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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E7868FFB7FF013BD7ADA7ECD9" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Hylekobolus analavelona Wesener &amp; Enghoff &amp; Sierwald 2009, sp. n." docType="treatment" docUuid="769102C4-C297-4628-85AA-B87EEB70DF3E" docUuidSource="ZooBank" docVersion="11" lastPageNumber="114" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="112" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesener, Thomas</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierwald, Petra</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>19</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791473" ID-GBIF-Taxon="163747606" ID-Zenodo-Dep="3791473" LSID="urn:lsid:zoobank.org:act:769102C4-C297-4628-85AA-B87EEB70DF3E" httpUri="http://treatment.plazi.org/id/1772122E7868FFB7FF013BD7ADA7ECD9" lastPageId="113" lastPageNumber="114" pageId="111" pageNumber="112">
<subSubSection box="[140,623,1187,1214]" pageId="111" pageNumber="112" type="nomenclature">
<paragraph blockId="111.[140,969,1187,1249]" box="[140,623,1187,1214]" pageId="111" pageNumber="112">
<heading box="[140,623,1187,1214]" fontSize="11" level="2" pageId="111" pageNumber="112" reason="3">
<emphasis bold="true" box="[140,623,1187,1214]" pageId="111" pageNumber="112">
<taxonomicName authority="Wesener" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,546,1187,1214]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis bold="true" box="[140,438,1187,1214]" italics="true" pageId="111" pageNumber="112">Hylekobolus analavelona</emphasis>
Wesener
</taxonomicName>
,
<taxonomicNameLabel box="[558,623,1187,1213]" pageId="111" pageNumber="112" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,969,1222,1249]" pageId="111" pageNumber="112" type="description">
<paragraph blockId="111.[140,969,1187,1249]" box="[140,969,1222,1249]" pageId="111" pageNumber="112">
<uri box="[140,969,1222,1249]" pageId="111" pageNumber="112">
urn:lsid:zoobank.org:act:
<uuid box="[417,969,1222,1249]" pageId="111" pageNumber="112">769102C4-C297-4628-85AA-B87EEB70DF3E</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="111" pageNumber="112" type="materials_examined">
<paragraph blockId="111.[140,1108,1293,1672]" pageId="111" pageNumber="112">
<emphasis bold="true" box="[140,373,1293,1319]" pageId="111" pageNumber="112">
<emphasis box="[140,241,1293,1319]" inLineHeading="true" pageId="111" pageNumber="112" reason="1">Material</emphasis>
examined:
</emphasis>
<specimenCount box="[381,428,1293,1320]" pageId="111" pageNumber="112" type="male">4 ♂</specimenCount>
,
<specimenCount box="[443,487,1293,1320]" pageId="111" pageNumber="112" type="female">6 ♀</specimenCount>
,
<specimenCount box="[504,589,1293,1319]" pageId="111" pageNumber="112" type="generic">3 imm.</specimenCount>
<materialsCitation ID-GBIF-Occurrence="2620669317" collectingDate="2000-11-02" collectingDateMax="2000-11-08" collectingDateMin="2000-11-02" collectingMethod="pitfall trap" collectionCode="FMMC" collectorName="S. Goodman" county="Province" elevation="1250" latitude="-22.643333" location="Manasay River" longLatPrecision="125" longitude="44.171665" municipality="Foret de Analavelona" pageId="111" pageNumber="112" specimenCount="1" specimenCount-male="1" stateProvince="Toliara" typeStatus="holotype">
<emphasis box="[598,700,1293,1319]" italics="true" pageId="111" pageNumber="112">
<typeStatus box="[598,695,1293,1319]" pageId="111" pageNumber="112">Holotype</typeStatus>
:
</emphasis>
<specimenCount box="[709,755,1293,1320]" pageId="111" pageNumber="112" type="male">1 ♂</specimenCount>
(
<quantity box="[772,858,1293,1320]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="111" pageNumber="112" unit="mm" value="56.0">56 mm</quantity>
long),
<collectionCode box="[943,1033,1293,1319]" collectionName="FMMC" pageId="111" pageNumber="112">FMMC</collectionCode>
