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264 lines
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.222.99420" ID-Pensoft-Pub="1314-2003-222-69" ID-Pensoft-UUID="924E690E2AC95EB29964EB150D58096C" ModsDocID="1314-2003-222-69" checkinTime="1679436427559" checkinUser="pensoft" docAuthor="Xue, Bine, Huang, Erfeng, Yao, Gang, Zhao, Guohua & Huang, Jiuxiang" docDate="2023" docId="1E12F3CAA56A5AD18475E8DDE9E3156F" docLanguage="en" docName="PhytoKeys 222: 69-74" docOrigin="PhytoKeys 222" docPubDate="2023-03-21" docSource="http://dx.doi.org/10.3897/phytokeys.222.99420" docTitle="Hoya spectatissima B. Xue, E. F. Huang, Gang Yao & J. X. Huang 2023, sp. nov." docType="treatment" docVersion="1" id="924E690E2AC95EB29964EB150D58096C" lastPageNumber="69" masterDocId="924E690E2AC95EB29964EB150D58096C" masterDocTitle="Hoya spectatissima (Gentianales, Apocynaceae), a new species from Yunnan, China" masterLastPageNumber="74" masterPageNumber="69" pageNumber="69" updateTime="1679436427559" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Hoya spectatissima (Gentianales, Apocynaceae), a new species from Yunnan, China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Xue, Bine</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4515-4316</mods:nameIdentifier>
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<mods:affiliation>College of Horticulture and Landscape Architecture, Zhongkai University of Agriculture and Engineering, Guangzhou 510225, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huang, Erfeng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5493-172X</mods:nameIdentifier>
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<mods:affiliation>Guangxi Nanning Roy Garden Co., Ltd., Nanning 530227, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yao, Gang</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3101-8230</mods:nameIdentifier>
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<mods:affiliation>College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou 510642, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhao, Guohua</mods:namePart>
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<mods:affiliation>Fairy Lake Botanical Garden, Shenzhen & Chinese Academy of Sciences, Shenzhen 518004, Guangdong, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huang, Jiuxiang</mods:namePart>
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<mods:affiliation>College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou 510642, China</mods:affiliation>
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<mods:nameIdentifier type="email">jxhuang@scau.edu.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-21</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>222</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>69</mods:start>
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<mods:end>74</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.222.99420</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.222.99420</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-222-69</mods:identifier>
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<mods:identifier type="Pensoft-UUID">924E690E2AC95EB29964EB150D58096C</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:1E12F3CAA56A5AD18475E8DDE9E3156F" httpUri="http://treatment.plazi.org/id/1E12F3CAA56A5AD18475E8DDE9E3156F" lastPageNumber="69" pageId="0" pageNumber="69">
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<subSubSection pageId="0" pageNumber="69" type="nomenclature">
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<paragraph pageId="0" pageNumber="69">
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<taxonomicName LSID="1E12F3CA-A56A-5AD1-8475-E8DDE9E3156F" authority="B. Xue, E. F. Huang, Gang Yao & J. X. Huang" authorityName="B. Xue, E. F. Huang, Gang Yao & J. X. Huang" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya spectatissima" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="spectatissima" status="sp. nov.">Hoya spectatissima B.Xue, E.F.Huang, Gang Yao & J.X.Huang</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="69">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="description">
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<paragraph pageId="0" pageNumber="69">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya spectatissima A habit B cross section of lamina C leaves D, E inflorescence F flower (top view) G flower (bottom view) H corona (top view; Red arrowheads indicate the inner angle of the corona lobes and yellow arrowheads indicate the outer angle of the corona lobes) I corona (side view) J ovaries K pollinarium." figureDoi="10.3897/phytokeys.222.99420.figure1" httpUri="https://binary.pensoft.net/fig/827806" pageId="0" pageNumber="69">Fig. 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="diagnosis">
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<paragraph pageId="0" pageNumber="69">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<taxonomicName authorityName="B. Xue, E. F. Huang, Gang Yao & J. X. Huang" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya spectatissima" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="spectatissima">
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<emphasis italics="true" pageId="0" pageNumber="69">Hoya spectatissima</emphasis>
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</taxonomicName>
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is most similar to
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. lyi" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="lyi">
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<emphasis italics="true" pageId="0" pageNumber="69">H. lyi</emphasis>
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</taxonomicName>
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H.
