treatments-xml/data/03/FF/87/03FF878E8C0E103964EFFBD987E9F9A6.xml

<document id="C3B7E497EDCCDBC78A4CA619285B0B5F" ID-DOI="10.26107/RBZ-2021-0013" ID-ISSN="2345-7600" ID-Zenodo-Dep="5352036" ID-ZooBank="C6DD0960-51EE-40C6-B948-BFAEF4E9F304" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630207527469" checkinUser="diego" docAuthor="Komai, Tomoyuki &amp; Rahayu, Dwi Listyo" docDate="2021" docId="03FF878E8C0E103964EFFBD987E9F9A6" docLanguage="en" docName="RBZ-2021-0013.pdf" docOrigin="Raffles Bulletin of Zoology 69" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Catapagurus fimbriatus Komai &amp; Rahayu 2021, new species" docType="treatment" docVersion="3" lastPageNumber="167" masterDocId="FFC6FFF68C0810326771FF848470FFF3" masterDocTitle="Three new species of the pagurid hermit crab genus Catapagurus A. Milne-Edwards, 1880 (Decapoda: Anomura: Paguroidea) from the Bohol Sea, the Philippines" masterLastPageNumber="174" masterPageNumber="156" pageNumber="162" updateTime="1699185298364" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="34B9F2EA033C1F81CDCD19F9BBD3EDE6">Three new species of the pagurid hermit crab genus Catapagurus A. Milne-Edwards, 1880 (Decapoda: Anomura: Paguroidea) from the Bohol Sea, the Philippines</mods:title>
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<mods:namePart id="945553F5871E2658AE3CF6A88134F746">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="B9F21C804F9A6C55BA68D7A935936697">Rahayu, Dwi Listyo</mods:namePart>
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<mods:date id="737A0F8A361BC2848FE72EDA817F0B46">2021</mods:date>
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<mods:number id="DB641B2E06E423C944F62AC13ED1920A">2021-05-17</mods:number>
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<treatment id="03FF878E8C0E103964EFFBD987E9F9A6" LSID="urn:lsid:plazi:treatment:03FF878E8C0E103964EFFBD987E9F9A6" httpUri="http://treatment.plazi.org/id/03FF878E8C0E103964EFFBD987E9F9A6" lastPageId="11" lastPageNumber="167" pageId="6" pageNumber="162">
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<paragraph id="8BE936988C0E103464EFFBD9814AFB86" blockId="6.[926,1338,1117,1173]" box="[926,1338,1117,1141]" pageId="6" pageNumber="162">
<heading id="D0A181F48C0E103464EFFBD9814AFB86" bold="true" box="[926,1338,1117,1141]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="162" reason="2">
<emphasis id="B922EA8A8C0E103464EFFBD9814AFB86" bold="true" box="[926,1338,1117,1141]" pageId="6" pageNumber="162">
<taxonomicName id="4C564D1B8C0E103464EFFBD980D6FB86" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[926,1190,1117,1141]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B922EA8A8C0E103464EFFBD980D6FB86" bold="true" box="[926,1190,1117,1141]" italics="true" pageId="6" pageNumber="162">Catapagurus fimbriatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21157F18C0E103463C4FBD9814AFB86" box="[1205,1338,1117,1141]" pageId="6" pageNumber="162" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C34C65138C0E10346341FBF980D7FB66" box="[1072,1191,1149,1173]" pageId="6" pageNumber="162" type="description">
<paragraph id="8BE936988C0E10346341FBF980D7FB66" blockId="6.[926,1338,1117,1173]" box="[1072,1191,1149,1173]" pageId="6" pageNumber="162">
(
<figureCitation id="136D2A1D8C0E10346349FBF980EFFB66" box="[1080,1183,1149,1173]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="7.[136,171,1535,1557]" captionStartId-1="8.[113,148,1643,1665]" captionStartId-2="9.[136,171,1427,1449]" captionStartId-3="10.[113,148,2035,2057]" captionTargetBox-0="[137,1476,175,1519]" captionTargetBox-1="[116,1450,188,1627]" captionTargetBox-2="[133,1474,189,1413]" captionTargetBox-3="[156,1408,188,2019]" captionTargetId-0="figure-8@7.[124,1492,171,1530]" captionTargetId-1="figure-11@8.[101,1465,175,1638]" captionTargetId-2="figure-8@9.[125,1490,166,1429]" captionTargetId-3="figure-11@10.[127,1459,173,2023]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="Fig. 5. Catapagurus fimbriatus, new species. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view." captionText-1="Fig. 6. Catapagurus fimbriatus, new species, holotype, male (sl 1.5 mm), NMCR 50802. A, chela of right cheliped, dorsal view; B, right cheliped, mesial view; C, same, lateral view; D, carpus of right cheliped, dorsal view." captionText-2="Fig. 7. Catapagurus fimbriatus, new species, holotype, male (sl 1.5 mm), NMCR 50802. A, chela of left cheliped, dorsal view; B, left cheliped, mesial view; C, same, lateral view; D, carpus of left cheliped, dorsal view." captionText-3="Fig. 8. Catapagurus fimbriatus, new species. A–F, holotype, male (sl 1.5 mm), NMCR 50802; G, paratype, male (sl 1.4 mm), ZRC 2021.0414. A, right pereopod 2, lateral view; B, same, propodus and carpus, mesial view; C, same, dactylus, mesial view; D, left pereopod 3, lateral view; E, same, propodus and carpus, mesial view; F, same, dactylus, mesial view. G, left pereopod 3, merus, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.5352046" figureDoi-1="http://doi.org/10.5281/zenodo.5352048" figureDoi-2="http://doi.org/10.5281/zenodo.5352050" figureDoi-3="http://doi.org/10.5281/zenodo.5352052" httpUri-0="https://zenodo.org/record/5352046/files/figure.png" httpUri-1="https://zenodo.org/record/5352048/files/figure.png" httpUri-2="https://zenodo.org/record/5352050/files/figure.png" httpUri-3="https://zenodo.org/record/5352052/files/figure.png" pageId="6" pageNumber="162">Figs. 5–8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34C65138C0E1034645CFB39801AFA46" pageId="6" pageNumber="162" type="materials_examined">
<paragraph id="8BE936988C0E1034645CFB3987B6FAA6" blockId="6.[813,1452,1213,1365]" pageId="6" pageNumber="162">
<emphasis id="B922EA8A8C0E1034645CFB39807EFB26" bold="true" box="[813,1038,1213,1237]" pageId="6" pageNumber="162">Material examined.</emphasis>
<materialsCitation id="3B3E3CC58C0E10346369FB3987B6FAA6" collectingDate="2004-06-25" collectionCode="T" elevation="122" latitude="9.556666" location="Sea" longLatPrecision="129" longitude="123.85" pageId="6" pageNumber="162" specimenCode="T27" specimenCount="1" stateProvince="Bohol" typeStatus="holotype">
<typeStatus id="54ED883A8C0E10346369FB39800FFB26" box="[1048,1151,1213,1237]" pageId="6" pageNumber="162" type="holotype">Holotype</typeStatus>
: male (sl 
<quantity id="4CAE9B7D8C0E1034639EFB398137FB26" box="[1263,1351,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="162" unit="mm" value="1.5">1.5 mm</quantity>
), NMCR 50802, PANGLAO 2004, stn 
<specimenCode id="DBF09EE38C0E103463F6FB5980C6FB06" box="[1159,1206,1245,1269]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="162">T27</specimenCode>
, 
<collectingRegion id="4992F87A8C0E103463B7FB59817AFB06" box="[1222,1290,1245,1269]" country="Philippines" name="Bohol" pageId="6" pageNumber="162">Bohol</collectingRegion>
<location id="8E8960438C0E10346265FB59814FFB06" LSID="urn:lsid:plazi:treatment:03FF878E8C0E103964EFFBD987E9F9A6:8E8960438C0E10346265FB59814FFB06" box="[1300,1343,1245,1269]" latitude="9.556666" longLatPrecision="129" longitude="123.85" name="Sea" pageId="6" pageNumber="162" stateProvince="Bohol">Sea</location>
, 
<locationDeviation id="9A5B2FBA8C0E1034623EFB5980E4FAE6" firstLocation="Panglao" pageId="6" pageNumber="162" secondLocation="Pamilacan islands">between Panglao and Pamilacan islands</locationDeviation>
, 
<geoCoordinate id="EE62505F8C0E103463D2FB788167FAE6" box="[1187,1303,1276,1301]" degrees="09" direction="north" minutes="33.4" orientation="latitude" pageId="6" pageNumber="162" precision="92" value="9.556666">09°33.4′N</geoCoordinate>
, 
