1326,185,1911]" captionTargetId="figure@14.[190,1397,174,1918]" captionTargetPageId="14" captionText="FIGURES 31–36. Habitus. Megathous tannourinensis sp. nov., male, holotype, 31—dorsal view, 34—lateral view; M. lizlerwerneri Mertlik, 2005, male, holotype, 32—dorsal view, 35—lateral view; M. menetriesi (Reitter, 1890), male, holotype, 33—dorsal view, 36—lateral view. Scale bars=2.0 mm." httpUri="https://zenodo.org/record/3241202/files/figure.png" pageId="13" pageNumber="140">Figs 32, 35</figureCitation>
).
<taxonomicName id="51864D11737DFFF1FADC454CFE974968" authorityName="Németh" authorityYear="2019" class="Insecta" family="Elateridae" genus="Megathous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="tannourinensis">
<emphasis id="A4F2EA80737DFFF1FADC454CFE974968" italics="true" pageId="13" pageNumber="140">M. tannourinensis</emphasis>
</taxonomicName>
has pronotum more convex, with shorter and only slightly divergent posterior angles, antennae overreaching posterior angles by about two antennomeres (
<figureCitation id="0EBD2A17737DFFF1FCA14507FC2D498C" box="[771,903,621,647]" captionStart="FIGURES 31–36" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[227,1326,185,1911]" captionTargetId="figure@14.[190,1397,174,1918]" captionTargetPageId="14" captionText="FIGURES 31–36. Habitus. Megathous tannourinensis sp. nov., male, holotype, 31—dorsal view, 34—lateral view; M. lizlerwerneri Mertlik, 2005, male, holotype, 32—dorsal view, 35—lateral view; M. menetriesi (Reitter, 1890), male, holotype, 33—dorsal view, 36—lateral view. Scale bars=2.0 mm." httpUri="https://zenodo.org/record/3241202/files/figure.png" pageId="13" pageNumber="140">Figs 31, 34</figureCitation>
). Lower part of prosternal process of
<taxonomicName id="51864D11737DFFF1FA964504FF4449A0" authorityName="Mertlik" authorityYear="2005" class="Insecta" family="Elateridae" genus="Megathous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="lizlerwerneri">
<emphasis id="A4F2EA80737DFFF1FA964504FF4449A0" italics="true" pageId="13" pageNumber="140">M. lizlerwerneri</emphasis>
</taxonomicName>
almost straight in lateral view, emerges from its two-third (
<figureCitation id="0EBD2A17737DFFF1FC3645FBFC4349A0" box="[916,1001,657,683]" captionStart="FIGURES 37–45" captionStartId="15.[151,264,1326,1351]" captionTargetBox="[151,1434,186,1297]" captionTargetId="figure@15.[151,1436,181,1306]" captionTargetPageId="15" captionText="FIGURES 37–45. Antennomeres. 37—M. tannourinensis sp. nov., male, holotype; 38—M. lizlerwerneri Mertlik, 2005, male, holotype; 39—M. menetriesi (Reitter, 1890), male, holotype. Prosternal process, lateral view. 40—M. tannourinensis sp. nov., male, holotype; 41—M. lizlerwerneri Mertlik, 2005, male, holotype; 42—M. menetriesi (Reitter, 1890), male, holotype.Aedeagus. 43—M. tannourinensis sp. nov., male, holotype. 44—M. lizlerwerneri Mertlik, 2005 male, holotype. 45—M. menetriesi (Reitter, 1890), male, holotype. Scale bars=0.5 mm (Figs 43–45), 1.0 mm (Figs 37–42)." httpUri="https://zenodo.org/record/3241206/files/figure.png" pageId="13" pageNumber="140">Fig. 41</figureCitation>
), while the new species has prosternal process emerging from its half in lateral view (
