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<document id="2079366D92A64B401E12DD1D4157CA7B" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D059247FDF3FD53CAE129DC" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius babyrousa Telnov 2020, sp. nov." docType="treatment" docUuid="E9202A3D-800A-47B0-B9D0-884E101246ED" docUuidSource="ZooBank" docVersion="10" lastPageNumber="29" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="26" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="6BF4BB1FC92272F51D55EF57A2BE5DA7">
<mods:title id="109A5CEC583825C0963C876F55C660B9">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:role id="D5818A9BBDD1101BA47B63329E35B13B">
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<mods:namePart id="E9E4043CF26476E967D321A11A6C6D00">Telnov, Dmitry</mods:namePart>
</mods:name>
<mods:typeOfResource id="FAA46B447176B29B75078290F9FFB808">text</mods:typeOfResource>
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<mods:titleInfo id="BB8D0AAA7ABE52CAE95186E0ECAADE00">
<mods:title id="CF70C86DF0F27815EEB62EC22FC5C119">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="EF96645EB6E7A221EECC92CB0DDB958E">
<mods:date id="DCF817BCA46485E86C9F6F038C9CB77A">2020</mods:date>
<mods:detail id="2153F2A1671F6F9FEA0657EA05A8A597" type="pubDate">
<mods:number id="0E8AC40CC88EDC1A18966DC9AAF3F52D">2020-10-19</mods:number>
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<mods:detail id="3274E43671620AE55BC8BCCF74540443" type="volume">
<mods:number id="E77B04BA150D2D543F45E3996C43C190">721</mods:number>
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<mods:classification id="FE75C1720541728DF0021D9F867C9D9C">journal article</mods:classification>
<mods:identifier id="DB4BB491E758222E9E4232ADF799B4D3" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="D6DA49033572E72DF209EBA8AA42015C" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="062C57E0A27EF6F5EC2320A8BB3ECEA8" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
<mods:identifier id="F63019FAF008FDC45A7CD1FEE3CDDF13" type="Zenodo-Dep">4122118</mods:identifier>
<mods:identifier id="E8698988C211064B9EE91D3FCE2A7BD1" type="ZooBank">89E62EF8-2E45-4C59-94B7-6A5603E8939B</mods:identifier>
</mods:mods>
<treatment id="1A3787F69D059247FDF3FD53CAE129DC" ID-DOI="http://doi.org/10.5281/zenodo.4344381" ID-GBIF-Taxon="168648185" ID-Zenodo-Dep="4344381" LSID="urn:lsid:zoobank.org:act:E9202A3D-800A-47B0-B9D0-884E101246ED" httpUri="http://treatment.plazi.org/id/1A3787F69D059247FDF3FD53CAE129DC" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26">
<subSubSection id="DA84656B9D059242FDF3FD53CB822B6B" box="[608,980,749,777]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="922136E09D059242FDF3FD53CB822B6B" blockId="25.[339,1248,749,852]" box="[608,980,749,777]" pageId="25" pageNumber="26">
<heading id="C969818C9D059242FDF3FD53CB822B6B" box="[608,980,749,777]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
<taxonomicName id="559E4D639D059242FDF3FD53CB242B6A" ID-CoL="7X3W9" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[608,882,749,776]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="babyrousa" status="sp. nov.">
<emphasis id="A0EAEAF29D059242FDF3FD53CB242B6A" bold="true" box="[608,882,749,776]" italics="true" pageId="25" pageNumber="26">Maechidius babyrousa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D059242FCE9FD51CB822B6B" box="[890,980,751,777]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D059242FEFCFCADCCB62B36" pageId="25" pageNumber="26" type="description">
<paragraph id="922136E09D059242FEFCFCADCC932B4F" blockId="25.[339,1248,749,852]" box="[367,1221,787,813]" pageId="25" pageNumber="26">
<uri id="E60F3AE29D059242FEFCFCADCC932B4F" box="[367,1221,787,813]" pageId="25" pageNumber="26">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D059242FD1EFCADCC932B4F" box="[653,1221,787,813]" pageId="25" pageNumber="26">
E9202A3D-800A-47B0-B9D0-884E
<date id="E62010209D059242FBD3FCADCCCD2B4F" box="[1088,1179,787,813]" pageId="25" pageNumber="26" value="46-10-12">101246</date>
ED
</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D059242FEC0FC87CCB62B36" blockId="25.[339,1248,749,852]" box="[339,1248,825,852]" pageId="25" pageNumber="26">