5429,
<collectingCounty box="[140,238,1328,1354]" pageId="111" pageNumber="112">Province</collectingCounty>
<collectingRegion box="[247,329,1328,1354]" country="Madagascar" name="Toliara" pageId="111" pageNumber="112">Toliara</collectingRegion>
,
<collectingMunicipality box="[344,593,1328,1354]" pageId="111" pageNumber="112">Forêt de Analavelona</collectingMunicipality>
, near spring of
<location LSID="urn:lsid:plazi:treatment:1772122E7868FFB7FF013BD7ADA7ECD9:9A04F5E37868FFA9FC833A45ACE2EF1B" box="[782,955,1328,1355]" county="Province" latitude="-22.643333" longLatPrecision="125" longitude="44.171665" municipality="Foret de Analavelona" name="Manasay River" pageId="111" pageNumber="112" stateProvince="Toliara">Manasay River</location>
, undisturbed mid-elevation forest with dry and humid elements,
<quantity box="[734,828,1363,1390]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="111" pageNumber="112" unit="m" value="1250.0">
<elevation box="[734,828,1363,1390]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="111" pageNumber="112" unit="m" value="1250.0">1250 m</elevation>
</quantity>
,
<geoCoordinate box="[841,964,1363,1390]" degrees="22" direction="south" minutes="38.6" orientation="latitude" pageId="111" pageNumber="112" precision="92" value="-22.643333">22°38.6’ S</geoCoordinate>
,
<geoCoordinate box="[978,1103,1364,1390]" degrees="44" direction="east" minutes="10.3" orientation="longitude" pageId="111" pageNumber="112" precision="92" value="44.171665">44°10.3’ E</geoCoordinate>
, leg.
<collectorName box="[189,340,1399,1425]" pageId="111" pageNumber="112">S. Goodman</collectorName>
,
<date box="[354,503,1399,1425]" pageId="111" pageNumber="112" value="2000-11-02" valueMax="2000-11-08" valueMin="2000-11-02">
<collectingDate box="[354,503,1399,1425]" pageId="111" pageNumber="112" value="2000-11-02" valueMax="2000-11-08" valueMin="2000-11-02">2–8.XI.2000</collectingDate>
</date>
,
<collectingMethod box="[518,643,1399,1425]" pageId="111" pageNumber="112">pitfall trap</collectingMethod>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2620669315" collectingDate="2000-11-02" collectingDateMax="2000-11-08" collectingDateMin="2000-11-02" collectorName="de Analavelona &amp; S. Goodman" county="Province" elevation="1250" latitude="-22.643333" location="Manasay River" longLatPrecision="125" longitude="44.171665" municipality="Foret de Analavelona" pageId="111" pageNumber="112" specimenCount="6" specimenCount-female="4" stateProvince="Toliara" typeStatus="paratype">
<emphasis box="[657,764,1399,1424]" italics="true" pageId="111" pageNumber="112">
<typeStatus box="[657,759,1399,1424]" pageId="111" pageNumber="112">Paratypes</typeStatus>
:
</emphasis>
<specimenCount box="[774,820,1399,1425]" pageId="111" pageNumber="112" type="female">4 ♀</specimenCount>
,
<specimenCount box="[838,927,1399,1425]" pageId="111" pageNumber="112" type="generic">2 imm.</specimenCount>
, FMMC 5429, same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph blockId="111.[140,1108,1293,1672]" pageId="111" pageNumber="112">
<emphasis bold="true" box="[185,499,1469,1495]" pageId="111" pageNumber="112">
<emphasis box="[185,257,1469,1495]" inLineHeading="true" pageId="111" pageNumber="112" reason="1">Other</emphasis>
material examined:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2620669314" collectingDate="2003-11-18" collectingDateMax="2003-11-22" collectingDateMin="2003-11-18" collectorName="de Analavelona &amp; Griswold &amp; Fisher" elevation="1100" latitude="-22.675" location="Foret de Analavelona" longLatPrecision="21" longitude="44.190002" municipality="Province" pageId="111" pageNumber="112" specimenCount="6" specimenCount-female="2" specimenCount-male="3" stateProvince="Toliara">
<specimenCount box="[509,557,1469,1496]" pageId="111" pageNumber="112" type="male">3 ♂</specimenCount>
,
<specimenCount box="[573,620,1469,1496]" pageId="111" pageNumber="112" type="female">2 ♀</specimenCount>
