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378 lines
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<document id="1A1B4D718D7E59D46B7515B93D82981A" ID-DOI="10.6620/ZS.2017.56-24" ID-ISSN="1810-522X" ID-PMC="PMC6517722" ID-PubMed="31966223" ID-Zenodo-Dep="12824860" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721913874541" checkinUser="felipe" docAuthor="Ahyong, Shane T. & Ng, Peter K. L." docDate="2017" docId="03FB8D475F14FFEAFCD7FDA8FD66FB7F" docLanguage="en" docName="ZoolStud.56.24.1-20.pdf.imf" docOrigin="Zoological Studies (Zool. Stud.) 56 (24)" docSource="http://dx.doi.org/10.5281/zenodo.8060427" docStyle="DocumentStyle:CB122E4603BA284FA287D3847868CAF6.2:ZoolStud.2016-2018.journal_article" docStyleId="CB122E4603BA284FA287D3847868CAF6" docStyleName="ZoolStud.2016-2018.journal_article" docStyleVersion="2" docTitle="Cymonomus japonicus Balss 1922" docType="treatment" docVersion="3" lastPageNumber="15" masterDocId="FFC2F53F5F1FFFE4FFADFFF4FFAEFFA9" masterDocTitle="Zavreliella undetermined" masterLastPageNumber="20" masterPageNumber="1" pageNumber="12" updateTime="1721930283392" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="348A0F62470D8BBCBAEBF90D31703718">
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<mods:title id="5FF308E27A1212F35FD5603A3E28E2E9">Zavreliella undetermined</mods:title>
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</mods:titleInfo>
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<mods:name id="39562DA97E5503C2E467FD8489B813E9" type="personal">
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<mods:namePart id="B899F6367B15A6F0012922DEBFA18E55">Ahyong, Shane T.</mods:namePart>
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<mods:affiliation id="16F75E4C9D84467ACBBBFC2EE0208942">Australian Museum, 1 William St., Sydney, NSW 2010, Australia, and School of Biological, Earth & Environmental Sciences, University of New South Wales, Kensington, NSW 2052, Australia</mods:affiliation>
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<mods:namePart id="DFD24EBC50C3399B51A4F9AC4AABE496">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="B838D7AB0CF3134EFE6A1FCBF4A66240">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore. E-mail: peterng @ nus. edu. sg</mods:affiliation>
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<mods:nameIdentifier id="876F49A85AE43492BBD6369EABF79B04" type="email">peterng@nus.edu.sg</mods:nameIdentifier>
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<mods:typeOfResource id="8A68AB24E3AD2B08D0FC119F1611F9AB">text</mods:typeOfResource>
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<mods:title id="99EEE7B9B28F132A2B3293F87143A6B5">Zoological Studies</mods:title>
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<mods:part id="2B7F84D45A673FB4BF3FB4DB82ADFECC">
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<mods:date id="B891A138B4B3741689CFF001F2D1FB72">2017</mods:date>
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<mods:title id="0C57737F5442ECD345282CCC5FE69BE0">Zool. Stud.</mods:title>
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<mods:number id="5A7FF9E7DD27DC58E970F865171885E9">2017-08-18</mods:number>
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<mods:number id="B8B0D41DFD7EC1E44022E9137350CFDC">56</mods:number>
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<mods:number id="80EE9ECDA38BE49556DA0C19086ADA3C">24</mods:number>
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<mods:classification id="C08AFAB73BEAEEC2F5277F328B1626F2">journal article</mods:classification>
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<mods:identifier id="C73DF59C517AD394FA1888F14315A76C" type="DOI">10.6620/ZS.2017.56-24</mods:identifier>
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<treatment id="03FB8D475F14FFEAFCD7FDA8FD66FB7F" LSID="urn:lsid:plazi:treatment:03FB8D475F14FFEAFCD7FDA8FD66FB7F" httpUri="http://treatment.plazi.org/id/03FB8D475F14FFEAFCD7FDA8FD66FB7F" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
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<subSubSection id="C3486FDA5F14FFEFFCD7FDA8FA96FDDF" box="[890,1336,604,630]" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="8BED3C515F14FFEFFCD7FDA8FA96FDDF" blockId="11.[890,1336,604,662]" box="[890,1336,604,630]" pageId="11" pageNumber="12">
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<heading id="D0A58B3D5F14FFEFFCD7FDA8FA96FDDF" bold="true" box="[890,1336,604,630]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="12" reason="2">
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<taxonomicName id="4C5247D25F14FFEFFCD7FDA8FA96FDDF" ID-CoL="337VH" authority="Balss, 1922" authorityName="Balss" authorityYear="1922" box="[890,1336,604,630]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="japonicus">
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<emphasis id="B926E0435F14FFEFFCD7FDA8FA96FDDF" bold="true" box="[890,1336,604,630]" pageId="11" pageNumber="12">
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<emphasis id="B926E0435F14FFEFFCD7FDA8FB31FDDF" bold="true" box="[890,1183,604,630]" italics="true" pageId="11" pageNumber="12">Cymonomus japonicus</emphasis>
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<bibRefCitation id="EFC341A05F14FFEFFB0AFDA9FA96FDDF" author="Balss H." box="[1191,1336,605,630]" pageId="11" pageNumber="12" pagination="104 - 140" refId="ref12958" refString="Balss H. 1922. Ostasiatische Decapoden. III. Die Dromiaceen, Oxystomen und Parthenopiden. Archiv fur Naturgeschichte 88 (A): 104 - 140, figs 1 - 9." type="journal article" year="1922">Balss, 1922</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3486FDA5F14FFEFFB82FD89FB2DFD3F" box="[1071,1155,637,662]" pageId="11" pageNumber="12" type="description">
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<paragraph id="8BED3C515F14FFEFFB82FD89FB2DFD3F" blockId="11.[890,1336,604,662]" box="[1071,1155,637,662]" pageId="11" pageNumber="12">
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(
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<figureCitation id="136920D45F14FFEFFB9BFD89FBD2FD3F" box="[1078,1148,637,662]" captionStart="Fig" captionStartId="12.[174,209,1842,1862]" captionTargetBox="[196,1413,251,1817]" captionTargetId="figure-1@12.[192,1433,238,1830]" captionTargetPageId="12" captionText="Fig. 5. Cymonomus japonicus Balss, 1922, Japan, WPMNH #47. A-G: female neotype, cl 5.9 mm, pcl 4.6 mm, cw 5.0 mm; H-J: male, cl 4.8 mm, pcl 3.7 mm, cw 3.8 mm; K: male, cl 4.0 mm, pcl 3.9 mm, cw 3.8 mm; L, male, cl 5.0 mm, pcl 3.9 mm, cw 3.8 mm. A, dorsal habitus; B, K, L, fronto-orbital region; C, right basal antennal spine; D, carapace tubercles of right anterolateral corner; E, right maxilliped 3; F, thoracic sternite 3; G, abdomen, posterior; H, telson; I, right gonopod 1, abdominal view; J, right gonopod 2, abdominal view. Scale bars: A, G = 2.0 mm; B, E, F, H-L = 1.0 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12824870" httpUri="https://zenodo.org/record/12824870/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3486FDA5F14FFEFFC80FD43FB79FCC5" pageId="11" pageNumber="12" type="reference_group">
