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<document id="EC4B871D4F1B32CD5285CAB59D29445B" ID-CLB-Dataset="29146" ID-DOI="10.11646/zootaxa.4422.2.6" ID-GBIF-Dataset="47816d99-6645-43ad-a52d-33cc68a6a7c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1455619" ID-ZooBank="F6658C2B-9681-430A-8975-7B3AE2C233EE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539185629917" checkinUser="plazi" docAuthor="Shear, William A." docDate="2018" docId="03FF87B7FFDAFF8041EAFAF01956F80B" docLanguage="en" docName="zootaxa.4422.2.6.pdf" docOrigin="Zootaxa 4422 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Speleopsobius weaveri Shear 2018, new species" docType="treatment" docVersion="7" lastPageNumber="277" masterDocId="FFC6FFCFFFD6FF92417DFFAA1B3DFFD7" masterDocTitle="The centipede family Anopsobiidae new to North America, with the description of a new genus and species and notes on the Henicopidae of North America and the Anopsobiidae of the Northern Hemisphere (Chilopoda, Lithobiomorpha)" masterLastPageNumber="283" masterPageNumber="259" pageNumber="271" updateTime="1698573810455" updateUser="ExternalLinkService">
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<mods:title id="A60C42ED1055DB18CA050D924554E7D8">The centipede family Anopsobiidae new to North America, with the description of a new genus and species and notes on the Henicopidae of North America and the Anopsobiidae of the Northern Hemisphere (Chilopoda, Lithobiomorpha)</mods:title>
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<mods:namePart id="60B21E7D8E1122AF63FB558F1D7BEBAD">Shear, William A.</mods:namePart>
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<mods:typeOfResource id="9E04F317B5DF6A33F9C7C31CD006315E">text</mods:typeOfResource>
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<mods:title id="8A7701CB3207907384F08F85A76112B0">Zootaxa</mods:title>
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<mods:part id="622B7876F6FA604BEEBCEA7CE0FE38A3">
<mods:date id="C2EA64E22795F27CB44001BEBDF1FB91">2018</mods:date>
<mods:detail id="72FEC8B65621998BD558562BADA30DDC" type="volume">
<mods:number id="2F26F362CBC89CAE23838A843D4676B8">4422</mods:number>
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<mods:identifier id="F09E3BAD44D122767E54CAE1D7468291" type="DOI">10.11646/zootaxa.4422.2.6</mods:identifier>
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<treatment id="03FF87B7FFDAFF8041EAFAF01956F80B" ID-DOI="http://doi.org/10.5281/zenodo.5969837" ID-GBIF-Taxon="148671711" ID-Zenodo-Dep="5969837" LSID="urn:lsid:plazi:treatment:03FF87B7FFDAFF8041EAFAF01956F80B" httpUri="http://treatment.plazi.org/id/03FF87B7FFDAFF8041EAFAF01956F80B" lastPageId="18" lastPageNumber="277" pageId="12" pageNumber="271">
<subSubSection id="C34C652AFFDAFF9E41EAFAF01A30FA41" pageId="12" pageNumber="271" type="nomenclature">
<paragraph id="8BE936A1FFDAFF9E41EAFAF01979FAA2" blockId="12.[151,580,1370,1430]" box="[151,580,1370,1397]" pageId="12" pageNumber="271">
<heading id="D0A181CDFFDAFF9E41EAFAF01979FAA2" bold="true" box="[151,580,1370,1397]" fontSize="11" level="1" pageId="12" pageNumber="271" reason="1">
<emphasis id="B922EAB3FFDAFF9E41EAFAF01979FAA2" bold="true" box="[151,580,1370,1397]" pageId="12" pageNumber="271">
<emphasis id="B922EAB3FFDAFF9E41EAFAF01A90FAA3" bold="true" box="[151,429,1370,1396]" italics="true" pageId="12" pageNumber="271">
<taxonomicName id="4C564D22FFDAFF9E41EAFAF01A9AFAA3" authority="Shear, 2018" authorityName="Shear" authorityYear="2018" box="[151,423,1370,1396]" class="Chilopoda" family="Anopsobiidae" genus="Speleopsobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiida" pageId="12" pageNumber="271" phylum="Arthropoda" rank="species" species="weaveri" status="sp. nov.">Speleopsobius weaveri</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A21157C8FFDAFF9E40C9FAF11979FAA2" box="[436,580,1371,1397]" pageId="12" pageNumber="271" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE936A1FFDAFF9E41EAFAD71A30FA41" blockId="12.[151,580,1370,1430]" box="[151,269,1405,1430]" pageId="12" pageNumber="271">
<figureCitation id="136D2A24FFDAFF9E41EAFAD71A30FA41" box="[151,269,1405,1430]" captionStart-0="FIGURES 1, 2" captionStart-1="FIGURES 3–6" captionStart-2="FIGURES 7–9" captionStart-3="FIGURES 10–13" captionStart-4="FIGURES 14–17" captionStart-5="FIGURES 18–25" captionStartId-0="13.[151,264,1981,2004]" captionStartId-1="14.[151,264,1476,1499]" captionStartId-2="15.[151,264,1969,1992]" captionStartId-3="16.[151,264,1580,1603]" captionStartId-4="17.[151,264,1681,1704]" captionStartId-5="18.[151,264,1715,1738]" captionTargetBox-0="[151,1436,927,1957]" captionTargetBox-1="[151,1436,193,1455]" captionTargetBox-2="[166,1410,692,1942]" captionTargetBox-3="[151,1436,193,1559]" captionTargetBox-4="[151,1436,301,1660]" captionTargetBox-5="[330,1230,433,1688]" captionTargetId-0="figure@13.[151,1436,927,1957]" captionTargetId-1="figure@14.[151,1436,193,1455]" captionTargetId-2="figure@15.[166,1410,692,1942]" captionTargetId-3="figure@16.[151,1436,193,1559]" captionTargetId-4="figure@17.[151,1436,301,1660]" captionTargetId-5="figure@18.[330,1256,409,1694]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="16" captionTargetPageId-4="17" captionTargetPageId-5="18" captionText-0="FIGURES 1, 2. Speleopsobius weaveri, n. sp. 1. Female holotype from Spider Cave, dorsal view. 2. Male paratype from Tee Cave, dorsal view. Numerals indicate tergite numbers." captionText-1="FIGURES 3–6. Speleopsobius weaveri, n. sp. 3. Head of female, dorsal view. 4. Same, ventral view. 5. Head of male, dorsal view. 6. Same, ventral view." captionText-2="FIGURES 7–9. Speleopsobius weaveri, n. sp. 7. Mandible, ectal view. 8. Same, medial view. 9. First maxilla. a, aciculae; t, mandibular teeth; cx, coxosternum; cp, coxal process; t1, first article of telopodite; t2, second article of telopodite." captionText-3="FIGURES 10–13. Speleopsobius weaveri, n. sp. 10. Plumose setae of epistome. 11. Telopodite of second maxilla. 12. Tarsus of same. 13. Tip of tarsus." captionText-4="FIGURES 14–17. Speleopsobius weaveri, n. sp. 14. Posterior end of female, ventral view. 15. Female gonopods, ventral view. 16. Posterior end of male, ventral view. 17. Gonopods of male, ventral view. Cx14, coxa 14; cx15, coxa 15; s14, sternite 14; s15, sternite 15, gs, genital sternite; gp, gonopod; svo, seminal vesicle opening; glo, gland openings." captionText-5="FIGURES 18–25. Speleopsobius weaveri, n. sp. 18. Leg 12. 19. Claw of leg 12. 20. Leg 13. 21. Claw of leg 13. 22. Leg 14. 23. Claw of leg 13. 24. Leg 15. 25. Claw of leg 15." httpUri-0="https://zenodo.org/record/1455621/files/figure.png" httpUri-1="https://zenodo.org/record/1455623/files/figure.png" httpUri-2="https://zenodo.org/record/1455625/files/figure.png" httpUri-3="https://zenodo.org/record/1455627/files/figure.png" httpUri-4="https://zenodo.org/record/1455629/files/figure.png" httpUri-5="https://zenodo.org/record/1455631/files/figure.png" pageId="12" pageNumber="271">Figs. 1–25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34C652AFFDAFF9E41EAFA6F1910F961" pageId="12" pageNumber="271" type="materials_examined">
<paragraph id="8BE936A1FFDAFF9E41EAFA6F1910F961" blockId="12.[151,1437,1477,2006]" pageId="12" pageNumber="271">
<emphasis id="B922EAB3FFDAFF9E41EAFA6F1A42FA09" bold="true" box="[151,383,1477,1502]" pageId="12" pageNumber="271">Material examined</emphasis>
:
<typeStatus id="54ED8803FFDAFF9E40EEFA6F1AC6FA09" box="[403,507,1477,1502]" pageId="12" pageNumber="271" type="holotype">Holotype</typeStatus>
female from Spider Cave, Gooding Co.,