,
<specimenCount box="[638,727,1469,1495]" pageId="111" pageNumber="112" type="generic">1 imm.</specimenCount>
, CAS BLF 7819 (kmp-fm-068),
<collectingMunicipality box="[140,238,1504,1530]" pageId="111" pageNumber="112">Province</collectingMunicipality>
<collectingRegion box="[247,329,1504,1530]" country="Madagascar" name="Toliara" pageId="111" pageNumber="112">Toliara</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:1772122E7868FFB7FF013BD7ADA7ECD9:9A04F5E37868FFA9FED53A94AD0BEFAB" box="[344,594,1504,1530]" latitude="-22.675" longLatPrecision="21" longitude="44.190002" municipality="Province" name="Foret de Analavelona" pageId="111" pageNumber="112" stateProvince="Toliara">
Forêt
<collectorName box="[415,594,1504,1530]" pageId="111" pageNumber="112">de Analavelona</collectorName>
</location>
, montane rainforest,
<quantity box="[846,942,1504,1531]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="111" pageNumber="112" unit="m" value="1100.0">
<elevation box="[846,942,1504,1531]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="111" pageNumber="112" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<geoCoordinate box="[958,1103,1504,1530]" degrees="22" direction="south" minutes="40" orientation="latitude" pageId="111" pageNumber="112" precision="15" seconds="30" value="-22.675">22°40’30’’ S</geoCoordinate>
,
<geoCoordinate box="[140,283,1540,1566]" degrees="44" direction="east" minutes="11" orientation="longitude" pageId="111" pageNumber="112" precision="15" seconds="24" value="44.190002">44°11’24’’ E</geoCoordinate>
, leg.
<collectorName box="[342,449,1539,1565]" pageId="111" pageNumber="112">Griswold</collectorName>
,
<collectorName box="[460,528,1539,1565]" pageId="111" pageNumber="112">Fisher</collectorName>
et al.,
<date box="[605,784,1540,1566]" pageId="111" pageNumber="112" value="2003-11-18" valueMax="2003-11-22" valueMin="2003-11-18">
<collectingDate box="[605,784,1540,1566]" pageId="111" pageNumber="112" value="2003-11-18" valueMax="2003-11-22" valueMin="2003-11-18">18–22.XI.2003</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="112" lastPageNumber="113" pageId="111" pageNumber="112" type="diagnosis">
<paragraph blockId="111.[140,1108,1293,1672]" pageId="111" pageNumber="112">
<emphasis bold="true" box="[185,452,1574,1600]" pageId="111" pageNumber="112">
<emphasis box="[185,324,1574,1600]" inLineHeading="true" pageId="111" pageNumber="112" reason="1">Differential</emphasis>
diagnosis:
</emphasis>
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[461,629,1575,1601]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species">
<emphasis box="[461,629,1575,1601]" italics="true" pageId="111" pageNumber="112">H. analavelona</emphasis>
</taxonomicName>
has a unique green/brown colour pattern (
<figureCitation box="[147,283,1610,1636]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="111" pageNumber="112">Figs 62A, B</figureCitation>
). The colour pattern and gonopods clearly identify this species. The gonopods feature a unique combination of characters: the anterior gonopods carry a short
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3820836" ID-Zenodo-Dep="3820836" httpUri="https://zenodo.org/record/3820836/files/figure.png" pageId="112" pageNumber="113" startId="112.[140,212,1190,1212]" targetBox="[140,1108,165,1171]" targetPageId="112">
<paragraph blockId="112.[140,1108,1190,1340]" pageId="112" pageNumber="113">
<emphasis bold="true" box="[140,253,1190,1212]" pageId="112" pageNumber="113">Figure 61.</emphasis>
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[260,461,1190,1212]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species">
<emphasis box="[260,461,1190,1212]" italics="true" pageId="112" pageNumber="113">Hylekobolus montanus</emphasis>
</taxonomicName>
female
<typeStatus box="[536,618,1191,1212]" pageId="112" pageNumber="113">paratype</typeStatus>
, SEM.