<figureCitation id="0EBD2A17737DFFF1FD0045DFFD5C49C4" box="[674,758,693,719]" captionStart="FIGURES 37–45" captionStartId="15.[151,264,1326,1351]" captionTargetBox="[151,1434,186,1297]" captionTargetId="figure@15.[151,1436,181,1306]" captionTargetPageId="15" captionText="FIGURES 37–45. Antennomeres. 37—M. tannourinensis sp. nov., male, holotype; 38—M. lizlerwerneri Mertlik, 2005, male, holotype; 39—M. menetriesi (Reitter, 1890), male, holotype. Prosternal process, lateral view. 40—M. tannourinensis sp. nov., male, holotype; 41—M. lizlerwerneri Mertlik, 2005, male, holotype; 42—M. menetriesi (Reitter, 1890), male, holotype.Aedeagus. 43—M. tannourinensis sp. nov., male, holotype. 44—M. lizlerwerneri Mertlik, 2005 male, holotype. 45—M. menetriesi (Reitter, 1890), male, holotype. Scale bars=0.5 mm (Figs 43–45), 1.0 mm (Figs 37–42)." httpUri="https://zenodo.org/record/3241206/files/figure.png" pageId="13" pageNumber="140">Fig. 40</figureCitation>
). Apex of parameres of aedeagus are narrow and sharp in
<taxonomicName id="51864D11737DFFF1FADC45DCFE8249F8" authorityName="Mertlik" authorityYear="2005" class="Insecta" family="Elateridae" genus="Megathous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="lizlerwerneri">
<emphasis id="A4F2EA80737DFFF1FADC45DCFE8249F8" italics="true" pageId="13" pageNumber="140">M. lizlerwerneri</emphasis>
</taxonomicName>
(
<figureCitation id="0EBD2A17737DFFF1FE9B45B3FE2449F8" box="[313,398,729,755]" captionStart="FIGURES 37–45" captionStartId="15.[151,264,1326,1351]" captionTargetBox="[151,1434,186,1297]" captionTargetId="figure@15.[151,1436,181,1306]" captionTargetPageId="15" captionText="FIGURES 37–45. Antennomeres. 37—M. tannourinensis sp. nov., male, holotype; 38—M. lizlerwerneri Mertlik, 2005, male, holotype; 39—M. menetriesi (Reitter, 1890), male, holotype. Prosternal process, lateral view. 40—M. tannourinensis sp. nov., male, holotype; 41—M. lizlerwerneri Mertlik, 2005, male, holotype; 42—M. menetriesi (Reitter, 1890), male, holotype.Aedeagus. 43—M. tannourinensis sp. nov., male, holotype. 44—M. lizlerwerneri Mertlik, 2005 male, holotype. 45—M. menetriesi (Reitter, 1890), male, holotype. Scale bars=0.5 mm (Figs 43–45), 1.0 mm (Figs 37–42)." httpUri="https://zenodo.org/record/3241206/files/figure.png" pageId="13" pageNumber="140">Fig. 44</figureCitation>
), and widely rounded in
<taxonomicName id="51864D11737DFFF1FD0E45B0FCD349F8" authorityName="Németh" authorityYear="2019" box="[684,889,730,755]" class="Insecta" family="Elateridae" genus="Megathous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="tannourinensis">
<emphasis id="A4F2EA80737DFFF1FD0E45B0FCD349F8" box="[684,889,730,755]" italics="true" pageId="13" pageNumber="140">M. tannourinensis</emphasis>
</taxonomicName>
(
<figureCitation id="0EBD2A17737DFFF1FC2845B3FC4A49F8" box="[906,992,729,755]" captionStart="FIGURES 37–45" captionStartId="15.[151,264,1326,1351]" captionTargetBox="[151,1434,186,1297]" captionTargetId="figure@15.[151,1436,181,1306]" captionTargetPageId="15" captionText="FIGURES 37–45. Antennomeres. 37—M. tannourinensis sp. nov., male, holotype; 38—M. lizlerwerneri Mertlik, 2005, male, holotype; 39—M. menetriesi (Reitter, 1890), male, holotype. Prosternal process, lateral view. 40—M. tannourinensis sp. nov., male, holotype; 41—M. lizlerwerneri Mertlik, 2005, male, holotype; 42—M. menetriesi (Reitter, 1890), male, holotype.Aedeagus. 43—M. tannourinensis sp. nov., male, holotype. 44—M. lizlerwerneri Mertlik, 2005 male, holotype. 45—M. menetriesi (Reitter, 1890), male, holotype. Scale bars=0.5 mm (Figs 43–45), 1.0 mm (Figs 37–42)." httpUri="https://zenodo.org/record/3241206/files/figure.png" pageId="13" pageNumber="140">Fig. 43</figureCitation>
).