<figureCitation id="0AA52A659D059242FEC0FC87C9FB2B36" box="[339,429,825,852]" captionStart="Figs 25–27" captionStartId="26.[189,241,1928,1954]" captionTargetBox="[381,1192,273,1894]" captionTargetId="figure-49@26.[361,1255,262,1894]" captionTargetPageId="26" captionText="Figs 25–27. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal and lateral views. 25. M. babyrousa sp. nov., holotype, ♂ (BMNH). 26–27. M. bintang sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122134" httpUri="https://zenodo.org/record/4122134/files/figure.png" pageId="25" pageNumber="26">Figs 25</figureCitation>
,
<figureCitation id="0AA52A659D059242FE2AFC87CA752B31" box="[441,547,825,851]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="25" pageNumber="26">106–107</figureCitation>
,
<figureCitation id="0AA52A659D059242FDBCFC87CA082B31" box="[559,606,825,851]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="25" pageNumber="26">203</figureCitation>
,
<figureCitation id="0AA52A659D059242FDFAFC87CACE2B31" box="[617,664,825,851]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="25" pageNumber="26">293</figureCitation>
, 365, 438–440, 449–450, 458–459, 538, 582–585
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D059242FF2EFCC3CC352C3D" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="922136E09D059242FF2EFCC3C99F2BF5" blockId="25.[189,457,893,919]" box="[189,457,893,919]" pageId="25" pageNumber="26">
<heading id="C969818C9D059242FF2EFCC3C99F2BF5" bold="true" box="[189,457,893,919]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="A0EAEAF29D059242FF2EFCC3C99F2BF5" bold="true" box="[189,457,893,919]" pageId="25" pageNumber="26">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D059242FF2EFC10CC352C3D" blockId="25.[189,1399,942,1119]" pageId="25" pageNumber="26">
This species is most similar to
<taxonomicName id="559E4D639D059242FDBFFC11CB012BAA" authority="Lansberge" authorityName="Lansberge" authorityYear="1886" box="[556,855,942,968]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="A0EAEAF29D059242FDBFFC11CA842BAA" box="[556,722,943,968]" italics="true" pageId="25" pageNumber="26">M. peregrinus</emphasis>
Lansberge
</taxonomicName>
,
<taxonomicName id="559E4D639D059242FCF0FC11CBBE2BAA" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[867,1000,942,968]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="deltouri" status="sp. nov.">
<emphasis id="A0EAEAF29D059242FCF0FC11CBBE2BAA" box="[867,1000,942,968]" italics="true" pageId="25" pageNumber="26">M. deltouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D059242FC63FC10CC182BAA" box="[1008,1102,942,968]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="559E4D639D059242FBC9FC11CCB72BAA" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1114,1249,942,968]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="legalovi" status="sp. nov.">
<emphasis id="A0EAEAF29D059242FBC9FC11CCB72BAA" box="[1114,1249,942,968]" italics="true" pageId="25" pageNumber="26">M. legalovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D059242FB7AFC10CD122BAA" box="[1257,1348,942,968]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="559E4D639D059242FF2EFC6AC9142B8F" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[189,322,980,1005]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="suwawa" status="sp. nov.">
<emphasis id="A0EAEAF29D059242FF2EFC6AC9142B8F" box="[189,322,980,1005]" italics="true" pageId="25" pageNumber="26">M. suwawa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D059242FEDBFC6AC9F42B8C" box="[328,418,980,1006]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
(all from
<collectingRegion id="505AF8029D059242FD87FC6DCAD52B8C" box="[532,643,979,1006]" country="Indonesia" name="Sulawesi Utara" pageId="25" pageNumber="26">Sulawesi</collectingRegion>
). This new species readily differs from them and other congeners only in the shape of the male genital organs, labroclypeus, apically unidentate (males) or bidentate (females) protibiae, and the presence of a transverse median hump on the female pygidium. Two vague circular impressions on the pronotal disc are also present in
<taxonomicName id="559E4D639D059242FCEEFBFBCC572C3C" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[893,1025,1092,1118]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="deltouri" status="sp. nov.">
<emphasis id="A0EAEAF29D059242FCEEFBFBCC572C3C" box="[893,1025,1092,1118]" italics="true" pageId="25" pageNumber="26">M. deltouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D059242FB9BFBFBCC352C3D" box="[1032,1123,1093,1119]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D059242FF2EFB36C9F02C9A" pageId="25" pageNumber="26" type="etymology">
