251 lines
30 KiB
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251 lines
30 KiB
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<document id="78318F2A6560C155BEEFA3EAAAEC12E7" ID-DOI="10.11646/zootaxa.5538.5.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14239799" ID-ZooBank="7AA0B8F5-099A-42F0-AFAE-2F0EAC3CCEE0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732829966751" checkinUser="plazi" docAuthor="Villastrigo, Adrián & García-Esquivel, Eva" docDate="2024" docId="0C566E0B1D1ECA183CEDB915FD6AF807" docLanguage="en" docName="zootaxa.5538.5.3.pdf" docOrigin="Zootaxa 5538 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5538.5.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ochthebius (Cobalius) vilanovensis Villastrigo & García-Esquivel, 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="443" masterDocId="F06F16731D1DCA1C3C7AB865FFD2FFE4" masterDocTitle="Overlooked coastal habitats expose a new species: Ochthebius vilanovensis sp. nov. (Coleoptera, Hydraenidae)" masterLastPageNumber="447" masterPageNumber="439" pageNumber="442" updateTime="1732920605681" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="C6E7E8D4B9AA02AA19094D987B94246D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="58697B95D0A328652D120E0BDAF8F097">
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<mods:title id="0CBA473A6CAE0E1BE1DC3E7F0E8C67CF">Overlooked coastal habitats expose a new species: Ochthebius vilanovensis sp. nov. (Coleoptera, Hydraenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="FBC5C58935A884C67C4E3269DA68826F" type="personal">
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<mods:role id="F19218B5BDF928F1F649A6E11A23241C">
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<mods:roleTerm id="89BA9CBD22E9F4BF7BA88F0AE7A64937">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="689BB87A92CFAD58AA7DE4626A0EC51C">Villastrigo, Adrián</mods:namePart>
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<mods:affiliation id="981EE15CFCEFEB90EF035C19B42E3C15">Division of Entomology, SNSB-Zoologische Staatssammlung München, Münchhausenstrasse 21, 81247 München, Germany</mods:affiliation>
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</mods:name>
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<mods:name id="3D176310102DE7D536DB24D1CF63B039" type="personal">
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<mods:role id="B181A2A776DE61BD42911A6F2CB1ECAF">
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<mods:roleTerm id="8F3142E1EF5FF3C60F852FCE3B6856FA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5DB5C5BB7A4BE4122FAE288FD1CC324D">García-Esquivel, Eva</mods:namePart>
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<mods:affiliation id="657F0570585E0B3767D41F4F69722D0D">Independent researcher</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="A18F2576FB6EBA789837BEF9EE552502">text</mods:typeOfResource>
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<mods:titleInfo id="2AD4BC8476A49F831D4DF4ED383D71D7">
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<mods:title id="1B307C198042EBBFD15AD2CC7812C563">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9D68AA8F1446C1C93695AE87A8FE50FA">
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<mods:date id="2B57669ABBB09226AFEFC79FD1AE852A">2024</mods:date>
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<mods:detail id="B69E2595AFD30802CEE395459AA9A6F2" type="pubDate">
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<mods:number id="A532178D2790D525B3878BEC6CA9A3F5">2024-11-19</mods:number>
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</mods:detail>
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<mods:detail id="3BD0A85D3F6ED0DDAB5F7BF5767D19D7" type="volume">
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<mods:number id="28CD1E8F1B304D632003947148A2DB90">5538</mods:number>
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</mods:detail>
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<mods:detail id="E91AEF1658B5A7AB221FCEFA821A238C" type="issue">
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<mods:number id="707F7B0D4D50FD8F405A4C223F36CDE3">5</mods:number>
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</mods:detail>
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<mods:extent id="D5559314CFE1A6C2E52DF20498456A02" unit="page">
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<mods:start id="BF5749F88351850C29B870EB8371AF92">439</mods:start>
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<mods:end id="003E56B8AE4880A8E70CAC21DA3BE676">447</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="0C45841CAE3B8DDAD29B6757184D1E23">
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<mods:url id="FFD67272FD2769019DA70338A6A4A4BE">http://dx.doi.org/10.11646/zootaxa.5538.5.3</mods:url>
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</mods:location>