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<paragraph id="8BED3C515F14FFEFFC80FD43FCD1FC9F" blockId="11.[813,1415,695,876]" pageId="11" pageNumber="12">
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<treatmentCitationGroup id="AB421B7F5F14FFEFFC80FD43FCD1FC9F" pageId="11" pageNumber="12">
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<treatmentCitation id="0AF31A405F14FFEFFC80FD43FAF3FD65" author="Balss H." box="[813,1373,695,716]" page="118" pageId="11" pageNumber="12" year="1922">
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<taxonomicName id="4C5247D25F14FFEFFC80FD43FAF3FD65" ID-CoL="337VH" authority="Balss, 1922: 117 - 118" authorityName="Balss" authorityPageNumber="117 - 118" authorityYear="1922" box="[813,1373,695,716]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="granulatus" subSpecies="japonicus">
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<emphasis id="B926E0435F14FFEFFC80FD43FBD3FD65" box="[813,1149,695,716]" italics="true" pageId="11" pageNumber="12">Cymonomus granulatus japonicus</emphasis>
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<bibRefCitation id="EFC341A05F14FFEFFB2AFD4CFAF3FD65" author="Balss H." box="[1159,1373,696,716]" pageId="11" pageNumber="12" pagination="104 - 140" refId="ref12958" refString="Balss H. 1922. Ostasiatische Decapoden. III. Die Dromiaceen, Oxystomen und Parthenopiden. Archiv fur Naturgeschichte 88 (A): 104 - 140, figs 1 - 9." type="journal article" year="1922">Balss, 1922: 117-118</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, fig. 5 [type locality: off Shionomisaki, Kii Peninsula,
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<collectingCountry id="F3457CC15F14FFEFFAE9FD26FA2DFD4F" box="[1348,1411,722,742]" name="Japan" pageId="11" pageNumber="12">Japan</collectingCountry>
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; fixed by present
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<typeStatus id="54E982F35F14FFEFFBADFD19FBE2FCA8" box="[1024,1100,749,769]" pageId="11" pageNumber="12" type="neotype">neotype</typeStatus>
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designation]. -
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<treatmentCitation id="0AF31A405F14FFEFFB72FD19FC6AFCB5" author="Griffin DJG & Brown DE" page="252" pageId="11" pageNumber="12" year="1976">
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<bibRefCitation id="EFC341A05F14FFEFFB72FD19FC6AFCB5" author="Griffin DJG & Brown DE" pageId="11" pageNumber="12" pagination="248 - 271" refId="ref13141" refString="Griffin DJG, Brown DE. 1976. Deepwater decapod Crustacea from eastern Australia: brachyuran crabs. Records of the Australian Museum 30: 248 - 271." type="journal article" year="1976">Griffin and Brown 1976: 252</bibRefCitation>
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</treatmentCitation>
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. -
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<treatmentCitation id="0AF31A405F14FFEFFC76FCF3FBC3FCB5" author="Sakai T." box="[987,1133,775,796]" page="36" pageId="11" pageNumber="12" year="1976">
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<bibRefCitation id="EFC341A05F14FFEFFC76FCF3FBC3FCB5" author="Sakai T." box="[987,1133,775,796]" pageId="11" pageNumber="12" refId="ref13600" refString="Sakai T. 1976. Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo." type="book" year="1976">Sakai 1976: 36</bibRefCitation>
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</treatmentCitation>
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, fig. 20. -
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<treatmentCitation id="0AF31A405F14FFEFFB66FCFCFACEFCB5" author="Nagai S." box="[1227,1376,776,796]" page="32" pageId="11" pageNumber="12" year="1991">
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<bibRefCitation id="EFC341A05F14FFEFFB66FCFCFACEFCB5" author="Nagai S." box="[1227,1376,776,796]" pageId="11" pageNumber="12" pagination="31 - 34" refId="ref13497" refString="Nagai S. 1991. Some remarkable crabs of Wakayama Prefecture I. Nankiseibutu 33: 31 - 34." type="journal article" year="1991">Nagai 1991: 32</bibRefCitation>
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</treatmentCitation>
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, fig. 1b.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BED3C515F14FFEFFC80FCC9FB79FCC5" blockId="11.[813,1415,695,876]" pageId="11" pageNumber="12">
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<treatmentCitationGroup id="AB421B7F5F14FFEFFC80FCC9FB79FCC5" pageId="11" pageNumber="12">
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<taxonomicName id="4C5247D25F14FFEFFC80FCC9FBBDFCF8" ID-CoL="337VH" authorityName="Balss" authorityYear="1922" box="[813,1043,829,849]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="japonicus">
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<emphasis id="B926E0435F14FFEFFC80FCC9FBBDFCF8" box="[813,1043,829,849]" italics="true" pageId="11" pageNumber="12">Cymonomus japonicus</emphasis>
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</taxonomicName>
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. -
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<treatmentCitation id="0AF31A405F14FFEFFB9EFCC9FB5EFCF8" author="Takeda M." box="[1075,1264,829,849]" page="224" pageId="11" pageNumber="12" year="2001">
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<bibRefCitation id="EFC341A05F14FFEFFB9EFCC9FB5EFCF8" author="Takeda M." box="[1075,1264,829,849]" pageId="11" pageNumber="12" pagination="217 - 262" refId="ref13750" refString="Takeda M. 2001. Annotated list of crabs from Tosa Bay, southwest Japan, collected by the R / V Kotaka Maru during the years 1997 - 2000. In: Fujita, T., Saito, H., & Takeda, M. (eds.), Deep-Sea Fauna and Pollutants in Tosa Bay. National Science Museum Monographs 20: 217 - 262." type="journal article" year="2001">Takeda 2001: 224</bibRefCitation>
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</treatmentCitation>
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. -
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<treatmentCitation id="0AF31A405F14FFEFFAA6FCC9FBBCFCC5" author="Ahyong ST & Brown DE" page="1372" pageId="11" pageNumber="12" year="2003">
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<bibRefCitation id="EFC341A05F14FFEFFAA6FCC9FBBCFCC5" author="Ahyong ST & Brown DE" pageId="11" pageNumber="12" pagination="1363 - 1374" refId="ref12549" refString="Ahyong ST, Brown DE. 2003. New species of Cymonomus from southeastern Australia (Brachyura, Cymonomidae) with a key to the Indo-West Pacific species. Crustaceana 75: 1363 - 1374." type="journal article" year="2003">Ahyong and Brown 2003: 1372</bibRefCitation>
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</treatmentCitation>
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. -