<emphasis bold="true" box="[690,709,1190,1212]" pageId="112" pageNumber="113">A</emphasis>
antenna, overview
<emphasis bold="true" box="[893,910,1190,1212]" pageId="112" pageNumber="113">B</emphasis>
antennomere 5 with sensilla basiconica
<emphasis bold="true" box="[314,333,1222,1244]" pageId="112" pageNumber="113">C</emphasis>
antennomere 6 with sensilla basiconica
<emphasis bold="true" box="[710,729,1222,1244]" pageId="112" pageNumber="113">D</emphasis>
gnathochilarium, part of endochilarium
<emphasis bold="true" box="[140,155,1254,1276]" pageId="112" pageNumber="113">E</emphasis>
left mandible, mesal view
<emphasis bold="true" box="[403,417,1254,1276]" pageId="112" pageNumber="113">F</emphasis>
left vulva, lateral view. 4iT = 4-combed internal teeth; av = anterior valve; cp = central pads; Endo = endochilarium; eT = external tooth; LT = lateral tooth; mp = molar plate; O = operculum; pL = pectinate lamella; pv = posterior valve; sb = sensilla basiconica.
</paragraph>
</caption>
<paragraph blockId="112.[140,1108,1434,1672]" pageId="112" pageNumber="113">
mesal coxite process (
<figureCitation box="[382,489,1434,1461]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="112" pageNumber="113">Fig. 62D</figureCitation>
), the retrorse telopodite process is wide and projecting (
<figureCitation box="[148,248,1469,1496]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="112" pageNumber="113">Fig. 62F</figureCitation>
), the apical process (
<emphasis box="[478,490,1470,1495]" italics="true" pageId="112" pageNumber="113">x</emphasis>
) of the posterior gonopods is 4–5 times wider than the sperm canal (
<figureCitation box="[289,392,1504,1531]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="112" pageNumber="113">Fig. 62E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="113" lastPageNumber="114" pageId="112" pageNumber="113" type="description">
<paragraph blockId="112.[140,1108,1434,1672]" pageId="112" pageNumber="113">
<emphasis bold="true" box="[185,335,1539,1565]" pageId="112" pageNumber="113">
<emphasis box="[185,330,1539,1565]" inLineHeading="true" pageId="112" pageNumber="113" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[344,496,1540,1565]" italics="true" pageId="112" pageNumber="113">Measurements</emphasis>
: male
<typeStatus box="[576,675,1539,1565]" pageId="112" pageNumber="113">holotype</typeStatus>
with 45 rings,
<quantity box="[855,939,1539,1566]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="112" pageNumber="113" unit="mm" value="56.0">56 mm</quantity>
long,
<quantity box="[1015,1107,1539,1566]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="112" pageNumber="113" unit="mm" value="3.6">3.6 mm</quantity>
wide. Females with 45 rings,
<quantity box="[470,589,1574,1601]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="5.6" metricValueMin="4.9" pageId="112" pageNumber="113" unit="mm" value="52.5" valueMax="56.0" valueMin="49.0">49–56mm</quantity>
long,
<quantity box="[661,817,1574,1601]" metricMagnitude="-3" metricUnit="m" metricValue="4.875" metricValueMax="5.1" metricValueMin="4.65" pageId="112" pageNumber="113" unit="mm" value="4.875" valueMax="5.1" valueMin="4.65">4.65–5.1 mm</quantity>
wide.
