380 lines
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380 lines
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.148.50042" ID-GBIF-Dataset="1224cf7b-1d2d-4657-89e9-11fee1f5ea0b" ID-PMC="PMC7266835" ID-Pensoft-Pub="1314-2003-148-21" ID-Pensoft-UUID="6E288E0A8F725B2787799DD17F2B1642" ID-PubMed="32523392" ModsDocID="1314-2003-148-21" checkinTime="1590560163243" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Peterson, Paul M., Romaschenko, Konstantin, Bravo-Pedraza, William J., Cuta-Alarcon, Lia E. & Soreng, Robert J." docDate="2020" docId="1819DFBEDDA25CD8BEF5C4583457B9CE" docLanguage="en" docName="PhytoKeys 148: 21-50" docOrigin="PhytoKeys 148" docSource="http://dx.doi.org/10.3897/phytokeys.148.50042" docTitle="Podagrostis liebmannii Sylvester & Soreng 2020, comb. nov." docType="treatment" docVersion="4" id="6E288E0A8F725B2787799DD17F2B1642" lastPageNumber="21" masterDocId="6E288E0A8F725B2787799DD17F2B1642" masterDocTitle="New combinations and updated descriptions in Podagrostis (Agrostidinae, Poaceae) from the Neotropics and Mexico" masterLastPageNumber="50" masterPageNumber="21" pageNumber="21" updateTime="1668139801695" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New combinations and updated descriptions in Podagrostis (Agrostidinae, Poaceae) from the Neotropics and Mexico</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sylvester, Steven P.</mods:namePart>
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<mods:affiliation>College of Biology and the Environment, Nanjing Forestry University, Long Pan Road No. 159, Nanjing, 210037, China & Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5577-8782</mods:nameIdentifier>
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<mods:nameIdentifier type="email">steven_sylvester@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Peterson, Paul M.</mods:namePart>
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<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Romaschenko, Konstantin</mods:namePart>
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<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bravo-Pedraza, William J.</mods:namePart>
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<mods:affiliation>Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cuta-Alarcon, Lia E.</mods:namePart>
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<mods:affiliation>Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soreng, Robert J.</mods:namePart>
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<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-8358-4915</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>148</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>21</mods:start>
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<mods:end>50</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.148.50042</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.148.50042</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-148-21</mods:identifier>
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<mods:identifier type="Pensoft-UUID">6E288E0A8F725B2787799DD17F2B1642</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="164360815" LSID="urn:lsid:plazi:treatment:1819DFBEDDA25CD8BEF5C4583457B9CE" httpUri="http://treatment.plazi.org/id/1819DFBEDDA25CD8BEF5C4583457B9CE" lastPageNumber="21" pageId="0" pageNumber="21">
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<subSubSection pageId="0" pageNumber="21" type="nomenclature">
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<paragraph pageId="0" pageNumber="21">
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<taxonomicName LSID="1819DFBE-DDA2-5CD8-BEF5-C4583457B9CE" authority="(E. Fourn.) Sylvester & Soreng" authorityName="Sylvester & Soreng" authorityYear="2020" baseAuthorityName="Sylvester & Soreng" baseAuthorityYear="2020" class="Liliopsida" family="Poaceae" genus="Podagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Podagrostis liebmannii" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="liebmannii" status="comb. nov.">Podagrostis liebmannii (E. Fourn.) Sylvester & Soreng</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="21">comb. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Podagrostis liebmannii. A Whole plant B ligular area of flag leaf C section of inflorescence showing spikelets D floret, ventral view, with the rachilla prolongation indicated with a red arrow. Images of epitype specimen Liebmann s. n. (US 00595641) courtesy of the United States National Herbarium (US)." figureDoi="10.3897/phytokeys.148.50042.figure3" httpUri="https://binary.pensoft.net/fig/413784" pageId="0" pageNumber="21">Fig. 3</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="21" type="reference_group">
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<paragraph pageId="0" pageNumber="21">
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis liebmannii" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="liebmannii">Agrostis liebmannii</taxonomicName>
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(E. Fourn.) Hitchc., N. Amer. Fl. 17(7): 519. 1937.