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<treatmentCitation id="0AF31A405F14FFEFFB85FCACFB7DFCC5" author="Ng PKL & Guinot D & Davie PJF" box="[1064,1235,856,876]" page="32" pageId="11" pageNumber="12" year="2008">
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<bibRefCitation id="EFC341A05F14FFEFFB85FCACFB7DFCC5" author="Ng PKL & Guinot D & Davie PJF" box="[1064,1235,856,876]" pageId="11" pageNumber="12" pagination="1 - 286" refId="ref13537" refString="Ng PKL, Guinot D, Davie PJF. 2008. Systema Brachyurorum: part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al. 2008: 32</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3486FDA5F14FFE9FCC4FC67FB86F83F" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="materials_examined">
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<paragraph id="8BED3C515F14FFEFFCC4FC67FB04FBA5" blockId="11.[813,1414,915,1964]" pageId="11" pageNumber="12">
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<emphasis id="B926E0435F14FFEFFCC4FC67FB8FFC05" box="[873,1057,915,940]" italics="true" pageId="11" pageNumber="12">Type material</emphasis>
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:
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<materialsCitation id="3B3A360C5F14FFEFFB95FC67FB04FBA5" collectionCode="WPMNH" country="Japan" elevation="500" location="Shionomisaki" municipality="Kii Peninsula" pageId="11" pageNumber="12" specimenCode="WPMNH #47" specimenCount="1" stateProvince="Wakayama" typeStatus="neotype">
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<typeStatus id="54E982F35F14FFEFFB95FC67FB66FC04" box="[1080,1224,915,941]" pageId="11" pageNumber="12" type="neotype">NEOTYPE</typeStatus>
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:
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<specimenCode id="DBF4942A5F14FFEFFB74FC67FA2AFC05" box="[1241,1412,915,940]" collectionCode="WPMNH" pageId="11" pageNumber="12">WPMNH #47</specimenCode>
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(part), female (cl
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<quantity id="4CAA91B45F14FFEFFC55FC47FBFBFC65" box="[1016,1109,947,972]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="11" pageNumber="12" unit="mm" value="5.9">5.9 mm</quantity>
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, pcl
|
|||
|
|
<quantity id="4CAA91B45F14FFEFFB26FC47FB46FC65" box="[1163,1256,947,972]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="11" pageNumber="12" unit="mm" value="4.6">4.6 mm</quantity>
|
|||
|
|
, cw 5.0 mm), off
|
|||
|
|
<location id="8E8D6A8A5F14FFEFFCF1FC27FBB9FC44" LSID="urn:lsid:plazi:treatment:03FB8D475F14FFEAFCD7FDA8FD66FB7F:8E8D6A8A5F14FFEFFCF1FC27FBB9FC44" box="[860,1047,979,1005]" country="Japan" municipality="Kii Peninsula" name="Shionomisaki" pageId="11" pageNumber="12" stateProvince="Wakayama">Shionomisaki</location>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="6B89A62B5F14FFEFFB8AFC27FB4CFC45" box="[1063,1250,979,1004]" pageId="11" pageNumber="12">Kii Peninsula</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="4996F2B35F14FFEFFB5EFC27FC02FBA5" country="Japan" name="Wakayama" pageId="11" pageNumber="12">Wakayama Prefecture</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F3457CC15F14FFEFFC1BFC07FBADFBA5" box="[950,1027,1011,1036]" name="Japan" pageId="11" pageNumber="12">Japan</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<quantity id="4CAA91B45F14FFEFFBA3FC07FBF2FBA5" box="[1038,1116,1011,1036]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="m" value="500.0">
|
|||
|
|
<elevation id="007FDB625F14FFEFFBA3FC07FBF2FBA5" box="[1038,1116,1011,1036]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="m" value="500.0">500 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
, 1990.
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BED3C515F14FFEFFCC4FBE7FB95FB25" blockId="11.[813,1414,915,1964]" pageId="11" pageNumber="12">
|
|||
|
|
<emphasis id="B926E0435F14FFEFFCC4FBE7FB31FB85" box="[873,1183,1043,1068]" italics="true" pageId="11" pageNumber="12">Other material examined</emphasis>
|
|||
|
|
:
|
|||
|
|
<collectingCountry id="F3457CC15F14FFEFFB1FFBE7FAA3FB85" box="[1202,1293,1043,1068]" name="Japan" pageId="11" pageNumber="12">JAPAN</collectingCountry>
|
|||
|
|
:
|
|||
|
|
<collectionCode id="ED43A4945F14FFEFFAB6FBE7FA2AFB85" box="[1307,1412,1043,1068]" pageId="11" pageNumber="12">WPMNH</collectionCode>
|
|||
|
|
#47 (part),
|
|||
|
|
<specimenCount id="9D54F7D85F14FFEFFC16FBC7FBB1FBE5" box="[955,1055,1075,1100]" count="4" pageId="11" pageNumber="12" type="male">4 males</specimenCount>
|
|||
|
|
(cl
|
|||
|
|
<quantity id="4CAA91B45F14FFEFFBFEFBC7FB18FBE5" box="[1107,1206,1075,1100]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="11" pageNumber="12" unit="mm" value="4.9">4.9 mm</quantity>
|
|||
|
|
, pcl
|
|||
|
|
<quantity id="4CAA91B45F14FFEFFB5EFBC7FAF8FBE5" box="[1267,1366,1075,1100]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="12" unit="mm" value="3.8">3.8 mm</quantity>
|
|||
|
|
, cw
|
|||
|
|
<quantity id="4CAA91B45F14FFEFFC80FBA7FC25FBC5" box="[813,907,1107,1132]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="12" unit="mm" value="3.8">3.8 mm</quantity>
|
|||
|
|
to cl
|
|||
|
|
<quantity id="4CAA91B45F14FFEFFC7AFBA7FB95FBC5" box="[983,1083,1107,1132]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="11" pageNumber="12" unit="mm" value="5.2">5.2 mm</quantity>
|
|||
|
|
, pcl
|
|||
|
|
<quantity id="4CAA91B45F14FFEFFBDAFBA7FB75FBC5" box="[1143,1243,1107,1132]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="11" pageNumber="12" unit="mm" value="3.9">3.9 mm</quantity>
|
|||
|
|
, cw
|
|||
|
|
<quantity id="4CAA91B45F14FFEFFAB8FBA7FAD4FBC5" box="[1301,1402,1107,1132]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="12" unit="mm" value="3.8">3.8 mm</quantity>
|
|||
|
|
), collected with
|
|||
|
|
<typeStatus id="54E982F35F14FFEFFC79FB87FB99FB25" box="[980,1079,1139,1164]" pageId="11" pageNumber="12" type="neotype">neotype</typeStatus>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BED3C515F14FFE9FCC4FB67FDEDFE7F" blockId="11.[813,1414,915,1964]" lastBlockId="13.[136,738,253,1974]" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
|
|||
|
|
<emphasis id="B926E0435F14FFEFFCC4FB67FC51FB05" box="[873,1023,1171,1196]" italics="true" pageId="11" pageNumber="12">Description</emphasis>
|
|||
|
|