</paragraph>
<paragraph blockId="112.[140,1108,1434,1672]" lastBlockId="113.[140,1108,1117,1672]" lastPageId="113" lastPageNumber="114" pageId="112" pageNumber="113">
<emphasis box="[185,308,1610,1636]" italics="true" pageId="112" pageNumber="113">Coloration:</emphasis>
collum and posterior half of body rings light brown (
<figureCitation box="[923,1027,1610,1637]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="112" pageNumber="113">Fig. 62A</figureCitation>
). Body rings in anterior half only laterally with dark green colour, dorsally light brown. Each body ring dorsally with an isolated dark green spot. Legs and antennae light brown (
<figureCitation box="[148,251,1152,1179]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="113" pageNumber="114">Fig. 62B</figureCitation>
).
<emphasis box="[270,313,1153,1178]" italics="true" pageId="113" pageNumber="114">Eyes</emphasis>
with 32–34 ocelli (
<figureCitation box="[536,640,1152,1179]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="113" pageNumber="114">Fig. 62A</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767360" ID-Zenodo-Dep="3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="113" pageNumber="114" startId="113.[140,212,856,878]" targetBox="[140,1108,165,833]" targetPageId="113">
<paragraph blockId="113.[140,1108,856,1007]" pageId="113" pageNumber="114">
<emphasis bold="true" box="[140,251,856,878]" pageId="113" pageNumber="114">Figure 62.</emphasis>
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[258,478,857,878]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[258,478,857,878]" italics="true" pageId="113" pageNumber="114">Hylekobolus analavelona</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[484,536,857,878]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
, male holotype.
<emphasis bold="true" box="[691,710,856,878]" pageId="113" pageNumber="114">A</emphasis>
head, lateral view
<emphasis bold="true" box="[884,901,856,878]" pageId="113" pageNumber="114">B</emphasis>
legs 1–7, ventral view
<emphasis bold="true" box="[140,159,888,910]" pageId="113" pageNumber="114">C</emphasis>
telson
<emphasis bold="true" box="[225,244,888,910]" pageId="113" pageNumber="114">D</emphasis>
anterior gonopod, anterior view
<emphasis bold="true" box="[552,567,888,910]" pageId="113" pageNumber="114">E</emphasis>
left posterior gonopod, anterior view
<emphasis bold="true" box="[919,933,888,910]" pageId="113" pageNumber="114">F</emphasis>
anterior gonopod, posterior view
<emphasis bold="true" box="[278,298,920,942]" pageId="113" pageNumber="114">G</emphasis>
left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph blockId="113.[140,1108,1117,1672]" pageId="113" pageNumber="114">
<emphasis box="[185,372,1187,1214]" italics="true" pageId="113" pageNumber="114">Anterior gonopod</emphasis>
sternite basally wide, apically elevated into a wide, well-rounded rectangular process (
<figureCitation box="[368,474,1223,1250]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="113" pageNumber="114">Fig. 62D</figureCitation>
). Mesal process of coxite wide and only slightly protruding (
<figureCitation box="[196,307,1258,1285]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="113" pageNumber="114">Fig. 62D</figureCitation>
). Apical telopodite process large and well-rounded. Retrorse margin wide, projecting above telopodite margin (
<figureCitation box="[625,727,1293,1320]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="113" pageNumber="114">Fig. 62F</figureCitation>
); margin basally of apical process not projecting (
<figureCitation box="[316,417,1328,1355]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="113" pageNumber="114">Fig. 62F</figureCitation>
).