<paragraph id="922136E09D059242FF2EFB36C9122CC0" blockId="25.[189,324,1160,1186]" box="[189,324,1160,1186]" pageId="25" pageNumber="26">
<heading id="C969818C9D059242FF2EFB36C9122CC0" bold="true" box="[189,324,1160,1186]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="A0EAEAF29D059242FF2EFB36C9122CC0" bold="true" box="[189,324,1160,1186]" pageId="25" pageNumber="26">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D059242FF2EFB07C9F02C9A" blockId="25.[189,1399,1209,1273]" pageId="25" pageNumber="26">
Named after
<emphasis id="A0EAEAF29D059242FEC1FB07C9862CB1" box="[338,464,1209,1235]" italics="true" pageId="25" pageNumber="26">Babyrousa</emphasis>
Perry, 1811, a genus of enigmatic Suidae endemic to
<collectingRegion id="505AF8029D059242FBA6FB07CCC92CB1" box="[1077,1183,1209,1235]" country="Indonesia" name="Sulawesi Utara" pageId="25" pageNumber="26">Sulawesi</collectingRegion>
and the
<collectingCountry id="EA8976709D059242FB6EFB07CD252CB1" box="[1277,1395,1209,1235]" name="Indonesia" pageId="25" pageNumber="26">Moluccas</collectingCountry>
. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D059242FF2EFA9CCC5A2EBC" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="922136E09D059242FF2EFA9CC93A2D5E" blockId="25.[189,364,1314,1340]" box="[189,364,1314,1340]" pageId="25" pageNumber="26">
<heading id="C969818C9D059242FF2EFA9CC93A2D5E" bold="true" box="[189,364,1314,1340]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="A0EAEAF29D059242FF2EFA9CC93A2D5E" bold="true" box="[189,364,1314,1340]" pageId="25" pageNumber="26">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D059242FF79FAEDC90A2D0F" blockId="25.[189,1398,1363,1503]" box="[234,348,1363,1389]" pageId="25" pageNumber="26">
<emphasis id="A0EAEAF29D059242FF79FAEDC90A2D0F" bold="true" box="[234,348,1363,1389]" pageId="25" pageNumber="26">
<typeStatus id="4D2588429D059242FF79FAEDC90A2D0F" box="[234,348,1363,1389]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D059242FF2EFAC7CB092DBC" blockId="25.[189,1398,1363,1503]" pageId="25" pageNumber="26">
<materialsCitation id="22F63CBD9D059242FF2EFAC7CB092DBC" ID-GBIF-Occurrence="2898878564" collectingDate="1985-05-30" collectingDateMax="1985-06-02" collectingDateMin="1985-05-30" collectionCode="BMNH" country="Indonesia" elevation="1780" location="Kotamobagu" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="SULAWESI UTARA" typeStatus="holotype">
<collectingCountry id="EA8976709D059242FF2EFAC7C90D2DF1" box="[189,347,1401,1427]" name="Indonesia" pageId="25" pageNumber="26">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D059242FEE0FAC7C9DD2DF1" box="[371,395,1401,1427]" pageId="25" pageNumber="26" type="male">♂</specimenCount>
; “
<collectingCountry id="EA8976709D059242FE31FAC7CA1F2DF1" box="[418,585,1401,1427]" name="Indonesia" pageId="25" pageNumber="26">INDONESIA</collectingCountry>
:
<collectingRegion id="505AF8029D059242FDC0FAC7CB1B2DF1" box="[595,845,1401,1427]" country="Indonesia" name="Sulawesi Utara" pageId="25" pageNumber="26">SULAWESI UTARA</collectingRegion>
Gng. Ambang F. R. nr
<location id="9741603B9D059242FBCFFAC7CCA22DF1" LSID="urn:lsid:plazi:treatment:1A3787F69D059247FDF3FD53CAE129DC:9741603B9D059242FBCFFAC7CCA22DF1" box="[1116,1268,1401,1427]" country="Indonesia" name="Kotamobagu" pageId="25" pageNumber="26" stateProvince="SULAWESI UTARA">Kotamobagu</location>
<date id="E62010209D059242FB69FAC7C8AD2DDB" pageId="25" pageNumber="26" value="1985-05-30" valueMax="1985-06-02" valueMin="1985-05-30">
<collectingDate id="F664E9C89D059242FB69FAC7C8AD2DDB" pageId="25" pageNumber="26" value="1985-05-30" valueMax="1985-06-02" valueMin="1985-05-30">30.v – 2.vi 1985</collectingDate>
</date>
. // Gng Muajat summit area ca
<quantity id="55669B059D059242FDFFFA20CAE82DDA" box="[620,702,1438,1464]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="25" pageNumber="26" unit="m" value="1780.0">
<elevation id="19B3D1D39D059242FDFFFA20CAE82DDA" box="[620,702,1438,1464]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="25" pageNumber="26" unit="m" value="1780.0">1780m</elevation>
</quantity>
// light [h, upper side of label] R.Ent.Soc.Lond. PROJECT WALLACE B.M. 1985-10 [underside of label]”;
<collectionCode id="F48FAE259D059242FD6CFA7ACB0D2DBC" box="[767,859,1476,1502]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="25" pageNumber="26">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D059242FF79F9B6CA742E40" blockId="25.[189,1399,1543,1758]" box="[234,546,1543,1570]" pageId="25" pageNumber="26">