<taxonomicName id="51864D11737DFFF1FC5745B3FA3649F8" authority="(Reitter, 1890)" baseAuthorityName="Reitter" baseAuthorityYear="1890" box="[1013,1436,729,755]" class="Insecta" family="Elateridae" genus="Megathous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="menetriesi">
<emphasis id="A4F2EA80737DFFF1FC5745B3FB4549F8" box="[1013,1263,729,755]" italics="true" pageId="13" pageNumber="140">Megathous menetriesi</emphasis>
(
<bibRefCitation id="F2174B63737DFFF1FB5C45B3FA3E49F8" author="Reitter, E." box="[1278,1428,729,755]" pageId="13" pageNumber="140" pagination="165 - 176" refId="ref22892" refString="Reitter, E. (1890) Neue Coleopteren aus Europa, den angrenzenden Landern und Sibirien, mit Bemerkungen uber bekannte Arten. Zehnter Theil. Deutsche Entomologische Zeitschrift, 34, 165 - 176. https: // doi. org / 10.1002 / mmnd. 48018900324" type="journal article" year="1890">Reitter, 1890</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="0EBD2A17737DFFF1FF3D4597FE8B481C" box="[159,289,765,791]" captionStart="FIGURES 31–36" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[227,1326,185,1911]" captionTargetId="figure@14.[190,1397,174,1918]" captionTargetPageId="14" captionText="FIGURES 31–36. Habitus. Megathous tannourinensis sp. nov., male, holotype, 31—dorsal view, 34—lateral view; M. lizlerwerneri Mertlik, 2005, male, holotype, 32—dorsal view, 35—lateral view; M. menetriesi (Reitter, 1890), male, holotype, 33—dorsal view, 36—lateral view. Scale bars=2.0 mm." httpUri="https://zenodo.org/record/3241202/files/figure.png" pageId="13" pageNumber="140">Figs 33, 36</figureCitation>
,
<figureCitation id="0EBD2A17737DFFF1FE8F4597FE0B4813" box="[301,417,765,792]" captionStart="FIGURES 37–45" captionStartId="15.[151,264,1326,1351]" captionTargetBox="[151,1434,186,1297]" captionTargetId="figure@15.[151,1436,181,1306]" captionTargetPageId="15" captionText="FIGURES 37–45. Antennomeres. 37—M. tannourinensis sp. nov., male, holotype; 38—M. lizlerwerneri Mertlik, 2005, male, holotype; 39—M. menetriesi (Reitter, 1890), male, holotype. Prosternal process, lateral view. 40—M. tannourinensis sp. nov., male, holotype; 41—M. lizlerwerneri Mertlik, 2005, male, holotype; 42—M. menetriesi (Reitter, 1890), male, holotype.Aedeagus. 43—M. tannourinensis sp. nov., male, holotype. 44—M. lizlerwerneri Mertlik, 2005 male, holotype. 45—M. menetriesi (Reitter, 1890), male, holotype. Scale bars=0.5 mm (Figs 43–45), 1.0 mm (Figs 37–42)." httpUri="https://zenodo.org/record/3241206/files/figure.png" pageId="13" pageNumber="140">39, 42, 45</figureCitation>
), occurs in
<collectingCountry id="EE917602737DFFF1FD814597FD34481C" box="[547,670,765,791]" name="Azerbaijan" pageId="13" pageNumber="140">Azerbaijan</collectingCountry>
and
<collectingCountry id="EE917602737DFFF1FD774594FCA84813" box="[725,770,766,792]" name="Iran" pageId="13" pageNumber="140">Iran</collectingCountry>
(
<bibRefCitation id="F2174B63737DFFF1FCB34597FC20481C" author="Cate, P." box="[785,906,765,791]" pageId="13" pageNumber="140" pagination="89 - 209" refId="ref18761" refString="Cate, P. (2007) Elateridae. In: Lobl, I. &amp; Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 89 - 209." type="book chapter" year="2007">Cate 2007</bibRefCitation>
), has elytra 3 times longer than pronotum (