<geoCoordinate id="EE62505F8C0E10346256FB7881D6FAE6" box="[1319,1446,1276,1301]" degrees="123" direction="east" minutes="51.0" orientation="longitude" pageId="6" pageNumber="162" precision="92" value="123.85">123°51.0′E</geoCoordinate>
, 
<quantity id="4CAE9B7D8C0E1034645CFA9987DDFAC6" box="[813,941,1309,1333]" metricMagnitude="2" metricUnit="m" metricValue="1.215" metricValueMax="1.3699999999999999" metricValueMin="1.06" pageId="6" pageNumber="162" unit="m" value="121.5" valueMax="137.0" valueMin="106.0">
<elevation id="007BD1AB8C0E1034645CFA9987DDFAC6" box="[813,941,1309,1333]" metricMagnitude="2" metricUnit="m" metricValue="1.215" metricValueMax="1.3699999999999999" metricValueMin="1.06" pageId="6" pageNumber="162" unit="m" value="121.5" valueMax="137.0" valueMin="106.0">106–137 m</elevation>
</quantity>
, fine sand and mud with echinoderms, trawl, 
<date id="FFE810588C0E1034645CFAB987B3FAA6" box="[813,963,1341,1365]" pageId="6" pageNumber="162" value="2004-06-25">
<collectingDate id="EFACE9B08C0E1034645CFAB987B3FAA6" box="[813,963,1341,1365]" pageId="6" pageNumber="162" value="2004-06-25">25 June 2004</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE936988C0E1034645CFAFA801AFA46" blockId="6.[813,1451,1405,1461]" pageId="6" pageNumber="162">
<materialsCitation id="3B3E3CC58C0E1034645CFAFA8016FA46" collectingDate="2004-06-25" collectionCode="ZRC" elevation="122" latitude="9.556666" location="Sea" longLatPrecision="129" longitude="123.85" pageId="6" pageNumber="162" specimenCode="ZRC 2021.0414" specimenCount="1" specimenCount-male="1" stateProvince="Bohol" typeStatus="paratype">
<typeStatus id="54ED883A8C0E1034645CFAFA87FFFA65" box="[813,911,1406,1430]" pageId="6" pageNumber="162" type="paratype">Paratype</typeStatus>
: 
<specimenCount id="9D50FD118C0E103464E9FAF98791FA66" box="[920,993,1405,1429]" pageId="6" pageNumber="162" type="male">1 male</specimenCount>
(sl 
<quantity id="4CAE9B7D8C0E10346378FAF9802FFA66" box="[1033,1119,1405,1429]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="162" unit="mm" value="1.4">1.4 mm</quantity>
), 
<specimenCode id="DBF09EE38C0E10346303FAF98151FA66" box="[1138,1313,1405,1429]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="162">ZRC 2021.0414</specimenCode>
, PANGLAO 2004, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C65138C0E10396700FA5980D8FEC6" lastPageId="11" lastPageNumber="167" pageId="6" pageNumber="162" type="discussion">
<paragraph id="8BE936988C0E10346700FA598012F886" blockId="6.[112,1454,1501,2133]" pageId="6" pageNumber="162">
<emphasis id="B922EA8A8C0E10346700FA598491FA06" bold="true" box="[113,225,1501,1525]" pageId="6" pageNumber="162">Remarks.</emphasis>
There were no female specimens available for 
<emphasis id="B922EA8A8C0E1034645CFA5987C8FA06" bold="true" box="[813,952,1501,1525]" pageId="6" pageNumber="162">Description.</emphasis>
Shield (
<figureCitation id="136D2A1D8C0E10346365FA598019FA06" box="[1044,1129,1501,1525]" captionStart="Fig" captionStartId="7.[136,171,1535,1557]" captionTargetBox="[137,1476,175,1519]" captionTargetId="figure-8@7.[124,1492,171,1530]" captionTargetPageId="7" captionText="Fig. 5. Catapagurus fimbriatus, new species. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352046" httpUri="https://zenodo.org/record/5352046/files/figure.png" pageId="6" pageNumber="162">Fig. 5A</figureCitation>
) 1.2 times broader than long; study. The present new species is assigned to 
<taxonomicName id="4C564D1B8C0E10346517FA7A8681F9E6" authorityName="A. Milne-Edwards" authorityYear="1880" box="[614,753,1534,1557]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="B922EA8A8C0E10346517FA7A8681F9E6" box="[614,753,1534,1557]" italics="true" pageId="6" pageNumber="162">Catapagurus</emphasis>
</taxonomicName>
anterior margin between rostral lobe and lateral projections without hesitation because it agrees with the generic features concave; anterolateral margins sloping; posterior margin of 
<taxonomicName id="4C564D1B8C0E103467FEF9BA856AF9A6" authorityName="A. Milne-Edwards" authorityYear="1880" box="[143,282,1598,1621]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="B922EA8A8C0E103467FEF9BA856AF9A6" box="[143,282,1598,1621]" italics="true" pageId="6" pageNumber="162">Catapagurus</emphasis>
</taxonomicName>
, including the gill formula consisting of 11 truncate; dorsal surface with longitudinal rows of tufts of pairs of biserial gills, the somewhat reduced crista dentata short setae on midline and posterior to lateral projections; on the maxilliped 3 ischium, the presence of a prominent carapace lateral lobes narrow, well calcified. Rostral lobe preungual process on the pereopod 4 dactyli, and the broadly and roundly triangular, usually produced as far as possession of a long sexual tube arising from the right lateral projections. Lateral projections broadly and roundly pereopod 5 coxa in males, curving exteriorly and over the triangular, each usually with terminal, submarginal spinule. lateral side of the body (
<bibRefCitation id="EFC74B698C0E1034660FF979863AF8E6" author="McLaughlin PA" box="[382,586,1789,1813]" pageId="6" pageNumber="162" pagination="111 - 144" refId="ref9987" refString="McLaughlin PA (2003) Illustrated keys to families and genera of the superfamily Paguroidea (Crustacea: Decapoda: Anomura), with diagnoses of genera of Paguridae. Memoirs of Museum Victoria, 60: 111 - 144." type="journal article" year="2003">McLaughlin, 2003</bibRefCitation>
, 
<bibRefCitation id="EFC74B698C0E10346526F97986E1F8E6" author="McLaughlin PA" box="[599,657,1789,1813]" pageId="6" pageNumber="162" pagination="1 - 16" refId="ref10028" refString="McLaughlin PA (2004) A reappraisal of the hermit crab genera Catapagurus A. Milne-Edwards and Hemipagurus Smith (Crustacea: Decapoda: Anomura: Paguridae), with the description of a new species. Zootaxa, 433: 1 - 16." type="journal article" year="2004">2004</bibRefCitation>
). Posterior carapace 0.5 length of shield; posterolateral plates moderately wide anteriorly, drawn out into relatively thin 
<taxonomicName id="4C564D1B8C0E10346700F8BA850DF8A6" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[113,381,1853,1877]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="exilidigitus" status="sp. nov.">
<emphasis id="B922EA8A8C0E10346700F8BA850DF8A6" box="[113,381,1853,1877]" italics="true" pageId="6" pageNumber="162">Catapagurus exilidigitus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21157F18C0E103466FDF8BA8662F8A6" box="[396,530,1853,1877]" pageId="6" pageNumber="162" rank="species">new species</taxonomicNameLabel>
, appears close to 
<emphasis id="B922EA8A8C0E103465A9F8BA8680F8A6" box="[728,752,1854,1877]" italics="true" pageId="6" pageNumber="162">C.</emphasis>
bands reaching to posterior margins, posterior median plate 
<taxonomicName id="4C564D1B8C0E10346700F8D9859BF886" authority="McLaughlin, 1997" authorityName="McLaughlin" authorityYear="1997" box="[113,491,1885,1909]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="tanimbarensis">
<emphasis id="B922EA8A8C0E10346700F8D9857DF886" box="[113,269,1885,1909]" italics="true" pageId="6" pageNumber="162">tanimbarensis</emphasis>
<bibRefCitation id="EFC74B698C0E10346669F8D9859BF886" author="McLaughlin PA" box="[280,491,1885,1909]" pageId="6" pageNumber="162" pagination="433 - 572" refId="ref9882" refString="McLaughlin PA (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier A &amp; Bouchet P (eds.) Resultats des Campagnes MUSORSTOM, Volume 16. Memoires du Museum national d'Histoire naturelle, Paris, 172: 433 - 572." type="journal article" year="1997">McLaughlin, 1997</bibRefCitation>
</taxonomicName>
, presently known only short, only weakly calcified.