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Apera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Apera liebmannii" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="liebmannii">Apera liebmannii</taxonomicName>
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E. Fourn., Mexic. Pl. 2: 97. 1886.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="21" type="materials_examined">
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<paragraph pageId="0" pageNumber="21">Type.</paragraph>
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<paragraph pageId="0" pageNumber="21">
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Mexico. [
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<emphasis bold="true" pageId="0" pageNumber="21">Veracruz</emphasis>
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:] Orizaba, M. Botteri 93 in part (
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">lectotype</emphasis>
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, designated here: P (P00740547 [image!]); isolectotypes: MSC fragm. ex P;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">epitype</emphasis>
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, designated here: Mexico. [
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<emphasis bold="true" pageId="0" pageNumber="21">Puebla</emphasis>
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:] Chinantla, May 1841, F.M. Liebmann s.n. "
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<taxonomicName kingdom="Plantae" lsidName="" pageId="0" pageNumber="21" rank="kingdom">Plantae</taxonomicName>
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mexicanae Liebmann #12591" {handwritten on label} (US (US00595641!)).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Syntype</emphasis>
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: Mexico. [
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<emphasis bold="true" pageId="0" pageNumber="21">Puebla</emphasis>
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:] Museum Botanicum Hauniense |
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<taxonomicName kingdom="Plantae" lsidName="" pageId="0" pageNumber="21" rank="kingdom">Plantae</taxonomicName>
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mexicanae Liebmann | 1841-43 {printed label},
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<taxonomicName family="Gramineae" lsidName="" pageId="0" pageNumber="21" rank="family">Gramineae</taxonomicName>
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N. 710 |
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Apera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Apera liebmannii" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="liebmannii">
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<emphasis italics="true" pageId="0" pageNumber="21">Apera liebmannii</emphasis>
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</taxonomicName>
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Fourn. | [determ.] Fournier | Chinantla. May 1841, {handwritten on label} (NY (NY00345814 [image!]), US [not seen]).
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</paragraph>
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<caption doi="10.3897/phytokeys.148.50042.figure3" httpUri="https://binary.pensoft.net/fig/413784" pageId="0" pageNumber="21" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="21">
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<emphasis bold="true" pageId="0" pageNumber="21">Figure 3.</emphasis>
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<taxonomicName authorityName="Sylvester & Soreng" authorityYear="2020" baseAuthorityName="Sylvester & Soreng" baseAuthorityYear="2020" class="Liliopsida" family="Poaceae" genus="Podagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Podagrostis liebmannii" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="liebmannii">
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<emphasis italics="true" pageId="0" pageNumber="21">Podagrostis liebmannii</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="0" pageNumber="21">A</emphasis>
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Whole plant
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<emphasis bold="true" pageId="0" pageNumber="21">B</emphasis>
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ligular area of flag leaf
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<emphasis bold="true" pageId="0" pageNumber="21">C</emphasis>
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section of inflorescence showing spikelets
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<emphasis bold="true" pageId="0" pageNumber="21">D</emphasis>
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floret, ventral view, with the rachilla prolongation indicated with a red arrow. Images of epitype specimen Liebmann s.n. (US00595641) courtesy of the United States National Herbarium (US).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="21" type="description">
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<paragraph pageId="0" pageNumber="21">Description.</paragraph>
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<paragraph pageId="0" pageNumber="21">
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<taxonomicName genus="Tufted" lsidName="Tufted perennial" pageId="0" pageNumber="21" rank="species" species="perennial">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Tufted perennial</emphasis>
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</taxonomicName>