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<mods:classification id="7EED3E199295CEC962E3061B2B490AAB">journal article</mods:classification>
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<mods:identifier id="01F673F521C6564228B927E5630E2392" type="DOI">10.11646/zootaxa.5538.5.3</mods:identifier>
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<mods:identifier id="13381B0058143AEEE1CB9A878012E1F5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="97118080C7F1552820AB52BBE5009EDC" type="Zenodo-Dep">14239799</mods:identifier>
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<mods:identifier id="7890E2999E173759A6E726D75A415B3A" type="ZooBank">7AA0B8F5-099A-42F0-AFAE-2F0EAC3CCEE0</mods:identifier>
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</mods:mods>
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<treatment id="0C566E0B1D1ECA183CEDB915FD6AF807" ID-DOI="http://doi.org/10.5281/zenodo.14248173" ID-Zenodo-Dep="14248173" LSID="urn:lsid:plazi:treatment:0C566E0B1D1ECA183CEDB915FD6AF807" httpUri="http://treatment.plazi.org/id/0C566E0B1D1ECA183CEDB915FD6AF807" lastPageId="4" lastPageNumber="443" pageId="3" pageNumber="442">
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<subSubSection id="CCE58C961D1ECA1F3CEDB915FD7BFE6F" box="[151,681,368,395]" pageId="3" pageNumber="442" type="nomenclature">
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<paragraph id="8440DF1D1D1ECA1F3CEDB915FD7BFE6F" blockId="3.[151,681,368,395]" box="[151,681,368,395]" pageId="3" pageNumber="442">
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<heading id="DF0868711D1ECA1F3CEDB915FD7BFE6F" bold="true" box="[151,681,368,395]" fontSize="11" level="1" pageId="3" pageNumber="442" reason="1">
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<emphasis id="B68B030F1D1ECA1F3CEDB915FD7BFE6F" bold="true" box="[151,681,368,395]" pageId="3" pageNumber="442">
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<taxonomicName id="43FFA49E1D1ECA1F3CEDB915FD96FE6F" authorityName="Villastrigo & García-Esquivel" authorityYear="2024" box="[151,580,368,395]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="vilanovensis" status="sp. nov." subGenus="Cobalius">
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<emphasis id="B68B030F1D1ECA1F3CEDB915FEF0FE6F" bold="true" box="[151,290,368,395]" italics="true" pageId="3" pageNumber="442">Ochthebius</emphasis>
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(
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<emphasis id="B68B030F1D1ECA1F3D49B915FE4DFE6F" bold="true" box="[307,415,368,395]" italics="true" pageId="3" pageNumber="442">Cobalius</emphasis>
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)
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<emphasis id="B68B030F1D1ECA1F3DCAB915FD96FE6F" bold="true" box="[432,580,368,395]" italics="true" pageId="3" pageNumber="442">vilanovensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="ADB8BE741D1ECA1F3E31B914FD7BFE6F" box="[587,681,369,395]" pageId="3" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CCE58C961D1ECA1F3CEDB9DCFC09FDDB" pageId="3" pageNumber="442" type="materials_examined">
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<paragraph id="8440DF1D1D1ECA1F3CEDB9DCFBB6FE37" blockId="3.[151,1437,441,756]" box="[151,1124,441,467]" pageId="3" pageNumber="442">
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<materialsCitation id="3497D5401D1ECA1F3CEDB9DCFBB2FE37" box="[151,1120,441,467]" collectorName="la Geltru" country="Spain" latitude="41.21786" location="Barcelona province" longLatPrecision="1" longitude="1.749349" pageId="3" pageNumber="442" specimenCount="1" typeStatus="holotype">
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<emphasis id="B68B030F1D1ECA1F3CEDB9DCFEE4FE37" bold="true" box="[151,310,441,467]" pageId="3" pageNumber="442">
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<typeStatus id="5B4461BF1D1ECA1F3CEDB9DCFF00FE37" box="[151,210,441,467]" pageId="3" pageNumber="442">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="FCE89F8D1D1ECA1F3D46B9DCFEADFE37" box="[316,383,441,467]" name="Spain" pageId="3" pageNumber="442">Spain</collectingCountry>
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,
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<location id="812089C61D1ECA1F3DF0B9DCFDBAFE37" LSID="urn:lsid:plazi:treatment:0C566E0B1D1ECA183CEDB915FD6AF807:812089C61D1ECA1F3DF0B9DCFDBAFE37" box="[394,616,441,467]" country="Spain" latitude="41.21786" longLatPrecision="1" longitude="1.749349" name="Barcelona province" pageId="3" pageNumber="442">Barcelona province</location>
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, Vilanova i
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<collectorName id="290ABACB1D1ECA1F3E97B9DCFC80FE37" box="[749,850,441,467]" pageId="3" pageNumber="442">la Geltrú</collectorName>
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,