<collectingRegion id="4992F843FFDAFF9E428CFA6F1F09FA09" box="[1009,1076,1477,1502]" country="United States of America" name="Idaho" pageId="12" pageNumber="271">Idaho</collectingRegion>
,
<geoCoordinate id="EE625066FFDAFF9E4539FA6F1FE3FA09" box="[1092,1246,1477,1502]" direction="north" orientation="latitude" pageId="12" pageNumber="271" precision="1" value="43.017506">43.017506°N</geoCoordinate>
,
<geoCoordinate id="EE625066FFDAFF9E4593FA6F1EA5FA09" box="[1262,1432,1477,1502]" direction="west" orientation="longitude" pageId="12" pageNumber="271" precision="1" value="-114.8354">114.835403°W</geoCoordinate>
, collected
<date id="FFE81061FFDAFF9E407AFA401A9DF9D5" box="[263,416,1513,1538]" pageId="12" pageNumber="271" value="2000-07-17">17 July 2000</date>
by D. A. Hubbard, Jr.;
<typeStatus id="54ED8803FFDAFF9E43BDFA40181DF9D5" box="[704,800,1514,1538]" pageId="12" pageNumber="271" type="paratype">paratype</typeStatus>
male from Tee Cave, Lincoln Co.,
<collectingRegion id="4992F843FFDAFF9E45B4FA431E31F9D5" box="[1225,1292,1513,1538]" country="United States of America" name="Idaho" pageId="12" pageNumber="271">Idaho</collectingRegion>
,
<geoCoordinate id="EE625066FFDAFF9E4466FA401EA5F9D5" box="[1307,1432,1514,1538]" direction="north" orientation="latitude" pageId="12" pageNumber="271" precision="5" value="43.0812">43.0812°N</geoCoordinate>
,
<geoCoordinate id="EE625066FFDAFF9E41EAF9A41A19F9F1" box="[151,292,1550,1574]" direction="west" orientation="longitude" pageId="12" pageNumber="271" precision="5" value="-114.4099">114.4099°W</geoCoordinate>
, collected
<date id="FFE81061FFDAFF9E40E3F9A4190DF9F1" box="[414,560,1549,1574]" pageId="12" pageNumber="271" value="2007-07-18">18 July 2007</date>
by D. A. Hubbard, Jr.;
<typeStatus id="54ED8803FFDAFF9E4246F9A418A1F9F1" box="[827,924,1550,1574]" pageId="12" pageNumber="271" type="paratype">paratype</typeStatus>
male from Tee Cave, collected
<date id="FFE81061FFDAFF9E447BF9A41EA1F9F1" box="[1286,1436,1550,1574]" pageId="12" pageNumber="271" value="2007-06-30">30 June 2007</date>
by C. Richart and A. Fusek;
<typeStatus id="54ED8803FFDAFF9E40A8F9981908F99D" box="[469,565,1586,1610]" pageId="12" pageNumber="271" type="paratype">paratype</typeStatus>
female from Tee Cave, collected
<date id="FFE81061FFDAFF9E42D0F99B1F00F99D" box="[941,1085,1585,1610]" pageId="12" pageNumber="271" value="1999-07-16">16 July 1999</date>
by D. A. Hubbard, Jr. The
<typeStatus id="54ED8803FFDAFF9E4411F9981EA1F99D" box="[1388,1436,1586,1610]" pageId="12" pageNumber="271">type</typeStatus>
material includes SEM stub WS 29-8. All specimens deposited in the
<collectionCode id="ED47AE64FFDAFF9E42D6F9FC18D8F9B9" box="[939,997,1622,1646]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="12" pageNumber="271">CAS</collectionCode>
. As the largest (and thus most likely to be fully mature) and only complete specimen, the Spider Cave female was selected as the
<typeStatus id="54ED8803FFDAFF9E45F4F9D31FD2F945" box="[1161,1263,1657,1682]" pageId="12" pageNumber="271" type="holotype">holotype</typeStatus>
. Spider Cave is sometimes known as Octopus Cave.
</paragraph>
</subSubSection>
<subSubSection id="C34C652AFFDAFF9E41BAF96B19B9F90D" box="[199,644,1729,1754]" pageId="12" pageNumber="271" type="diagnosis">
<paragraph id="8BE936A1FFDAFF9E41BAF96B19B9F90D" blockId="12.[151,1437,1477,2006]" box="[199,644,1729,1754]" pageId="12" pageNumber="271">
<emphasis id="B922EAB3FFDAFF9E41BAF96B1A07F90D" bold="true" box="[199,314,1729,1754]" pageId="12" pageNumber="271">Diagnosis</emphasis>
: As for the genus, see above.
</paragraph>
</subSubSection>
<subSubSection id="C34C652AFFDAFF9E41BAF94F1897F891" pageId="12" pageNumber="271" type="etymology">
<paragraph id="8BE936A1FFDAFF9E41BAF94F1897F891" blockId="12.[151,1437,1477,2006]" pageId="12" pageNumber="271">
<emphasis id="B922EAB3FFDAFF9E41BAF94F1A75F929" bold="true" box="[199,328,1765,1790]" pageId="12" pageNumber="271">Etymology</emphasis>
: The species epithet thanks and honors Dr. Andrew A. Weaver, Professor of Biology Emeritus, College of Wooster, Wooster,
<collectingRegion id="4992F843FFDAFF9E4099F8A31922F8F5" box="[484,543,1801,1826]" country="United States of America" name="Ohio" pageId="12" pageNumber="271">Ohio</collectingRegion>
, whose extensive unpublished work on North American centipedes inspired this study and who was an early mentor (from 1959 onward) of the author.
</paragraph>
</subSubSection>
<subSubSection id="C34C652AFFDAFF8041BAF8FB1AD2FEB2" lastPageId="18" lastPageNumber="277" pageId="12" pageNumber="271" type="description">
<paragraph id="8BE936A1FFDAFF9D41BAF8FB1929FEB2" blockId="12.[151,1437,1477,2006]" lastBlockId="15.[151,1436,151,357]" lastPageId="15" lastPageNumber="274" pageId="12" pageNumber="271">
<emphasis id="B922EAB3FFDAFF9E41BAF8FB1A6CF8BD" bold="true" box="[199,337,1873,1898]" pageId="12" pageNumber="271">Description</emphasis>
: Based on female
<typeStatus id="54ED8803FFDAFF9E4353F8FB19ADF8BD" box="[558,656,1873,1898]" pageId="12" pageNumber="271" type="holotype">holotype</typeStatus>
from Spider Cave: Length 20.5 mm. Color uniformly pale yelloworange (
<figureCitation id="136D2A24FFDAFF9E418BF8DF1A06F859" box="[246,315,1909,1934]" captionStart="FIGURES 1, 2" captionStartId="13.[151,264,1981,2004]" captionTargetBox="[151,1436,927,1957]" captionTargetId="figure@13.[151,1436,927,1957]" captionTargetPageId="13" captionText="FIGURES 1, 2. Speleopsobius weaveri, n. sp. 1. Female holotype from Spider Cave, dorsal view. 2. Male paratype from Tee Cave, dorsal view. Numerals indicate tergite numbers." httpUri="https://zenodo.org/record/1455621/files/figure.png" pageId="12" pageNumber="271">Fig. 1</figureCitation>
).
<emphasis id="B922EAB3FFDAFF9E402AF8DF1AABF859" bold="true" box="[343,406,1909,1934]" pageId="12" pageNumber="271">Head</emphasis>
. Length of head shield 1.8 mm. Transverse suture of head (
<figureCitation id="136D2A24FFDAFF9E4526F8DF1F9CF859" box="[1115,1185,1909,1934]" captionStart="FIGURES 3–6" captionStartId="14.[151,264,1476,1499]" captionTargetBox="[151,1436,193,1455]" captionTargetId="figure@14.[151,1436,193,1455]" captionTargetPageId="14" captionText="FIGURES 3–6. Speleopsobius weaveri, n. sp. 3. Head of female, dorsal view. 4. Same, ventral view. 5. Head of male, dorsal view. 6. Same, ventral view." httpUri="https://zenodo.org/record/1455623/files/figure.png" pageId="12" pageNumber="271">Fig. 3</figureCitation>
) parabolic, anteriorly slightly sinuate, extending back 42% of head length. Median notch shallow, median furrow (suture) absent. Posterior margin of head strongly rebordered, rebordering interrupted in posterior median line by short suture or notch. Head depigmented anterior to antennocellar sutures. Setation of head sparse, some setae rubbed off; two pairs of setae near median notch; at least four long setae on each side of head posterior to transverse suture; setae of head shield disc all very small, probably four pairs arranged symmetrically. Antennae (
<figureCitation id="136D2A24FFDBFF9F451AFF751F95FF2F" box="[1127,1192,223,248]" captionStart="FIGURES 1, 2" captionStartId="13.[151,264,1981,2004]" captionTargetBox="[151,1436,927,1957]" captionTargetId="figure@13.[151,1436,927,1957]" captionTargetPageId="13" captionText="FIGURES 1, 2. Speleopsobius weaveri, n. sp. 1. Female holotype from Spider Cave, dorsal view. 2. Male paratype from Tee Cave, dorsal view. Numerals indicate tergite numbers." httpUri="https://zenodo.org/record/1455621/files/figure.png" pageId="13" pageNumber="272">Fig. 1</figureCitation>
) long, approximately half as long as body, with 60 articles, the two basal articles larger than more distal ones, well-furnished with slit sense organs around distal rim. Tömösvàry organ small, oval, on cephalic pleurite.