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forming lax tufts, with the basal foliage reaching c. 11 cm tall and inflorescences well-exserted from the basal foliage.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Tillers</emphasis>
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extravaginal.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Culms</emphasis>
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35-80 cm tall, erect, simple, delicate;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">nodes and internodes</emphasis>
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terete, nodes smooth, internodes and segment below the panicle smooth throughout (or smooth proximally and lightly scaberulous towards their apices in Moore 3339), usually with at least 1 or 2 elongated internodes visible, with 1-2 nodes exposed at flowering, uppermost internode c. 5.5-11.2 cm long, longer than the sheath.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Leaves</emphasis>
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basal and cauline;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">sheaths</emphasis>
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terete, glabrous, lower sheaths tending to be smooth, upper sheaths lightly to densely scabrous with short hooks; flag sheath 8-11.5 cm long; basal sheaths 0.5-1 cm long, striate, becoming fibrous, smooth;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">ligules</emphasis>
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c. 0.5-4 mm long, membranaceous or scareous, usually strongly decurrent with the sheath, abaxially scabrous; upper culm ligules 1.5-4 mm long, obtuse to acute, sometimes slightly erose towards the apex; ligules of tillers shorter to those of the culm, c. 0.5-1 mm long, truncate to acute;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">blades</emphasis>
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2-8.6 cm long, (0.5-)1-3.5 mm wide in diameter, flat, flaccid to firm, basal blades sometimes very narrow, abaxial surfaces glabrous, smooth to lightly scabrous, or usually more densely scabrous further up the culm, adaxial surfaces glabrous, smooth to lightly scabrous on the veins, scaberulous further up the culm, edges scaberulous.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Panicles</emphasis>
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7-18
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<normalizedToken originalValue="×">x</normalizedToken>
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(1-)2.5-7 cm, open to slightly congested following anthesis, usually ovoid;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">panicle branches</emphasis>
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ascendant to patent, branched above the middle, filiform, with spikelets not present near the base, smooth, longest branches 2-4.7 cm long;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">pedicels</emphasis>
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1.5-5 mm long, usually longer than the length of the spikelets, divaricate, smooth or lightly scaberulous.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Spikelets</emphasis>
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(1.8-)2-2.1 mm long;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">glumes</emphasis>
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remaining on the inflorescence at maturity, equal or subequal, the lower usually slightly longer than the upper, subequaling the length of the floret to 0.4 mm longer, lanceolate, slightly keeled, apices acute, glabrous, keels lightly scaberulous just in the distal 1/3, surfaces smooth; lower glume 1- (or 3-)veined, lateral veins, if present, vestigial; upper glume 1- (or 3-)veined, lateral veins, if present, vestigial;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">lemmas</emphasis>
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1.7-2 mm long, glabrous, smooth, strongly 5-veined with excurrent prominent veins, apex broadly acute, awn absent;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">paleas</emphasis>
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well-developed, 1.4-1.9 mm long, usually reaching from
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<normalizedToken originalValue="¾">3/4</normalizedToken>
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to subequaling the lemma, keels obscure, smooth, apex bifid and sometimes erose;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">rachilla</emphasis>
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prolonged from the base of the floret, 0.3-0.5 mm long, glabrous, smooth.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Calluses</emphasis>
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not or slightly elongated, 0.05-0.1 mm long, glabrous, smooth.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Flowers; lodicules</emphasis>
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c.0.3-0.4 mm long, lanceolate with acuminate apices, not lobed;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">anthers</emphasis>
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3 in number, 0.6-1 mm long.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Caryopses</emphasis>
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not seen. 2
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<emphasis italics="true" pageId="0" pageNumber="21">n</emphasis>
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= unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="21" type="distribution">
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<paragraph pageId="0" pageNumber="21">Distribution and ecology.</paragraph>