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<normalizedToken originalValue="Lév">Lev</normalizedToken>
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, but differs from the latter by its succulent leaves (vs. coriaceous leaves) that are 2-4.5 cm in length (vs. up to 9 cm in length) and rounded or rarely obtuse at the apex (vs. mostly acute at the apex), pink to pale pink corolla (vs. yellowish-white corolla) that is densely sericeous adaxially (vs. densely pubescent adaxially), ivory white corona lobes (vs. purple towards the inner angle and ivory white towards the outer angle of corona lobes) that are sub-rhombic in top view (vs. ovoid in top view) and have an acute outer angle (vs. truncate to shallowly concave outer angle).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="materials_examined">
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<paragraph pageId="0" pageNumber="69">
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<typeStatus>Type</typeStatus>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<materialsCitation collectingDate="2021-07-07" country="China" county="Gengma Hsien" location="Guanlong Village" municipality="Sipaishan Town" specimenCount="1" stateProvince="Yunnan Province" typeStatus="Holotype">
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<collectingCountry name="China">China</collectingCountry>
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,
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<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
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,
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<collectingCounty>Gengma Hsien</collectingCounty>
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,
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<collectingMunicipality>Sipaishan Town</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:1E12F3CAA56A5AD18475E8DDE9E3156F:5B4CAF50182431D05D05A82CFC72A3B2" country="China" county="Gengma Hsien" municipality="Sipaishan Town" name="Guanlong Village" stateProvince="Yunnan Province">Guanlong Village</location>
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,
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<location LSID="urn:lsid:plazi:treatment:1E12F3CAA56A5AD18475E8DDE9E3156F:6D917F65FB487C7EF6065AECB9BAE40F" country="China" county="Gengma Hsien" municipality="Sipaishan Town" name="Luoshuidong" stateProvince="Yunnan Province">Luoshuidong</location>
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, on rocks in karst habitat,
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<collectingDate value="2021-07-07">7 July 2021</collectingDate>
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,
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<emphasis italics="true" pageId="0" pageNumber="69">E.F. Huang-RG0017</emphasis>
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(
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<typeStatus>Holotype</typeStatus>
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: IBSC!)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="description">
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<paragraph pageId="0" pageNumber="69">Description.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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Lithophytic climber. Stems rounded in cross section, up to 1 m in length, 1.5-2 mm in diameter, densely pubescent when young, subglabrous when mature, sometimes 3-5 branches at nodes, internodes 3-8 cm long. Leaves opposite, wine-red and pubescent when young; mature lamina subglabrous, succulent, elliptic or oblong, 2-4.5
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<normalizedToken originalValue="×">x</normalizedToken>
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1.3-1.8 cm, 2.5-3 mm in thickness, bright green adaxially, pale green abaxially, base rounded, apex rounded or rarely obtuse; mid-vein slightly elevated or obscure adaxially, not elevated abaxially, lateral veins 2-3 pairs or sometimes invisible in adaxial surface, invisible in abaxial surface; petioles sometimes recurved, 3-7 mm long, 1-1.5 mm in diameter. Inflorescences axillary pseudo-umbels, many-flowered, globose; peduncle longer than pedicels, 4.5-5 cm long, papillose to sparsely pubescent; pedicels 2.5-3 cm long, yellow-green, papillose to sparsely pubescent. Calyx lobes triangular to narrowly triangular, ca. 1.5
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<normalizedToken originalValue="×">x</normalizedToken>
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2 mm, purplish-red, sparely pubescent or glabrous. Corolla rotate, pink in early flowering phase, pale pink in late flowering phase, ca. 2 cm in diameter, densely sericeous adaxially, sub-glabrous abaxially, lobes triangular, 5-6 mm long and 5-6 mm wide, margin recurved, apex rounded, revolute. Corona ivory white, 7-8 mm in diameter, 2.0-2.4 mm high, lobes 5, stellate spreading, ca. 3.5