<figureCitation id="0EBD2A17737DFFF1FACD4597FF1E4830" captionStart="FIGURES 31–36" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[227,1326,185,1911]" captionTargetId="figure@14.[190,1397,174,1918]" captionTargetPageId="14" captionText="FIGURES 31–36. Habitus. Megathous tannourinensis sp. nov., male, holotype, 31—dorsal view, 34—lateral view; M. lizlerwerneri Mertlik, 2005, male, holotype, 32—dorsal view, 35—lateral view; M. menetriesi (Reitter, 1890), male, holotype, 33—dorsal view, 36—lateral view. Scale bars=2.0 mm." httpUri="https://zenodo.org/record/3241202/files/figure.png" pageId="13" pageNumber="140">Fig. 33</figureCitation>
) and slender antennomeres (
<figureCitation id="0EBD2A17737DFFF1FE5B444BFDE44830" box="[505,590,801,827]" captionStart="FIGURES 37–45" captionStartId="15.[151,264,1326,1351]" captionTargetBox="[151,1434,186,1297]" captionTargetId="figure@15.[151,1436,181,1306]" captionTargetPageId="15" captionText="FIGURES 37–45. Antennomeres. 37—M. tannourinensis sp. nov., male, holotype; 38—M. lizlerwerneri Mertlik, 2005, male, holotype; 39—M. menetriesi (Reitter, 1890), male, holotype. Prosternal process, lateral view. 40—M. tannourinensis sp. nov., male, holotype; 41—M. lizlerwerneri Mertlik, 2005, male, holotype; 42—M. menetriesi (Reitter, 1890), male, holotype.Aedeagus. 43—M. tannourinensis sp. nov., male, holotype. 44—M. lizlerwerneri Mertlik, 2005 male, holotype. 45—M. menetriesi (Reitter, 1890), male, holotype. Scale bars=0.5 mm (Figs 43–45), 1.0 mm (Figs 37–42)." httpUri="https://zenodo.org/record/3241206/files/figure.png" pageId="13" pageNumber="140">Fig. 39</figureCitation>
).
<taxonomicName id="51864D11737DFFF1FDC0444BFC264830" authority="Németh, 2019" authorityName="Németh" authorityYear="2019" box="[610,908,801,827]" class="Insecta" family="Elateridae" genus="Megathous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="tannourinensis" status="sp. nov.">
<emphasis id="A4F2EA80737DFFF1FDC0444BFC264830" box="[610,908,801,827]" italics="true" pageId="13" pageNumber="140">Megathous tannourinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFC157FB737DFFF1FC364448FC444837" box="[916,1006,802,828]" pageId="13" pageNumber="140" rank="species">
<emphasis id="A4F2EA80737DFFF1FC364448FC444837" bold="true" box="[916,1006,802,828]" pageId="13" pageNumber="140">sp. nov.</emphasis>
</taxonomicNameLabel>
has elytra only up to 2.7 times longer than pronotum (
<figureCitation id="0EBD2A17737DFFF1FEE9442FFE374854" box="[331,413,837,863]" captionStart="FIGURES 31–36" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[227,1326,185,1911]" captionTargetId="figure@14.[190,1397,174,1918]" captionTargetPageId="14" captionText="FIGURES 31–36. Habitus. Megathous tannourinensis sp. nov., male, holotype, 31—dorsal view, 34—lateral view; M. lizlerwerneri Mertlik, 2005, male, holotype, 32—dorsal view, 35—lateral view; M. menetriesi (Reitter, 1890), male, holotype, 33—dorsal view, 36—lateral view. Scale bars=2.0 mm." httpUri="https://zenodo.org/record/3241202/files/figure.png" pageId="13" pageNumber="140">Fig. 31</figureCitation>
), and antennomeres relatively wider (
<figureCitation id="0EBD2A17737DFFF1FCE6442FFC324854" box="[836,920,837,863]" captionStart="FIGURES 37–45" captionStartId="15.[151,264,1326,1351]" captionTargetBox="[151,1434,186,1297]" captionTargetId="figure@15.[151,1436,181,1306]" captionTargetPageId="15" captionText="FIGURES 37–45. Antennomeres. 37—M. tannourinensis sp. nov., male, holotype; 38—M. lizlerwerneri Mertlik, 2005, male, holotype; 39—M. menetriesi (Reitter, 1890), male, holotype. Prosternal process, lateral view. 40—M. tannourinensis sp. nov., male, holotype; 41—M. lizlerwerneri Mertlik, 2005, male, holotype; 42—M. menetriesi (Reitter, 1890), male, holotype.Aedeagus. 