<emphasis id="B922EAB3FFDBFF9F45E6FE8D1E17FE97" bold="true" box="[1179,1322,295,320]" pageId="13" pageNumber="272">Mouthparts</emphasis>
. Clypeus projecting, rounded, with single large seta medially, two smaller setae on either side. Labrum deeply notched, with single median hastate tooth; labral fringed setae narrow, feathery, with branches all along length of setae. Mandibles (
<figureCitation id="136D2A24FFDBFF9F4061FE3E1AB9FE7B" box="[284,388,404,429]" captionStart="FIGURES 7–9" captionStartId="15.[151,264,1969,1992]" captionTargetBox="[166,1410,692,1942]" captionTargetId="figure@15.[166,1410,692,1942]" captionTargetPageId="15" captionText="FIGURES 7–9. Speleopsobius weaveri, n. sp. 7. Mandible, ectal view. 8. Same, medial view. 9. First maxilla. a, aciculae; t, mandibular teeth; cx, coxosternum; cp, coxal process; t1, first article of telopodite; t2, second article of telopodite." httpUri="https://zenodo.org/record/1455625/files/figure.png" pageId="13" pageNumber="272">Figs. 7, 8</figureCitation>
) with about 12 slightly curved aciculae, each minutely dentate along both sides of distal third. Four mandibular teeth, distal the largest, decreasing in size ectally, with three cusps (outer cusp obselete on ectalmost tooth); accessory denticles as small acute nodules decreasing in size proximally. Fringe of about 10 branching bristles, decreasing in size mesally, with broad bases, fringe beginning at base of each bristle, extending to tip; bristles abruptly transitioning to field of finely branched scales extending to furry pad. Furry pad not clearly delimited from scale field, consisting of finely branched, narrow scales. First maxillae (
<figureCitation id="136D2A24FFDBFF9F4515FDED1F96FDB7" box="[1128,1195,583,608]" captionStart="FIGURES 7–9" captionStartId="15.[151,264,1969,1992]" captionTargetBox="[166,1410,692,1942]" captionTargetId="figure@15.[166,1410,692,1942]" captionTargetPageId="15" captionText="FIGURES 7–9. Speleopsobius weaveri, n. sp. 7. Mandible, ectal view. 8. Same, medial view. 9. First maxilla. a, aciculae; t, mandibular teeth; cx, coxosternum; cp, coxal process; t1, first article of telopodite; t2, second article of telopodite." httpUri="https://zenodo.org/record/1455625/files/figure.png" pageId="13" pageNumber="272">Fig. 9</figureCitation>
) with broad but short sternite, sternite fused to coxae. Coxal process with about six simple setae. Telopodite with inner row of at least eight plumose setae, numerous simple setae form irregular row lateral of plumose setae. Both telopodite articles with numerous simple setae laterally. Second maxillae (
<figureCitation id="136D2A24FFDBFF9F426BFD1E18A1FD1B" box="[790,924,692,717]" captionStart="FIGURES 10–13" captionStartId="16.[151,264,1580,1603]" captionTargetBox="[151,1436,193,1559]" captionTargetId="figure@16.[151,1436,193,1559]" captionTargetPageId="16" captionText="FIGURES 10–13. Speleopsobius weaveri, n. sp. 10. Plumose setae of epistome. 11. Telopodite of second maxilla. 12. Tarsus of same. 13. Tip of tarsus." httpUri="https://zenodo.org/record/1455627/files/figure.png" pageId="13" pageNumber="272">
Figs. 11
<emphasis id="B922EAB3FFDBFF9F420FFD1E18BDFD1A" bold="true" box="[882,896,692,717]" pageId="13" pageNumber="272">–</emphasis>
13
</figureCitation>
) with short, subtriangular coxae; telopodites with four articles, the trochanter short, both joints present; femur curved, about twice as long as tibia; tarsus slightly longer than tibia. Inner surface of tarsus (
<figureCitation id="136D2A24FFDBFF9F43AFFD561818FCC3" box="[722,805,764,789]" captionStart="FIGURES 10–13" captionStartId="16.[151,264,1580,1603]" captionTargetBox="[151,1436,193,1559]" captionTargetId="figure@16.[151,1436,193,1559]" captionTargetPageId="16" captionText="FIGURES 10–13. Speleopsobius weaveri, n. sp. 10. Plumose setae of epistome. 11. Telopodite of second maxilla. 12. Tarsus of same. 13. Tip of tarsus." httpUri="https://zenodo.org/record/1455627/files/figure.png" pageId="13" pageNumber="272">Fig. 12</figureCitation>
) concave (perhaps poorly sclerotized) densely set with plumose setae. Claw with perhaps five subequal branches, claw subtended laterally by two long setae plumose in their distal halves (
<figureCitation id="136D2A24FFDBFF9F4013FCEE1AFCFC8B" box="[366,449,836,861]" captionStart="FIGURES 10–13" captionStartId="16.[151,264,1580,1603]" captionTargetBox="[151,1436,193,1559]" captionTargetId="figure@16.[151,1436,193,1559]" captionTargetPageId="16" captionText="FIGURES 10–13. Speleopsobius weaveri, n. sp. 10. Plumose setae of epistome. 11. Telopodite of second maxilla. 12. Tarsus of same. 13. Tip of tarsus." httpUri="https://zenodo.org/record/1455627/files/figure.png" pageId="13" pageNumber="272">Fig. 13</figureCitation>
).
<emphasis id="B922EAB3FFDBFF9F40AAFCEE1932FC8A" bold="true" box="[471,527,836,861]" pageId="13" pageNumber="272">Legs</emphasis>
. Forcipules robust (
<figureCitation id="136D2A24FFDBFF9F438FFCEE180AFC8B" box="[754,823,836,861]" captionStart="FIGURES 3–6" captionStartId="14.[151,264,1476,1499]" captionTargetBox="[151,1436,193,1455]" captionTargetId="figure@14.[151,1436,193,1455]" captionTargetPageId="14" captionText="FIGURES 3–6. Speleopsobius weaveri, n. sp. 3. Head of female, dorsal view. 4. Same, ventral view. 5. Head of male, dorsal view. 6. Same, ventral view." httpUri="https://zenodo.org/record/1455623/files/figure.png" pageId="13" pageNumber="272">Fig. 4</figureCitation>
). Pleural collar continuous, without median sclerite or median suture. Coxosternite broad, coxal lobes set off by shallow grooves, lateral margins of sternal part gently, evenly curved, distal margin slightly indented, disc with four pairs of prominent setae, scattered setae distally. Coxosternal teeth 3 + 3; porodonts absent or indistinguishable from setae. Prefemur of telopodite about twice as long as broad, sparsely setose. Femur and tibia of telopodite subequal, broader than long, with few setae. Tarsungulum (“claw”) about 5/6ths as long as prefemur, tarsal section set off from pretarsal section by small median nodule, pretarsal section gently curving, about three times longer than tarsal section.
<emphasis id="B922EAB3FFD8FF9C45A7F91A1E04F91E" bold="true" box="[1242,1337,1712,1737]" pageId="14" pageNumber="273">Tergites</emphasis>
(
<figureCitation id="136D2A24FFD8FF9C4431F91A1EADF91E" box="[1356,1424,1712,1737]" captionStart="FIGURES 1, 2" captionStartId="13.[151,264,1981,2004]" captionTargetBox="[151,1436,927,1957]" captionTargetId="figure@13.[151,1436,927,1957]" captionTargetPageId="13" captionText="FIGURES 1, 2. Speleopsobius weaveri, n. sp. 1. Female holotype from Spider Cave, dorsal view. 2. Male paratype from Tee Cave, dorsal view. Numerals indicate tergite numbers." httpUri="https://zenodo.org/record/1455621/files/figure.png" pageId="14" pageNumber="273">Fig. 1</figureCitation>
) smooth and shining, with few scattered very small setae across surface, with three or four marginal setae including one at posteriolateral angle, posterior margins with paired paramedian setae. All tergites narrowly rebordered laterally and posteriorly. Tergites 3, 5 and 8 with median embayment, deepest and most obvious on tergite 8. Tergite 14 with slightly extended, triangular posteriolateral corners. Other tergites with straight or slightly convex posterior margins.
<emphasis id="B922EAB3FFD8FF9C4013F8CE1AEBF8AA" bold="true" box="[366,470,1892,1917]" pageId="14" pageNumber="273">Sternites</emphasis>
sparsely setose, posterior margins straight, with posterior rows of four to six setae. Spiracles on segments 3, 5, 8, 10, 12 and 14.