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<geoCoordinate id="E1CBB9DA1D1ECA1F3F24B9DCFC09FE37" box="[862,987,441,467]" degrees="41.21786" direction="north" orientation="latitude" pageId="3" pageNumber="442" precision="1" value="41.21786">41.21786N</geoCoordinate>
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<geoCoordinate id="E1CBB9DA1D1ECA1F3F98B9DCFBB2FE37" box="[994,1120,441,467]" degrees="1.749349" direction="east" orientation="longitude" pageId="3" pageNumber="442" precision="1" value="1.749349">1.749349E</geoCoordinate>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8440DF1D1D1ECA1F3CBDB9B8FC09FDDB" blockId="3.[151,1437,441,756]" pageId="3" pageNumber="442">
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<emphasis id="B68B030F1D1ECA1F3CBDB9B8FEA0FE13" bold="true" box="[199,370,477,503]" pageId="3" pageNumber="442">Type material.</emphasis>
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<materialsCitation id="3497D5401D1ECA1F3D0DB9B8FAF1FDFF" collectingDate="2024-05-26" collectionCode="ZSM" collectorName="E. Garcia & A. Villastrigo" country="Spain" latitude="41.21786" location="Barcelona" longLatPrecision="1" longitude="1.749349" municipality="Vilanova" pageId="3" pageNumber="442" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="5B4461BF1D1ECA1F3D0DB9B8FE0DFE13" box="[375,479,477,503]" pageId="3" pageNumber="442" type="holotype">Holotype</typeStatus>
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<specimenCount id="92F914941D1ECA1F3D9EB9BBFE2BFE1C" box="[484,505,478,504]" count="1" pageId="3" pageNumber="442" type="male">♂</specimenCount>
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(
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<collectionCode id="E2EE47D81D1ECA1F3E7FB9B8FD97FE13" box="[517,581,477,503]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="442" type="Museum">ZSM</collectionCode>
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): “
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<collectingCountry id="FCE89F8D1D1ECA1F3E1AB9B8FD74FE13" box="[608,678,477,503]" name="Spain" pageId="3" pageNumber="442">Spain</collectingCountry>
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:
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<location id="812089C61D1ECA1F3ED4B9B8FCF0FE13" LSID="urn:lsid:plazi:treatment:0C566E0B1D1ECA183CEDB915FD6AF807:812089C61D1ECA1F3ED4B9B8FCF0FE13" box="[686,802,477,503]" country="Spain" latitude="41.21786" longLatPrecision="1" longitude="1.749349" municipality="Vilanova" name="Barcelona" pageId="3" pageNumber="442">Barcelona</location>
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,
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<collectingMunicipality id="642445671D1ECA1F3F51B9B8FC42FE13" box="[811,912,477,503]" pageId="3" pageNumber="442">Vilanova</collectingMunicipality>
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i / la Geltrú
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<date id="F041F9DD1D1ECA1F386BB9B8FB5DFE1C" box="[1041,1167,477,504]" pageId="3" pageNumber="442" value="2024-05-26">
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<collectingDate id="E00500351D1ECA1F386BB9B8FB5DFE1C" box="[1041,1167,477,504]" pageId="3" pageNumber="442" value="2024-05-26">26.05.2024</collectingDate>
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</date>
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/
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<geoCoordinate id="E1CBB9DA1D1ECA1F38DAB9B8FACCFE13" box="[1184,1310,477,503]" degrees="41.21786" direction="north" orientation="latitude" pageId="3" pageNumber="442" precision="1" value="41.21786">41.21786N</geoCoordinate>
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<geoCoordinate id="E1CBB9DA1D1ECA1F3958B9B8FA4EFE13" box="[1314,1436,477,503]" degrees="1.749349" direction="east" orientation="longitude" pageId="3" pageNumber="442" precision="1" value="1.749349">1.749349E</geoCoordinate>
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/
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<collectorName id="290ABACB1D1ECA1F3CD2BA67FEC6FDFF" box="[168,276,513,540]" pageId="3" pageNumber="442">E. García</collectorName>
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&
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<collectorName id="290ABACB1D1ECA1F3D41BA64FE09FDFF" box="[315,475,513,539]" pageId="3" pageNumber="442">A. Villastrigo</collectorName>
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”; with
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<typeStatus id="5B4461BF1D1ECA1F3E57BA64FD5DFDFF" box="[557,655,513,539]" pageId="3" pageNumber="442" type="holotype">holotype</typeStatus>
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label. Aedeagus dissected and mounted in the same card
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</materialsCitation>
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.