</paragraph>
<paragraph id="8BE936988C0E10346700F8F987E9F7C6" blockId="6.[112,1454,1501,2133]" pageId="6" pageNumber="162">
from the Banda Sea, 
<collectingCountry id="F34176088C0E10346625F8F985CEF866" box="[340,446,1917,1941]" name="Indonesia" pageId="6" pageNumber="162">Indonesia</collectingCountry>
, in the general armature and slenderness of the pereopod 2 and 3 dactyli. In particular, in Ocular peduncles (
<figureCitation id="136D2A1D8C0E10346377F8198012F846" box="[1030,1122,1949,1973]" captionStart="Fig" captionStartId="7.[136,171,1535,1557]" captionTargetBox="[137,1476,175,1519]" captionTargetId="figure-8@7.[124,1492,171,1530]" captionTargetPageId="7" captionText="Fig. 5. Catapagurus fimbriatus, new species. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352046" httpUri="https://zenodo.org/record/5352046/files/figure.png" pageId="6" pageNumber="162">Fig. 5A</figureCitation>
) short (0.7 times as long as these two taxa, the dorsal spiniform or bristle-like setae on the shield), stout, widened distally; corneal width 0.7 peduncular mesial surfaces of the ambulatory dactyli are relatively few length. Ocular acicles narrowly triangular, slender, reaching compared to other species in the 
<taxonomicName id="4C564D1B8C0E103466BDF87A8615F7E6" baseAuthorityName="Terao" baseAuthorityYear="1914" box="[460,613,2045,2069]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="162" phylum="Arthropoda" rank="species" species="misakiensis">
<emphasis id="B922EA8A8C0E103466BDF87A8615F7E6" box="[460,613,2045,2069]" italics="true" pageId="6" pageNumber="162">C. misakiensis</emphasis>
</taxonomicName>
group (20 or 0.3 length of ocular peduncle, terminating acutely; widely less versus more than 30) (de Saint Laurent &amp; McLaughlin, separated.
</paragraph>
<paragraph id="8BE936988C0E10346703F7B9869FF7A6" blockId="6.[112,1454,1501,2133]" box="[114,751,2109,2133]" pageId="6" pageNumber="162">
2000; 
<bibRefCitation id="EFC74B698C0E103467B3F7B98507F7A6" author="Asakura A" box="[194,375,2109,2133]" pageId="6" pageNumber="162" pagination="823 - 891" refId="ref9098" refString="Asakura A (2001) A revision of the hermit crab genera Catapagurus A. Milne-Edwards and Hemipagurus Smith from the Indo- West Pacific (Crustacea: Decapoda: Anomura: Paguridae). Invertebrate Taxonomy, 15: 823 - 891." type="journal article" year="2001">Asakura, 2001</bibRefCitation>
; 
<bibRefCitation id="EFC74B698C0E103466F8F7B9861EF7A6" author="McLaughlin PA" box="[393,622,2109,2133]" pageId="6" pageNumber="162" pagination="385 - 460" refId="ref9937" refString="McLaughlin PA (2002) A review of the hermit-crab (Decapoda: Anomura: Paguridea) fauna of southern Thailand, with particular emphasis on the Andaman Sea, and descriptions of three new species. Phuket Marine Biological Center Special Publication, 23: 385 - 460." type="journal article" year="2002">McLaughlin, 2002</bibRefCitation>
; Komai &amp;
</paragraph>
<caption id="DF2966108C0F103567F9FA7B858FF97B" ID-DOI="http://doi.org/10.5281/zenodo.5352046" ID-Zenodo-Dep="5352046" httpUri="https://zenodo.org/record/5352046/files/figure.png" pageId="7" pageNumber="163" startId="7.[136,171,1535,1557]" targetBox="[137,1476,175,1519]" targetPageId="7">
<paragraph id="8BE936988C0F103567F9FA7B858FF97B" blockId="7.[136,1475,1535,1672]" pageId="7" pageNumber="163">
Fig. 5. 
<taxonomicName id="4C564D1B8C0F103567BDFA7B85C0F9E7" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[204,432,1535,1556]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="163" phylum="Arthropoda" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B922EA8A8C0F103567BDFA7B85C0F9E7" box="[204,432,1535,1556]" italics="true" pageId="7" pageNumber="163">Catapagurus fimbriatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21157F18C0F103566CCFA7B8643F9E6" box="[445,563,1535,1557]" pageId="7" pageNumber="163" rank="species">new species</taxonomicNameLabel>
. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view.
</paragraph>
</caption>
<paragraph id="8BE936988C0F103567F9F93B8507F844" blockId="7.[135,775,1727,1975]" pageId="7" pageNumber="163">
Antennular peduncle (
<figureCitation id="136D2A1D8C0F10356601F93B85AEF924" box="[368,478,1727,1751]" captionStart="Fig" captionStartId="7.[136,171,1535,1557]" captionTargetBox="[137,1476,175,1519]" captionTargetId="figure-8@7.[124,1492,171,1530]" captionTargetPageId="7" captionText="Fig. 5. Catapagurus fimbriatus, new species. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352046" httpUri="https://zenodo.org/record/5352046/files/figure.png" pageId="7" pageNumber="163">Fig. 5A, B</figureCitation>
) overreaching distal corneal margin by full length of ultimate article. Ultimate article 0.8 times as long as shield, 5.3 times as long as distal height, with tuft of long setae at dorsodistal margin and some short setae on dorsal surface. Penultimate article with few short setae. Basal article with very slightly produced, subacute ventrodistal margin; statocyst lobe gently inflated in proximal half of lateral margin.
</paragraph>
<paragraph id="8BE936988C0F103567F9F85B8094F864" blockId="7.[136,775,2015,2071]" lastBlockId="7.[834,1474,1727,1943]" pageId="7" pageNumber="163">
Antennal peduncle (
<figureCitation id="136D2A1D8C0F1035661DF85B85B4F804" box="[364,452,2015,2039]" captionStart="Fig" captionStartId="7.[136,171,1535,1557]" captionTargetBox="[137,1476,175,1519]" captionTargetId="figure-8@7.[124,1492,171,1530]" captionTargetPageId="7" captionText="Fig. 5. Catapagurus fimbriatus, new species. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352046" httpUri="https://zenodo.org/record/5352046/files/figure.png" pageId="7" pageNumber="163">Fig. 5A</figureCitation>
) overreaching distal corneal margin by 0.7 length of ultimate article. Articles 5 and 4 with few, scattered short setae. Article 3 with few setae on unarmed ventrodistal angle. Article 2 with produced dorsolateral distal angle reaching to or beyond base of article 4, terminating in spine; dorsomesial distal angle with prominent spine. Article 1 with small spine at dorsolateral distal angle. Antennal acicle reaching distal margin of article 4 and reaching corneal base, terminating in spine. Flagella missing.
</paragraph>
<paragraph id="8BE936988C0F10356432F83B81FFF7E4" blockId="7.[834,1474,1983,2072]" pageId="7" pageNumber="163">
Maxilliped 3 endopod (
<figureCitation id="136D2A1D8C0F10356347F83B80F5F824" box="[1078,1157,1983,2007]" captionStart="Fig" captionStartId="7.[136,171,1535,1557]" captionTargetBox="[137,1476,175,1519]" captionTargetId="figure-8@7.[124,1492,171,1530]" captionTargetPageId="7" captionText="Fig. 5. Catapagurus fimbriatus, new species. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352046" httpUri="https://zenodo.org/record/5352046/files/figure.png" pageId="7" pageNumber="163">Fig. 5C</figureCitation>
) moderately stout; merus with small dorsodistal spine; ischium with crista dentata consisting of 5 small teeth; accessory tooth prominent (
<figureCitation id="136D2A1D8C0F1035625AF87B81F2F7E4" box="[1323,1410,2047,2071]" captionStart="Fig" captionStartId="7.[136,171,1535,1557]" captionTargetBox="[137,1476,175,1519]" captionTargetId="figure-8@7.[124,1492,171,1530]" captionTargetPageId="7" captionText="Fig. 5. Catapagurus fimbriatus, new species. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352046" httpUri="https://zenodo.org/record/5352046/files/figure.png" pageId="7" pageNumber="163">Fig. 5D</figureCitation>
).