</paragraph>
<paragraph blockId="113.[140,1108,1117,1672]" pageId="113" pageNumber="114">
<emphasis box="[185,376,1363,1389]" italics="true" pageId="113" pageNumber="114">Posterior gonopods</emphasis>
telopodite bent 90° (
<figureCitation box="[605,709,1363,1390]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="113" pageNumber="114">Fig. 62G</figureCitation>
). Projection absent from lateral margin (
<emphasis box="[190,209,1399,1424]" italics="true" pageId="113" pageNumber="114">w</emphasis>
,
<figureCitation box="[222,319,1399,1426]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="113" pageNumber="114">Fig. 62E</figureCitation>
). Tip of apical process (
<emphasis box="[572,584,1399,1424]" italics="true" pageId="113" pageNumber="114">x</emphasis>
) sharp-edged, basally slender, 4–5 times as wide as sperm canal (
<figureCitation box="[312,412,1434,1461]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="113" pageNumber="114">Fig. 62E</figureCitation>
). Membranous lobe (
<emphasis box="[648,658,1435,1460]" italics="true" pageId="113" pageNumber="114">y</emphasis>
) large and well-rounded (
<figureCitation box="[942,1041,1434,1461]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="113" pageNumber="114">Fig. 62E</figureCitation>
). Projection of sperm canal (
<emphasis box="[398,410,1470,1495]" italics="true" pageId="113" pageNumber="114">z</emphasis>
) long and slender, running parallel to apical process (
<figureCitation box="[996,1095,1469,1496]" captionStart="Figure 62" captionStartId="113.[140,212,856,878]" captionTargetBox="[140,1108,165,833]" captionTargetId="figure@113.[140,1108,445,838]" captionTargetPageId="113" captionText="Figure 62. Hylekobolus analavelona sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767360" httpUri="https://zenodo.org/record/3767360/files/figure.png" pageId="113" pageNumber="114">Fig. 62E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="113" pageNumber="114" type="distribution">
<paragraph blockId="113.[140,1108,1117,1672]" pageId="113" pageNumber="114">
<emphasis bold="true" box="[185,484,1504,1530]" pageId="113" pageNumber="114">
<emphasis box="[185,336,1504,1530]" inLineHeading="true" pageId="113" pageNumber="114" reason="1">Distribution</emphasis>
and ecology
</emphasis>
: only known from the Analavelona Mountain (
<figureCitation box="[1011,1093,1504,1531]" captionStart="Figure 54" captionStartId="97.[138,210,1653,1675]" captionTargetBox="[130,1107,1100,1636]" captionTargetId="figure@97.[138,1107,1109,1633]" captionTargetPageId="97" captionText="Figure 54. Distribution map of the genus Hylekobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3767346" httpUri="https://zenodo.org/record/3767346/files/figure.png" pageId="113" pageNumber="114">Fig. 54</figureCitation>
), an isolated hilltop with rainforest in the otherwise dry southwestern region of
<collectingCountry name="Madagascar" pageId="113" pageNumber="114">Madagascar</collectingCountry>
(DuPuy and
<bibRefCitation author="Moat" box="[362,493,1575,1601]" firstAuthor="Moat" pageId="113" pageNumber="114" refId="ref57536" refString="Moat J, Smith P (2007) Atlas of the Vegetation of Madagascar. Kew Publishing, London, 124 pp." type="book" year="2007">Moat 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="113" pageNumber="114" type="etymology">
<paragraph blockId="113.[140,1108,1117,1672]" pageId="113" pageNumber="114">
<emphasis bold="true" box="[185,323,1610,1636]" pageId="113" pageNumber="114">
<emphasis box="[185,318,1610,1636]" inLineHeading="true" pageId="113" pageNumber="114" reason="1">Etymology</emphasis>
:
</emphasis>
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[331,461,1610,1636]" class="Diplopoda" family="Spirobolellidae" genus="Hylekobolus" kingdom="Animalia" order="Spirobolida" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species">
<emphasis box="[331,461,1610,1636]" italics="true" pageId="113" pageNumber="114">analavelona</emphasis>
</taxonomicName>
, noun in apposition, after the only place where this species can be found, the Analavelona Mountain (
<figureCitation box="[668,752,1645,1672]" captionStart="Figure 54" captionStartId="97.[138,210,1653,1675]" captionTargetBox="[130,1107,1100,1636]" captionTargetId="figure@97.[138,1107,1109,1633]" captionTargetPageId="97" captionText="Figure 54. Distribution map of the genus Hylekobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3767346" httpUri="https://zenodo.org/record/3767346/files/figure.png" pageId="113" pageNumber="114">Fig. 54</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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