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<materialsCitation id="3497D5401D1ECA1F3955BA67FEA6FDDB" collectionCode="ZSM, CHC, AVC" country="Spain" pageId="3" pageNumber="442" specimenCount="1" typeStatus="paratype">
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<typeStatus id="5B4461BF1D1ECA1F3955BA67FA4EFDF8" box="[1327,1436,514,540]" pageId="3" pageNumber="442" type="paratype">Paratypes</typeStatus>
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(
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<collectionCode id="E2EE47D81D1ECA1F3CE5BA40FF0FFDDB" box="[159,221,549,575]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="442" type="Museum">ZSM</collectionCode>
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,
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<collectionCode id="E2EE47D81D1ECA1F3C92BA40FEF6FDDB" box="[232,292,549,575]" pageId="3" pageNumber="442">CHC</collectionCode>
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,
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<collectionCode id="E2EE47D81D1ECA1F3D54BA40FEBEFDDB" box="[302,364,549,575]" pageId="3" pageNumber="442">AVC</collectionCode>
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)
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</materialsCitation>
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:
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<materialsCitation id="3497D5401D1ECA1F3D04BA40FC05FDDB" box="[382,983,549,576]" collectingDate="2024-05-26" collectionCode="ZSM" collectorName="E. Garcia & A. Villastrigo" country="Spain" latitude="41.21786" location="Barcelona" longLatPrecision="1" longitude="1.749349" municipality="Vilanova" pageId="3" pageNumber="442" specimenCount="9" specimenCount-female="7" specimenCount-male="2" typeStatus="paratype">
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<specimenCount id="92F914941D1ECA1F3D04BA40FE78FDA4" box="[382,426,549,576]" count="2" pageId="3" pageNumber="442" type="male">2 ♂</specimenCount>
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,
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<specimenCount id="92F914941D1ECA1F3DCCBA43FE32FDA4" box="[438,480,550,576]" count="7" pageId="3" pageNumber="442" type="female">7 ♀</specimenCount>
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, same data as holotype, with
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<typeStatus id="5B4461BF1D1ECA1F3F51BA43FC59FDA4" box="[811,907,550,576]" pageId="3" pageNumber="442" type="paratype">paratype</typeStatus>
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labels
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="CCE58C961D1ECA183CBDBA2CFBD3FA47" lastPageId="4" lastPageNumber="443" pageId="3" pageNumber="442" type="description">
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<paragraph id="8440DF1D1D1ECA1F3CBDBA2CFC3AFD63" blockId="3.[151,1437,441,756]" pageId="3" pageNumber="442">
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<emphasis id="B68B030F1D1ECA1F3CBDBA2CFE8BFD87" bold="true" box="[199,345,585,611]" pageId="3" pageNumber="442">Description.</emphasis>
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Habitus as in
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<figureCitation id="1CC4C3981D1ECA1F3E71BA2CFDA3FD87" box="[523,625,585,611]" captionStart="FIGURE 2" captionStartId="3.[152,255,1592,1616]" captionTargetBox="[158,1429,787,1562]" captionTargetId="figure-182@3.[151,1436,779,1569]" captionTargetPageId="3" captionText="FIGURE 2. Ochthebius vilanovensis sp. nov.: (A) male habitus, (B) female habitus and (C) aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.14239805" httpUri="https://zenodo.org/record/14239805/files/figure.png" pageId="3" pageNumber="442">Figure 2</figureCitation>
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. Length:
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<quantity id="430772F81D1ECA1F3E9EBA2CFCA0FD80" box="[740,882,585,612]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="3" pageNumber="442" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6–1.9 mm</quantity>
|
|||
|
|
; maximum width:
|
|||
|
|
<quantity id="430772F81D1ECA1F3829BA2CFB2FFD80" box="[1107,1277,585,612]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" metricValueMax="7.0" metricValueMin="5.6" pageId="3" pageNumber="442" unit="mm" value="0.63" valueMax="0.7" valueMin="0.56">0.56–0.70 mm</quantity>
|
|||
|
|
. Body colour uniformly black, appendages from brown to dark brown, with lighter tibiae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8440DF1D1D1ECA1F3CBDBAF4FD83FD17" blockId="3.[151,1437,441,756]" pageId="3" pageNumber="442">Head microreticulated and rugose; with short whitish hook-like setae; frontoclypeal suture impressed; clypeus and labrum shiny and dark brown with transparent setae; anterior margin of labrum slightly emarginated; with two fovea on vertex. Eyes large, prominent.</paragraph>
|
|||
|
|
<caption id="D0808F951D1ECA1F3CE2BE5DFA93F9B5" ID-DOI="http://doi.org/10.5281/zenodo.14239805" ID-Zenodo-Dep="14239805" box="[152,1345,1592,1617]" httpUri="https://zenodo.org/record/14239805/files/figure.png" pageId="3" pageNumber="442" startId="3.[152,255,1592,1616]" targetBox="[158,1429,787,1562]" targetPageId="3" targetType="figure">
|
|||
|
|
<paragraph id="8440DF1D1D1ECA1F3CE2BE5DFA93F9B5" blockId="3.[152,1345,1592,1617]" box="[152,1345,1592,1617]" pageId="3" pageNumber="442">
|
|||
|
|
<emphasis id="B68B030F1D1ECA1F3CE2BE5DFECAF9B4" bold="true" box="[152,280,1592,1616]" pageId="3" pageNumber="442">FIGURE 2.</emphasis>
|
|||
|
|
<taxonomicName id="43FFA49E1D1ECA1F3D65BE5CFDC7F9B5" authorityName="Villastrigo & García-Esquivel" authorityYear="2024" box="[287,533,1593,1617]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="vilanovensis" status="sp. nov.">
|
|||
|
|
<emphasis id="B68B030F1D1ECA1F3D65BE5CFDC7F9B5" box="[287,533,1593,1617]" italics="true" pageId="3" pageNumber="442">Ochthebius vilanovensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B68B030F1D1ECA1F3E66BE5CFDBFF9B5" bold="true" box="[540,621,1593,1617]" pageId="3" pageNumber="442">
|
|||
|
|
<taxonomicNameLabel id="ADB8BE741D1ECA1F3E66BE5CFDBFF9B5" box="[540,621,1593,1617]" pageId="3" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
: (A) male habitus, (B) female habitus and (C) aedeagus in lateral view.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8440DF1D1D1ECA1F3CBDBEFEFE73F919" blockId="3.[151,1437,1691,2041]" pageId="3" pageNumber="442">Pronotum with very convex side margins with dented lateral margins and anterior margin straightened; dense irregular setiferous punctures; densely covered by whitish hook-like setae; anterior and posterior margins with an expanded hyaline band.</paragraph>
|
|||
|
|
<paragraph id="8440DF1D1D1ECA1F3CBDBF62FCEEF8A1" blockId="3.[151,1437,1691,2041]" pageId="3" pageNumber="442">Elytra elongated, oval; serrated margins laterally, less serrated in the distal section; regular series of punctures with short whitish hook-like setae as in pronotum. Apterous.</paragraph>
|
|||
|
|
<paragraph id="8440DF1D1D1ECA1F3CBDBF2AFE95F869" blockId="3.[151,1437,1691,2041]" pageId="3" pageNumber="442">Short, robust legs with rows of spine-like setae plus an external row of hook-like setae on the tibiae; without natatorial setae.</paragraph>
|
|||
|
|
<paragraph id="8440DF1D1D1ECA1F3CBDBFF2FE2BF81D" blockId="3.[151,1437,1691,2041]" pageId="3" pageNumber="442">Metaventrite weakly microreticulate. Abdominal ventrites (i-v) densely pubescent with thin recumbent setae and well-impressed rugose microreticulation; ventrites vi-vii with longer brownish and thicker setae on posterior margin; finer microreticulation.</paragraph>
|
|||
|
|
<paragraph id="8440DF1D1D19CA183CBDB8F2FBADFF1D" blockId="4.[151,1436,151,249]" pageId="4" pageNumber="443">
|
|||
|
|
Aedeagus (
|
|||
|
|