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<normalizedToken originalValue="×">x</normalizedToken>
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2.5 mm, sub-rhombic in top view, outer angle acute, inner angle acuminate, the inner tips convex and spreading obviously towards the centre of the flower, ivory white or rarely shallowly pink. Pollinia clavate, ca. 0.73
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<normalizedToken originalValue="×">x</normalizedToken>
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0.26 mm, base cuneate, apex truncate, narrowing towards the base, sterile edge all along the outer edge of the pollinium, translator arms attached at the centre of the corpusculum. Ovaries 2, triangular-ovate, attached to each other, ca. 2 mm long and 1.5 mm wide, green, glabrous. Fruit and seed not seen.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="distribution">
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<paragraph pageId="0" pageNumber="69">Distribution and habit.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<taxonomicName authorityName="B. Xue, E. F. Huang, Gang Yao & J. X. Huang" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya spectatissima" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="spectatissima">
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<emphasis italics="true" pageId="0" pageNumber="69">Hoya spectatissima</emphasis>
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</taxonomicName>
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is known from at least two localities in Cangyuan Hsien and Gengma Hsien, Yunnan Province, China. It is a lithophytic liana that climbs on rocks in karst habitat. It is a common species in Guanlong Village, Gengma Hsien, but an occasional species in Menglai Town, Cangyuan Hsien.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="etymology">
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<paragraph pageId="0" pageNumber="69">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<taxonomicName authorityName="B. Xue, E. F. Huang, Gang Yao & J. X. Huang" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya spectatissima" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="spectatissima">
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<emphasis italics="true" pageId="0" pageNumber="69">Hoya spectatissima</emphasis>
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</taxonomicName>
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is named to reflect its beautiful flowers.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="taxonomic discussion">
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<paragraph pageId="0" pageNumber="69">Taxonomic discussion.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<taxonomicName authorityName="B. Xue, E. F. Huang, Gang Yao & J. X. Huang" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya spectatissima" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="spectatissima">
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<emphasis italics="true" pageId="0" pageNumber="69">Hoya spectatissima</emphasis>
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</taxonomicName>
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has evidently succulent leaves that are 2.5-3 mm thick (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya spectatissima A habit B cross section of lamina C leaves D, E inflorescence F flower (top view) G flower (bottom view) H corona (top view; Red arrowheads indicate the inner angle of the corona lobes and yellow arrowheads indicate the outer angle of the corona lobes) I corona (side view) J ovaries K pollinarium." figureDoi="10.3897/phytokeys.222.99420.figure1" httpUri="https://binary.pensoft.net/fig/827806" pageId="0" pageNumber="69">1B</figureCitation>
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); this trait is also found in
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. pandurata" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="pandurata">
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<emphasis italics="true" pageId="0" pageNumber="69">H. pandurata</emphasis>
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</taxonomicName>
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Tsiang, which is also endemic in Yunnan Province, China. However,
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. pandurata" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="pandurata">
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<emphasis italics="true" pageId="0" pageNumber="69">H. pandurata</emphasis>
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</taxonomicName>
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is a subshrub and is epiphytic on trees in open or mixed woods and it further differs from
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. spectatissima" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="spectatissima">
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<emphasis italics="true" pageId="0" pageNumber="69">H. spectatissima</emphasis>