: Carapace quadrate, almost square, lateral margins almost parallel; regions weakly indicated, cervical groove more pronounced in males than females, broadly V-shaped; with slender conical anteriorly directed anterolateral spine and similar anterolaterally directed spine on lateral margin behind anterolateral spine; lower pterygostomian region swollen; anterolateral surfaces with few scattered setae. Dorsal and lateral surfaces entirely covered with minute rounded granules, with granules becoming larger more elongate anterolaterally, conical or subcylindrical, not globose. Fronto-orbital margin (excluding rostrum and lateral projections) slightly advanced beyond anterolateral margins, more pronounced in males than females; exceeding half anterior carapace width; outer orbital processes slender, elongate, directed anteriorly, situated below plane of rostrum, laterally spinulate, with acute apices, up to half rostral length. Rostrum slightly longer than eyestalks; 0.26- 0.32 pcl; slender, tapering to acute apex, granular dorsally, minutely spinular laterally. Eyestalks distinctly divergent, stout, flattened, granular and prominently spinulate, fused to carapace below rostral base but demarcation distinct, reaching anteriorly to midlength of antennular peduncle article 1; cornea apparently vestigial, not pigmented. Epistome surface granulate, with blunt tubercle mesial to base of antennules, small spine mesial to base of antenna, with small cluster of elongate granules at base of rostrum.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF2D6CD95F13FFE8FF03F8C6FCFCF81F" ID-DOI="http://doi.org/10.5281/zenodo.12824870" ID-Zenodo-Dep="12824870" httpUri="https://zenodo.org/record/12824870/files/figure.png" pageId="12" pageNumber="13" startId="12.[174,209,1842,1862]" targetBox="[196,1413,251,1817]" targetPageId="12" targetType="figure">
|
|||
|
|
<paragraph id="8BED3C515F13FFE8FF03F8C6FCFCF81F" blockId="12.[174,1453,1841,1974]" pageId="12" pageNumber="13">
|
|||
|
|
<emphasis id="B926E0435F13FFE8FF03F8C6FF41F8EF" bold="true" box="[174,239,1842,1862]" pageId="12" pageNumber="13">Fig. 5.</emphasis>
|
|||
|
|
<taxonomicName id="4C5247D25F13FFE8FF52F8C5FDF0F8EF" authority="Balss, 1922" authorityName="Balss" authorityYear="1922" box="[255,606,1841,1862]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="japonicus">
|
|||
|
|
<emphasis id="B926E0435F13FFE8FF52F8C5FE73F8EF" box="[255,477,1841,1862]" italics="true" pageId="12" pageNumber="13">Cymonomus japonicus</emphasis>
|
|||
|
|
<bibRefCitation id="EFC341A05F13FFE8FE4BF8C6FDF0F8EF" author="Balss H." box="[486,606,1842,1862]" pageId="12" pageNumber="13" pagination="104 - 140" refId="ref12958" refString="Balss H. 1922. Ostasiatische Decapoden. III. Die Dromiaceen, Oxystomen und Parthenopiden. Archiv fur Naturgeschichte 88 (A): 104 - 140, figs 1 - 9." type="journal article" year="1922">Balss, 1922</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, Japan, WPMNH #47. A-G: female neotype, cl 5.9 mm, pcl 4.6 mm, cw 5.0 mm; H-J: male, cl 4.8 mm, pcl 3.7 mm, cw 3.8 mm; K: male, cl 4.0 mm, pcl 3.9 mm, cw 3.8 mm; L, male, cl 5.0 mm, pcl 3.9 mm, cw 3.8 mm. A, dorsal habitus; B, K, L, fronto-orbital region; C, right basal antennal spine; D, carapace tubercles of right anterolateral corner; E, right maxilliped 3; F, thoracic sternite 3; G, abdomen, posterior; H, telson; I, right gonopod 1, abdominal view; J, right gonopod 2, abdominal view. Scale bars: A, G = 2.0 mm; B, E, F, H-L = 1.0 mm; C, D = 0.5 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BED3C515F12FFE9FF68FE29FEDBFDDF" blockId="13.[136,738,253,1974]" pageId="13" pageNumber="14">Antennular peduncle 0.97-1.02 pcl (male), 0.79 pcl (female); articles 1 and 2 minutely spinular; article 3 minutely granular. Basal antennal article fused to epistome; articles 2-5 irregularly granular or spinular.</paragraph>
|
|||
|
|
<paragraph id="8BED3C515F12FFE9FF68FD89FD2EFC1F" blockId="13.[136,738,253,1974]" pageId="13" pageNumber="14">Maxilliped 3 ischiobasis subquadrate, sparsely granular and spinular, with longitudinal sublateral groove; ischium and basis demarcated by faint groove. Merus slightly shorter than ischiobasis, length about twice width, tapering distally to rounded apex; surface and margins spinulate. Dactylus conical, unarmed; propodus and carpus sparsely spinulate. Exopod sparsely granular, reaching beyond carpo-meral articulation but not reaching end of merus of endopod.</paragraph>
|
|||
|
|
<paragraph id="8BED3C515F12FFE9FF68FC49FD4CFB5F" blockId="13.[136,738,253,1974]" pageId="13" pageNumber="14">Chelipeds (pereopod 1) equal in size and ornamentation, setose. Merus finely granular proximally, spinose distally. Carpus granular, dorsal margin with 3 spines. Palm surfaces granular, flexor and extensor margins with tubular and club-shaped tubercles. Dactylus slightly longer than upper palm length; proximal dorsal half with club-shaped tubercles; with faint longitudinal carina on outer surface, occlusal surfaces of dactylus and pollex crenulate, without gape when fingers closed.</paragraph>
|
|||
|
|
<paragraph id="8BED3C515F12FFE9FF68FB09FD2FF97F" blockId="13.[136,738,253,1974]" pageId="13" pageNumber="14">Pereopods 2 and 3 sparsely setose; all articles except for dactylus finely granular; propodus and carpus with blunt spinular extensor margins; merus with blunt spinular extensor and flexor margins. Pereopod 3 longest, merus 1.15- 1.28 pcl (male), 1.12 pcl (female). Dactyli broadly curved, smooth, with longitudinal rib. Pereopod 3 dactylus longer than combined length of propodus and carpus. Pereopods 4 and 5 coarsely granular, some small spines, sparsely setose; shorter than merus of pereopod 3; dactyli markedly shorter than propodi, falcate, with corneous apex and 2 or 3 obliquely inclined, corneous spines on flexor margin. Pereopod 5 merus, when folded against carapace, reaching midlength of carapace.</paragraph>
|
|||
|
|
<paragraph id="8BED3C515F12FFE9FF68F929FEE4F8FF" blockId="13.[136,738,253,1974]" pageId="13" pageNumber="14">Thoracic sternite 3 pentagonal, about 1.5 × wider than long; lateral margins subparallel, irregular, surface granulate. Margins of sternites 4 and 5 granulate.</paragraph>
|
|||
|
|
<paragraph id="8BED3C515F12FFE9FF68F8A9FAEEFEBF" blockId="13.[136,738,253,1974]" lastBlockId="13.[813,1415,253,1974]" pageId="13" pageNumber="14">Abdomen of both sexes with margins and surface finely granular or minutely spinulate; pleotelson or both sexes distally obtuse, bluntly rounded, length slightly exceeding half width.</paragraph>
|
|||
|
|
<paragraph id="8BED3C515F12FFE9FCC4FEE9FB26FE3F" blockId="13.[813,1415,253,1974]" pageId="13" pageNumber="14">Gonopod 1 distal article cannulate, forming copulatory tube, with moderately long distal setae. Gonopod 2 with articles fused; distomesial margin slightly hollowed, apex acute.</paragraph>
|
|||
|
|
<paragraph id="8BED3C515F12FFE9FCC4FE69FB8BF93F" blockId="13.[813,1415,253,1974]" pageId="13" pageNumber="14">
|
|||
|
|
<emphasis id="B926E0435F12FFE9FCC4FE69FC4AFE1F" box="[873,996,413,438]" italics="true" pageId="13" pageNumber="14">Remarks</emphasis>
|
|||
|
|
:
|
|||
|
|
<taxonomicName id="4C5247D25F12FFE9FC53FE69FAEFFE1F" authorityName="Balss" authorityYear="1922" box="[1022,1345,413,438]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="japonicus">
|
|||
|
|
<emphasis id="B926E0435F12FFE9FC53FE69FAEFFE1F" box="[1022,1345,413,438]" italics="true" pageId="13" pageNumber="14">Cymonomus japonicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
was originally described as a subspecies of
|
|||
|
|