</paragraph>
<caption id="DF2966108C00103A6700F9EF87B1F96D" ID-DOI="http://doi.org/10.5281/zenodo.5352048" ID-Zenodo-Dep="5352048" httpUri="https://zenodo.org/record/5352048/files/figure.png" pageId="8" pageNumber="164" startId="8.[113,148,1643,1665]" targetBox="[116,1450,188,1627]" targetPageId="8">
<paragraph id="8BE936988C00103A6700F9EF87B1F96D" blockId="8.[113,1451,1643,1694]" pageId="8" pageNumber="164">
Fig. 6. 
<taxonomicName id="4C564D1B8C00103A67C8F9E885EEF973" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[185,414,1643,1665]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="164" phylum="Arthropoda" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B922EA8A8C00103A67C8F9E885EEF973" box="[185,414,1643,1665]" italics="true" pageId="8" pageNumber="164">Catapagurus fimbriatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21157F18C00103A66DAF9E88653F972" box="[427,547,1643,1665]" pageId="8" pageNumber="164" rank="species">new species</taxonomicNameLabel>
, holotype, male (sl 1.5 mm), NMCR 50802. A, chela of right cheliped, dorsal view; B, right cheliped, mesial view; C, same, lateral view; D, carpus of right cheliped, dorsal view.
</paragraph>
</caption>
<paragraph id="8BE936988C00103B6700F9518543F966" blockId="8.[112,754,1749,2093]" lastBlockId="9.[136,775,1533,1685]" lastPageId="9" lastPageNumber="165" pageId="8" pageNumber="164">
Right cheliped (
<figureCitation id="136D2A1D8C00103A666DF95185FFF91E" box="[284,399,1749,1773]" captionStart="Fig" captionStartId="8.[113,148,1643,1665]" captionTargetBox="[116,1450,188,1627]" captionTargetId="figure-11@8.[101,1465,175,1638]" captionTargetPageId="8" captionText="Fig. 6. Catapagurus fimbriatus, new species, holotype, male (sl 1.5 mm), NMCR 50802. A, chela of right cheliped, dorsal view; B, right cheliped, mesial view; C, same, lateral view; D, carpus of right cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352048" httpUri="https://zenodo.org/record/5352048/files/figure.png" pageId="8" pageNumber="164">Fig. 6A–D</figureCitation>
) long, moderately slender. Chela elongate subovate (
<figureCitation id="136D2A1D8C00103A663FF97185DAF8FE" box="[334,426,1781,1805]" captionStart="Fig" captionStartId="8.[113,148,1643,1665]" captionTargetBox="[116,1450,188,1627]" captionTargetId="figure-11@8.[101,1465,175,1638]" captionTargetPageId="8" captionText="Fig. 6. Catapagurus fimbriatus, new species, holotype, male (sl 1.5 mm), NMCR 50802. A, chela of right cheliped, dorsal view; B, right cheliped, mesial view; C, same, lateral view; D, carpus of right cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352048" httpUri="https://zenodo.org/record/5352048/files/figure.png" pageId="8" pageNumber="164">Fig. 6A</figureCitation>
), 2.7 times as long as wide. Dactylus 0.9 length of palm; weakly convex dorsal surface almost smooth and with sparse tufts of setae; dorsomesial margin not delimited, rounded; occlusal margin with 2 blunt calcareous teeth, slightly overlapped by fixed finger. Palm subequal in length to carpus, 1.6 times as long as wide; dorsomesial and dorsolateral margins not delimited, dorsal and mesial surfaces minutely granular; ventral surface nearly smooth; fixed finger with sparse tufts of setae in distal half, occlusal margin with row of low, blunt calcareous teeth over entire length. Carpus (
<figureCitation id="136D2A1D8C00103A636CF9518002F91E" box="[1053,1138,1749,1773]" captionStart="Fig" captionStartId="8.[113,148,1643,1665]" captionTargetBox="[116,1450,188,1627]" captionTargetId="figure-11@8.[101,1465,175,1638]" captionTargetPageId="8" captionText="Fig. 6. Catapagurus fimbriatus, new species, holotype, male (sl 1.5 mm), NMCR 50802. A, chela of right cheliped, dorsal view; B, right cheliped, mesial view; C, same, lateral view; D, carpus of right cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352048" httpUri="https://zenodo.org/record/5352048/files/figure.png" pageId="8" pageNumber="164">Fig. 6D</figureCitation>
) slightly shorter than merus, 2.7 times as long as distal width; dorsal surface with sparse granules, dorsomesial margin delimited with row of tiny spines in distal 0.3 and of minute granules proximally, and with few setae, dorsodistal margin unarmed; dorsolateral margin delimited with irregular row of spinules, and with few setae; lateral and mesial surfaces with sparse minute granules, ventral surface nearly smooth; all with covering of tiny or minute spinules, tubercles or granules. Merus subtriangular in cross section; dorsodistal margin with small spine; dorsal surface with widely spaced short setae; mesial and lateral faces almost smooth; ventrolateral and ventromesial margins each with small distal spine; ventral surface smooth, only with few short setae. Ischium unarmed on distolateral margin. Coxa with small ventromesial distal spine (
<figureCitation id="136D2A1D8C01103B67A3F9F98555F966" box="[210,293,1661,1685]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="9" pageNumber="165">Fig. 1E</figureCitation>
).
</paragraph>
<caption id="DF2966108C01103B67F9FA1787B9FA35" ID-DOI="http://doi.org/10.5281/zenodo.5352050" ID-Zenodo-Dep="5352050" httpUri="https://zenodo.org/record/5352050/files/figure.png" pageId="9" pageNumber="165" startId="9.[136,171,1427,1449]" targetBox="[133,1474,189,1413]" targetPageId="9">
<paragraph id="8BE936988C01103B67F9FA1787B9FA35" blockId="9.[136,1474,1427,1478]" pageId="9" pageNumber="165">
Fig. 7. 
<taxonomicName id="4C564D1B8C01103B67A0FA1785C8FA5B" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[209,440,1427,1448]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="165" phylum="Arthropoda" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B922EA8A8C01103B67A0FA1785C8FA5B" box="[209,440,1427,1448]" italics="true" pageId="9" pageNumber="165">Catapagurus fimbriatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21157F18C01103B66B6FA10864FFA5A" box="[455,575,1427,1449]" pageId="9" pageNumber="165" rank="species">new species</taxonomicNameLabel>
, holotype, male (sl 1.5 mm), NMCR 50802. A, chela of left cheliped, dorsal view; B, left cheliped, mesial view; C, same, lateral view; D, carpus of left cheliped, dorsal view.