<emphasis id="B922EAB3FFD8FF9C43D9F82218BFF876" bold="true" box="[676,898,1928,1953]" pageId="14" pageNumber="273">Posteriormost legs</emphasis>
. Coxal pores in single series, round to broadly oval, distribution 3, 5, 5, 6 on coxae 12
<emphasis id="B922EAB3FFD8FF9C4337F8061965F812" bold="true" box="[586,600,1964,1989]" pageId="14" pageNumber="273">–</emphasis>
15, first in series on coxa 15 about ½ the size of more distal pores. Coxa 15 (
<figureCitation id="136D2A24FFD8FF9C41E2F87A1BCEF83E" box="[159,243,2000,2025]" captionStart="FIGURES 14–17" captionStartId="17.[151,264,1681,1704]" captionTargetBox="[151,1436,301,1660]" captionTargetId="figure@17.[151,1436,301,1660]" captionTargetPageId="17" captionText="FIGURES 14–17. Speleopsobius weaveri, n. sp. 14. Posterior end of female, ventral view. 15. Female gonopods, ventral view. 16. Posterior end of male, ventral view. 17. Gonopods of male, ventral view. Cx14, coxa 14; cx15, coxa 15; s14, sternite 14; s15, sternite 15, gs, genital sternite; gp, gonopod; svo, seminal vesicle opening; glo, gland openings." httpUri="https://zenodo.org/record/1455629/files/figure.png" pageId="14" pageNumber="273">Fig. 14</figureCitation>
) with small, but distinct, coxal spine bearing dark, sclerotized tip without distinct articulation. Coxa 14 distally rounded, spine absent. More anterior coxae without spines. Proportions of legs 12–15 as in
<figureCitation id="136D2A24FFD9FF9D4472FF3D1EAAFF67" box="[1295,1431,151,176]" captionStart="FIGURES 18–25" captionStartId="18.[151,264,1715,1738]" captionTargetBox="[330,1230,433,1688]" captionTargetId="figure@18.[330,1256,409,1694]" captionTargetPageId="18" captionText="FIGURES 18–25. Speleopsobius weaveri, n. sp. 18. Leg 12. 19. Claw of leg 12. 20. Leg 13. 21. Claw of leg 13. 22. Leg 14. 23. Claw of leg 13. 24. Leg 15. 25. Claw of leg 15." httpUri="https://zenodo.org/record/1455631/files/figure.png" pageId="15" pageNumber="274">
Figs. 18
<emphasis id="B922EAB3FFD9FF9D4416FF3D1E44FF67" bold="true" box="[1387,1401,151,176]" pageId="15" pageNumber="274">–</emphasis>
25
</figureCitation>
,
<tableCitation id="C6D4031AFFD9FF9D41EAFF161BCFFF03" box="[151,242,188,213]" captionStart="TABLE 2" captionStartId="15.[151,239,404,427]" captionText="TABLE 2. Lengths of Podomeres in mm." pageId="15" pageNumber="274">Table 2</tableCitation>
. Tibia of legs 1–13 with distal spine. Legs 13–15 with biarticulate tarsi. Tarsi of more anterior legs proportionally similar, with definite flexion point at same relative position as articulations of tarsi 13–15, but without distinct joint. Claws of legs 1–12 (
<figureCitation id="136D2A24FFD9FF9D43F9FEAE19EBFECB" box="[644,726,260,285]" captionStart="FIGURES 18–25" captionStartId="18.[151,264,1715,1738]" captionTargetBox="[330,1230,433,1688]" captionTargetId="figure@18.[330,1256,409,1694]" captionTargetPageId="18" captionText="FIGURES 18–25. Speleopsobius weaveri, n. sp. 18. Leg 12. 19. Claw of leg 12. 20. Leg 13. 21. Claw of leg 13. 22. Leg 14. 23. Claw of leg 13. 24. Leg 15. 25. Claw of leg 15." httpUri="https://zenodo.org/record/1455631/files/figure.png" pageId="15" pageNumber="274">Fig. 19</figureCitation>
) with basal, anterior accessory branch, basal, posterior bristle little more than half length of claw, claws 13–14 with posterior bristle only (
<figureCitation id="136D2A24FFD9FF9D4295FE8D1F52FE97" box="[1000,1135,295,320]" captionStart="FIGURES 18–25" captionStartId="18.[151,264,1715,1738]" captionTargetBox="[330,1230,433,1688]" captionTargetId="figure@18.[330,1256,409,1694]" captionTargetPageId="18" captionText="FIGURES 18–25. Speleopsobius weaveri, n. sp. 18. Leg 12. 19. Claw of leg 12. 20. Leg 13. 21. Claw of leg 13. 22. Leg 14. 23. Claw of leg 13. 24. Leg 15. 25. Claw of leg 15." httpUri="https://zenodo.org/record/1455631/files/figure.png" pageId="15" pageNumber="274">Figs. 21, 23</figureCitation>
), claw 15 (
<figureCitation id="136D2A24FFD9FF9D4597FE8D1E06FE97" box="[1258,1339,295,320]" captionStart="FIGURES 18–25" captionStartId="18.[151,264,1715,1738]" captionTargetBox="[330,1230,433,1688]" captionTargetId="figure@18.[330,1256,409,1694]" captionTargetPageId="18" captionText="FIGURES 18–25. Speleopsobius weaveri, n. sp. 18. Leg 12. 19. Claw of leg 12. 20. Leg 13. 21. Claw of leg 13. 22. Leg 14. 23. Claw of leg 13. 24. Leg 15. 25. Claw of leg 15." httpUri="https://zenodo.org/record/1455631/files/figure.png" pageId="15" pageNumber="274">Fig. 25</figureCitation>
) lacking both accessory branch and bristle.
</paragraph>
<caption id="DF296629FFDBFF9F41EAF8171981F825" httpUri="https://zenodo.org/record/1455621/files/figure.png" pageId="13" pageNumber="272" startId="13.[151,264,1981,2004]" targetBox="[151,1436,927,1957]" targetPageId="13">
<paragraph id="8BE936A1FFDBFF9F41EAF8171981F825" blockId="13.[151,1436,1981,2034]" pageId="13" pageNumber="272">
<emphasis id="B922EAB3FFDBFF9F41EAF8171A01F804" bold="true" box="[151,316,1981,2004]" pageId="13" pageNumber="272">FIGURES 1, 2.</emphasis>
<taxonomicName id="4C564D22FFDBFF9F403EF8171916F803" authority="Shear, 2018" authorityName="Shear" authorityYear="2018" box="[323,555,1981,2004]" class="Chilopoda" family="Anopsobiidae" genus="Speleopsobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiida" pageId="13" pageNumber="272" phylum="Arthropoda" rank="species" species="weaveri" status="sp. nov.">
<emphasis id="B922EAB3FFDBFF9F403EF8171916F803" box="[323,555,1981,2004]" italics="true" pageId="13" pageNumber="272">Speleopsobius weaveri</emphasis>
</taxonomicName>
,
<emphasis id="B922EAB3FFDBFF9F4344F81419B0F804" bold="true" box="[569,653,1982,2003]" pageId="13" pageNumber="272">
<taxonomicNameLabel id="A21157C8FFDBFF9F4344F814194FF804" box="[569,626,1982,2003]" pageId="13" pageNumber="272" rank="species">n. sp.</taxonomicNameLabel>
1.
</emphasis>
Female holotype from Spider Cave, dorsal view.
<emphasis id="B922EAB3FFDBFF9F45F1F8141FA3F804" bold="true" box="[1164,1182,1982,2003]" pageId="13" pageNumber="272">2.</emphasis>
Male paratype from Tee Cave, dorsal view. Numerals indicate tergite numbers.
</paragraph>
</caption>
<caption id="DF296629FFD8FF9C41EAFA6E1A88FA2E" httpUri="https://zenodo.org/record/1455623/files/figure.png" pageId="14" pageNumber="273" startId="14.[151,264,1476,1499]" targetBox="[151,1436,193,1455]" targetPageId="14">
<paragraph id="8BE936A1FFD8FF9C41EAFA6E1A88FA2E" blockId="14.[151,1436,1476,1529]" pageId="14" pageNumber="273">
<emphasis id="B922EAB3FFD8FF9C41EAFA6E1A09FA0D" bold="true" box="[151,308,1476,1499]" pageId="14" pageNumber="273">FIGURES 3–6</emphasis>
.
<emphasis id="B922EAB3FFD8FF9C403FFA6E190CFA0C" box="[322,561,1476,1499]" italics="true" pageId="14" pageNumber="273">
<taxonomicName id="4C564D22FFD8FF9C403FFA6E1910FA0C" authority="Shear, 2018" authorityName="Shear" authorityYear="2018" box="[322,557,1476,1499]" class="Chilopoda" family="Anopsobiidae" genus="Speleopsobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiida" pageId="14" pageNumber="273" phylum="Arthropoda" rank="species" species="weaveri" status="sp. nov.">Speleopsobius weaveri</taxonomicName>
,
</emphasis>
<emphasis id="B922EAB3FFD8FF9C4345FA6F19BBFA0D" bold="true" box="[568,646,1477,1498]" pageId="14" pageNumber="273">
<taxonomicNameLabel id="A21157C8FFD8FF9C4345FA6F194FFA0D" box="[568,626,1477,1498]" pageId="14" pageNumber="273" rank="species">n. sp.</taxonomicNameLabel>
3
</emphasis>
. Head of female, dorsal view.