43—M. tannourinensis sp. nov., male, holotype. 44—M. lizlerwerneri Mertlik, 2005 male, holotype. 45—M. menetriesi (Reitter, 1890), male, holotype. Scale bars=0.5 mm (Figs 43–45), 1.0 mm (Figs 37–42)." httpUri="https://zenodo.org/record/3241206/files/figure.png" pageId="13" pageNumber="140">Fig. 37</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DE9C6519737DFFF1FF654403FE344D90" pageId="13" pageNumber="140" type="description">
<paragraph id="96393692737DFFF1FF654403FCD94FAF" blockId="13.[151,1437,261,1944]" pageId="13" pageNumber="140">
<emphasis id="A4F2EA80737DFFF1FF654403FEF3488F" bold="true" box="[199,345,873,900]" pageId="13" pageNumber="140">Description.</emphasis>
<typeStatus id="493D8830737DFFF1FEFD4403FE604888" box="[351,458,873,899]" pageId="13" pageNumber="140" type="holotype">Holotype</typeStatus>
, male. Body length
<quantity id="517E9B77737DFFF1FD054403FCA6488F" box="[679,780,873,900]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="13" pageNumber="140" unit="mm" value="14.3">14.3 mm</quantity>
, width
<quantity id="517E9B77737DFFF1FCFF4403FC1F488F" box="[861,949,873,900]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="13" pageNumber="140" unit="mm" value="3.9">3.9 mm</quantity>
, convex, elongate, shiny black, legs and antennae ferruginous, pubescence dark-brown to black, long, dense (
<figureCitation id="0EBD2A17737DFFF1FCDC44E7FBAB48AC" box="[894,1025,909,935]" captionStart="FIGURES 31–36" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[227,1326,185,1911]" captionTargetId="figure@14.[190,1397,174,1918]" captionTargetPageId="14" captionText="FIGURES 31–36. Habitus. Megathous tannourinensis sp. nov., male, holotype, 31—dorsal view, 34—lateral view; M. lizlerwerneri Mertlik, 2005, male, holotype, 32—dorsal view, 35—lateral view; M. menetriesi (Reitter, 1890), male, holotype, 33—dorsal view, 36—lateral view. Scale bars=2.0 mm." httpUri="https://zenodo.org/record/3241202/files/figure.png" pageId="13" pageNumber="140">Figs 31, 34</figureCitation>
). Head including eyes narrower than pronotum, frons with distinct V-shaped depression medially; punctation umbilicated, dense and coarse, intervals between punctures 0.2–0.6 times as puncture diameter. Antenna (
<figureCitation id="0EBD2A17737DFFF1FCDA44BFFC7A48E4" box="[888,976,981,1007]" captionStart="FIGURES 31–36" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[227,1326,185,1911]" captionTargetId="figure@14.[190,1397,174,1918]" captionTargetPageId="14" captionText="FIGURES 31–36. Habitus. Megathous tannourinensis sp. nov., male, holotype, 31—dorsal view, 34—lateral view; M. lizlerwerneri Mertlik, 2005, male, holotype, 32—dorsal view, 35—lateral view; M. menetriesi (Reitter, 1890), male, holotype, 33—dorsal view, 36—lateral view. Scale bars=2.0 mm." httpUri="https://zenodo.org/record/3241202/files/figure.png" pageId="13" pageNumber="140">Figs 31</figureCitation>
,
<figureCitation id="0EBD2A17737DFFF1FC7F44BFFC5048E4" box="[989,1018,981,1007]" captionStart="FIGURES 37–45" captionStartId="15.[151,264,1326,1351]" captionTargetBox="[151,1434,186,1297]" captionTargetId="figure@15.[151,1436,181,1306]" captionTargetPageId="15" captionText="FIGURES 37–45. Antennomeres. 37—M. tannourinensis sp. nov., male, holotype; 38—M. lizlerwerneri Mertlik, 2005, male, holotype; 39—M. menetriesi (Reitter, 1890), male, holotype. Prosternal process, lateral view. 40—M. tannourinensis sp. nov., male, holotype; 41—M. lizlerwerneri Mertlik, 2005, male, holotype; 42—M. menetriesi (Reitter, 1890), male, holotype.Aedeagus. 43—M. tannourinensis sp. nov., male, holotype. 44—M. lizlerwerneri Mertlik, 2005 male, holotype. 45—M. menetriesi (Reitter, 1890), male, holotype. Scale bars=0.5 mm (Figs 43–45), 1.0 mm (Figs 37–42)." httpUri="https://zenodo.org/record/3241206/files/figure.png" pageId="13" pageNumber="140">37</figureCitation>