</paragraph>
</caption>
<paragraph id="8BE936988C01103B67F9F9398174F886" blockId="9.[135,777,1725,2101]" lastBlockId="9.[834,1475,1533,1909]" pageId="9" pageNumber="165">
Left cheliped (
<figureCitation id="136D2A1D8C01103B6651F93985E2F926" box="[288,402,1725,1749]" captionStart="Fig" captionStartId="9.[136,171,1427,1449]" captionTargetBox="[133,1474,189,1413]" captionTargetId="figure-8@9.[125,1490,166,1429]" captionTargetPageId="9" captionText="Fig. 7. Catapagurus fimbriatus, new species, holotype, male (sl 1.5 mm), NMCR 50802. A, chela of left cheliped, dorsal view; B, left cheliped, mesial view; C, same, lateral view; D, carpus of left cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352050" httpUri="https://zenodo.org/record/5352050/files/figure.png" pageId="9" pageNumber="165">Fig. 7A–D</figureCitation>
) slender, subequal in length to right cheliped; dactyl and fixed finger nearly straight. Chela (
<figureCitation id="136D2A1D8C01103B65ABF95984DAF8E6" captionStart="Fig" captionStartId="9.[136,171,1427,1449]" captionTargetBox="[133,1474,189,1413]" captionTargetId="figure-8@9.[125,1490,166,1429]" captionTargetPageId="9" captionText="Fig. 7. Catapagurus fimbriatus, new species, holotype, male (sl 1.5 mm), NMCR 50802. A, chela of left cheliped, dorsal view; B, left cheliped, mesial view; C, same, lateral view; D, carpus of left cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352050" httpUri="https://zenodo.org/record/5352050/files/figure.png" pageId="9" pageNumber="165">Fig. 7A</figureCitation>
) 4.8 times as long as wide. Dactylus 1.8 times as long as palm; surfaces almost smooth, with sparse tufts of setae; dorsomesial margin not delimited; occlusal margin with row of tiny corneous teeth in distal half, otherwise unarmed. Palm 0.5 length of carpus, 1.8 times as long as wide; dorsomesial and dorsolateral margins not delimited, dorsal, lateral and mesial surfaces with few setae and sparse minute granules; ventral surface slightly convex, smooth; fixed finger with smooth surfaces, few scattered long setae; occlusal margin with row of minute calcareous teeth in distal half, unarmed in proximal half. Carpus (
<figureCitation id="136D2A1D8C01103B6300FA7980BBF9E6" box="[1137,1227,1533,1557]" captionStart="Fig" captionStartId="9.[136,171,1427,1449]" captionTargetBox="[133,1474,189,1413]" captionTargetId="figure-8@9.[125,1490,166,1429]" captionTargetPageId="9" captionText="Fig. 7. Catapagurus fimbriatus, new species, holotype, male (sl 1.5 mm), NMCR 50802. A, chela of left cheliped, dorsal view; B, left cheliped, mesial view; C, same, lateral view; D, carpus of left cheliped, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352050" httpUri="https://zenodo.org/record/5352050/files/figure.png" pageId="9" pageNumber="165">Fig. 7D</figureCitation>
) slightly shorter than merus, 4.0 times as long as distal width; dorsomesial margin with row of spinules and few setae, dorsolateral margin with row of granules, becoming spinulose distally, dorsal surface with few granules distally, otherwise smooth; mesial and lateral surfaces with sparse granules, ventral surface smooth. Merus with small spine on dorsodistal margin, dorsal surface with row of widely spaced setae; lateral surface with sparse granules, mesial and ventral surfaces almost smooth, ventromesial and ventrolateral distal angles each with small spine. Ischium with small spine distolaterally. Coxa with small ventromesial distal spine (
<figureCitation id="136D2A1D8C01103B63D2F8D98087F886" box="[1187,1271,1885,1909]" captionStart="Fig" captionStartId="2.[113,148,1627,1649]" captionTargetBox="[116,1450,188,1609]" captionTargetId="figure-11@2.[92,1469,169,1620]" captionTargetPageId="2" captionText="Fig. 1. Catapagurus exilidigitus, new species, holotype, male (sl 1.7 mm), NMCR 50801. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, ischium and basis, ventral view; D, right pereopod 4, lateral view; E, coxae of pereopods 5 and thoracic sternite 8, ventral view; F, distal part of right sexual tube, dorsal view; G, thoracic sternite 6, ventral view; H, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352038" httpUri="https://zenodo.org/record/5352038/files/figure.png" pageId="9" pageNumber="165">Fig. 1E</figureCitation>
).
</paragraph>
<paragraph id="8BE936988C0110396432F8198778FDA6" blockId="9.[835,1475,1949,2101]" lastBlockId="11.[135,777,189,597]" lastPageId="11" lastPageNumber="167" pageId="9" pageNumber="165">
Pereopods 2 (
<figureCitation id="136D2A1D8C01103B64A6F819803BF846" box="[983,1099,1949,1973]" captionStart="Fig" captionStartId="10.[113,148,2035,2057]" captionTargetBox="[156,1408,188,2019]" captionTargetId="figure-11@10.[127,1459,173,2023]" captionTargetPageId="10" captionText="Fig. 8. Catapagurus fimbriatus, new species. A–F, holotype, male (sl 1.5 mm), NMCR 50802; G, paratype, male (sl 1.4 mm), ZRC 2021.0414. A, right pereopod 2, lateral view; B, same, propodus and carpus, mesial view; C, same, dactylus, mesial view; D, left pereopod 3, lateral view; E, same, propodus and carpus, mesial view; F, same, dactylus, mesial view. G, left pereopod 3, merus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5352052" httpUri="https://zenodo.org/record/5352052/files/figure.png" pageId="9" pageNumber="165">Fig. 8A–C</figureCitation>
) considerably overreaching tips of outstretched chelipeds. Dactyli narrowly blade-shaped, 1.1 times as long as propodi; in dorsal view, straight; in lateral view, gently curved ventrally; dorsal margins each with row of moderately spaced setae, becoming longer and more stiff distally; mesial faces (
<figureCitation id="136D2A1D8C031039660AFF3985BEFF26" box="[379,462,189,213]" captionStart="Fig" captionStartId="10.[113,148,2035,2057]" captionTargetBox="[156,1408,188,2019]" captionTargetId="figure-11@10.[127,1459,173,2023]" captionTargetPageId="10" captionText="Fig. 8. Catapagurus fimbriatus, new species. A–F, holotype, male (sl 1.5 mm), NMCR 50802; G, paratype, male (sl 1.4 mm), ZRC 2021.0414. A, right pereopod 2, lateral view; B, same, propodus and carpus, mesial view; C, same, dactylus, mesial view; D, left pereopod 3, lateral view; E, same, propodus and carpus, mesial view; F, same, dactylus, mesial view. G, left pereopod 3, merus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5352052" httpUri="https://zenodo.org/record/5352052/files/figure.png" pageId="11" pageNumber="167">Fig. 8C</figureCitation>
) slightly concave, each with row of about 20 moderately spaced, stiff setae adjacent to ventral margin; lateral faces and ventral margins unarmed. Propodi approximately twice length of carpi; dorsal margins and lateral surface smooth, almost glabrous; mesial surfaces each with longitudinal rows of setae adjacent to dorsal and ventral margins. Carpi 0.6 length of meri; dorsal surfaces each with row of very small spinules, strongest distally; lateral, mesial and ventral surfaces almost smooth. Meri each with 1 small dorsodistal spine; dorsal surface with 2 small subdistal spines and widely spaced stiff setae; lateral and mesial faces glabrous; ventral surfaces with few short setae, ventrolateral distal angle with small spine. Ischia unarmed.
</paragraph>
<caption id="DF2966108C0210386700F87781F2F7B0" ID-DOI="http://doi.org/10.5281/zenodo.5352052" ID-Zenodo-Dep="5352052" httpUri="https://zenodo.org/record/5352052/files/figure.png" pageId="10" pageNumber="166" startId="10.[113,148,2035,2057]" targetBox="[156,1408,188,2019]" targetPageId="10">
<paragraph id="8BE936988C0210386700F87781F2F7B0" blockId="10.[113,1452,2035,2115]" pageId="10" pageNumber="166">
Fig. 8. 
<taxonomicName id="4C564D1B8C02103867CDF87785D8F7FB" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[188,424,2035,2056]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="166" phylum="Arthropoda" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B922EA8A8C02103867CDF87785D8F7FB" box="[188,424,2035,2056]" italics="true" pageId="10" pageNumber="166">Catapagurus fimbriatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21157F18C02103866C6F8708642F7FA" box="[439,562,2035,2057]" pageId="10" pageNumber="166" rank="species">new species</taxonomicNameLabel>
. A–F, holotype, male (sl 1.5 mm), NMCR 50802; G, paratype, male (sl 1.4 mm), ZRC 2021.0414. A, right pereopod 2, lateral view; B, same, propodus and carpus, mesial view; C, same, dactylus, mesial view; D, left pereopod 3, lateral view; E, same, propodus and carpus, mesial view; F, same, dactylus, mesial view. G, left pereopod 3, merus, lateral view.