<emphasis id="B922EAB3FFD8FF9C42BCFA6F18F3FA0D" bold="true" box="[961,974,1477,1498]" pageId="14" pageNumber="273">4</emphasis>
. Same, ventral view.
<emphasis id="B922EAB3FFD8FF9C45D1FA6F1F84FA0D" bold="true" box="[1196,1209,1477,1498]" pageId="14" pageNumber="273">5</emphasis>
. Head of male, dorsal view.
<emphasis id="B922EAB3FFD8FF9C41AEFA491BDDFA2F" bold="true" box="[211,224,1507,1528]" pageId="14" pageNumber="273">6</emphasis>
. Same, ventral view.
</paragraph>
</caption>
<caption id="DF296629FFD9FF9D41EAFE3E196EFE7C" box="[151,595,404,427]" pageId="15" pageNumber="274">
<paragraph id="8BE936A1FFD9FF9D41EAFE3E196EFE7C" blockId="15.[151,595,404,427]" box="[151,595,404,427]" pageId="15" pageNumber="274">
<emphasis id="B922EAB3FFD9FF9D41EAFE3E1A34FE7C" bold="true" box="[151,265,404,427]" pageId="15" pageNumber="274">TABLE 2.</emphasis>
Lengths of Podomeres in mm.
</paragraph>
</caption>
<paragraph id="8BE936A1FFD9FF9D41E2FE6D1E18FD55" pageId="15" pageNumber="274">
<table id="F956C401FFD9006D41E2FE6C1E58FD55" box="[159,1381,454,642]" gridcols="7" gridrows="5" pageId="15" pageNumber="274">
<tr id="356634E3FFD9006D41E2FE6C1E58FE0B" box="[159,1381,454,476]" gridrow="0" pageId="15" pageNumber="274">
<th id="76B75D9FFFD9006D41E2FE6C1BFBFE0B" box="[159,198,454,476]" gridcol="0" gridrow="0" pageId="15" pageNumber="274">Leg</th>
<th id="76B75D9FFFD9006D402BFE6C1A89FE0B" box="[342,436,454,476]" gridcol="1" gridrow="0" pageId="15" pageNumber="274">Prefemur</th>
<th id="76B75D9FFFD9006D4373FE6C196DFE0B" box="[526,592,454,476]" gridcol="2" gridrow="0" pageId="15" pageNumber="274">Femur</th>
<th id="76B75D9FFFD9006D43B8FE6C19C4FE0B" box="[709,761,454,476]" gridcol="3" gridrow="0" pageId="15" pageNumber="274">Tibia</th>
<th id="76B75D9FFFD9006D4200FE6C18EFFE0B" box="[893,978,454,476]" gridcol="4" gridrow="0" pageId="15" pageNumber="274">Tarsus 1</th>
<th id="76B75D9FFFD9006D4549FE6C1FB4FE0B" box="[1076,1161,454,476]" gridcol="5" gridrow="0" pageId="15" pageNumber="274">Tarsus 2</th>
<th id="76B75D9FFFD9006D4591FE6C1E58FE0B" box="[1260,1381,454,476]" gridcol="6" gridrow="0" pageId="15" pageNumber="274">Total length</th>
</tr>
<tr id="356634E3FFD9006D41E2FE5A1E58FDD1" box="[159,1381,496,518]" gridrow="1" pageId="15" pageNumber="274">
<th id="76B75D9FFFD9006D41E2FE5A1BFBFDD1" box="[159,198,496,518]" gridcol="0" gridrow="1" pageId="15" pageNumber="274">12</th>
<td id="76B75D9FFFD9006D402BFE5A1A89FDD1" box="[342,436,496,518]" gridcol="1" gridrow="1" pageId="15" pageNumber="274">1.43</td>
<td id="76B75D9FFFD9006D4373FE5A196DFDD1" box="[526,592,496,518]" gridcol="2" gridrow="1" pageId="15" pageNumber="274">1.83</td>
<td id="76B75D9FFFD9006D43B8FE5A19C4FDD1" box="[709,761,496,518]" gridcol="3" gridrow="1" pageId="15" pageNumber="274">2.46</td>
<td id="76B75D9FFFD9006D4200FE5A18EFFDD1" box="[893,978,496,518]" gridcol="4" gridrow="1" pageId="15" pageNumber="274">2.54</td>
<td id="76B75D9FFFD9006D4549FE5A1FB4FDD1" box="[1076,1161,496,518]" gridcol="5" gridrow="1" pageId="15" pageNumber="274">-</td>
<td id="76B75D9FFFD9006D4591FE5A1E58FDD1" box="[1260,1381,496,518]" gridcol="6" gridrow="1" pageId="15" pageNumber="274">8.26</td>
</tr>
<tr id="356634E3FFD9006D41E2FDB31E58FDF8" box="[159,1381,537,559]" gridrow="2" pageId="15" pageNumber="274">
<th id="76B75D9FFFD9006D41E2FDB31BFBFDF8" box="[159,198,537,559]" gridcol="0" gridrow="2" pageId="15" pageNumber="274">13</th>
<td id="76B75D9FFFD9006D402BFDB31A89FDF8" box="[342,436,537,559]" gridcol="1" gridrow="2" pageId="15" pageNumber="274">1.59</td>
<td id="76B75D9FFFD9006D4373FDB3196DFDF8" box="[526,592,537,559]" gridcol="2" gridrow="2" pageId="15" pageNumber="274">1.90</td>
<td id="76B75D9FFFD9006D43B8FDB319C4FDF8" box="[709,761,537,559]" gridcol="3" gridrow="2" pageId="15" pageNumber="274">2.46</td>
<td id="76B75D9FFFD9006D4200FDB318EFFDF8" box="[893,978,537,559]" gridcol="4" gridrow="2" pageId="15" pageNumber="274">2.38</td>
<td id="76B75D9FFFD9006D4549FDB31FB4FDF8" box="[1076,1161,537,559]" gridcol="5" gridrow="2" pageId="15" pageNumber="274">1,14</td>
<td id="76B75D9FFFD9006D4591FDB31E58FDF8" box="[1260,1381,537,559]" gridcol="6" gridrow="2" pageId="15" pageNumber="274">9.47</td>
</tr>
<tr id="356634E3FFD9006D41E2FDE81E58FD8F" box="[159,1381,578,600]" gridrow="3" pageId="15" pageNumber="274">
<th id="76B75D9FFFD9006D41E2FDE81BFBFD8F" box="[159,198,578,600]" gridcol="0" gridrow="3" pageId="15" pageNumber="274">14</th>
<td id="76B75D9FFFD9006D402BFDE81A89FD8F" box="[342,436,578,600]" gridcol="1" gridrow="3" pageId="15" pageNumber="274">1.90</td>
<td id="76B75D9FFFD9006D4373FDE8196DFD8F" box="[526,592,578,600]" gridcol="2" gridrow="3" pageId="15" pageNumber="274">2.30</td>
<td id="76B75D9FFFD9006D43B8FDE819C4FD8F" box="[709,761,578,600]" gridcol="3" gridrow="3" pageId="15" pageNumber="274">2.78</td>
<td id="76B75D9FFFD9006D4200FDE818EFFD8F" box="[893,978,578,600]" gridcol="4" gridrow="3" pageId="15" pageNumber="274">2.67</td>
<td id="76B75D9FFFD9006D4549FDE81FB4FD8F" box="[1076,1161,578,600]" gridcol="5" gridrow="3" pageId="15" pageNumber="274">1.35</td>
<td id="76B75D9FFFD9006D4591FDE81E58FD8F" box="[1260,1381,578,600]" gridcol="6" gridrow="3" pageId="15" pageNumber="274">11.00</td>
</tr>
<tr id="356634E3FFD9006D41E2FDC61E58FD55" box="[159,1381,620,642]" gridrow="4" pageId="15" pageNumber="274">
<th id="76B75D9FFFD9006D41E2FDC61BFBFD55" box="[159,198,620,642]" gridcol="0" gridrow="4" pageId="15" pageNumber="274">15</th>
<td id="76B75D9FFFD9006D402BFDC61A89FD55" box="[342,436,620,642]" gridcol="1" gridrow="4" pageId="15" pageNumber="274">2.49</td>
<td id="76B75D9FFFD9006D4373FDC6196DFD55" box="[526,592,620,642]" gridcol="2" gridrow="4" pageId="15" pageNumber="274">2.84</td>
<td id="76B75D9FFFD9006D43B8FDC619C4FD55" box="[709,761,620,642]" gridcol="3" gridrow="4" pageId="15" pageNumber="274">3.17</td>
<td id="76B75D9FFFD9006D4200FDC618EFFD55" box="[893,978,620,642]" gridcol="4" gridrow="4" pageId="15" pageNumber="274">2.95</td>
<td id="76B75D9FFFD9006D4549FDC61FB4FD55" box="[1076,1161,620,642]" gridcol="5" gridrow="4" pageId="15" pageNumber="274">1.71</td>
<td id="76B75D9FFFD9006D4591FDC61E58FD55" box="[1260,1381,620,642]" gridcol="6" gridrow="4" pageId="15" pageNumber="274">13.16</td>
</tr>
</table>
</paragraph>
<caption id="DF296629FFD9FF9D41EAF81B1E1DF830" httpUri="https://zenodo.org/record/1455625/files/figure.png" pageId="15" pageNumber="274" startId="15.[151,264,1969,1992]" targetBox="[166,1410,692,1942]" targetPageId="15">
<paragraph id="8BE936A1FFD9FF9D41EAF81B1E1DF830" blockId="15.[151,1435,1969,2023]" pageId="15" pageNumber="274">
<emphasis id="B922EAB3FFD9FF9D41EAF81B1A01F810" bold="true" box="[151,316,1969,1992]" pageId="15" pageNumber="274">FIGURES 7–9.</emphasis>
<emphasis id="B922EAB3FFD9FF9D403EF818190FF81F" box="[323,562,1970,1992]" italics="true" pageId="15" pageNumber="274">
<taxonomicName id="4C564D22FFD9FF9D403EF8181912F81F" authority="Shear, 2018" authorityName="Shear" authorityYear="2018" box="[323,559,1970,1992]" class="Chilopoda" family="Anopsobiidae" genus="Speleopsobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiida" pageId="15" pageNumber="274" phylum="Arthropoda" rank="species" species="weaveri" status="sp. nov.">Speleopsobius weaveri</taxonomicName>
,
</emphasis>
<emphasis id="B922EAB3FFD9FF9D4347F81819B5F810" bold="true" box="[570,648,1970,1992]" pageId="15" pageNumber="274">
<taxonomicNameLabel id="A21157C8FFD9FF9D4347F8181949F81F" box="[570,628,1970,1992]" pageId="15" pageNumber="274" rank="species">n. sp.</taxonomicNameLabel>
7
</emphasis>
. Mandible, ectal view.