<taxonomicName id="4C5247D25F12FFE9FAC4FE49FC16FE5F" baseAuthorityName="Norman in Thomson" baseAuthorityYear="1873" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="granulatus">
|
|||
|
|
<emphasis id="B926E0435F12FFE9FAC4FE49FC16FE5F" italics="true" pageId="13" pageNumber="14">C. granulatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
based on a single male collected from Haidashi Bank, Sagami Bay (
|
|||
|
|
<quantity id="4CAA91B45F12FFE9FB58FE09FAE1FDBF" box="[1269,1359,509,534]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="14" unit="m" value="600.0">600 m</quantity>
|
|||
|
|
). To date,
|
|||
|
|
<taxonomicName id="4C5247D25F12FFE9FCD8FDE9FBA1FD9F" authorityName="Balss" authorityYear="1922" box="[885,1039,541,566]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="japonicus">
|
|||
|
|
<emphasis id="B926E0435F12FFE9FCD8FDE9FBA1FD9F" box="[885,1039,541,566]" italics="true" pageId="13" pageNumber="14">C. japonicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has been reported only twice since it was described in 1922:
|
|||
|
|
<bibRefCitation id="EFC341A05F12FFE9FB60FDC9FA2AFDFF" author="Takeda M." box="[1229,1412,573,598]" pageId="13" pageNumber="14" pagination="217 - 262" refId="ref13750" refString="Takeda M. 2001. Annotated list of crabs from Tosa Bay, southwest Japan, collected by the R / V Kotaka Maru during the years 1997 - 2000. In: Fujita, T., Saito, H., & Takeda, M. (eds.), Deep-Sea Fauna and Pollutants in Tosa Bay. National Science Museum Monographs 20: 217 - 262." type="journal article" year="2001">Takeda (2001)</bibRefCitation>
|
|||
|
|
from Tosa Bay (
|
|||
|
|
<quantity id="4CAA91B45F12FFE9FC54FDA9FB22FDDF" box="[1017,1164,605,630]" metricMagnitude="2" metricUnit="m" metricValue="5.325" metricValueMax="5.37" metricValueMin="5.279999999999999" pageId="13" pageNumber="14" unit="m" value="532.5" valueMax="537.0" valueMin="528.0">528-537 m</quantity>
|
|||
|
|
), and
|
|||
|
|
<bibRefCitation id="EFC341A05F12FFE9FB73FDA9FA2BFDDF" author="Nagai S." box="[1246,1413,605,630]" pageId="13" pageNumber="14" pagination="31 - 34" refId="ref13497" refString="Nagai S. 1991. Some remarkable crabs of Wakayama Prefecture I. Nankiseibutu 33: 31 - 34." type="journal article" year="1991">Nagai (1991)</bibRefCitation>
|
|||
|
|
from off Shionomisaki (
|
|||
|
|
<quantity id="4CAA91B45F12FFE9FBC4FD89FB69FD3F" box="[1129,1223,637,662]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="m" value="500.0">500 m</quantity>
|
|||
|
|
).
|
|||
|
|
<taxonomicName id="4C5247D25F12FFE9FB4FFD89FC0BFD1F" authorityName="Balss" authorityYear="1922" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="japonicus">
|
|||
|
|
<emphasis id="B926E0435F12FFE9FB4FFD89FC0BFD1F" italics="true" pageId="13" pageNumber="14">Cymonomus japonicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is the only known Japanese species with a ‘long’ rostrum, and on this basis (together with agreement with the brief type description and figure), the present specimens are identified with Balss’ species. Other similar ‘long’ rostrum species, however, occur in the western Pacific, such as
|
|||
|
|
<taxonomicName id="4C5247D25F12FFE9FC33FCA9FB1AFCDF" authority="Ihle, 1916" authorityName="Ihle" authorityYear="1916" box="[926,1204,861,886]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="indicus">
|
|||
|
|
<emphasis id="B926E0435F12FFE9FC33FCA9FB8EFCDF" box="[926,1056,861,886]" italics="true" pageId="13" pageNumber="14">C. indicus</emphasis>
|
|||
|
|
<bibRefCitation id="EFC341A05F12FFE9FB80FCA9FB1AFCDF" author="Ihle JEW" box="[1069,1204,861,886]" pageId="13" pageNumber="14" pagination="97 - 158" refId="ref13221" refString="Ihle JEW. 1916. Die Decapoda Brachyura der Siboga- Expedition. II. Oxystomata, Dorippidae. Siboga-Expeditie Monograph 39 B 1: 97 - 158." type="journal article" year="1916">Ihle, 1916</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, from
|
|||
|
|
<collectingCountry id="F3457CC15F12FFE9FAA4FCA9FA2BFCDF" box="[1289,1413,861,886]" name="Indonesia" pageId="13" pageNumber="14">Indonesia</collectingCountry>
|
|||
|
|
and a possibly undescribed species from the
|
|||
|
|
<collectingCountry id="F3457CC15F12FFE9FC80FC69FBA8FC1F" box="[813,1030,925,951]" name="Solomon Islands" pageId="13" pageNumber="14">Solomon Islands</collectingCountry>
|
|||
|
|
(Ahyong, unpublished data); although distinct from the present Japanese material, the Indonesian and the
|
|||
|
|
<collectingCountry id="F3457CC15F12FFE9FB13FC29FA28FC5F" box="[1214,1414,989,1015]" name="Solomon Islands" pageId="13" pageNumber="14">Solomon Islands</collectingCountry>
|
|||
|
|
specimens also fit Balss’ account of
|
|||
|
|
<taxonomicName id="4C5247D25F12FFE9FB45FC09FAD0FBBF" authorityName="Balss" authorityYear="1922" box="[1256,1406,1021,1046]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="japonicus">
|
|||
|
|
<emphasis id="B926E0435F12FFE9FB45FC09FAD0FBBF" box="[1256,1406,1021,1046]" italics="true" pageId="13" pageNumber="14">C. japonicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Extensive searches over the past 15 years of museum collections in which Balss’ material may be housed (Naturalis, Leiden; the Natural History Museum, London; Muséum national d’Histoire Naturelle, Paris; Zoologisches Museum an der Humboldt-Universität, Berlin; Natur-Museum und Forschungsinstitut Senckenberg, Frankfurt am Main; Zoologisches Institut and Zoologisches Museum, Hamburg; Zoologische Staatssammlung, Munich) failed to locate the
|
|||
|
|
<typeStatus id="54E982F35F12FFE9FB12FAC9FA9DFAFF" box="[1215,1331,1341,1366]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C5247D25F12FFE9FAC4FAC9FC0AFADF" authorityName="Balss" authorityYear="1922" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="japonicus">
|
|||
|
|
<emphasis id="B926E0435F12FFE9FAC4FAC9FC0AFADF" italics="true" pageId="13" pageNumber="14">C. japonicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; it must therefore be considered lost. To fix the identity of the species and allow us to clarify the taxonomy of allied taxa, we designate the largest specimen in the present series as the
|
|||
|
|
<typeStatus id="54E982F35F12FFE9FC80FA29FC21FA5F" box="[813,911,1501,1526]" pageId="13" pageNumber="14" type="neotype">neotype</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C5247D25F12FFE9FC11FA29FBF9FA5F" authorityName="Balss" authorityYear="1922" box="[956,1111,1501,1526]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="japonicus">
|
|||
|
|
<emphasis id="B926E0435F12FFE9FC11FA29FBF9FA5F" box="[956,1111,1501,1526]" italics="true" pageId="13" pageNumber="14">C. japonicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(female, cl
|
|||
|
|
<quantity id="4CAA91B45F12FFE9FB5CFA29FAFAFA5F" box="[1265,1364,1501,1526]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="13" pageNumber="14" unit="mm" value="5.9">5.9 mm</quantity>
|
|||
|
|
, pcl
|
|||
|
|
<quantity id="4CAA91B45F12FFE9FC80FA09FC23F9BF" box="[813,909,1533,1558]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="13" pageNumber="14" unit="mm" value="4.6">4.6 mm</quantity>
|