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</taxonomicName>
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by its pandurate or oblong leaves (vs. elliptic or oblong) with acuminate apex (vs. rounded or rarely obtuse), shorter peduncle and pedicels that are ca. 3 mm long (vs. 4.5-5 cm long) and 1.5 cm long (vs. 2.5-3 cm long) respectively and yellow or reddish corolla (vs. pink or pale pink) that is 0.8-1 cm (vs. ca. 2 cm) in diameter (
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<bibRefCitation author="Li, PT" editor="Wu, CY" journalOrPublisher="16). Science Press & Missouri Botanical Garden Press, Beijing, China / St. Louis" pageId="0" pageNumber="69" pagination="228 - 236" refId="B6" refString="Li, PT, Gilbert, MG, Stevens, WD, 1995. Hoya. In: Wu, CY, Raven, PH, Eds., Flora of China (Vol. 16). Science Press & Missouri Botanical Garden Press, Beijing, China / St. Louis: 228 - 236" title="Hoya." volumeTitle="Flora of China (Vol." year="1995">Li et al. 1995</bibRefCitation>
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). Additionally,
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. spectatissima" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="spectatissima">
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<emphasis italics="true" pageId="0" pageNumber="69">H. spectatissima</emphasis>
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</taxonomicName>
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is morphologically similar to
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. lyi" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="lyi">
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<emphasis italics="true" pageId="0" pageNumber="69">H. lyi</emphasis>
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</taxonomicName>
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, a species recorded from China, Laos and Vietnam (
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<bibRefCitation DOI="https://doi.org/10.1017/S0960428611000412" author="Rodda, M" journalOrPublisher="Edinburgh Journal of Botany" pageId="0" pageNumber="69" pagination="83 - 93" refId="B9" refString="Rodda, M, 2012. Taxonomy of Hoya lyi, Hoya yuennanensis and Hoya mekongensis (Apocynaceae - Asclepiadoideae). Edinburgh Journal of Botany 69 (1): 83 - 93, DOI: https://doi.org/10.1017/S0960428611000412" title="Taxonomy of Hoya lyi, Hoya yuennanensis and Hoya mekongensis (Apocynaceae - Asclepiadoideae)." url="https://doi.org/10.1017/S0960428611000412" volume="69" year="2012">Rodda 2012</bibRefCitation>
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). Both species have elliptic or oblong leaves (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya spectatissima A habit B cross section of lamina C leaves D, E inflorescence F flower (top view) G flower (bottom view) H corona (top view; Red arrowheads indicate the inner angle of the corona lobes and yellow arrowheads indicate the outer angle of the corona lobes) I corona (side view) J ovaries K pollinarium." figureDoi="10.3897/phytokeys.222.99420.figure1" httpUri="https://binary.pensoft.net/fig/827806" pageId="0" pageNumber="69">1A, C</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hoya lyi A isosynytpe (E. Bodinier 1853, P 00634485) B inflorescence C habit." figureDoi="10.3897/phytokeys.222.99420.figure2" httpUri="https://binary.pensoft.net/fig/827807" pageId="0" pageNumber="69">2A, C</figureCitation>
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), triangular to narrowly triangular calyx lobes, corolla ca. 2 cm in diameter and clavate pollinia. However,
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<taxonomicName authorityName="B. Xue, E. F. Huang, Gang Yao & J. X. Huang" authorityYear="2023" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya spectatissima" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="spectatissima">
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<emphasis italics="true" pageId="0" pageNumber="69">Hoya spectatissima</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" lsidName="H. lyi" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="lyi">
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<emphasis italics="true" pageId="0" pageNumber="69">H. lyi</emphasis>
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</taxonomicName>
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by the following characters: succulent leaves that are 2.5-3 mm thick (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya spectatissima A habit B cross section of lamina C leaves D, E inflorescence F flower (top view) G flower (bottom view) H corona (top view; Red arrowheads indicate the inner angle of the corona lobes and yellow arrowheads indicate the outer angle of the corona lobes) I corona (side view) J ovaries K pollinarium." figureDoi="10.3897/phytokeys.222.99420.figure1" httpUri="https://binary.pensoft.net/fig/827806" pageId="0" pageNumber="69">1B</figureCitation>