<emphasis id="B922EAB3FFD9FF9D4206F81818B5F810" bold="true" box="[891,904,1970,1991]" pageId="15" pageNumber="274">8</emphasis>
. Same, medial view.
<emphasis id="B922EAB3FFD9FF9D451BF8181F4EF810" bold="true" box="[1126,1139,1970,1991]" pageId="15" pageNumber="274">9</emphasis>
. First maxilla.
<emphasis id="B922EAB3FFD9FF9D446CF8181E23F810" bold="true" box="[1297,1310,1970,1991]" pageId="15" pageNumber="274">a</emphasis>
, aciculae;
<emphasis id="B922EAB3FFD9FF9D44F0F8181EA8F810" bold="true" box="[1421,1429,1970,1991]" pageId="15" pageNumber="274">t</emphasis>
, mandibular teeth;
<emphasis id="B922EAB3FFD9FF9D4032F87B1A5AF831" bold="true" box="[335,359,2001,2022]" pageId="15" pageNumber="274">cx</emphasis>
, coxosternum;
<emphasis id="B922EAB3FFD9FF9D4379F87B1920F831" bold="true" box="[516,541,2001,2022]" pageId="15" pageNumber="274">cp</emphasis>
, coxal process;
<emphasis id="B922EAB3FFD9FF9D43BDF87B19EBF831" bold="true" box="[704,726,2001,2022]" pageId="15" pageNumber="274">t1</emphasis>
, first article of telopodite;
<emphasis id="B922EAB3FFD9FF9D4299F87B18C7F831" bold="true" box="[996,1018,2001,2022]" pageId="15" pageNumber="274">t2</emphasis>
, second article of telopodite.
</paragraph>
</caption>
<caption id="DF296629FFC6FF8241EAF9861AB9F9B2" httpUri="https://zenodo.org/record/1455627/files/figure.png" pageId="16" pageNumber="275" startId="16.[151,264,1580,1603]" targetBox="[151,1436,193,1559]" targetPageId="16">
<paragraph id="8BE936A1FFC6FF8241EAF9861AB9F9B2" blockId="16.[151,1436,1580,1637]" pageId="16" pageNumber="275">
<emphasis id="B922EAB3FFC6FF8241EAF9861A6EF996" bold="true" box="[151,339,1580,1603]" pageId="16" pageNumber="275">FIGURES 10–13.</emphasis>
<emphasis id="B922EAB3FFC6FF824025F986197BF995" box="[344,582,1580,1602]" italics="true" pageId="16" pageNumber="275">
<taxonomicName id="4C564D22FFC6FF824025F986197FF995" authority="Shear, 2018" authorityName="Shear" authorityYear="2018" box="[344,578,1580,1602]" class="Chilopoda" family="Anopsobiidae" genus="Speleopsobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiida" pageId="16" pageNumber="275" phylum="Arthropoda" rank="species" species="weaveri" status="sp. nov.">Speleopsobius weaveri</taxonomicName>
,
</emphasis>
<emphasis id="B922EAB3FFC6FF824336F9871994F996" bold="true" box="[587,681,1580,1602]" pageId="16" pageNumber="275">
<taxonomicNameLabel id="A21157C8FFC6FF824336F98719BEF995" box="[587,643,1581,1602]" pageId="16" pageNumber="275" rank="species">n. sp.</taxonomicNameLabel>
10.
</emphasis>
Plumose setae of epistome.
<emphasis id="B922EAB3FFC6FF8242BBF98618DEF996" bold="true" box="[966,995,1580,1601]" pageId="16" pageNumber="275">11.</emphasis>
Telopodite of second maxilla.
<emphasis id="B922EAB3FFC6FF824467F9861E07F996" bold="true" box="[1306,1338,1580,1601]" pageId="16" pageNumber="275">12.</emphasis>
Tarsus of same.
<emphasis id="B922EAB3FFC6FF8241AAF9E51BCAF9B3" bold="true" box="[215,247,1615,1636]" pageId="16" pageNumber="275">13.</emphasis>
Tip of tarsus.
</paragraph>
</caption>
<paragraph id="8BE936A1FFC6FF8241BAF93F18F7F8B5" blockId="16.[151,1437,1685,2034]" pageId="16" pageNumber="275">
Sternite of segment 15 narrowed posteriorly, posterior margin straight, four prominent setae, several thinner ones along posterior margin (
<figureCitation id="136D2A24FFC6FF824098F913194CF905" box="[485,625,1721,1746]" captionStart="FIGURES 14–17" captionStartId="17.[151,264,1681,1704]" captionTargetBox="[151,1436,301,1660]" captionTargetId="figure@17.[151,1436,301,1660]" captionTargetPageId="17" captionText="FIGURES 14–17. Speleopsobius weaveri, n. sp. 14. Posterior end of female, ventral view. 15. Female gonopods, ventral view. 16. Posterior end of male, ventral view. 17. Gonopods of male, ventral view. Cx14, coxa 14; cx15, coxa 15; s14, sternite 14; s15, sternite 15, gs, genital sternite; gp, gonopod; svo, seminal vesicle opening; glo, gland openings." httpUri="https://zenodo.org/record/1455629/files/figure.png" pageId="16" pageNumber="275">Figs. 14, 15</figureCitation>
). First genital sternite with median sulcus, distomesally with four large setae, large seta at posteriolateral angle.
<emphasis id="B922EAB3FFC6FF82430EF97719D1F921" bold="true" box="[627,748,1757,1782]" pageId="16" pageNumber="275">Gonopods</emphasis>
. Gonopodal (
<figureCitation id="136D2A24FFC6FF8242F3F97718DEF921" box="[910,995,1757,1782]" captionStart="FIGURES 14–17" captionStartId="17.[151,264,1681,1704]" captionTargetBox="[151,1436,301,1660]" captionTargetId="figure@17.[151,1436,301,1660]" captionTargetPageId="17" captionText="FIGURES 14–17. Speleopsobius weaveri, n. sp. 14. Posterior end of female, ventral view. 15. Female gonopods, ventral view. 16. Posterior end of male, ventral view. 17. Gonopods of male, ventral view. Cx14, coxa 14; cx15, coxa 15; s14, sternite 14; s15, sternite 15, gs, genital sternite; gp, gonopod; svo, seminal vesicle opening; glo, gland openings." httpUri="https://zenodo.org/record/1455629/files/figure.png" pageId="16" pageNumber="275">Fig. 15</figureCitation>
) first article densely setose on lateral surface, mesal surface glabrous, distoventrally with two subequal acuminate spurs. Second article about half length of first, 5
<emphasis id="B922EAB3FFC6FF824183F88F1A31F8E9" bold="true" box="[254,268,1829,1854]" pageId="16" pageNumber="275">–</emphasis>
6 setae on lateral surface, mesal surface glabrous; terminal article shorter than second article, with single large lateral seta, one or two much smaller ones, claw simple, acuminate.