<emphasis id="A0EAEAF29D059242FF79F9B6C9312E40" bold="true" box="[234,359,1544,1570]" pageId="25" pageNumber="26">
<typeStatus id="4D2588429D059242FF79F9B6C9312E40" box="[234,359,1544,1570]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="8498FD699D059242FEEBF9B6CA4C2E40" box="[376,538,1543,1570]" pageId="25" pageNumber="26" type="generic">13 specimens</specimenCount>
)
</paragraph>
<paragraph id="922136E09D059242FF2EF993CC5A2EBC" blockId="25.[189,1399,1543,1758]" pageId="25" pageNumber="26">
<materialsCitation id="22F63CBD9D059242FF2EF993CBBD2E2A" ID-GBIF-Occurrence="2898878392" box="[189,1003,1581,1608]" collectionCode="BMNH" country="Indonesia" location="INDONESIA" pageId="25" pageNumber="26" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="holotype">
<collectingCountry id="EA8976709D059242FF2EF993C90D2E25" box="[189,347,1581,1607]" name="Indonesia" pageId="25" pageNumber="26">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D059242FE13F993C99D2E2A" box="[384,459,1581,1608]" pageId="25" pageNumber="26" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="8498FD699D059242FE4EF993CA472E2A" box="[477,529,1581,1608]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
; same labels as for holotype;
<collectionCode id="F48FAE259D059242FC00F990CBBD2E2A" box="[915,1003,1582,1608]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="25" pageNumber="26">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D059242FB83F993CB262EF1" ID-GBIF-Occurrence="2898878486" collectingDate="1985-05-18" collectingDateMax="1985-05-24" collectingDateMin="1985-05-18" collectingMethod="Yellow pan trap" collectionCode="BMNH" country="Indonesia" location="Kotamobagu" municipality="Kotamobagu" pageId="25" pageNumber="26" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="SULAWESI UTARA" typeStatus="holotype">
<specimenCount id="8498FD699D059242FB83F993CC0D2E2A" box="[1040,1115,1581,1608]" pageId="25" pageNumber="26" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="8498FD699D059242FBFEF993CCE12E2A" box="[1133,1207,1581,1608]" pageId="25" pageNumber="26" type="female">6 ♀♀</specimenCount>
; “
<collectingCountry id="EA8976709D059242FB46F993CD202E25" box="[1237,1398,1581,1607]" name="Indonesia" pageId="25" pageNumber="26">INDONESIA</collectingCountry>
<collectingRegion id="505AF8029D059242FF2EF9EDC9EF2E0F" box="[189,441,1619,1645]" country="Indonesia" name="Sulawesi Utara" pageId="25" pageNumber="26">SULAWESI UTARA</collectingRegion>
Gng. Ambang F.R. nr.
<collectingMunicipality id="7245AC9A9D059242FD42F9EDCB3F2E0F" box="[721,873,1619,1645]" pageId="25" pageNumber="26">Kotamobagu</collectingMunicipality>
<date id="E62010209D059242FCE1F9EDCC6A2E0F" box="[882,1084,1618,1645]" pageId="25" pageNumber="26" value="1985-05-18" valueMax="1985-05-24" valueMin="1985-05-18">
<collectingDate id="F664E9C89D059242FCE1F9EDCC6A2E0F" box="[882,1084,1618,1645]" pageId="25" pageNumber="26" value="1985-05-18" valueMax="1985-05-24" valueMin="1985-05-18">18-24 May 1985</collectingDate>
</date>
. //
<collectingMethod id="4BDF4EF79D059242FBC4F9EDCD422E0F" box="[1111,1300,1619,1645]" pageId="25" pageNumber="26">Yellow pan trap</collectingMethod>
// R.Ent. Soc.Lond. PROJECT WALLACE B.M. 1985-10”;
<collectionCode id="F48FAE259D059242FC8BF9C7CB262EF1" box="[792,880,1657,1683]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="25" pageNumber="26">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D059242FC18F9C6CC5E2EBC" ID-GBIF-Occurrence="2898878328" collectingDate="1985-05-18" collectingDateMax="1985-05-24" collectingDateMin="1985-05-18" collectingMethod="Yellow pan trap" collectionCode="BMNH" country="Indonesia" elevation="1765" isTailEnumeration="true" location="Kotamobagu" municipality="Kotamobagu" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="SULAWESI UTARA" typeStatus="holotype">
<specimenCount id="8498FD699D059242FC18F9C6CBEF2EF1" box="[907,953,1656,1683]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
; “
<collectingCountry id="EA8976709D059242FC41F9C6CC252EF0" box="[978,1139,1656,1682]" name="Indonesia" pageId="25" pageNumber="26">INDONESIA</collectingCountry>
<collectingRegion id="505AF8029D059242FBE8F9C6CD202EF0" box="[1147,1398,1656,1682]" country="Indonesia" name="Sulawesi Utara" pageId="25" pageNumber="26">SULAWESI UTARA</collectingRegion>
Gng. Ambang F.R. nr.