|||
|
|
, cw 5.0 mm;
|
|||
|
|
<collectionCode id="ED43A4945F12FFE9FB9EFA09FB35F9BF" box="[1075,1179,1533,1558]" pageId="13" pageNumber="14">WPMNH</collectionCode>
|
|||
|
|
#47). Although not from the original type locality (Sagami Bay), the
|
|||
|
|
<typeStatus id="54E982F35F12FFE9FC80F9C9FC20F9FF" box="[813,910,1597,1622]" pageId="13" pageNumber="14" type="neotype">neotype</typeStatus>
|
|||
|
|
locality (Kii Peninsula) is geographically close to Sagami Bay and well within the expected range of the species.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BED3C515F12FFE9FCC4F969FB86F83F" blockId="13.[813,1415,253,1974]" pageId="13" pageNumber="14">Aside from normal sexual dimorphism, the most notable variation in the present series is the proportional length of the eyestalks (proportionally shortest in the largest specimens). The eyestalks anteriorly exceed the outer orbital processes by half the eyestalk length in the smallest individual to only slightly in the largest individual; the specimens otherwise agree well.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3486FDA5F12FFEAFCC4F869FDB1FBDF" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="reference_group">
|
|||
|
|
<paragraph id="8BED3C515F12FFEAFCC4F869FDB1FBDF" blockId="13.[813,1415,253,1974]" lastBlockId="14.[174,775,253,1238]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
|||
|
|
<taxonomicName id="4C5247D25F12FFE9FCC4F869FB06F81F" ID-CoL="337VH" authorityName="Balss" authorityYear="1922" box="[873,1192,1949,1974]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="japonicus">
|
|||
|
|
<emphasis id="B926E0435F12FFE9FCC4F869FB06F81F" box="[873,1192,1949,1974]" italics="true" pageId="13" pageNumber="14">Cymonomus japonicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
belongs to the
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFF03FF09FECEFEBF" ID-CoL="337VC" baseAuthorityName="Norman in Thomson" baseAuthorityYear="1873" box="[174,352,253,278]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="granulatus">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFF03FF09FECEFEBF" box="[174,352,253,278]" italics="true" pageId="14" pageNumber="15">C. granulatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
group (
|
|||
|
|
<bibRefCitation id="EFC341A05F11FFEAFE66FF09FDE0FEBF" author="Dell RK" box="[459,590,253,278]" pageId="14" pageNumber="15" pagination="55 - 64" refId="ref13107" refString="Dell RK. 1971. Two new species of crabs of the genus Cymonomus from New Zealand (Crustacea: Brachyura). Records of the Dominion Museum 7: 55 - 64." type="journal article" year="1971">Dell 1971</bibRefCitation>
|
|||
|
|
) in having an elongate rostrum that distinctly overreaches the eyes, and prominently developed, spinose outer orbital processes. Other species in the
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFF03FE89FEF9FE3F" ID-CoL="337VC" baseAuthorityName="Norman in Thomson" baseAuthorityYear="1873" box="[174,343,381,406]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="granulatus">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFF03FE89FEF9FE3F" box="[174,343,381,406]" italics="true" pageId="14" pageNumber="15">C. granulatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
group are as
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFDA2FE89FF42FE1F" ID-CoL="6C6YM" authority="Dell, 1971" authorityName="Dell" authorityYear="1971" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aequilonius">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFDA2FE89FD6AFE3F" box="[527,708,381,406]" italics="true" pageId="14" pageNumber="15">C. aequilonius</emphasis>
|
|||
|
|
<bibRefCitation id="EFC341A05F11FFEAFD62FE89FF42FE1F" author="Dell RK" pageId="14" pageNumber="15" pagination="55 - 64" refId="ref13107" refString="Dell RK. 1971. Two new species of crabs of the genus Cymonomus from New Zealand (Crustacea: Brachyura). Records of the Dominion Museum 7: 55 - 64." type="journal article" year="1971">Dell, 1971</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<collectingCountry id="F3457CC15F11FFEAFF50FE69FE01FE1F" box="[253,431,413,438]" name="New Zealand" pageId="14" pageNumber="15">New Zealand</collectingCountry>
|
|||
|
|
),
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFE69FE69FDC0FE1F" ID-CoL="337VC" baseAuthorityName="Norman in Thomson" baseAuthorityYear="1873" box="[452,622,413,438]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="granulatus">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFE69FE69FDC0FE1F" box="[452,622,413,438]" italics="true" pageId="14" pageNumber="15">C. granulatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(Norman in
|
|||
|
|
<bibRefCitation id="EFC341A05F11FFEAFF03FE49FE76FE7F" author="Wyville Thomson C." box="[174,472,445,470]" pageId="14" pageNumber="15" refId="ref14051" refString="Wyville Thomson C. 1873. The depths of the sea. London, 176 pp." type="book" year="1873">Wyville Thomson, 1873</bibRefCitation>
|
|||
|
|
) (northeast Atlantic),
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFD46FE49FED1FE5F" ID-CoL="337VG" authority="Ihle, 1916" authorityName="Ihle" authorityYear="1916" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="indicus">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFD46FE49FEAFFE5F" italics="true" pageId="14" pageNumber="15">C. indicus</emphasis>
|
|||
|
|
<bibRefCitation id="EFC341A05F11FFEAFEA4FE29FED1FE5F" author="Ihle JEW" box="[265,383,477,502]" pageId="14" pageNumber="15" pagination="97 - 158" refId="ref13221" refString="Ihle JEW. 1916. Die Decapoda Brachyura der Siboga- Expedition. II. Oxystomata, Dorippidae. Siboga-Expeditie Monograph 39 B 1: 97 - 158." type="journal article" year="1916">Ihle, 1916</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<collectingCountry id="F3457CC15F11FFEAFE22FE29FDA5FE5F" box="[399,523,477,502]" name="Indonesia" pageId="14" pageNumber="15">Indonesia</collectingCountry>
|
|||
|
|
), and
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFDFFFE29FEC8FDBF" ID-CoL="337VM" authority="Tavares, 1991" authorityName="Tavares" authorityYear="1991" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="magnirostris">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFDFFFE29FCA9FE5F" box="[594,775,477,502]" italics="true" pageId="14" pageNumber="15">C. magnirostris</emphasis>
|
|||
|
|
<bibRefCitation id="EFC341A05F11FFEAFF03FE09FEC8FDBF" author="Tavares M." box="[174,358,509,534]" pageId="14" pageNumber="15" pagination="3 - 4" refId="ref13898" refString="Tavares M. 1991. Especes nouvelles de Cyclodorippoidea Ortmann et remarques sur les genres Tymolus Stimpson et Cyclodorippe A. Milne Edwards (Crustacea, Decapoda, Brachyura). Bulletin du Museum national d'Histoire naturelle 4 A, 12 (3 - 4): 623 - 648 (dated 1990, published 1991)." type="journal article" year="1991">Tavares, 1991</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<collectingCountry id="F3457CC15F11FFEAFED4FE09FE65FDBF" box="[377,459,509,534]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
|
|||
|
|
).