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<paragraph pageId="0" pageNumber="21">
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Mexico, endemic. The authors have only verified specimens from Hidalgo, Puebla and Veracruz states of central Mexico, with
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<bibRefCitation author="Beetle, AA" pageId="0" pageNumber="21" refId="B1" refString="Beetle, AA, 1983. Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." title="Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." year="1983">Beetle (1983)</bibRefCitation>
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,
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<bibRefCitation DOI="https://doi.org/10.17129/botsci.1894" author="Davila, P" journalOrPublisher="Botanical Sciences" pageId="0" pageNumber="21" pagination="462 - 514" refId="B4" refString="Davila, P, Mejia-Saules, MT, Soriano-Martinez, AM, Herrera-Arrieta, Y, 2018. Conocimiento taxonomico de la familia Poaceae en Mexico. Botanical Sciences 96 (3): 462 - 514, DOI: https://doi.org/10.17129/botsci.1894" title="Conocimiento taxonomico de la familia Poaceae en Mexico." url="https://doi.org/10.17129/botsci.1894" volume="96" year="2018">
|
|||
|
|
<normalizedToken originalValue="Dávila">Davila</normalizedToken>
|
|||
|
|
et al. (2018)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
, and
|
|||
|
|
<bibRefCitation DOI="https://doi.org/10.21829/abm126.2019.1379" author="Sanchez-Ken, JG" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="21" refId="B25" refString="Sanchez-Ken, JG, 2019. Riqueza de especies, clasificacion y listado de las gramineas (Poaceae) de Mexico. Acta Botanica Mexicana 126: e1379. https://doi.org/10.21829/abm126.2019.1379" title="Riqueza de especies, clasificacion y listado de las gramineas (Poaceae) de Mexico. Acta Botanica Mexicana 126: e 1379." url="https://doi.org/10.21829/abm126.2019.1379" year="2019">
|
|||
|
|
<normalizedToken originalValue="Sánchez-Ken">Sanchez-Ken</normalizedToken>
|
|||
|
|
(2019)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
mentioning the species to range from Durango state in the north to Oaxaca state in southern Mexico. Found in humid areas of pine and fir forests,
|
|||
|
|
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sphagnum" order="Sphagnales" pageId="0" pageNumber="21" phylum="Bryophyta" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">Sphagnum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
bogs, and by streams, 2100-2300 m (
|
|||
|
|
<bibRefCitation author="Beetle, AA" pageId="0" pageNumber="21" refId="B1" refString="Beetle, AA, 1983. Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." title="Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." year="1983">Beetle 1983</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="21" type="materials_examined">
|
|||
|
|
<paragraph pageId="0" pageNumber="21">Other specimens examined.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="21">
|
|||
|
|
Mexico.
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="21">Hidalgo</emphasis>
|
|||
|
|
: Distrito Zacualtipan, pine woods and
|
|||
|
|
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sphagnum" order="Sphagnales" pageId="0" pageNumber="21" phylum="Bryophyta" rank="genus">Sphagnum</taxonomicName>
|
|||
|
|
bogs about 3 miles from Zacualtipan on road to Tianguistengo, 2100 m alt., 4 July 1947, H.E. Moore, Jr. 3339 (US00486609).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="21" type="notes">
|
|||
|
|
<paragraph pageId="0" pageNumber="21">Notes.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="21">
|
|||
|
|
<bibRefCitation author="Beetle, AA" pageId="0" pageNumber="21" refId="B1" refString="Beetle, AA, 1983. Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." title="Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." year="1983">Beetle (1983)</bibRefCitation>
|
|||
|
|
appears to consider
|
|||
|
|
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis durangensis" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="durangensis">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">Agrostis durangensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Mez a synonym of
|
|||
|
|
<taxonomicName lsidName="A. liebmannii" pageId="0" pageNumber="21" rank="species" species="liebmannii">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">A. liebmannii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and states
|
|||
|
|
<taxonomicName lsidName="A. liebmannii" pageId="0" pageNumber="21" rank="species" species="liebmannii">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">A. liebmannii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
to be distributed as far north as Durango state based on the type locality of
|
|||
|
|
<taxonomicName lsidName="A. durangensis" pageId="0" pageNumber="21" rank="species" species="durangensis">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">A. durangensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. However, we consider
|
|||
|
|
<taxonomicName lsidName="A. durangensis" pageId="0" pageNumber="21" rank="species" species="durangensis">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">A. durangensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
to be a synonym of
|
|||
|
|
<taxonomicName lsidName="A. exarata" pageId="0" pageNumber="21" rank="species" species="exarata">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">A. exarata</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Trin. and have only verified specimens from as far north as Hidalgo state.