<figureCitation id="1CC4C3981D19CA183D3DB8F2FE66FF55" box="[327,436,151,177]" captionStart="FIGURE 2" captionStartId="3.[152,255,1592,1616]" captionTargetBox="[158,1429,787,1562]" captionTargetId="figure-182@3.[151,1436,779,1569]" captionTargetPageId="3" captionText="FIGURE 2. Ochthebius vilanovensis sp. nov.: (A) male habitus, (B) female habitus and (C) aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.14239805" httpUri="https://zenodo.org/record/14239805/files/figure.png" pageId="4" pageNumber="443">Figures 2</figureCitation>
|
|||
|
|
and
|
|||
|
|
<figureCitation id="1CC4C3981D19CA183D95B8F2FE2FFF55" box="[495,509,151,177]" captionStart="FIGURE 3" captionStartId="4.[152,255,1203,1227]" captionTargetBox="[162,1425,284,1169]" captionTargetId="figure-85@4.[151,1436,273,1180]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of the male genitalia of Ochthebius evae Villastrigo, Hernando, Millán & Ribera, 2020 (A and C) and Ochthebius vilanovensis sp. nov. (B and D)." figureDoi="http://doi.org/10.5281/zenodo.14239807" httpUri="https://zenodo.org/record/14239807/files/figure.png" pageId="4" pageNumber="443">3</figureCitation>
|
|||
|
|
) with curved median lobe in lateral view, middle part slightly compressed, apex pointed. Distal lobe wide, with an approximately 90º angle on the dorsal margin and ventral margin convex, apex hyaline. Parameres inserted near the basal third of the median lobe, not reaching its apex.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="D0808F951D19CA183CE2BCD6FD8BFB0B" ID-DOI="http://doi.org/10.5281/zenodo.14239807" ID-Zenodo-Dep="14239807" httpUri="https://zenodo.org/record/14239807/files/figure.png" pageId="4" pageNumber="443" startId="4.[152,255,1203,1227]" targetBox="[162,1425,284,1169]" targetPageId="4" targetType="figure">
|
|||
|
|
<paragraph id="8440DF1D1D19CA183CE2BCD6FD8BFB0B" blockId="4.[151,1437,1203,1263]" pageId="4" pageNumber="443">
|
|||
|
|
<emphasis id="B68B030F1D19CA183CE2BCD6FECAFB2F" bold="true" box="[152,280,1203,1227]" pageId="4" pageNumber="443">FIGURE 3.</emphasis>
|
|||
|
|
Comparison of the male genitalia of
|
|||
|
|
<taxonomicName id="43FFA49E1D19CA183EF5BCD6FADBFB2F" authority="Villastrigo, Hernando, Millan & Ribera, 2020" authorityName="Villastrigo, Hernando, Millan & Ribera" authorityYear="2020" box="[655,1289,1203,1227]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="443" phylum="Arthropoda" rank="species" species="evae">
|
|||
|
|
<emphasis id="B68B030F1D19CA183EF5BCD6FCE4FB2F" box="[655,822,1203,1227]" italics="true" pageId="4" pageNumber="443">Ochthebius evae</emphasis>
|
|||
|
|
<bibRefCitation id="E06EA2EC1D19CA183F46BCD6FADBFB2F" author="Villastrigo, A. & Hernando, C. & Millan, A. & Ribera, I." box="[828,1289,1203,1227]" pageId="4" pageNumber="443" pagination="785 - 801" refId="ref5537" refString="Villastrigo, A., Hernando, C., Millan, A. & Ribera, I. (2020 b) The neglected diversity of the Ochthebius fauna from Eastern Atlantic and Central and Western Mediterranean coastal rockpools (Coleoptera, Hydraenidae). Organisms, diversity & evolution, 20, 785 - 801. https: // doi. org / 10.1007 / s 13127 - 020 - 00463 - y" type="journal article" year="2020">Villastrigo, Hernando, Millán & Ribera, 2020</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(A and C) and
|
|||
|
|
<taxonomicName id="43FFA49E1D19CA183CEDBCB2FE5CFB0B" authorityName="Villastrigo & García-Esquivel" authorityYear="2024" box="[151,398,1239,1263]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="443" phylum="Arthropoda" rank="species" species="vilanovensis" status="sp. nov.">
|
|||
|
|
<emphasis id="B68B030F1D19CA183CEDBCB2FE5CFB0B" box="[151,398,1239,1263]" italics="true" pageId="4" pageNumber="443">Ochthebius vilanovensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="ADB8BE741D19CA183DEEBCB2FE35FB0B" box="[404,487,1239,1263]" pageId="4" pageNumber="443" rank="species">
|
|||
|
|
<emphasis id="B68B030F1D19CA183DEEBCB2FE32FB0B" bold="true" box="[404,480,1239,1263]" pageId="4" pageNumber="443">sp. nov</emphasis>
|
|||
|
|
.