) long, surpassing posterior angles of pronotum of about 2 antennomeres when directed backwards, obtusely serrate from antennomere III. Antennomere I robust, shiny, 3 times longer than II; II short, cylindrical, slightly longer than wide, III triangular, 2 times longer than II, slightly serrate, IV the widest, 1.3 times longer than wide, more serrated than III, V triangular, VI to X gradually widened apically, about 2 times longer than wide; apical antennomere longest, subacute apically. Surface of antennomeres rugosed, covered with moderately long, semierect setae.
</paragraph>
<paragraph id="96393692737DFFF1FF6543C4FA9E4E70" blockId="13.[151,1437,261,1944]" pageId="13" pageNumber="140">
Pronotum 1.1 times longer than wide, widest at middle, flattened dorsally, with shiny, shallow, partly visible longitudinal keel, without punctures (
<figureCitation id="0EBD2A17737DFFF1FD9E43BBFD154FE0" box="[572,703,1233,1259]" captionStart="FIGURES 31–36" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[227,1326,185,1911]" captionTargetId="figure@14.[190,1397,174,1918]" captionTargetPageId="14" captionText="FIGURES 31–36. Habitus. Megathous tannourinensis sp. nov., male, holotype, 31—dorsal view, 34—lateral view; M. lizlerwerneri Mertlik, 2005, male, holotype, 32—dorsal view, 35—lateral view; M. menetriesi (Reitter, 1890), male, holotype, 33—dorsal view, 36—lateral view. Scale bars=2.0 mm." httpUri="https://zenodo.org/record/3241202/files/figure.png" pageId="13" pageNumber="140">Figs 31, 34</figureCitation>
). Posterior angles slightly divergent, with short, indistinct sublateral carina; sides almost parallel, slightly narrowed from anterior third. Lateral carina distinct, complete anteriorly, visible from dorsal view. Punctation dense and coarse, uniformly distributed, punctures rather umbilicated, on disk with intervals on average equal to their diameters, slightly denser toward sides and base. Pubescence moderately long, semierect. Ventral side of prosternal process gradually sloping towards body from its base (
<figureCitation id="0EBD2A17737DFFF1FB71420BFA8D4E70" box="[1235,1319,1377,1403]" captionStart="FIGURES 37–45" captionStartId="15.[151,264,1326,1351]" captionTargetBox="[151,1434,186,1297]" captionTargetId="figure@15.[151,1436,181,1306]" captionTargetPageId="15" captionText="FIGURES 37–45. Antennomeres. 37—M. tannourinensis sp. nov., male, holotype; 38—M. lizlerwerneri Mertlik, 2005, male, holotype; 39—M. menetriesi (Reitter, 1890), male, holotype. Prosternal process, lateral view. 40—M. tannourinensis sp. nov., male, holotype; 41—M. lizlerwerneri Mertlik, 2005, male, holotype; 42—M. menetriesi (Reitter, 1890), male, holotype.Aedeagus. 43—M. tannourinensis sp. nov., male, holotype. 44—M. lizlerwerneri Mertlik, 2005 male, holotype. 45—M. menetriesi (Reitter, 1890), male, holotype. Scale bars=0.5 mm (Figs 43–45), 1.0 mm (Figs 37–42)." httpUri="https://zenodo.org/record/3241206/files/figure.png" pageId="13" pageNumber="140">Fig. 40</figureCitation>
).