</paragraph>
</caption>
<paragraph id="8BE936988C03103967F9FDF98657FAE6" blockId="11.[134,777,637,1301]" pageId="11" pageNumber="167">
Pereopods 3 (
<figureCitation id="136D2A1D8C031039666EFDF985E6FD66" box="[287,406,637,661]" captionStart="Fig" captionStartId="10.[113,148,2035,2057]" captionTargetBox="[156,1408,188,2019]" captionTargetId="figure-11@10.[127,1459,173,2023]" captionTargetPageId="10" captionText="Fig. 8. Catapagurus fimbriatus, new species. A–F, holotype, male (sl 1.5 mm), NMCR 50802; G, paratype, male (sl 1.4 mm), ZRC 2021.0414. A, right pereopod 2, lateral view; B, same, propodus and carpus, mesial view; C, same, dactylus, mesial view; D, left pereopod 3, lateral view; E, same, propodus and carpus, mesial view; F, same, dactylus, mesial view. G, left pereopod 3, merus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5352052" httpUri="https://zenodo.org/record/5352052/files/figure.png" pageId="11" pageNumber="167">Fig. 8D–G</figureCitation>
) distinctly longer than pereopods 2. Dactyli (
<figureCitation id="136D2A1D8C031039667BFD19852EFD46" box="[266,350,669,693]" captionStart="Fig" captionStartId="10.[113,148,2035,2057]" captionTargetBox="[156,1408,188,2019]" captionTargetId="figure-11@10.[127,1459,173,2023]" captionTargetPageId="10" captionText="Fig. 8. Catapagurus fimbriatus, new species. A–F, holotype, male (sl 1.5 mm), NMCR 50802; G, paratype, male (sl 1.4 mm), ZRC 2021.0414. A, right pereopod 2, lateral view; B, same, propodus and carpus, mesial view; C, same, dactylus, mesial view; D, left pereopod 3, lateral view; E, same, propodus and carpus, mesial view; F, same, dactylus, mesial view. G, left pereopod 3, merus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5352052" httpUri="https://zenodo.org/record/5352052/files/figure.png" pageId="11" pageNumber="167">Fig. 8F</figureCitation>
) narrowly blade-shaped, 1.3 times as long as propodi; in dorsal view, straight; in lateral view, gently curved ventrally; dorsal margins each with row of numerous, closely spaced setae, becoming longer and more stiff distally; mesial faces each with row of numerous setae adjacent to ventral margin; lateral surfaces and ventral margins unarmed. Propodi (
<figureCitation id="136D2A1D8C03103966B1FCD98664FC86" box="[448,532,861,885]" captionStart="Fig" captionStartId="10.[113,148,2035,2057]" captionTargetBox="[156,1408,188,2019]" captionTargetId="figure-11@10.[127,1459,173,2023]" captionTargetPageId="10" captionText="Fig. 8. Catapagurus fimbriatus, new species. A–F, holotype, male (sl 1.5 mm), NMCR 50802; G, paratype, male (sl 1.4 mm), ZRC 2021.0414. A, right pereopod 2, lateral view; B, same, propodus and carpus, mesial view; C, same, dactylus, mesial view; D, left pereopod 3, lateral view; E, same, propodus and carpus, mesial view; F, same, dactylus, mesial view. G, left pereopod 3, merus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5352052" httpUri="https://zenodo.org/record/5352052/files/figure.png" pageId="11" pageNumber="167">Fig. 8E</figureCitation>
) approximately twice length of carpi; dorsal margins unarmed; mesial surfaces each with sparse granules and row of short setae adjacent to dorsal margins; lateral surfaces smooth; ventral margins each with row of short setae in distal 0.7. Carpi (
<figureCitation id="136D2A1D8C03103965DCFC598770FC06" box="[685,768,989,1013]" captionStart="Fig" captionStartId="10.[113,148,2035,2057]" captionTargetBox="[156,1408,188,2019]" captionTargetId="figure-11@10.[127,1459,173,2023]" captionTargetPageId="10" captionText="Fig. 8. Catapagurus fimbriatus, new species. A–F, holotype, male (sl 1.5 mm), NMCR 50802; G, paratype, male (sl 1.4 mm), ZRC 2021.0414. A, right pereopod 2, lateral view; B, same, propodus and carpus, mesial view; C, same, dactylus, mesial view; D, left pereopod 3, lateral view; E, same, propodus and carpus, mesial view; F, same, dactylus, mesial view. G, left pereopod 3, merus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5352052" httpUri="https://zenodo.org/record/5352052/files/figure.png" pageId="11" pageNumber="167">Fig. 8E</figureCitation>
) 0.6 length of meri; dorsal surfaces each with row of very small spinules, strongest distally; lateral surface with sparse granules medially, mesial and ventral surfaces almost smooth. Meri each with 1 small dorsodistal spine; dorsal surface with 2 small subdistal spines and irregular rows of short stiff setae mesially; lateral surface with sparse granules; mesial surface smooth; ventral surfaces slightly granulate, bearing 3 spinules in 
<typeStatus id="54ED883A8C03103967D7FB5A8576FB05" box="[166,262,1246,1270]" pageId="11" pageNumber="167" type="paratype">paratype</typeStatus>
, no distinct spinule in 
<typeStatus id="54ED883A8C031039657DFB59861EFB06" box="[524,622,1245,1269]" pageId="11" pageNumber="167" type="holotype">holotype</typeStatus>
, ventrolateral distal angle unarmed. Ischia unarmed.
</paragraph>
<paragraph id="8BE936988C03103967F9FAB98672FA26" blockId="11.[135,777,1341,1493]" pageId="11" pageNumber="167">
Pereopods 4 (
<figureCitation id="136D2A1D8C0310396669FAB98517FAA6" box="[280,359,1341,1365]" captionStart="Fig" captionStartId="7.[136,171,1535,1557]" captionTargetBox="[137,1476,175,1519]" captionTargetId="figure-8@7.[124,1492,171,1530]" captionTargetPageId="7" captionText="Fig. 5. Catapagurus fimbriatus, new species. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352046" httpUri="https://zenodo.org/record/5352046/files/figure.png" pageId="11" pageNumber="167">Fig. 5F</figureCitation>
) weakly semichelate; dactyli each with well developed preungual process usually far extending beyond tip of claw; propodal rasp consisting of single row of corneous scales on distal half of ventral margin, distalmost scale longest. Pereopods 5 chelate.
</paragraph>
<paragraph id="8BE936988C03103967F9FA7986E9F986" blockId="11.[135,775,1533,1653]" pageId="11" pageNumber="167">
Males with long right sexual tube curved dorsally over pleon and reaching pleonal midline, terminating in 2 small, unequal lobes (
<figureCitation id="136D2A1D8C03103967BFF9B98535F9A6" box="[206,325,1597,1621]" captionStart="Fig" captionStartId="7.[136,171,1535,1557]" captionTargetBox="[137,1476,175,1519]" captionTargetId="figure-8@7.[124,1492,171,1530]" captionTargetPageId="7" captionText="Fig. 5. Catapagurus fimbriatus, new species. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352046" httpUri="https://zenodo.org/record/5352046/files/figure.png" pageId="11" pageNumber="167">Fig. 5G, H</figureCitation>
). Left coxa lacking gonopore, having row of setae on ventral surface posteriorly (
<figureCitation id="136D2A1D8C0310396544F9D986FCF986" box="[565,652,1629,1653]" captionStart="Fig" captionStartId="7.[136,171,1535,1557]" captionTargetBox="[137,1476,175,1519]" captionTargetId="figure-8@7.[124,1492,171,1530]" captionTargetPageId="7" captionText="Fig. 5. Catapagurus fimbriatus, new species. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352046" httpUri="https://zenodo.org/record/5352046/files/figure.png" pageId="11" pageNumber="167">Fig. 5G</figureCitation>
).
</paragraph>
<paragraph id="8BE936988C03103967F9F9198609F8C6" blockId="11.[134,777,1693,1845]" pageId="11" pageNumber="167">
Thoracic sternite 6 (pereopods 3) (
<figureCitation id="136D2A1D8C0310396570F919863CF946" box="[513,588,1693,1717]" captionStart="Fig" captionStartId="7.[136,171,1535,1557]" captionTargetBox="[137,1476,175,1519]" captionTargetId="figure-8@7.[124,1492,171,1530]" captionTargetPageId="7" captionText="Fig. 5. Catapagurus fimbriatus, new species. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352046" httpUri="https://zenodo.org/record/5352046/files/figure.png" pageId="11" pageNumber="167">Fig. 5I</figureCitation>
) with very wide, subtrapezoidal anterior lobe, much shorter than large posterior lobe; with row of short setae on anterior margin; posterior lobe much longer than anterior lobe. Sternite 8 consisting of two small rounded lobes widely separated.