|
|||
|
|
<bibRefCitation author="Herrera-Arrieta, Y" journalOrPublisher="Oxford University Press, New York" pageId="0" pageNumber="21" refId="B9" refString="Herrera-Arrieta, Y, 2001. Las gramineas de Durango. Instituto Politecnico Nacional Comision Nacional para el Conocimiento y Uso de la Biodiversidad, D.F., Mexico." title="Las gramineas de Durango. Instituto Politecnico Nacional Comision Nacional para el Conocimiento y Uso de la Biodiversidad, D. F., Mexico." year="2001">Herrera-Arrieta (2001)</bibRefCitation>
|
|||
|
|
cite
|
|||
|
|
<taxonomicName lsidName="A. liebmannii" pageId="0" pageNumber="21" rank="species" species="liebmannii">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">A. liebmannii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
for Durango state based on specimen Palmer 190 (US00486604), which is treated here as
|
|||
|
|
<taxonomicName lsidName="A. exarata" pageId="0" pageNumber="21" rank="species" species="exarata">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">A. exarata</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
<bibRefCitation DOI="https://doi.org/10.21829/abm106.2014.214" author="Herrera-Arrieta, Y" journalOrPublisher="Acta Botanica Mexicana" pageId="0" pageNumber="21" pagination="79 - 95" refId="B10" refString="Herrera-Arrieta, Y, 2014. Additions and updated names for grasses of Durango, Mexico. Acta Botanica Mexicana 106 (106): 79 - 95, DOI: https://doi.org/10.21829/abm106.2014.214" title="Additions and updated names for grasses of Durango, Mexico." url="https://doi.org/10.21829/abm106.2014.214" volume="106" year="2014">Herrera-Arrieta (2014)</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation DOI="https://doi.org/10.17129/botsci.1894" author="Davila, P" journalOrPublisher="Botanical Sciences" pageId="0" pageNumber="21" pagination="462 - 514" refId="B4" refString="Davila, P, Mejia-Saules, MT, Soriano-Martinez, AM, Herrera-Arrieta, Y, 2018. Conocimiento taxonomico de la familia Poaceae en Mexico. Botanical Sciences 96 (3): 462 - 514, DOI: https://doi.org/10.17129/botsci.1894" title="Conocimiento taxonomico de la familia Poaceae en Mexico." url="https://doi.org/10.17129/botsci.1894" volume="96" year="2018">
|
|||
|
|
<normalizedToken originalValue="Dávila">Davila</normalizedToken>
|
|||
|
|
et al. (2018)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
and
|
|||
|
|
<bibRefCitation DOI="https://doi.org/10.21829/abm126.2019.1379" author="Sanchez-Ken, JG" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="21" refId="B25" refString="Sanchez-Ken, JG, 2019. Riqueza de especies, clasificacion y listado de las gramineas (Poaceae) de Mexico. Acta Botanica Mexicana 126: e1379. https://doi.org/10.21829/abm126.2019.1379" title="Riqueza de especies, clasificacion y listado de las gramineas (Poaceae) de Mexico. Acta Botanica Mexicana 126: e 1379." url="https://doi.org/10.21829/abm126.2019.1379" year="2019">
|
|||
|
|
<normalizedToken originalValue="Sánchez-Ken">Sanchez-Ken</normalizedToken>
|
|||
|
|
(2019)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
also mention
|
|||
|
|
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis liebmannii" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="liebmannii">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">Agrostis liebmannii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
to be found as far north as Durango state, but do not include voucher specimens and may have based this on
|
|||
|
|
<bibRefCitation author="Beetle, AA" pageId="0" pageNumber="21" refId="B1" refString="Beetle, AA, 1983. Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." title="Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." year="1983">
|
|||
|
|
<normalizedToken originalValue="Beetle’s">Beetle's</normalizedToken>
|
|||
|
|
(1983)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
and
|
|||
|
|
<bibRefCitation author="Herrera-Arrieta, Y" journalOrPublisher="Oxford University Press, New York" pageId="0" pageNumber="21" refId="B9" refString="Herrera-Arrieta, Y, 2001. Las gramineas de Durango. Instituto Politecnico Nacional Comision Nacional para el Conocimiento y Uso de la Biodiversidad, D.F., Mexico." title="Las gramineas de Durango. Instituto Politecnico Nacional Comision Nacional para el Conocimiento y Uso de la Biodiversidad, D. F., Mexico." year="2001">
|
|||
|
|
<normalizedToken originalValue="Herrera-Arrieta’s">Herrera-Arrieta's</normalizedToken>
|
|||
|
|
(2001)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
treatments. Certain characteristics in
|
|||
|
|
<bibRefCitation author="Beetle, AA" pageId="0" pageNumber="21" refId="B1" refString="Beetle, AA, 1983. Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." title="Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." year="1983">
|
|||
|
|
<normalizedToken originalValue="Beetle’s">Beetle's</normalizedToken>
|
|||
|
|
(1983
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 82) description of
|
|||
|
|
<taxonomicName lsidName="A. liebmannii" pageId="0" pageNumber="21" rank="species" species="liebmannii">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">A. liebmannii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