</paragraph>
<paragraph id="8BE936A1FFC6FF8341BAF8C71BFEFF2F" blockId="16.[151,1437,1685,2034]" lastBlockId="17.[151,1436,151,248]" lastPageId="17" lastPageNumber="276" pageId="16" pageNumber="275">
<emphasis id="B922EAB3FFC6FF8241BAF8C71A47F851" bold="true" box="[199,378,1901,1926]" pageId="16" pageNumber="275">
Male
<typeStatus id="54ED8803FFC6FF824072F8C71A47F851" box="[271,378,1901,1926]" pageId="16" pageNumber="275" type="paratype">paratype</typeStatus>
</emphasis>
(
<figureCitation id="136D2A24FFC6FF8240F3F8C71ADFF851" box="[398,482,1901,1926]" captionStart="FIGURES 1, 2" captionStartId="13.[151,264,1981,2004]" captionTargetBox="[151,1436,927,1957]" captionTargetId="figure@13.[151,1436,927,1957]" captionTargetPageId="13" captionText="FIGURES 1, 2. Speleopsobius weaveri, n. sp. 1. Female holotype from Spider Cave, dorsal view. 2. Male paratype from Tee Cave, dorsal view. Numerals indicate tergite numbers." httpUri="https://zenodo.org/record/1455621/files/figure.png" pageId="16" pageNumber="275">Figs. 2</figureCitation>
,
<figureCitation id="136D2A24FFC6FF82408EF8C7191EF851" box="[499,547,1901,1926]" captionStart="FIGURES 3–6" captionStartId="14.[151,264,1476,1499]" captionTargetBox="[151,1436,193,1455]" captionTargetId="figure@14.[151,1436,193,1455]" captionTargetPageId="14" captionText="FIGURES 3–6. Speleopsobius weaveri, n. sp. 3. Head of female, dorsal view. 4. Same, ventral view. 5. Head of male, dorsal view. 6. Same, ventral view." httpUri="https://zenodo.org/record/1455623/files/figure.png" pageId="16" pageNumber="275">5, 6</figureCitation>
,
<figureCitation id="136D2A24FFC6FF824349F8C71943F851" box="[564,638,1901,1926]" captionStart="FIGURES 14–17" captionStartId="17.[151,264,1681,1704]" captionTargetBox="[151,1436,301,1660]" captionTargetId="figure@17.[151,1436,301,1660]" captionTargetPageId="17" captionText="FIGURES 14–17. Speleopsobius weaveri, n. sp. 14. Posterior end of female, ventral view. 15. Female gonopods, ventral view. 16. Posterior end of male, ventral view. 17. Gonopods of male, ventral view. Cx14, coxa 14; cx15, coxa 15; s14, sternite 14; s15, sternite 15, gs, genital sternite; gp, gonopod; svo, seminal vesicle opening; glo, gland openings." httpUri="https://zenodo.org/record/1455629/files/figure.png" pageId="16" pageNumber="275">16, 17</figureCitation>
) collected by D. A. Hubbard from Tee Cave differing from female
<typeStatus id="54ED8803FFC6FF8241EAF83B1BC4F87D" box="[151,249,1937,1962]" pageId="16" pageNumber="275" type="holotype">holotype</typeStatus>
as follows: length,
<quantity id="4CAE9B44FFC6FF8240ABF8381915F87E" box="[470,552,1938,1962]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="16" pageNumber="275" unit="mm" value="17.0">17 mm</quantity>
. Tergites 3, 5, 8, and 10 with triangular embayment posteriorly, tergite 12 with concave posterior margin (
<figureCitation id="136D2A24FFC6FF8240B4F81F192DF819" box="[457,528,1973,1998]" captionStart="FIGURES 1, 2" captionStartId="13.[151,264,1981,2004]" captionTargetBox="[151,1436,927,1957]" captionTargetId="figure@13.[151,1436,927,1957]" captionTargetPageId="13" captionText="FIGURES 1, 2. Speleopsobius weaveri, n. sp. 1. Female holotype from Spider Cave, dorsal view. 2. Male paratype from Tee Cave, dorsal view. Numerals indicate tergite numbers." httpUri="https://zenodo.org/record/1455621/files/figure.png" pageId="16" pageNumber="275">Fig. 2</figureCitation>
). Coxal pores 3, 4, 4/5, 5, proximal pore on coxa 15 less than half diameter of largest pore (Figs. 16,17). First genital sternite (
<figureCitation id="136D2A24FFC6FF8243D6F87319C6F825" box="[683,763,2009,2034]" captionStart="FIGURES 14–17" captionStartId="17.[151,264,1681,1704]" captionTargetBox="[151,1436,301,1660]" captionTargetId="figure@17.[151,1436,301,1660]" captionTargetPageId="17" captionText="FIGURES 14–17. Speleopsobius weaveri, n. sp. 14. Posterior end of female, ventral view. 15. Female gonopods, ventral view. 16. Posterior end of male, ventral view. 17. Gonopods of male, ventral view. Cx14, coxa 14; cx15, coxa 15; s14, sternite 14; s15, sternite 15, gs, genital sternite; gp, gonopod; svo, seminal vesicle opening; glo, gland openings." httpUri="https://zenodo.org/record/1455629/files/figure.png" pageId="16" pageNumber="275">Fig. 17</figureCitation>
) entire, densely setose. Gonopod (
<figureCitation id="136D2A24FFC6FF824503F8731FF3F825" box="[1150,1230,2009,2034]" captionStart="FIGURES 14–17" captionStartId="17.[151,264,1681,1704]" captionTargetBox="[151,1436,301,1660]" captionTargetId="figure@17.[151,1436,301,1660]" captionTargetPageId="17" captionText="FIGURES 14–17. Speleopsobius weaveri, n. sp. 14. Posterior end of female, ventral view. 15. Female gonopods, ventral view. 16. Posterior end of male, ventral view. 17. Gonopods of male, ventral view. Cx14, coxa 14; cx15, coxa 15; s14, sternite 14; s15, sternite 15, gs, genital sternite; gp, gonopod; svo, seminal vesicle opening; glo, gland openings." httpUri="https://zenodo.org/record/1455629/files/figure.png" pageId="16" pageNumber="275">Fig. 17</figureCitation>
) basal article with about eight setae, second article about half length of first, third article subconical, terminal filamentous part short, about half length of third article, sharply set off at base. Anal coxosternite with 6
<emphasis id="B922EAB3FFC7FF834555FF161F0BFF02" bold="true" box="[1064,1078,188,213]" pageId="17" pageNumber="276">–</emphasis>
8 setae, paired glands (
<figureCitation id="136D2A24FFC7FF834444FF161B8BFF2F" captionStart="FIGURES 14–17" captionStartId="17.[151,264,1681,1704]" captionTargetBox="[151,1436,301,1660]" captionTargetId="figure@17.[151,1436,301,1660]" captionTargetPageId="17" captionText="FIGURES 14–17. Speleopsobius weaveri, n. sp. 14. Posterior end of female, ventral view. 15. Female gonopods, ventral view. 16. Posterior end of male, ventral view. 17. Gonopods of male, ventral view. Cx14, coxa 14; cx15, coxa 15; s14, sternite 14; s15, sternite 15, gs, genital sternite; gp, gonopod; svo, seminal vesicle opening; glo, gland openings." httpUri="https://zenodo.org/record/1455629/files/figure.png" pageId="17" pageNumber="276">Figs. 16, 17</figureCitation>
).
</paragraph>
<caption id="DF296629FFC7FF8341EAF93B1FADF932" httpUri="https://zenodo.org/record/1455629/files/figure.png" pageId="17" pageNumber="276" startId="17.[151,264,1681,1704]" targetBox="[151,1436,301,1660]" targetPageId="17">
<paragraph id="8BE936A1FFC7FF8341EAF93B1FADF932" blockId="17.[151,1437,1681,1765]" pageId="17" pageNumber="276">
<emphasis id="B922EAB3FFC7FF8341EAF93B1A6EF970" bold="true" box="[151,339,1681,1704]" pageId="17" pageNumber="276">FIGURES 14–17.</emphasis>
<taxonomicName id="4C564D22FFC7FF834024F938197DF97F" authority="Shear, 2018" authorityName="Shear" authorityYear="2018" box="[345,576,1682,1704]" class="Chilopoda" family="Anopsobiidae" genus="Speleopsobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiida" pageId="17" pageNumber="276" phylum="Arthropoda" rank="species" species="weaveri" status="sp. nov.">
<emphasis id="B922EAB3FFC7FF834024F938197DF97F" box="[345,576,1682,1704]" italics="true" pageId="17" pageNumber="276">Speleopsobius weaveri</emphasis>
</taxonomicName>
,
<emphasis id="B922EAB3FFC7FF834330F9381991F970" bold="true" box="[589,684,1682,1703]" pageId="17" pageNumber="276">
<taxonomicNameLabel id="A21157C8FFC7FF834330F93819BBF970" box="[589,646,1682,1703]" pageId="17" pageNumber="276" rank="species">n. sp.</taxonomicNameLabel>
14.