<collectingMunicipality id="7245AC9A9D059242FE45F920CA382EDA" box="[470,622,1694,1720]" pageId="25" pageNumber="26">Kotamobagu</collectingMunicipality>
<date id="E62010209D059242FDEAF920CB1E2EDB" box="[633,840,1694,1721]" pageId="25" pageNumber="26" value="1985-05-18" valueMax="1985-05-24" valueMin="1985-05-18">
<collectingDate id="F664E9C89D059242FDEAF920CB1E2EDB" box="[633,840,1694,1721]" pageId="25" pageNumber="26" value="1985-05-18" valueMax="1985-05-24" valueMin="1985-05-18">18-24 May 1985</collectingDate>
</date>
.// Gng. Muajat
<quantity id="55669B059D059242FB9BF920CCFF2EDA" box="[1032,1193,1694,1721]" metricMagnitude="3" metricUnit="m" metricValue="1.765" metricValueMax="1.78" metricValueMin="1.75" pageId="25" pageNumber="26" unit="m" value="1765.0" valueMax="1780.0" valueMin="1750.0">
<elevation id="19B3D1D39D059242FB9BF920CCFF2EDA" box="[1032,1193,1694,1721]" metricMagnitude="3" metricUnit="m" metricValue="1.765" metricValueMax="1.78" metricValueMin="1.75" pageId="25" pageNumber="26" unit="m" value="1765.0" valueMax="1780.0" valueMin="1750.0">1750-1780 m</elevation>
</quantity>
[h]//
<collectingMethod id="4BDF4EF79D059242FB7EF920C8BD2EBC" pageId="25" pageNumber="26">Yellow pan trap</collectingMethod>
// R.Ent.Soc.Lond. PROJECT WALLACE B.M. 1985-10.”;
<collectionCode id="F48FAE259D059242FC3FF97ACC5E2EBC" box="[940,1032,1732,1758]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="25" pageNumber="26">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D059240FF2EF8B9CA282F8A" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="description">
<paragraph id="922136E09D059242FF2EF8B9C9182F43" blockId="25.[189,334,1799,1825]" box="[189,334,1799,1825]" pageId="25" pageNumber="26">
<heading id="C969818C9D059242FF2EF8B9C9182F43" bold="true" box="[189,334,1799,1825]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="A0EAEAF29D059242FF2EF8B9C9182F43" bold="true" box="[189,334,1799,1825]" pageId="25" pageNumber="26">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D059242FF2EF887CC062F8B" blockId="25.[189,1399,1848,2025]" pageId="25" pageNumber="26">
MEASUREMENTS (exposed abdominal ventrites not included). Male
<typeStatus id="4D2588429D059242FC50F886CC7B2F30" box="[963,1069,1848,1874]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
, total body length
<quantity id="55669B059D059242FA94F886CD242F31" box="[1287,1394,1848,1875]" metricMagnitude="-3" metricUnit="m" metricValue="6.95" pageId="25" pageNumber="26" unit="mm" value="6.95">6.95 mm</quantity>
. Head
<quantity id="55669B059D059242FE97F8E3C9382F1A" box="[260,366,1885,1912]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="25" pageNumber="26" unit="mm" value="1.25">1.25 mm</quantity>
long, across eyes
<quantity id="55669B059D059242FDD9F8E3CAE22F1A" box="[586,692,1885,1912]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="25" pageNumber="26" unit="mm" value="1.6">1.60 mm</quantity>
wide. Pronotum
<quantity id="55669B059D059242FC10F8E0CBBB2F1A" box="[899,1005,1886,1912]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="25" pageNumber="26" unit="mm" value="1.4">1.40 mm</quantity>
long, maximum width
<quantity id="55669B059D059242FA96F8E0CD252F1A" box="[1285,1395,1886,1912]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="25" pageNumber="26" unit="mm" value="2.4">2.40 mm</quantity>
. Elytral length
<quantity id="55669B059D059242FEFEF83DC98B2FFC" box="[365,477,1923,1950]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="25" pageNumber="26" unit="mm" value="4.3">4.30 mm</quantity>
, maximum combined width
<quantity id="55669B059D059242FCAFF83DCBFD2FFC" box="[828,939,1923,1950]" metricMagnitude="-3" metricUnit="m" metricValue="3.34" pageId="25" pageNumber="26" unit="mm" value="3.34">3.34 mm</quantity>
. Selected female
<typeStatus id="4D2588429D059242FB16F83ACCBB2FFC" box="[1157,1261,1924,1950]" pageId="25" pageNumber="26" type="paratype">paratype</typeStatus>
, total body length