also do not fit the specimens examined, which may be due to the
|
|||
|
|
<normalizedToken originalValue="author’s">author's</normalizedToken>
|
|||
|
|
inclusion of
|
|||
|
|
<taxonomicName lsidName="A. durangensis" pageId="0" pageNumber="21" rank="species" species="durangensis">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">A. durangensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in the species circumscription. Spikelet size of 2.5 mm mentioned by
|
|||
|
|
<bibRefCitation author="Beetle, AA" pageId="0" pageNumber="21" refId="B1" refString="Beetle, AA, 1983. Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." title="Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." year="1983">Beetle (1983</bibRefCitation>
|
|||
|
|
: 82) does not fit the specimens studied which had spikelets (1.8-)2-2.1 mm long, although
|
|||
|
|
<bibRefCitation author="Beetle, AA" pageId="0" pageNumber="21" refId="B1" refString="Beetle, AA, 1983. Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." title="Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." year="1983">
|
|||
|
|
<normalizedToken originalValue="Beetle’s">Beetle's</normalizedToken>
|
|||
|
|
(1983
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 73) key to species separates
|
|||
|
|
<taxonomicName lsidName="A. liebmannii" pageId="0" pageNumber="21" rank="species" species="liebmannii">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">A. liebmannii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from
|
|||
|
|
<taxonomicName lsidName="A. hiemalis subsp. var. var. laxiflora" pageId="0" pageNumber="21" rank="variety" species="hiemalis" subSpecies="var." variety="laxiflora">A. hiemalis var. laxiflora</taxonomicName>
|
|||
|
|
(Michx.) Beetle (=
|
|||
|
|
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis scabra" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="scabra">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">Agrostis scabra</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Willd.) based on spikelets being c. 2 mm long and shorter than 2.5 mm long.
|
|||
|
|
<bibRefCitation author="Beetle, AA" pageId="0" pageNumber="21" refId="B1" refString="Beetle, AA, 1983. Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." title="Las Gramineas de Mexico. Secretaria de Agricultura y Recursos Hidraulicos, COTECOCA." year="1983">
|
|||
|
|
<normalizedToken originalValue="Beetle’s">Beetle's</normalizedToken>
|
|||
|
|
(1983)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
mention that the tiller blades are more-or-less involute was also not seen, although specimen Moore 3339 (US00486609) was intermixed with another species with involute blades, which might explain this.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="21">
|
|||
|
|
<taxonomicName authorityName="Sylvester & Soreng" authorityYear="2020" baseAuthorityName="Sylvester & Soreng" baseAuthorityYear="2020" class="Liliopsida" family="Poaceae" genus="Podagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Podagrostis liebmannii" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="liebmannii">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">Podagrostis liebmannii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
bears close affinity to
|
|||
|
|
<taxonomicName lsidName="P. thurberiana" pageId="0" pageNumber="21" rank="species" species="thurberiana">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">P. thurberiana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, a North American species that is found as far south as California (
|
|||
|
|
<bibRefCitation author="Hitchcock, CL" journalOrPublisher="Contributions from the United States National Herbarium" pageId="0" pageNumber="21" refId="B13" refString="Hitchcock, CL, Cronquist, A, Ownbey, M, 1969. Vascular Plants of the Pacific Northwest, Part 1: Vascular Cryptogams, Gymnosperms, and Monocotyledons. University of Washington Press, Seattle." title="Vascular Plants of the Pacific Northwest, Part 1: Vascular Cryptogams, Gymnosperms, and Monocotyledons. University of Washington Press, Seattle." year="1969">Hitchcock et al. 1969</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation author="Harvey, MJ" editor="Barkworth, ME" journalOrPublisher="Oxford University Press, New York" pageId="0" pageNumber="21" pagination="693 - 695" refId="B8" refString="Harvey, MJ, 2007. 14.42 Podagrostis (Griseb.) Scribn. & Merr. In: Barkworth, ME, Capels, KM, Long, S, Anderton, LK, Piep, MB, Eds., Flora of North America Volume 24 Magnoliophyta: Commelinidae (in part): Poaceae, part 1. Oxford University Press, New York: 693 - 695" title="14.42 Podagrostis (Griseb.) Scribn. & Merr." volumeTitle="Flora of North America Volume 24 Magnoliophyta: Commelinidae (in part): Poaceae, part 1." year="2007">Harvey 2007</bibRefCitation>
|
|||
|
|
). These similarities include a) the overall habit, with tall culms, loosely tufted and subrhizomatous habit with extravaginal shoots; b) spikelet morphology, with spikelets usually <2.3 mm long, lemmas with excurrent prominent veins and paleas almost subequalling the lemma; c) panicles open and becoming slightly congested following anthesis; d) flat leaf blades.