</paragraph>
<paragraph id="8BE936988C03103967F9F8D9850EF846" blockId="11.[136,775,1885,1973]" pageId="11" pageNumber="167">Pleon in males with 3 unpaired, uniramous pleopods (pleopods 3–5); females unknown. Uropodal protopods not protruding posteriorly.</paragraph>
<paragraph id="8BE936988C03103967F9F8598077FF06" blockId="11.[136,775,2013,2102]" lastBlockId="11.[835,1473,189,245]" pageId="11" pageNumber="167">
Telson (
<figureCitation id="136D2A1D8C031039679DF8598532F806" box="[236,322,2013,2037]" captionStart="Fig" captionStartId="7.[136,171,1535,1557]" captionTargetBox="[137,1476,175,1519]" captionTargetId="figure-8@7.[124,1492,171,1530]" captionTargetPageId="7" captionText="Fig. 5. Catapagurus fimbriatus, new species. A–I, holotype, male (sl 1.5 mm), NMCR 50802; J, paratype, male (1.4 mm), ZRC 2021.0414. A, shield, carapace lateral lobes and cephalic appendages, dorsal view; B, left antennule, ultimate article of peduncle and flagella, lateral view; C, left maxilliped 3, endopod, lateral view; D, ischium, ventral view; E, coxae of chelipeds, ventral view; F, left pereopod 4, lateral view; G, coxae of pereopods 5 and thoracic sternite 8, ventral view; H, distal part of right sexual tube, dorsal view; I, thoracic sternite 6, ventral view; J, telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5352046" httpUri="https://zenodo.org/record/5352046/files/figure.png" pageId="11" pageNumber="167">Fig. 5J</figureCitation>
) with lateral indentations suggesting separation of anterior and posterior portions; triangular posterior lobes separated by very wide, trapezoidal medial cleft, each terminating in subacute tip; oblique terminal margins unarmed.
</paragraph>
<paragraph id="8BE936988C0310396432FE9980D8FEC6" blockId="11.[835,1192,285,309]" box="[835,1192,285,309]" pageId="11" pageNumber="167">
<emphasis id="B922EA8A8C0310396432FE99806FFEC6" bold="true" box="[835,1055,285,309]" pageId="11" pageNumber="167">Colouration in life.</emphasis>
<collectingRegion id="4992F87A8C0310396359FE9A8020FEC6" box="[1064,1104,286,309]" country="Maldives" name="Uthuru" pageId="11" pageNumber="167">Not</collectingRegion>
known.
</paragraph>
</subSubSection>
<subSubSection id="C34C65138C0310396432FED9816AFE66" pageId="11" pageNumber="167" type="distribution">
<paragraph id="8BE936988C0310396432FED9816AFE66" blockId="11.[835,1473,349,405]" pageId="11" pageNumber="167">
<emphasis id="B922EA8A8C0310396432FED987A8FE86" bold="true" box="[835,984,349,373]" pageId="11" pageNumber="167">Distribution.</emphasis>
Presently known only from the 
<collectingRegion id="4992F87A8C0310396239FED981FAFE86" box="[1352,1418,349,373]" country="Philippines" name="Bohol" pageId="11" pageNumber="167">Bohol</collectingRegion>
Sea, central 
<collectingCountry id="F34176088C03103964E5FEF98060FE66" box="[916,1040,381,405]" name="Philippines" pageId="11" pageNumber="167">Philippines</collectingCountry>
; at depths of 
<quantity id="4CAE9B7D8C03103963D7FEF98166FE66" box="[1190,1302,381,405]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.3699999999999999" metricValueMin="0.77" pageId="11" pageNumber="167" unit="m" value="107.0" valueMax="137.0" valueMin="77.0">
77– 
<specimenCount id="9D50FD118C03103963BEFEF98166FE66" box="[1231,1302,381,405]" pageId="11" pageNumber="167" type="generic">137 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C65138C0310396432FE398088FA26" pageId="11" pageNumber="167" type="discussion">
<paragraph id="8BE936988C0310396432FE398152FCC6" blockId="11.[834,1477,445,821]" pageId="11" pageNumber="167">
<emphasis id="B922EA8A8C0310396432FE3987C3FE26" bold="true" box="[835,947,445,469]" pageId="11" pageNumber="167">Remarks.</emphasis>
There were no female specimens available for study. None of the 
<specimenCount id="9D50FD118C0310396366FE5A800FFE05" box="[1047,1151,478,502]" pageId="11" pageNumber="167" type="generic">two types</specimenCount>
is intact: in the 
<typeStatus id="54ED883A8C0310396244FE5981E5FE06" box="[1333,1429,477,501]" pageId="11" pageNumber="167" type="holotype">holotype</typeStatus>
, the entire pleon is missing; in the 
<typeStatus id="54ED883A8C03103963FEFE7A809EFDE5" box="[1167,1262,510,534]" pageId="11" pageNumber="167" type="paratype">paratype</typeStatus>
, both chelipeds and pereopods 2 are missing. Nevertheless, the new species is assigned to 
<taxonomicName id="4C564D1B8C0310396499FDBA8006FDA6" authorityName="A. Milne-Edwards" authorityYear="1880" box="[1000,1142,574,597]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="167" phylum="Arthropoda" rank="genus">
<emphasis id="B922EA8A8C0310396499FDBA8006FDA6" box="[1000,1142,574,597]" italics="true" pageId="11" pageNumber="167">Catapagurus</emphasis>
</taxonomicName>
without hesitation because it agrees with the generic features, including the gill formula consisting of 11 pairs of biserial gills, the somewhat reduced crista dentata on the maxilliped 3 ischium, the presence of a prominent preungual process on the pereopods 4 dactyli and the possession of a long sexual tube arising from the right pereopod 5 coxa, curving exteriorly and over the lateral side of the body (
<bibRefCitation id="EFC74B698C0310396372FC9980A0FCC6" author="McLaughlin PA" box="[1027,1232,797,821]" pageId="11" pageNumber="167" pagination="111 - 144" refId="ref9987" refString="McLaughlin PA (2003) Illustrated keys to families and genera of the superfamily Paguroidea (Crustacea: Decapoda: Anomura), with diagnoses of genera of Paguridae. Memoirs of Museum Victoria, 60: 111 - 144." type="journal article" year="2003">McLaughlin, 2003</bibRefCitation>
, 
<bibRefCitation id="EFC74B698C03103963ADFC998166FCC6" author="McLaughlin PA" box="[1244,1302,797,821]" pageId="11" pageNumber="167" pagination="1 - 16" refId="ref10028" refString="McLaughlin PA (2004) A reappraisal of the hermit crab genera Catapagurus A. Milne-Edwards and Hemipagurus Smith (Crustacea: Decapoda: Anomura: Paguridae), with the description of a new species. Zootaxa, 433: 1 - 16." type="journal article" year="2004">2004</bibRefCitation>
).