<quantity id="55669B059D059242FE9CF817C92A2FA1" box="[271,380,1961,1987]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="25" pageNumber="26" unit="mm" value="8.2">8.20 mm</quantity>
. Head
<quantity id="55669B059D059242FE5DF817CA6E2FA1" box="[462,568,1961,1988]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="26" unit="mm" value="1.5">1.50 mm</quantity>
long, across eyes
<quantity id="55669B059D059242FC80F817CB2A2FA1" box="[787,892,1961,1987]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="25" pageNumber="26" unit="mm" value="1.7">1.70 mm</quantity>
wide. Pronotum
<quantity id="55669B059D059242FBD9F817CCE52FA1" box="[1098,1203,1961,1987]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="25" pageNumber="26" unit="mm" value="1.7">1.70 mm</quantity>
long, maximum width
<quantity id="55669B059D059242FE94F870C9252F8B" box="[263,371,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="25" pageNumber="26" unit="mm" value="2.7">2.70 mm</quantity>
. Elytral length 5.00 mm, maximum combined width
<quantity id="55669B059D059242FC73F870CC1A2F8B" box="[992,1100,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="3.52" pageId="25" pageNumber="26" unit="mm" value="3.52">3.52 mm</quantity>
.
</paragraph>
<caption id="C6E166689D069241FF2EF836C94B2F8A" ID-DOI="http://doi.org/10.5281/zenodo.4122134" ID-Zenodo-Dep="4122134" httpUri="https://zenodo.org/record/4122134/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,241,1928,1954]" targetBox="[381,1192,273,1894]" targetPageId="26">
<paragraph id="922136E09D069241FF2EF836C94B2F8A" blockId="26.[189,1399,1928,2025]" pageId="26" pageNumber="27">
<emphasis id="A0EAEAF29D069241FF2EF836C9192FC0" bold="true" box="[189,335,1928,1954]" pageId="26" pageNumber="27">Figs 25–27.</emphasis>
Papuan and Wallacean
<taxonomicName id="559E4D639D069241FDECF836CB9D2FC0" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[639,971,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D069241FDECF836CB5E2FC0" box="[639,776,1928,1954]" italics="true" pageId="26" pageNumber="27">Maechidius</emphasis>
<bibRefCitation id="F60F4B119D069241FC85F836CB9D2FC0" author="Macleay W. S." box="[790,971,1928,1954]" pageId="26" pageNumber="27" refId="ref98319" refString="Macleay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. Containing General Observations on the Geography, Manners, and the Affinities of the Insects which compose the Genus Scarabaeus of Linnaeus; to which are added a few incidental Remarks on the Genera Lucanus and Hister of the same Author. With Appendix and Plates. R. and A. Taylor, London." type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
, habitus, dorsal and lateral views.
<emphasis id="A0EAEAF29D069241FF2EF815C88C2FA7" bold="true" box="[189,218,1963,1989]" pageId="26" pageNumber="27">25</emphasis>
.
<taxonomicName id="559E4D639D069241FF7AF812C9DA2FA7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[233,396,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="babyrousa" status="sp. nov.">
<emphasis id="A0EAEAF29D069241FF7AF812C9DA2FA7" box="[233,396,1963,1989]" italics="true" pageId="26" pageNumber="27">M. babyrousa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D069241FE00F812C9A62FA4" box="[403,496,1964,1990]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (BMNH).
<emphasis id="A0EAEAF29D069241FC99F815CB052FA7" bold="true" box="[778,851,1963,1989]" pageId="26" pageNumber="27">26–27</emphasis>
.
<taxonomicName id="559E4D639D069241FCF1F812CBB52FA7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[866,995,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bintang" status="sp. nov.">
<emphasis id="A0EAEAF29D069241FCF1F812CBB52FA7" box="[866,995,1963,1989]" italics="true" pageId="26" pageNumber="27">M. bintang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D069241FC79F812CC1E2FA4" box="[1002,1096,1964,1990]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NME). Not to scale.