</emphasis>
Posterior end of female, ventral view.
<emphasis id="B922EAB3FFC7FF834549F9381F69F970" bold="true" box="[1076,1108,1682,1703]" pageId="17" pageNumber="276">15.</emphasis>
Female gonopods, ventral view.
<emphasis id="B922EAB3FFC7FF8341EAF91A1B8BF912" bold="true" box="[151,182,1712,1733]" pageId="17" pageNumber="276">16.</emphasis>
Posterior end of male, ventral view.
<emphasis id="B922EAB3FFC7FF834348F91A1969F912" bold="true" box="[565,596,1712,1733]" pageId="17" pageNumber="276">17.</emphasis>
Gonopods of male, ventral view.
<emphasis id="B922EAB3FFC7FF8342C9F91A18D1F911" bold="true" box="[948,1004,1712,1734]" pageId="17" pageNumber="276">Cx14</emphasis>
, coxa 14;
<emphasis id="B922EAB3FFC7FF834524F91A1FACF912" bold="true" box="[1113,1169,1712,1733]" pageId="17" pageNumber="276">cx15,</emphasis>
coxa 15;
<emphasis id="B922EAB3FFC7FF834585F91A1E1CF912" bold="true" box="[1272,1313,1712,1733]" pageId="17" pageNumber="276">s14,</emphasis>
sternite 14;
<emphasis id="B922EAB3FFC7FF8341EAF9651BFDF933" bold="true" box="[151,192,1743,1764]" pageId="17" pageNumber="276">s15,</emphasis>
sternite 15,
<emphasis id="B922EAB3FFC7FF834040F9651A67F933" bold="true" box="[317,346,1743,1764]" pageId="17" pageNumber="276">gs,</emphasis>
genital sternite;
<emphasis id="B922EAB3FFC7FF834379F9651918F933" bold="true" box="[516,549,1743,1764]" pageId="17" pageNumber="276">gp,</emphasis>
gonopod;
<emphasis id="B922EAB3FFC7FF8343ECF9651987F933" bold="true" box="[657,698,1743,1764]" pageId="17" pageNumber="276">svo,</emphasis>
seminal vesicle opening;
<emphasis id="B922EAB3FFC7FF8342BCF96518D5F933" bold="true" box="[961,1000,1743,1764]" pageId="17" pageNumber="276">glo,</emphasis>
gland openings.
</paragraph>
</caption>
<paragraph id="8BE936A1FFC7FF8041BAF8A51AD2FEB2" blockId="17.[151,1436,1804,2015]" lastBlockId="18.[151,1437,151,357]" lastPageId="18" lastPageNumber="277" pageId="17" pageNumber="276">
<emphasis id="B922EAB3FFC7FF8341BAF8A51A37F8FF" bold="true" box="[199,266,1807,1832]" pageId="17" pageNumber="276">Notes</emphasis>
: The podomere proportions of the last three pairs of legs contrast with the proportions of the 12th legs, and are remarkably consistant. Taking the length of the prefemur as 1.0, the respective proportions of the femur, tibia, first tarsus and second tarsus of the 13th legs are 1.2/1.5/1.5/0.7, of the 14th legs 1.2/1.5/1.4/0.6, and of the 15th legs 1.2/1.3/1.2/0.7. The Hubbard male
<typeStatus id="54ED8803FFC7FF834305F8D519E5F840" box="[632,728,1919,1943]" pageId="17" pageNumber="276" type="paratype">paratype</typeStatus>
may not be fully mature, perhaps being a maturus junior (epimorphic stage VI), but has 60 antennal articles. The difference in the male tergites may be an expression of sexual dimorphism, but could possibly indicate that the Tee Cave specimens are not the same species as the one in Spider Cave, since the male
<typeStatus id="54ED8803FFC4FF8040A8FF32190BFF67" box="[469,566,152,176]" pageId="18" pageNumber="277" type="paratype">paratype</typeStatus>
collected by C. Richart and the female
<typeStatus id="54ED8803FFC4FF80428DFF321F6CFF67" box="[1008,1105,152,176]" pageId="18" pageNumber="277" type="paratype">paratype</typeStatus>
have the same distribution of tergite embayments as the male collected by D. Hubbard. The Richart male’s antennae are not complete; coxal pores are 3, 4, 4, 4, so in comparison with the other male
<typeStatus id="54ED8803FFC4FF80420DFF4A18EEFF2F" box="[880,979,224,248]" pageId="18" pageNumber="277" type="paratype">paratype</typeStatus>
, the specimen might be a prematurus (epimorphic state V). The Tee Cave female specimen is likely also immature, based on the lower number (52/54) of antennal articles and coxal pores counting 3, 4, 4, 4. To fully resolve the identity of both of these populations, additional material is required.
</paragraph>
</subSubSection>
<caption id="DF296629FFC4FF8041EAF9191948F93F" httpUri="https://zenodo.org/record/1455631/files/figure.png" pageId="18" pageNumber="277" startId="18.[151,264,1715,1738]" targetBox="[330,1230,433,1688]" targetPageId="18">
<paragraph id="8BE936A1FFC4FF8041EAF9191948F93F" blockId="18.[151,1436,1715,1768]" pageId="18" pageNumber="277">
<emphasis id="B922EAB3FFC4FF8041EAF9191A6EF91E" bold="true" box="[151,339,1715,1738]" pageId="18" pageNumber="277">FIGURES 18–25.</emphasis>
<emphasis id="B922EAB3FFC4FF804025F91E197BF91D" box="[344,582,1716,1738]" italics="true" pageId="18" pageNumber="277">
<taxonomicName id="4C564D22FFC4FF804025F91E197FF91D" authority="Shear, 2018" authorityName="Shear" authorityYear="2018" box="[344,578,1716,1738]" class="Chilopoda" family="Anopsobiidae" genus="Speleopsobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiida" pageId="18" pageNumber="277" phylum="Arthropoda" rank="species" species="weaveri" status="sp. nov.">Speleopsobius weaveri</taxonomicName>
,
</emphasis>
<emphasis id="B922EAB3FFC4FF804336F91E1994F91E" bold="true" box="[587,681,1716,1737]" pageId="18" pageNumber="277">
<taxonomicNameLabel id="A21157C8FFC4FF804336F91E19BEF91E" box="[587,643,1716,1737]" pageId="18" pageNumber="277" rank="species">n. sp.</taxonomicNameLabel>
18.
</emphasis>
Leg 12.
<emphasis id="B922EAB3FFC4FF80427CF91E181CF91E" bold="true" box="[769,801,1716,1737]" pageId="18" pageNumber="277">19.</emphasis>
Claw of leg 12.
<emphasis id="B922EAB3FFC4FF8042BAF91E18DAF91E" bold="true" box="[967,999,1716,1737]" pageId="18" pageNumber="277">20.</emphasis>
Leg 13.
<emphasis id="B922EAB3FFC4FF804542F91E1F62F91E" bold="true" box="[1087,1119,1716,1737]" pageId="18" pageNumber="277">21.</emphasis>
Claw of leg 13.
<emphasis id="B922EAB3FFC4FF804479F91E1E19F91E" bold="true" box="[1284,1316,1716,1737]" pageId="18" pageNumber="277">22.</emphasis>
Leg 14.
<emphasis id="B922EAB3FFC4FF804401F91E1EA1F91E" bold="true" box="[1404,1436,1716,1737]" pageId="18" pageNumber="277">23.</emphasis>
Claw of leg 13.
<emphasis id="B922EAB3FFC4FF804044F9781A64F930" bold="true" box="[313,345,1746,1767]" pageId="18" pageNumber="277">24.</emphasis>
Leg 15.
<emphasis id="B922EAB3FFC4FF8040CEF9781AEEF930" bold="true" box="[435,467,1746,1767]" pageId="18" pageNumber="277">25.</emphasis>
Claw of leg 15.
</paragraph>
</caption>
<subSubSection id="C34C652AFFC4FF8041BAF8A51956F80B" pageId="18" pageNumber="277" type="biology_ecology">
<paragraph id="8BE936A1FFC4FF8041BAF8A51956F80B" blockId="18.[151,1436,1807,2012]" pageId="18" pageNumber="277">
<emphasis id="B922EAB3FFC4FF8041BAF8A51A1BF8FF" bold="true" box="[199,294,1807,1832]" pageId="18" pageNumber="277">Ecology</emphasis>
: Troglobiosis is indicated in this species by the extremely long antennae with many additional anntennomeres, the long, thin legs and the large size (troglobiotic gigantism). Via Google Earth, I examined the area immediately around Spider Cave, which is situated in a lava outcrop just off an unpaved road. Multiple tire tracks suggest the cave is frequently visited. The surroundings appear to be typical desert or steppe grasslands, with considerable bare soil and scattered outcrops of black lava. I am certain that no henicopid or anopsobiid centipede could be supported in this surface habitat.
</paragraph>
</subSubSection>
</treatment>
</document>
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