</paragraph>
<paragraph id="96393692737DFFF1FF6542EFFDDF4EC8" blockId="13.[151,1437,261,1944]" pageId="13" pageNumber="140">Scutellar shield tongue-shaped, elongate, 1.8 times longer than wide, flattened, shiny, punctation dense and shallow, covered with greyish pubescence.</paragraph>
<paragraph id="96393692737DFFF1FF6542A7FB544EEC" blockId="13.[151,1437,261,1944]" box="[199,1278,1485,1512]" pageId="13" pageNumber="140">Elytra 2.7 times longer than pronotum and 3.4 times longer than wide, moderately convex; sides</paragraph>
<paragraph id="96393692737DFFF1FF65429BFC414D24" blockId="13.[151,1437,261,1944]" pageId="13" pageNumber="140">subparallel from base to beyond middle, then gradually narrowing to apices; striae regularly punctate; interstriae flat, densely and finely punctate. Pubescence decumbent to semierect.</paragraph>
<paragraph id="96393692737DFFF1FF654153FCA94D7C" blockId="13.[151,1437,261,1944]" pageId="13" pageNumber="140">
Aedeagus (
<figureCitation id="0EBD2A17737DFFF1FEE74153FE3C4D58" box="[325,406,1593,1619]" captionStart="FIGURES 37–45" captionStartId="15.[151,264,1326,1351]" captionTargetBox="[151,1434,186,1297]" captionTargetId="figure@15.[151,1436,181,1306]" captionTargetPageId="15" captionText="FIGURES 37–45. Antennomeres. 37—M. tannourinensis sp. nov., male, holotype; 38—M. lizlerwerneri Mertlik, 2005, male, holotype; 39—M. menetriesi (Reitter, 1890), male, holotype. Prosternal process, lateral view. 40—M. tannourinensis sp. nov., male, holotype; 41—M. lizlerwerneri Mertlik, 2005, male, holotype; 42—M. menetriesi (Reitter, 1890), male, holotype.Aedeagus. 43—M. tannourinensis sp. nov., male, holotype. 44—M. lizlerwerneri Mertlik, 2005 male, holotype. 45—M. menetriesi (Reitter, 1890), male, holotype. Scale bars=0.5 mm (Figs 43–45), 1.0 mm (Figs 37–42)." httpUri="https://zenodo.org/record/3241206/files/figure.png" pageId="13" pageNumber="140">Fig. 43</figureCitation>
) elongated, about three times as long as wide, with median lobe surpassing tip of paramere, paramere with widened apical lobe, rounded anteriorly.
</paragraph>
<paragraph id="96393692737DFFF1FF6541EBFE344D90" blockId="13.[151,1437,261,1944]" box="[199,414,1665,1691]" pageId="13" pageNumber="140">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="DE9C6519737DFFF1FF6541CFFE7B4DB4" box="[199,465,1701,1728]" pageId="13" pageNumber="140" type="distribution">
<paragraph id="96393692737DFFF1FF6541CFFE7B4DB4" blockId="13.[151,1437,261,1944]" box="[199,465,1701,1728]" pageId="13" pageNumber="140">
<emphasis id="A4F2EA80737DFFF1FF6541CFFECB4DCB" bold="true" box="[199,353,1701,1728]" pageId="13" pageNumber="140">Distribution.</emphasis>
<collectingCountry id="EE917602737DFFF1FEC541CFFE674DB4" box="[359,461,1701,1727]" name="Lebanon" pageId="13" pageNumber="140">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DE9C6519737DFFF1FF6541A3FEBB4C78" pageId="13" pageNumber="140" type="biology_ecology">
<paragraph id="96393692737DFFF1FF6541A3FEBB4C78" blockId="13.[151,1437,261,1944]" pageId="13" pageNumber="140">
<emphasis id="A4F2EA80737DFFF1FF6541A3FE8D4DEF" bold="true" box="[199,295,1737,1764]" pageId="13" pageNumber="140">Biology.</emphasis>
The two known males were reared from an old hollow
<taxonomicName id="51864D11737DFFF1FC0541A0FABC4DE8" authority=", Tannourine" authorityName="Tannourine" box="[935,1302,1737,1764]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="140" phylum="Tracheophyta" rank="species" species="pinnatifida">