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFE4FFE09FCA9FDBF" ID-CoL="337VH" authorityName="Balss" authorityYear="1922" box="[482,775,509,535]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="japonicus">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFE4FFE09FCA9FDBF" box="[482,775,509,535]" italics="true" pageId="14" pageNumber="15">Cymonomus japonicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is readily separated from
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFDA2FDE9FD72FD9F" ID-CoL="337VM" authorityName="Tavares" authorityYear="1991" box="[527,732,541,566]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="magnirostris">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFDA2FDE9FD72FD9F" box="[527,732,541,566]" italics="true" pageId="14" pageNumber="15">C. magnirostris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the shorter maxilliped 3 exopod (
|
|||
|
|
<figureCitation id="136920D45F11FFEAFDCDFDC9FD6CFDFF" box="[608,706,573,598]" captionStart="Fig" captionStartId="12.[174,209,1842,1862]" captionTargetBox="[196,1413,251,1817]" captionTargetId="figure-1@12.[192,1433,238,1830]" captionTargetPageId="12" captionText="Fig. 5. Cymonomus japonicus Balss, 1922, Japan, WPMNH #47. A-G: female neotype, cl 5.9 mm, pcl 4.6 mm, cw 5.0 mm; H-J: male, cl 4.8 mm, pcl 3.7 mm, cw 3.8 mm; K: male, cl 4.0 mm, pcl 3.9 mm, cw 3.8 mm; L, male, cl 5.0 mm, pcl 3.9 mm, cw 3.8 mm. A, dorsal habitus; B, K, L, fronto-orbital region; C, right basal antennal spine; D, carapace tubercles of right anterolateral corner; E, right maxilliped 3; F, thoracic sternite 3; G, abdomen, posterior; H, telson; I, right gonopod 1, abdominal view; J, right gonopod 2, abdominal view. Scale bars: A, G = 2.0 mm; B, E, F, H-L = 1.0 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12824870" httpUri="https://zenodo.org/record/12824870/files/figure.png" pageId="14" pageNumber="15">Fig. 5E</figureCitation>
|
|||
|
|
) (not overreaching the distal end of the merus versus distinctly overreaching the merus;
|
|||
|
|
<bibRefCitation id="EFC341A05F11FFEAFDF4FD89FCADFD3F" author="Tavares M." box="[601,771,637,662]" pageId="14" pageNumber="15" pagination="3 - 4" refId="ref13898" refString="Tavares M. 1991. Especes nouvelles de Cyclodorippoidea Ortmann et remarques sur les genres Tymolus Stimpson et Cyclodorippe A. Milne Edwards (Crustacea, Decapoda, Brachyura). Bulletin du Museum national d'Histoire naturelle 4 A, 12 (3 - 4): 623 - 648 (dated 1990, published 1991)." type="journal article" year="1991">Tavares 1991</bibRefCitation>
|
|||
|
|
: fig. 8E) and shorter pereopod 5 (
|
|||
|
|
<figureCitation id="136920D45F11FFEAFDE8FD69FD0FFD1F" box="[581,673,669,694]" captionStart="Fig" captionStartId="12.[174,209,1842,1862]" captionTargetBox="[196,1413,251,1817]" captionTargetId="figure-1@12.[192,1433,238,1830]" captionTargetPageId="12" captionText="Fig. 5. Cymonomus japonicus Balss, 1922, Japan, WPMNH #47. A-G: female neotype, cl 5.9 mm, pcl 4.6 mm, cw 5.0 mm; H-J: male, cl 4.8 mm, pcl 3.7 mm, cw 3.8 mm; K: male, cl 4.0 mm, pcl 3.9 mm, cw 3.8 mm; L, male, cl 5.0 mm, pcl 3.9 mm, cw 3.8 mm. A, dorsal habitus; B, K, L, fronto-orbital region; C, right basal antennal spine; D, carapace tubercles of right anterolateral corner; E, right maxilliped 3; F, thoracic sternite 3; G, abdomen, posterior; H, telson; I, right gonopod 1, abdominal view; J, right gonopod 2, abdominal view. Scale bars: A, G = 2.0 mm; B, E, F, H-L = 1.0 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12824870" httpUri="https://zenodo.org/record/12824870/files/figure.png" pageId="14" pageNumber="15">Fig. 5A</figureCitation>
|
|||
|
|
) (merus reaching to midlength of carapace instead of anterior one-fourth;
|
|||
|
|
<bibRefCitation id="EFC341A05F11FFEAFE08FD29FDFFFD5F" author="Tavares M." box="[421,593,733,758]" pageId="14" pageNumber="15" pagination="3 - 4" refId="ref13898" refString="Tavares M. 1991. Especes nouvelles de Cyclodorippoidea Ortmann et remarques sur les genres Tymolus Stimpson et Cyclodorippe A. Milne Edwards (Crustacea, Decapoda, Brachyura). Bulletin du Museum national d'Histoire naturelle 4 A, 12 (3 - 4): 623 - 648 (dated 1990, published 1991)." type="journal article" year="1991">Tavares 1991</bibRefCitation>
|
|||
|
|
: fig. 10F); and from
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFF46FD09FE24FCBF" ID-CoL="337VC" baseAuthorityName="Norman in Thomson" baseAuthorityYear="1873" box="[235,394,765,790]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="granulatus">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFF46FD09FE24FCBF" box="[235,394,765,790]" italics="true" pageId="14" pageNumber="15">C. granulatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the slender versus triangular rostrum with basal width of about half the length (
|
|||
|
|
<figureCitation id="136920D45F11FFEAFF18FCC9FEC1FCFF" box="[181,367,829,854]" captionStart="Fig" captionStartId="12.[174,209,1842,1862]" captionTargetBox="[196,1413,251,1817]" captionTargetId="figure-1@12.[192,1433,238,1830]" captionTargetPageId="12" captionText="Fig. 5. Cymonomus japonicus Balss, 1922, Japan, WPMNH #47. A-G: female neotype, cl 5.9 mm, pcl 4.6 mm, cw 5.0 mm; H-J: male, cl 4.8 mm, pcl 3.7 mm, cw 3.8 mm; K: male, cl 4.0 mm, pcl 3.9 mm, cw 3.8 mm; L, male, cl 5.0 mm, pcl 3.9 mm, cw 3.8 mm. A, dorsal habitus; B, K, L, fronto-orbital region; C, right basal antennal spine; D, carapace tubercles of right anterolateral corner; E, right maxilliped 3; F, thoracic sternite 3; G, abdomen, posterior; H, telson; I, right gonopod 1, abdominal view; J, right gonopod 2, abdominal view. Scale bars: A, G = 2.0 mm; B, E, F, H-L = 1.0 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12824870" httpUri="https://zenodo.org/record/12824870/files/figure.png" pageId="14" pageNumber="15">Fig. 5A, B, K, L</figureCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFC341A05F11FFEAFED6FCC9FDC2FCFF" author="Mura M & Cau A." box="[379,620,829,855]" pageId="14" pageNumber="15" pagination="1133 - 1139" refId="ref13438" refString="Mura M, Cau A. 2003. Occurrence of a rare deep-sea crab, Cymonomus granulatus (Norman, 1873) (Decapoda, Brachyura), in the Sardinian channel. Crustaceana 75: 1133 - 1139." type="journal article" year="2003">Mura and Cau 2003</bibRefCitation>