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) (vs. coriaceous leaves that are less than 1.5 mm thick, Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hoya lyi A isosynytpe (E. Bodinier 1853, P 00634485) B inflorescence C habit." figureDoi="10.3897/phytokeys.222.99420.figure2" httpUri="https://binary.pensoft.net/fig/827807" pageId="0" pageNumber="69">2A, C</figureCitation>
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); shorter leaves that are 2-4.5 cm long (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya spectatissima A habit B cross section of lamina C leaves D, E inflorescence F flower (top view) G flower (bottom view) H corona (top view; Red arrowheads indicate the inner angle of the corona lobes and yellow arrowheads indicate the outer angle of the corona lobes) I corona (side view) J ovaries K pollinarium." figureDoi="10.3897/phytokeys.222.99420.figure1" httpUri="https://binary.pensoft.net/fig/827806" pageId="0" pageNumber="69">1C</figureCitation>
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) (vs. up to 9 cm long, Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hoya lyi A isosynytpe (E. Bodinier 1853, P 00634485) B inflorescence C habit." figureDoi="10.3897/phytokeys.222.99420.figure2" httpUri="https://binary.pensoft.net/fig/827807" pageId="0" pageNumber="69">2A</figureCitation>
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); apex of leaves rounded or rarely obtuse (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya spectatissima A habit B cross section of lamina C leaves D, E inflorescence F flower (top view) G flower (bottom view) H corona (top view; Red arrowheads indicate the inner angle of the corona lobes and yellow arrowheads indicate the outer angle of the corona lobes) I corona (side view) J ovaries K pollinarium." figureDoi="10.3897/phytokeys.222.99420.figure1" httpUri="https://binary.pensoft.net/fig/827806" pageId="0" pageNumber="69">1A, C</figureCitation>
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) (vs. mostly acute, Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hoya lyi A isosynytpe (E. Bodinier 1853, P 00634485) B inflorescence C habit." figureDoi="10.3897/phytokeys.222.99420.figure2" httpUri="https://binary.pensoft.net/fig/827807" pageId="0" pageNumber="69">2A, C</figureCitation>
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); 2-3 paired lateral veins in leaves (vs. 3-5 paired lateral veins); pink to pale pink corolla (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya spectatissima A habit B cross section of lamina C leaves D, E inflorescence F flower (top view) G flower (bottom view) H corona (top view; Red arrowheads indicate the inner angle of the corona lobes and yellow arrowheads indicate the outer angle of the corona lobes) I corona (side view) J ovaries K pollinarium." figureDoi="10.3897/phytokeys.222.99420.figure1" httpUri="https://binary.pensoft.net/fig/827806" pageId="0" pageNumber="69">1D, E</figureCitation>
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) (vs. yellowish-white, Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hoya lyi A isosynytpe (E. Bodinier 1853, P 00634485) B inflorescence C habit." figureDoi="10.3897/phytokeys.222.99420.figure2" httpUri="https://binary.pensoft.net/fig/827807" pageId="0" pageNumber="69">2B</figureCitation>
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); corolla lobes densely sericeous adaxially (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya spectatissima A habit B cross section of lamina C leaves D, E inflorescence F flower (top view) G flower (bottom view) H corona (top view; Red arrowheads indicate the inner angle of the corona lobes and yellow arrowheads indicate the outer angle of the corona lobes) I corona (side view) J ovaries K pollinarium." figureDoi="10.3897/phytokeys.222.99420.figure1" httpUri="https://binary.pensoft.net/fig/827806" pageId="0" pageNumber="69">1F, G</figureCitation>
|
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) (vs. densely pubescent adaxially, Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hoya lyi A isosynytpe (E. Bodinier 1853, P 00634485) B inflorescence C habit." figureDoi="10.3897/phytokeys.222.99420.figure2" httpUri="https://binary.pensoft.net/fig/827807" pageId="0" pageNumber="69">2B</figureCitation>
|
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); corona lobes sub-rhombic in top view (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya spectatissima A habit B cross section of lamina C leaves D, E inflorescence F flower (top view) G flower (bottom view) H corona (top view; Red arrowheads indicate the inner angle of the corona lobes and yellow arrowheads indicate the outer angle of the corona lobes) I corona (side view) J ovaries K pollinarium." figureDoi="10.3897/phytokeys.222.99420.figure1" httpUri="https://binary.pensoft.net/fig/827806" pageId="0" pageNumber="69">1F, H</figureCitation>
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|
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) (vs. ovoid, Fig.