|
|||
|
|
<taxonomicName authorityName="Sylvester & Soreng" authorityYear="2020" baseAuthorityName="Sylvester & Soreng" baseAuthorityYear="2020" class="Liliopsida" family="Poaceae" genus="Podagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Podagrostis liebmannii" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="liebmannii">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">Podagrostis liebmannii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
can be distinguished from the aforementioned species by a) panicles generally much larger, 8-18
|
|||
|
|
<normalizedToken originalValue="×">x</normalizedToken>
|
|||
|
|
(1-)2.5-7 cm, with patent panicle branches at anthesis (vs. 5-14
|
|||
|
|
<normalizedToken originalValue="×">x</normalizedToken>
|
|||
|
|
0.2-3 cm, panicle branches usually ascending at anthesis in
|
|||
|
|
<taxonomicName lsidName="P. thurberiana" pageId="0" pageNumber="21" rank="species" species="thurberiana">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">P. thurberiana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
); b) callus glabrous (vs. with short hairs to 0.5 mm long emerging from the basal side-ridges of the callus in
|
|||
|
|
<taxonomicName lsidName="P. thurberiana" pageId="0" pageNumber="21" rank="species" species="thurberiana">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">P. thurberiana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
); c) rachilla glabrous, smooth (vs. short hairs to 0.3 mm long emerging from the apex of the rachilla in
|
|||
|
|
<taxonomicName lsidName="P. thurberiana" pageId="0" pageNumber="21" rank="species" species="thurberiana">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">P. thurberiana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="21">
|
|||
|
|
<bibRefCitation author="Fournier, EPN" journalOrPublisher="Botanical Sciences" pageId="0" pageNumber="21" refId="B5" refString="Fournier, EPN, 1886. Mexicanas Plantas 2. Ex Typographeo Reipublicae, Paris." title="Mexicanas Plantas 2. Ex Typographeo Reipublicae, Paris." year="1886">Fournier (1886</bibRefCitation>
|
|||
|
|
: 97) cited two specimens, Liebmann 710 and Botteri 93, in the protologue but only indicated the P herbarium for the Botteri 93 specimen. As we are sure that Fournier saw the Botteri 93 specimen at P, we lectotypify on this collection. Because both the Botteri 93 and Liebmann 710 material included just an inflorescence culm and part of the flag leaf and did not show the basal parts of the plant, we also epitypify the lectotype on Liebmann s.n. (US00595641) that was collected on the same date and at the same locality as the Liebmann 710 collection and includes the basal portion of the plant to help interpret the lectotype. The epitype Liebmann s.n. (US00595641) cited here may in fact be a duplicate of the Liebmann 710 syntype that Fournier (if he ever saw it) did not annotate at the Copenhagen herbarium. The US specimen identified as
|
|||
|
|
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis liebmannii" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="liebmannii">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">Agrostis liebmannii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
("
|
|||
|
|
<taxonomicName class="Liliopsida" family="Poaceae" genus="Apera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Apera" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="21">Apera</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
" not mentioned) was catalogued as
|
|||
|
|
<taxonomicName kingdom="Plantae" lsidName="" pageId="0" pageNumber="21" rank="kingdom">Plantae</taxonomicName>
|
|||
|
|
mexicanae Liebmann #12591; these catalogue numbers are often followed by
|
|||
|
|
<normalizedToken originalValue="Liebmann’s">Liebmann's</normalizedToken>
|
|||
|
|
field numbers such as the handwritten
|
|||
|
|
<normalizedToken originalValue="“710”">"710"</normalizedToken>
|
|||
|
|
. The Copenhagen herbarium does not have a Liebmann duplicate bearing either of these numbers, or either of the synonyms.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|