</paragraph>
<paragraph id="8BE936988C0310396432FCDA8088FA26" blockId="11.[834,1476,861,1493]" pageId="11" pageNumber="167">
<taxonomicName id="4C564D1B8C0310396432FCDA804DFC86" authority="Komai &amp; Rahayu, 2021" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[835,1085,861,885]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="167" phylum="Arthropoda" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B922EA8A8C0310396432FCDA804DFC86" box="[835,1085,861,885]" italics="true" pageId="11" pageNumber="167">Catapagurus fimbriatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21157F18C0310396339FCDA80B8FC86" box="[1096,1224,861,885]" pageId="11" pageNumber="167" rank="species">new species</taxonomicNameLabel>
, is readily distinguished from all other congeneric species in the characteristic setation of the pereopod 3 dactyli and propodi: the dactylus has a row of dense setae on the dorsal margin and on the mesial face adjacent to the ventral margin; the merus is also provided with setal rows on the mesial face adjacent to the dorsal margin and on the distal 0.7 of the ventral margin. Such setation is not seen in the 24 other species in the genus (
<bibRefCitation id="EFC74B698C031039643AFBD9806AFB86" author="McLaughlin PA" box="[843,1050,1117,1141]" pageId="11" pageNumber="167" pagination="433 - 572" refId="ref9882" refString="McLaughlin PA (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier A &amp; Bouchet P (eds.) Resultats des Campagnes MUSORSTOM, Volume 16. Memoires du Museum national d'Histoire naturelle, Paris, 172: 433 - 572." type="journal article" year="1997">McLaughlin, 1997</bibRefCitation>
, 
<bibRefCitation id="EFC74B698C0310396358FBD98011FB86" author="McLaughlin PA" box="[1065,1121,1117,1141]" pageId="11" pageNumber="167" pagination="385 - 460" refId="ref9937" refString="McLaughlin PA (2002) A review of the hermit-crab (Decapoda: Anomura: Paguridea) fauna of southern Thailand, with particular emphasis on the Andaman Sea, and descriptions of three new species. Phuket Marine Biological Center Special Publication, 23: 385 - 460." type="journal article" year="2002">2002</bibRefCitation>
, 
<bibRefCitation id="EFC74B698C031039631EFBD980D8FB86" author="McLaughlin PA" box="[1135,1192,1117,1141]" pageId="11" pageNumber="167" pagination="1 - 16" refId="ref10028" refString="McLaughlin PA (2004) A reappraisal of the hermit crab genera Catapagurus A. Milne-Edwards and Hemipagurus Smith (Crustacea: Decapoda: Anomura: Paguridae), with the description of a new species. Zootaxa, 433: 1 - 16." type="journal article" year="2004">2004</bibRefCitation>
; 
<bibRefCitation id="EFC74B698C03103963C6FBD981EBFB86" author="Hogarth P &amp; Gherardi F &amp; McLaughlin PA" box="[1207,1435,1117,1141]" pageId="11" pageNumber="167" pagination="149 - 175" refId="ref9273" refString="Hogarth P, Gherardi F &amp; McLaughlin PA (1998) Hermit crabs of the Maldives, with the description of a new species of Catapagurus. Tropical Zoology, 11: 149 - 175." type="journal article" year="1998">Hogarth et al., 1998</bibRefCitation>
; de 
<bibRefCitation id="EFC74B698C0310396432FBF980B3FB66" author="Saint Laurent M de &amp; McLaughlin PA" box="[835,1219,1149,1173]" pageId="11" pageNumber="167" pagination="104 - 209" refId="ref10625" refString="Saint Laurent M de &amp; McLaughlin PA (2000) Superfamily Paguroidea, Family Paguridae. In: Forest J, Saint Laurent M de, McLaughlin PA &amp; Lemaitre R (eds.) The marine fauna of New Zealand: Paguridea (Decapoda: Anomura) exclusive of the Lithodidae. NIWA Biodiversity Memoir, 114: 104 - 209." type="journal article" year="2000">Saint Laurent &amp; McLaughlin, 2000</bibRefCitation>
; 
<bibRefCitation id="EFC74B698C03103963BFFBF9811EFB66" author="Asakura A" box="[1230,1390,1149,1173]" pageId="11" pageNumber="167" pagination="823 - 891" refId="ref9098" refString="Asakura A (2001) A revision of the hermit crab genera Catapagurus A. Milne-Edwards and Hemipagurus Smith from the Indo- West Pacific (Crustacea: Decapoda: Anomura: Paguridae). Invertebrate Taxonomy, 15: 823 - 891." type="journal article" year="2001">Asakura, 2001</bibRefCitation>
; 
<bibRefCitation id="EFC74B698C0310396208FBF9879CFB46" author="Komai T &amp; Osawa M" pageId="11" pageNumber="167" pagination="57 - 68" refId="ref9346" refString="Komai T &amp; Osawa M (2009) A new hermit crab species of the genus Catapagurus (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southern Japan. Zootaxa, 2211: 57 - 68." type="journal article" year="2009">Komai &amp; Osawa, 2009</bibRefCitation>
; 
<bibRefCitation id="EFC74B698C0310396487FB1980A3FB46" author="Nucci PR &amp; Melo GAS de" box="[1014,1235,1181,1205]" pageId="11" pageNumber="167" pagination="81 - 85" refId="ref10435" refString="Nucci PR &amp; Melo GAS de (2012) A new species of Catapagurus A. Milne-Edwards (Crustacea: Decapoda: Anomura: Paguridae) from Brazil, with a key to the western Atlantic species. Zootaxa, 3499: 81 - 85." type="journal article" year="2012">Nucci &amp; Melo, 2012</bibRefCitation>
; this study). The shape of the ambulatory dactyli of the present new species is also notable: the dactyli are slender as in species referred to the 
<taxonomicName id="4C564D1B8C0310396434FB7A87ACFAE6" baseAuthorityName="Terao" baseAuthorityYear="1914" box="[837,988,1277,1301]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="167" phylum="Arthropoda" rank="species" species="misakiensis">
<emphasis id="B922EA8A8C0310396434FB7A87ACFAE6" box="[837,988,1277,1301]" italics="true" pageId="11" pageNumber="167">C. misakiensis</emphasis>
</taxonomicName>
species group as diagnosed by 
<bibRefCitation id="EFC74B698C031039626EFB7981CEFAE6" author="Asakura A" box="[1311,1470,1277,1301]" pageId="11" pageNumber="167" pagination="823 - 891" refId="ref9098" refString="Asakura A (2001) A revision of the hermit crab genera Catapagurus A. Milne-Edwards and Hemipagurus Smith from the Indo- West Pacific (Crustacea: Decapoda: Anomura: Paguridae). Invertebrate Taxonomy, 15: 823 - 891." type="journal article" year="2001">Asakura (2001</bibRefCitation>
; as 
<taxonomicName id="4C564D1B8C0310396415FA9A879EFAC6" authorityName="Yokoya" authorityYear="1933" box="[868,1006,1310,1333]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="167" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B922EA8A8C0310396415FA9A879EFAC6" box="[868,1006,1310,1333]" italics="true" pageId="11" pageNumber="167">C. japonicus</emphasis>
</taxonomicName>
group), but in this new species the shape is similar to the blade-shaped dactyli with shallow concave mesial faces of the 
<taxonomicName id="4C564D1B8C0310396360FADA8009FA86" authorityName="Henderson" authorityYear="1888" box="[1041,1145,1373,1397]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="167" phylum="Arthropoda" rank="species" species="ensifer">
<emphasis id="B922EA8A8C0310396360FADA8009FA86" box="[1041,1145,1373,1397]" italics="true" pageId="11" pageNumber="167">C. ensifer</emphasis>
</taxonomicName>
species of 
<bibRefCitation id="EFC74B698C0310396381FAD981EAFA86" author="Asakura A" box="[1264,1434,1373,1397]" pageId="11" pageNumber="167" pagination="823 - 891" refId="ref9098" refString="Asakura A (2001) A revision of the hermit crab genera Catapagurus A. Milne-Edwards and Hemipagurus Smith from the Indo- West Pacific (Crustacea: Decapoda: Anomura: Paguridae). Invertebrate Taxonomy, 15: 823 - 891." type="journal article" year="2001">Asakura (2001)</bibRefCitation>
. As a result, this new species appears to be intermediate between the two species groups as proposed by 
<bibRefCitation id="EFC74B698C0310396397FA1981FFFA46" author="Asakura A" box="[1254,1423,1437,1461]" pageId="11" pageNumber="167" pagination="823 - 891" refId="ref9098" refString="Asakura A (2001) A revision of the hermit crab genera Catapagurus A. Milne-Edwards and Hemipagurus Smith from the Indo- West Pacific (Crustacea: Decapoda: Anomura: Paguridae). Invertebrate Taxonomy, 15: 823 - 891." type="journal article" year="2001">Asakura (2001)</bibRefCitation>
, and cannot be satisfactorily placed in either.
</paragraph>
</subSubSection>
<subSubSection id="C34C65138C0310396432FA7987E9F9A6" pageId="11" pageNumber="167" type="etymology">
<paragraph id="8BE936988C0310396432FA7987E9F9A6" blockId="11.[835,1474,1533,1621]" pageId="11" pageNumber="167">
<emphasis id="B922EA8A8C0310396432FA7987B5F9E6" bold="true" box="[835,965,1533,1557]" pageId="11" pageNumber="167">Etymology.</emphasis>
The Latin 
<taxonomicName id="4C564D1B8C0310396335FA7980C0F9E6" authorityName="Komai &amp; Rahayu" authorityYear="2021" box="[1092,1200,1533,1557]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="167" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="B922EA8A8C0310396335FA7980C0F9E6" box="[1092,1200,1533,1557]" italics="true" pageId="11" pageNumber="167">fimbriatus</emphasis>
</taxonomicName>
(= fringed) refers to the characteristic setation of the ambulatory dactyli of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>