</paragraph>
</caption>
<paragraph id="922136E09D079240FF2EFEB7C9912F6A" blockId="27.[189,1399,265,1800]" pageId="27" pageNumber="28">
Dorsum uniformly black to black-brown (elytra somewhat paler brown than forebody) with castaneous labroclypeus, mouthparts, legs and venter. Head transverse, trapezoid, slightly convex dorsally between eyes, glossy dorsally and ventrally, with large not prominent eyes occupying over half side of head. Male labroclypeus (
<figureCitation id="0AA52A659D079240FEF5FEC9C99A29F0" box="[358,460,375,402]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="27" pageNumber="28">Fig. 106</figureCitation>
) slightly emarginate on anterior margin, with lateral margins smooth, slightly sinuous in both dorsal and lateral views. Anterolateral angles not protruding anteriad, strongly obtuse, bent up almost perpendicularly to axis of frons in lateral view. Female labroclypeus (
<figureCitation id="0AA52A659D079240FB29FE7FCD7629B9" box="[1210,1312,449,475]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="27" pageNumber="28">Fig. 107</figureCitation>
) rather deeply emarginate on anterior margin, with slightly protruding, strongly obtuse anterolateral angles. Punctures circular to hexagonal. Intervening spaces glossy, smaller than up to as large as punctures. Pubescence dirty yellowish, moderately long and suberect, each seta rises from anterior margin of corresponding puncture. Labroclypeus laterally and frontally with more delicate setae. Few longest setae present near eyes. Male antenna 9-segmented, club 3-lamellate. Scape large, bulbous dorsally in distal half, provided with few long erect setae on posterior and distal margins. Antennomere 2 short, slightly transverse. Pronotum transverse, glossy dorsally and laterally. Anterior margin slightly sinuous with anterolateral angles (stronger) and mesal portion (slightly) protruding anteriad. Basal margin broadly rounded. Lateral margin of pronotum evenly broadly rounded, delicately crenulate all along (
<figureCitation id="0AA52A659D079240FF55FC8FC9792B29" box="[198,303,817,843]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="27" pageNumber="28">Fig. 203</figureCitation>
). Pronotal disc with two vague circular postmedian impressions on either side of disc (this feature not visible in
<typeStatus id="4D2588429D079240FE2EFCEBCA7F2B12" box="[445,553,853,880]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
). Lateral margin of pronotum hardly sinuous in lateral view. Punctures of pronotum ovoid to circular, deep and coarse. Intervening spaces glossy and glabrous, smaller than punctures on disc, becoming larger along lateral margins. Setae shorter than on head, stronger curved. Clavate scale-like seta present between every two crenulae on lateral margin of pronotum. Lateral and basal margins, antero- and posterolateral angles partly covered with microscopical velvety pubescence. Hypomeron smooth, sparsely covered by circular punctures. Hypomeron flange-like, with large emargination opposite to eye (to receive canthus separating compound eye) covered with long setae. Antennal pocket deep. Scutellar shield triangular, narrow apically. Elytra cylindrical, maximum width in median third, glossy and slightly convex dorsally, with rounded humeri and slightly elevated omoplates. Indistinct track of one–two flat longitudinal carinae on each elytron. Sutural carinae not indicated. Rather large obtuse hump on each elytron near apex. Disc and lateral sides of elytra irregularly densely punctured, punctures of irregular size and form (circular to ovoid) (
<figureCitation id="0AA52A659D079240FC58FB54CC672D66" box="[971,1073,1258,1284]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="27" pageNumber="28">Fig. 293</figureCitation>
). Intervening spaces glossy, smaller than punctures. Moderately long suberect setae arranged in irregular longitudinal paired rows. Inconspicuous very short setae not surpassing length of corresponding punctures positioned between longitudinal rows of longer setae. Each seta rises from anterior margin of each puncture. Epipleuron with row or two (in broad anterior part) of suberect setae. Male pygidium with shallow annular punctures and glossy intervening spaces (
<figureCitation id="0AA52A659D079240FDB8FA1CCAC52DDF" box="[555,659,1442,1469]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="27" pageNumber="28">Fig. 458</figureCitation>
). Female pygidium similarly punctured as in male, with obtuse transverse median hump (
<figureCitation id="0AA52A659D079240FE61FA79CA0A2D83" box="[498,604,1478,1505]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="27" pageNumber="28">Fig. 459</figureCitation>