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
(B and D).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8440DF1D1D19CA183CBDBD24FBD3FA47" blockId="4.[151,1437,1345,2020]" pageId="4" pageNumber="443">Sexual dimorphism: Almost identical morphologies and body sizes in males and females. Females have a more strongly emarginated labrum than males, although the character is slightly variable and requires further study with a longer series of specimens before this apparent difference can be confirmed.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="CCE58C961D19CA183CBDBDC8FEAEFA0F" pageId="4" pageNumber="443" type="etymology">
|
|||
|
|
<paragraph id="8440DF1D1D19CA183CBDBDC8FEAEFA0F" blockId="4.[151,1437,1345,2020]" pageId="4" pageNumber="443">
|
|||
|
|
<emphasis id="B68B030F1D19CA183CBDBDC8FE9CFA23" bold="true" box="[199,334,1453,1479]" pageId="4" pageNumber="443">Etymology.</emphasis>
|
|||
|
|
Named after the
|
|||
|
|
<typeStatus id="5B4461BF1D19CA183E59BDCBFD81FA2C" box="[547,595,1454,1480]" pageId="4" pageNumber="443">type</typeStatus>
|
|||
|
|
locality, Vilanova i la Geltrú. The specific name is an adjective in the nominative singular.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="CCE58C961D19CA183CBDBD90FE74F9B3" pageId="4" pageNumber="443" type="distribution">
|
|||
|
|
<paragraph id="8440DF1D1D19CA183CBDBD90FCB8F9EB" blockId="4.[151,1437,1345,2020]" box="[199,874,1525,1552]" pageId="4" pageNumber="443">
|
|||
|
|
<emphasis id="B68B030F1D19CA183CBDBD90FEB3F9EB" bold="true" box="[199,353,1525,1551]" pageId="4" pageNumber="443">Distribution.</emphasis>
|
|||
|
|
Only known from the
|
|||
|
|
<typeStatus id="5B4461BF1D19CA183E1ABD93FD42F9F4" box="[608,656,1526,1552]" pageId="4" pageNumber="443">type</typeStatus>
|
|||
|
|
locality (
|
|||
|
|
<figureCitation id="1CC4C3981D19CA183E80BD90FC8FF9EB" box="[762,861,1525,1551]" captionStart="FIGURE 4" captionStartId="5.[152,255,1559,1583]" captionTargetBox="[151,1436,181,1535]" captionTargetId="figure-17@5.[151,1436,181,1535]" captionTargetPageId="5" captionText="FIGURE 4. Distribution map (A) and habitat of Ochthebius vilanovensisis sp. nov. (B and C) plus the known localities of Ochthebius evae Villastrigo, Hernando, Millán & Ribera, 2020 in Spain (red dots in Figure 3A)." figureDoi="http://doi.org/10.5281/zenodo.14239811" httpUri="https://zenodo.org/record/14239811/files/figure.png" pageId="4" pageNumber="443">Figure 4</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8440DF1D1D19CA183CBDBE7CFE74F9B3" blockId="4.[151,1437,1345,2020]" pageId="4" pageNumber="443">
|
|||
|
|
<emphasis id="B68B030F1D19CA183CBDBE7CFEF8F9D7" bold="true" box="[199,298,1561,1587]" pageId="4" pageNumber="443">Habitat.</emphasis>
|
|||
|
|
Specimens were found in crevices close to rockpools in the supralittoral zone, often in vertical crevices with a fine humus layer.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="CCE58C961D19CA183CBDBE04FD6AF807" pageId="4" pageNumber="443" type="diagnosis">
|
|||
|
|
<paragraph id="8440DF1D1D19CA183CBDBE04FD6AF807" blockId="4.[151,1437,1345,2020]" pageId="4" pageNumber="443">
|
|||
|
|
<emphasis id="B68B030F1D19CA183CBDBE04FE00F99F" bold="true" box="[199,466,1633,1659]" pageId="4" pageNumber="443">Differential diagnosis.</emphasis>
|
|||
|
|
Members of the
|
|||
|
|
<taxonomicName id="43FFA49E1D19CA183EDBBE04FCA6F99F" authorityName="Jach & Delgado" authorityYear="2017" box="[673,884,1633,1659]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="443" phylum="Arthropoda" rank="species" species="biltoni">
|
|||
|
|
<emphasis id="B68B030F1D19CA183EDBBE04FCA6F99F" box="[673,884,1633,1659]" italics="true" pageId="4" pageNumber="443">Ochthebius biltoni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species group are morphologically cryptic, but rather strongly divergent genetically. Most have geographically isolated distributions (on the Mediterranean islands of Sicily,
|
|||
|
|
<collectingCountry id="FCE89F8D1D19CA183D7BBECCFE90F927" box="[257,322,1705,1731]" name="Malta" pageId="4" pageNumber="443">Malta</collectingCountry>
|
|||
|
|
or Sardinia), with the only previously described species inhabiting mainland areas being