<emphasis id="A4F2EA80737DFFF1FC0541A0FB224DE8" box="[935,1160,1737,1764]" italics="true" pageId="13" pageNumber="140">Quercus pinnatifida</emphasis>
, Tannourine
</taxonomicName>
Nature Reserve in Northern
<collectingCountry id="EE917602737DFFF1FEC44187FE664C0C" box="[358,460,1773,1799]" name="Lebanon" pageId="13" pageNumber="140">Lebanon</collectingCountry>
. Larvae were found in red-rotten wood of the cavity, together with adults of
<taxonomicName id="51864D11737DFFF1FA944187FE654C20" authority="(Fabricius, 1792)" baseAuthorityName="Fabricius" baseAuthorityYear="1792" class="Insecta" family="Elateridae" genus="Ampedus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="praeustus">
<emphasis id="A4F2EA80737DFFF1FA944187FEAE4C20" italics="true" pageId="13" pageNumber="140">Ampedus praeustus</emphasis>
(
<bibRefCitation id="F2174B63737DFFF1FEB6407BFE6C4C20" author="Fabricius, J. C." box="[276,454,1809,1835]" pageId="13" pageNumber="140" refId="ref19322" refString="Fabricius, J. C. (1792) Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonymis, locis, descriptionibus, observationibus. Tom I. Pars II. Christ. Gottl. Proft, Hafniae, 538 pp. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1792">Fabricius, 1792</bibRefCitation>
)
</taxonomicName>
and larvae of
<emphasis id="A4F2EA80737DFFF1FDD6407BFD714C20" box="[628,731,1809,1835]" italics="true" pageId="13" pageNumber="140">Ampedus</emphasis>
sp. and
<taxonomicName id="51864D11737DFFF1FC9F407BFC4C4C20" authority="Németh, 2019" authorityName="Németh" authorityYear="2019" box="[829,998,1809,1835]" class="Insecta" family="Elateridae" genus="Elathous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="species" species="agilis" status="sp. nov.">
<emphasis id="A4F2EA80737DFFF1FC9F407BFC4C4C20" box="[829,998,1809,1835]" italics="true" pageId="13" pageNumber="140">Elathous agilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFC157FB737DFFF1FC4D4078FBE04C27" box="[1007,1098,1810,1836]" pageId="13" pageNumber="140" rank="species">
<emphasis id="A4F2EA80737DFFF1FC4D4078FBE04C27" bold="true" box="[1007,1098,1810,1836]" pageId="13" pageNumber="140">sp. nov.</emphasis>
</taxonomicNameLabel>
Larval specimens were taken home alive in plastic boxes with parts of the rotten wood, adults emerged in the
<taxonomicName id="51864D11737DFFF1FBB7405FFB3A4C44" authorityName="Linnaeus" authorityYear="1758" box="[1045,1168,1845,1871]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
collection of HNHM in
<date id="E2381052737DFFF1FF354030FEA44C78" box="[151,270,1881,1908]" pageId="13" pageNumber="140" value="2017-06">June 2017</date>
.
</paragraph>
</subSubSection>
<subSubSection id="DE9C6519737DFFF1FF654014FB294C9C" box="[199,1155,1917,1944]" pageId="13" pageNumber="140" type="etymology">
<paragraph id="96393692737DFFF1FF654014FB294C9C" blockId="13.[151,1437,261,1944]" box="[199,1155,1917,1944]" pageId="13" pageNumber="140">
<emphasis id="A4F2EA80737DFFF1FF654014FEE44C93" bold="true" box="[199,334,1918,1944]" pageId="13" pageNumber="140">Etymology.</emphasis>
This species is named after the town of Tannourine in northern
<collectingCountry id="EE917602737DFFF1FBBB4017FBD54C9C" box="[1049,1151,1917,1943]" name="Lebanon" pageId="13" pageNumber="140">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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