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|
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hoya lyi A isosynytpe (E. Bodinier 1853, P 00634485) B inflorescence C habit." figureDoi="10.3897/phytokeys.222.99420.figure2" httpUri="https://binary.pensoft.net/fig/827807" pageId="0" pageNumber="69">2B</figureCitation>
|
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|
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) and ivory white (Fig.
|
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|
|
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya spectatissima A habit B cross section of lamina C leaves D, E inflorescence F flower (top view) G flower (bottom view) H corona (top view; Red arrowheads indicate the inner angle of the corona lobes and yellow arrowheads indicate the outer angle of the corona lobes) I corona (side view) J ovaries K pollinarium." figureDoi="10.3897/phytokeys.222.99420.figure1" httpUri="https://binary.pensoft.net/fig/827806" pageId="0" pageNumber="69">1F, H</figureCitation>
|
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|
|
) (vs. purple towards the inner angle and ivory white towards the outer angle, Fig.
|
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|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hoya lyi A isosynytpe (E. Bodinier 1853, P 00634485) B inflorescence C habit." figureDoi="10.3897/phytokeys.222.99420.figure2" httpUri="https://binary.pensoft.net/fig/827807" pageId="0" pageNumber="69">2B</figureCitation>
|
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); and outer angle of corona lobes acute (Fig.
|
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|
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hoya spectatissima A habit B cross section of lamina C leaves D, E inflorescence F flower (top view) G flower (bottom view) H corona (top view; Red arrowheads indicate the inner angle of the corona lobes and yellow arrowheads indicate the outer angle of the corona lobes) I corona (side view) J ovaries K pollinarium." figureDoi="10.3897/phytokeys.222.99420.figure1" httpUri="https://binary.pensoft.net/fig/827806" pageId="0" pageNumber="69">1H</figureCitation>
|
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|
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) (vs. truncate to shallowly concave, Fig.
|
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|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Hoya lyi A isosynytpe (E. Bodinier 1853, P 00634485) B inflorescence C habit." figureDoi="10.3897/phytokeys.222.99420.figure2" httpUri="https://binary.pensoft.net/fig/827807" pageId="0" pageNumber="69">2B</figureCitation>
|
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).
|
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|
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</paragraph>
|
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|
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<caption doi="10.3897/phytokeys.222.99420.figure2" httpUri="https://binary.pensoft.net/fig/827807" pageId="0" pageNumber="69" start="Figure 2" startId="F2">
|
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|
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<paragraph pageId="0" pageNumber="69">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="69">Figure 2.</emphasis>
|
|||
|
|
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Hoya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoya lyi" order="Gentianales" pageId="0" pageNumber="69" phylum="Tracheophyta" rank="species" species="lyi">
|
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|
|
<emphasis italics="true" pageId="0" pageNumber="69">Hoya lyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="69">A</emphasis>
|
|||
|
|
isosynytpe (
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="69">E. Bodinier 1853</emphasis>
|
|||
|
|
, P00634485)
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="69">B</emphasis>
|
|||
|
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inflorescence
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="69">C</emphasis>
|
|||
|
|
habit.
|
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|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
</subSubSection>
|
|||
|
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<subSubSection pageId="0" pageNumber="69" type="paratype">
|
|||
|
|
<paragraph pageId="0" pageNumber="69">Paratype.</paragraph>
|
|||
|
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<paragraph pageId="0" pageNumber="69">
|
|||
|
|
China. Yunnan Province, Cangyuan Hsien, Menglai Town, on rocks in karst habitat, 22 August 2021,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="69">E.F. Huang-RG0033</emphasis>
|
|||
|
|
(IBSC).
|
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|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
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|
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</treatment>
|
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|
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</document>
|