). Seta rises from centre of each puncture of pygidium. Abdominal ventrites glossy, covered with large annular punctures, each provided with short seta. Legs slender, femora and tibiae covered with delicate whitish setae. Outer margins of meso- and metatibiae densely denticulate. Male protibia with one (apical) strong tooth on external margin, female protibia with two teeth (
<figureCitation id="0AA52A659D079240FE95F9E4C9F82E17" box="[262,430,1626,1653]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="27" pageNumber="28">Figs 449–450</figureCitation>
). Male lower meso- and metatibial terminal spurs strongly curved (
<figureCitation id="0AA52A659D079240FB54F9E4CD3C2E16" box="[1223,1386,1626,1652]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="27" pageNumber="28">Fig. 439–440</figureCitation>
), female ones straight (
<figureCitation id="0AA52A659D079240FE57F9C0CA792EFB" box="[452,559,1662,1689]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="27" pageNumber="28">Fig. 438</figureCitation>
). Distal margin of metatibia in both sexes with projection over basal metatarsomere. Metatibial terminal spurs paired, shorter than basal metatarsomere in male but almost as long as in female. Tarsal claws with pulvilli in both sexes. Male aedeagus as in
<figureCitation id="0AA52A659D079240FBCBF976CCAA2E81" box="[1112,1276,1736,1763]" captionStart="Figs 570–585" captionStartId="170.[189,241,1892,1918]" captionTargetBox="[425,1155,265,1858]" captionTargetId="figure-74@170.[416,1172,265,1864]" captionTargetPageId="170" captionText="Figs 570–585. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 570– 572. M. aiyura sp. nov., holotype, ♂ (BMNH). 573–575. M. alesbezdeki sp. nov., holotype, ♂ (NME). 576–578. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 579–581. M. awu sp. nov., holotype, ♂ (IECA). 582–585. M. babyrousa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122232" httpUri="https://zenodo.org/record/4122232/files/figure.png" pageId="27" pageNumber="28">Figs 582–585</figureCitation>
. Spiculum gastrale as in Fig. 538.
</paragraph>
<paragraph id="922136E09D079240FF2EF88EC9E62F28" blockId="27.[189,432,1840,1866]" box="[189,432,1840,1866]" pageId="27" pageNumber="28">
<heading id="C969818C9D079240FF2EF88EC9E62F28" bold="true" box="[189,432,1840,1866]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="A0EAEAF29D079240FF2EF88EC9E62F28" bold="true" box="[189,432,1840,1866]" pageId="27" pageNumber="28">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D079240FF2EF8DECA282F8A" blockId="27.[189,1399,1888,2025]" pageId="27" pageNumber="28">Female is generally larger, with shorter lamellae of antennal club; protibial spur slightly curved distally in male, straight in female; metatibial spurs somewhat longer in female than in male; protibia bidentate in female; female pygidium with modifications (see above); female lower meso- and metatibial terminal spurs slender, slightly curved apically.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D009247FF2EFEB7CD7E2932" box="[189,1320,265,337]" pageId="28" pageNumber="29" type="biology_ecology">
<paragraph id="922136E09D009247FF2EFEB7C9762941" blockId="28.[189,288,265,291]" box="[189,288,265,291]" pageId="28" pageNumber="29">
<heading id="C969818C9D009247FF2EFEB7C9762941" bold="true" box="[189,288,265,291]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="A0EAEAF29D009247FF2EFEB7C9762941" bold="true" box="[189,288,265,291]" pageId="28" pageNumber="29">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D009247FF2EFE88CD7E2932" blockId="28.[189,1320,310,337]" box="[189,1320,310,337]" pageId="28" pageNumber="29">Occurs in lowland rainforests. Attracted to yellow pan traps (anthophilous?). Possibly nocturnal.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D009247FF2EFEC8CAE129DC" box="[189,695,374,446]" pageId="28" pageNumber="29" type="distribution">
<paragraph id="922136E09D009247FF2EFEC8C90029F2" blockId="28.[189,342,374,400]" box="[189,342,374,400]" pageId="28" pageNumber="29">
<heading id="C969818C9D009247FF2EFEC8C90029F2" bold="true" box="[189,342,374,400]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="A0EAEAF29D009247FF2EFEC8C90029F2" bold="true" box="[189,342,374,400]" pageId="28" pageNumber="29">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D009247FF2EFE1ACAE129DC" blockId="28.[189,695,420,446]" box="[189,695,420,446]" pageId="28" pageNumber="29">
Hitherto only known from
<collectingRegion id="505AF8029D009247FE6AFE1ACAE529DC" box="[505,691,420,446]" country="Indonesia" name="Sulawesi Utara" pageId="28" pageNumber="29">North Sulawesi</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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