|
|||
|
|
: fig. 2). From
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFF03FCA9FE82FCDF" ID-CoL="337VG" authorityName="Ihle" authorityYear="1916" box="[174,300,861,886]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="indicus">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFF03FCA9FE82FCDF" box="[174,300,861,886]" italics="true" pageId="14" pageNumber="15">C. indicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFEDCFCA9FD88FCDF" ID-CoL="6C6YM" authorityName="Dell" authorityYear="1971" box="[369,550,861,886]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aequilonius">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFEDCFCA9FD88FCDF" box="[369,550,861,886]" italics="true" pageId="14" pageNumber="15">C. aequilonius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFD97FCA9FD61FCDF" ID-CoL="337VH" authorityName="Balss" authorityYear="1922" box="[570,719,861,886]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="japonicus">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFD97FCA9FD61FCDF" box="[570,719,861,886]" italics="true" pageId="14" pageNumber="15">C japonicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
can be separated by the differences in the shape of the dorsal carapace tubercles (rounded versus polygonal or stellate in
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFE78FC49FDFFFC7F" ID-CoL="337VG" authorityName="Ihle" authorityYear="1916" box="[469,593,957,982]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="indicus">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFE78FC49FDFFFC7F" box="[469,593,957,982]" italics="true" pageId="14" pageNumber="15">C. indicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
).
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFDC0FC49FE88FC5F" ID-CoL="337VH" authorityName="Balss" authorityYear="1922" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="japonicus">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFDC0FC49FE88FC5F" italics="true" pageId="14" pageNumber="15">Cymonomus japonicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="4C5247D25F11FFEAFE62FC29FD28FC5F" ID-CoL="6C6YM" authorityName="Dell" authorityYear="1971" box="[463,646,989,1014]" class="Malacostraca" family="Cymonomidae" genus="Cymonomus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aequilonius">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFE62FC29FD28FC5F" box="[463,646,989,1014]" italics="true" pageId="14" pageNumber="15">C. aequilonius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in having more pronounced, coarser surface granulation and in the shorter pereopod 4, reaching to the carapace midlength rather than anterior one-fourth (
|
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|
|
<figureCitation id="136920D45F11FFEAFF18FBA9FEA1FBDF" box="[181,271,1117,1142]" captionStart="Fig" captionStartId="12.[174,209,1842,1862]" captionTargetBox="[196,1413,251,1817]" captionTargetId="figure-1@12.[192,1433,238,1830]" captionTargetPageId="12" captionText="Fig. 5. Cymonomus japonicus Balss, 1922, Japan, WPMNH #47. A-G: female neotype, cl 5.9 mm, pcl 4.6 mm, cw 5.0 mm; H-J: male, cl 4.8 mm, pcl 3.7 mm, cw 3.8 mm; K: male, cl 4.0 mm, pcl 3.9 mm, cw 3.8 mm; L, male, cl 5.0 mm, pcl 3.9 mm, cw 3.8 mm. A, dorsal habitus; B, K, L, fronto-orbital region; C, right basal antennal spine; D, carapace tubercles of right anterolateral corner; E, right maxilliped 3; F, thoracic sternite 3; G, abdomen, posterior; H, telson; I, right gonopod 1, abdominal view; J, right gonopod 2, abdominal view. Scale bars: A, G = 2.0 mm; B, E, F, H-L = 1.0 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12824870" httpUri="https://zenodo.org/record/12824870/files/figure.png" pageId="14" pageNumber="15">Fig. 5A</figureCitation>
|
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|
|
;
|
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|
|
<bibRefCitation id="EFC341A05F11FFEAFEB4FBA9FE14FBDF" author="Ahyong ST" box="[281,442,1117,1142]" pageId="14" pageNumber="15" pagination="1 - 72" refId="ref12494" refString="Ahyong ST. 2008. Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura). Zootaxa 1708: 1 - 72." type="journal article" year="2008">Ahyong 2008</bibRefCitation>
|
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|
: fig. 1F).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3486FDA5F11FFEAFF47FB89FD66FB7F" pageId="14" pageNumber="15" type="distribution">
|
|||
|
|
<paragraph id="8BED3C515F11FFEAFF47FB89FD66FB7F" blockId="14.[174,775,253,1238]" pageId="14" pageNumber="15">
|
|||
|
|
<emphasis id="B926E0435F11FFEAFF47FB89FEDBFB3F" box="[234,373,1149,1174]" italics="true" pageId="14" pageNumber="15">Distribution</emphasis>
|
|||
|
|
: Known only from southeastern
|
|||
|
|
<collectingCountry id="F3457CC15F11FFEAFF03FB69FF55FB1F" box="[174,251,1181,1206]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
|
|||
|
|
: Sagami Bay (
|
|||
|
|
<quantity id="4CAA91B45F11FFEAFE0FFB69FE5CFB1F" box="[418,498,1181,1206]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="15" unit="m" value="600.0">600 m</quantity>
|
|||
|
|
), Shionomisaki (
|
|||
|
|
<quantity id="4CAA91B45F11FFEAFD1CFB69FCAEFB1F" box="[689,768,1181,1206]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="15" unit="m" value="500.0">500 m</quantity>
|
|||
|
|
) and Tosa Bay (
|
|||
|
|
<quantity id="4CAA91B45F11FFEAFEF0FB49FE4DFB7F" box="[349,483,1213,1238]" metricMagnitude="2" metricUnit="m" metricValue="5.325" metricValueMax="5.37" metricValueMin="5.279999999999999" pageId="14" pageNumber="15" unit="m" value="532.5" valueMax="537.0" valueMin="528.0">528-537 m</quantity>
|
|||
|
|
);
|
|||
|
|
<quantity id="4CAA91B45F11FFEAFE5BFB49FDD8FB7F" box="[502,630,1213,1238]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="14" pageNumber="15" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">500-600 m</quantity>
|
|||
|
|
depth.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|