|
|||
|
|
<taxonomicName id="43FFA49E1D19CA183967BECCFF18F903" authorityName="Villastrigo, Hernando, Millan & Ribera" authorityYear="2020" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="443" phylum="Arthropoda" rank="species" species="evae">
|
|||
|
|
<emphasis id="B68B030F1D19CA183967BECCFF18F903" italics="true" pageId="4" pageNumber="443">Ochthebius evae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, along the Mediterranean coast of the Iberian Peninsula. Despite its external morphological resemblance to
|
|||
|
|
<taxonomicName id="43FFA49E1D19CA183CEDBE94FE81F8EF" authorityName="Villastrigo, Hernando, Millan & Ribera" authorityYear="2020" box="[151,339,1777,1803]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="443" phylum="Arthropoda" rank="species" species="evae">
|
|||
|
|
<emphasis id="B68B030F1D19CA183CEDBE94FE81F8EF" box="[151,339,1777,1803]" italics="true" pageId="4" pageNumber="443">Ochthebius evae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the distal lobe of the aedeagus of
|
|||
|
|
<taxonomicName id="43FFA49E1D19CA183E93BE94FC2EF8EF" authorityName="Villastrigo & García-Esquivel" authorityYear="2024" box="[745,1020,1777,1803]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="443" phylum="Arthropoda" rank="species" species="vilanovensis" status="sp. nov.">
|
|||
|
|
<emphasis id="B68B030F1D19CA183E93BE94FC2EF8EF" box="[745,1020,1777,1803]" italics="true" pageId="4" pageNumber="443">Ochthebius vilanovensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B68B030F1D19CA18387FBE94FBB3F8EF" bold="true" box="[1029,1121,1777,1803]" pageId="4" pageNumber="443">
|
|||
|
|
<taxonomicNameLabel id="ADB8BE741D19CA18387FBE94FBB3F8EF" box="[1029,1121,1777,1803]" pageId="4" pageNumber="443" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is distinctly larger, with its dorsal margin forming a pronounced 90º angle, and a slight middle compression on the main piece of the aedeagus (for direct comparison see
|
|||
|
|
<figureCitation id="1CC4C3981D19CA183DB6BF5CFDFCF8B7" box="[460,558,1849,1875]" captionStart="FIGURE 3" captionStartId="4.[152,255,1203,1227]" captionTargetBox="[162,1425,284,1169]" captionTargetId="figure-85@4.[151,1436,273,1180]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of the male genitalia of Ochthebius evae Villastrigo, Hernando, Millán & Ribera, 2020 (A and C) and Ochthebius vilanovensis sp. nov. (B and D)." figureDoi="http://doi.org/10.5281/zenodo.14239807" httpUri="https://zenodo.org/record/14239807/files/figure.png" pageId="4" pageNumber="443">Figure 3</figureCitation>
|
|||
|
|
). Additionally, molecular evidence delineates this species, with a divergence greater than 7.63% for COI-3, 7.44% for COI-5 and 2.46% in 16S. Nuclear genes also display differences despite their slow evolutionary pace; 5 differences for 28S and 2 differences for 18S. The new species has only 1 difference in 18S with respect to
|
|||
|
|
<taxonomicName id="43FFA49E1D19CA183DE9BFC0FD78F85B" authorityName="Villastrigo, Hernando, Millan & Ribera" authorityYear="2020" box="[403,682,1957,1983]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="443" phylum="Arthropoda" rank="species" species="cortomaltese">
|
|||
|
|
<emphasis id="B68B030F1D19CA183DE9BFC0FD78F85B" box="[403,682,1957,1983]" italics="true" pageId="4" pageNumber="443">Ochthebius cortomaltese</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, which may point to a closer relationship, although the two species are strongly differentiated overall (see
|
|||
|
|
<figureCitation id="1CC4C3981D19CA183E33BFACFD74F807" box="[585,678,1993,2019]" captionStart="FIGURE 1" captionStartId="2.[152,255,1845,1869]" captionTargetBox="[157,1424,925,1815]" captionTargetId="figure-320@2.[151,1436,921,1821]" captionTargetPageId="2" captionText="FIGURE 1. Selected Bayesian analysis of the subgenus Cobalius. White circles on nodes for support values higher than 0.95. Blue branches represent the Ochthebius biltoni species group." figureDoi="http://doi.org/10.5281/zenodo.14239803" httpUri="https://zenodo.org/record/14239803/files/figure.png" pageId="4" pageNumber="443">Figure 1</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|