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<document id="C755E7154C056EE3FBAD75C33C427D3E" ID-CLB-Dataset="56315" ID-DOI="10.11646/zootaxa.5244.3.4" ID-GBIF-Dataset="10c70395-38c4-4847-b4bf-86522cbd3baf" ID-ISSN="1175-5326" ID-Zenodo-Dep="7656363" ID-ZooBank="4212B66F-DE64-4425-A4DE-674A36DDDCC8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676883040364" checkinUser="plazi" docAuthor="Jaschhof, Mathias &amp; Jaschhof, Catrin" docDate="2023" docId="03F687AEEC61390619A5FEE9FDB1F91E" docLanguage="en" docName="zootaxa.5244.3.4.pdf" docOrigin="Zootaxa 5244 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5244.3.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Asycampta Mamaev &amp; Zaitzev 2023" docType="treatment" docVersion="7" lastPageNumber="265" masterDocId="FFCFFFD6EC6239021932FFACFE2CFFC5" masterDocTitle="A review of Afrotropical Porricondylinae (Diptera: Cecidomyiidae), with descriptions of five new species of Asynaptini from KwaZulu-Natal Province South Africa" masterLastPageNumber="275" masterPageNumber="261" pageNumber="264" updateTime="1699475464623" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D9F4DC637244FC27BC34B02D5082C4B8">A review of Afrotropical Porricondylinae (Diptera: Cecidomyiidae), with descriptions of five new species of Asynaptini from KwaZulu-Natal Province South Africa</mods:title>
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<mods:namePart id="EDB37FED085704AB611E11332438ECB6">Jaschhof, Mathias</mods:namePart>
<mods:nameIdentifier id="C055F306C59772A41DAC01944543867B" type="ORCID">0000-0003-3447-1620</mods:nameIdentifier>
<mods:affiliation id="430CDB8DFC3E435DA90A333731DDB263">Station Linné, Ölands Skogsby 161, SE- 38693 Färjestaden, Sweden. &amp; mjaschhof @ yahoo. de; https: // orcid. org / 0000 - 0003 - 3447 - 1620</mods:affiliation>
<mods:nameIdentifier id="48DDD3C8DBEE41E7CCAA2211F3B58A8A" type="email">mjaschhof@yahoo.de</mods:nameIdentifier>
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<mods:namePart id="9972F330C5F5008EA14E07924C6ED818">Jaschhof, Catrin</mods:namePart>
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<mods:affiliation id="71BA37FCAD807CFC4A38CEA292D73012">Station Linné, Ölands Skogsby 161, SE- 38693 Färjestaden, Sweden. &amp; https: // orcid. org / 0000 - 0002 - 1030 - 0934</mods:affiliation>
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<mods:title id="D5E621ED444836AC65098128AAE1381B">Zootaxa</mods:title>
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<mods:date id="B9128EEA49BB28F38524F33AFC595693">2023</mods:date>
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<mods:number id="09074C47C5B912693D9291514EBE6166">2023-02-20</mods:number>
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<treatment id="03F687AEEC61390619A5FEE9FDB1F91E" ID-DOI="http://doi.org/10.5281/zenodo.7674090" ID-GBIF-Taxon="209343093" ID-Zenodo-Dep="7674090" LSID="urn:lsid:plazi:treatment:03F687AEEC61390619A5FEE9FDB1F91E" httpUri="http://treatment.plazi.org/id/03F687AEEC61390619A5FEE9FDB1F91E" lastPageId="4" lastPageNumber="265" pageId="3" pageNumber="264">
<subSubSection id="C3456533EC61390119A5FEE9FFE2FE9A" box="[151,462,325,351]" pageId="3" pageNumber="264" type="nomenclature">
<paragraph id="8BE036B8EC61390119A5FEE9FFE2FE9A" blockId="3.[151,1437,325,1756]" box="[151,462,325,351]" pageId="3" pageNumber="264">
<heading id="D0A881D4EC61390119A5FEE9FFE2FE9A" bold="true" box="[151,462,325,351]" fontSize="11" level="1" pageId="3" pageNumber="264" reason="1">
<emphasis id="B92BEAAAEC61390119A5FEE9FFE2FE9A" bold="true" box="[151,462,325,351]" pageId="3" pageNumber="264">
Redefinition of
<taxonomicName id="4C5F4D3BEC6139011862FEE9FFE2FE9A" ID-CoL="B944K" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[336,462,325,351]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011862FEE9FFE2FE9A" bold="true" box="[336,462,325,351]" italics="true" pageId="3" pageNumber="264">Asycampta</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3456533EC61390118E7FEE9FDD5FE9A" box="[469,1017,325,352]" pageId="3" pageNumber="264" type="reference_group">
<paragraph id="8BE036B8EC61390118E7FEE9FDD5FE9A" blockId="3.[151,1437,325,1756]" box="[469,1017,325,352]" pageId="3" pageNumber="264">
<heading id="D0A881D4EC61390118E7FEE9FDD5FE9A" bold="true" box="[469,1017,325,352]" fontSize="11" level="1" pageId="3" pageNumber="264" reason="1">
<emphasis id="B92BEAAAEC61390118E7FEE9FDD5FE9A" bold="true" box="[469,1017,325,352]" pageId="3" pageNumber="264">
within the
<taxonomicName id="4C5F4D3BEC6139011B67FEE9FD19FEA5" ID-CoL="6ZF3" authorityName="Parnell" authorityYear="1971" box="[597,821,325,352]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011B67FEE9FD19FEA5" bold="true" box="[597,821,325,352]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
group of genera
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3456533EC61390619A5FEC5FCC0FD34" lastPageId="4" lastPageNumber="265" pageId="3" pageNumber="264" type="description">
<paragraph id="8BE036B8EC61390119A5FEC5FF13FB91" blockId="3.[151,1437,325,1756]" pageId="3" pageNumber="264">
(A) Context. Although not explicitely stated in the publication,
<bibRefCitation id="EFCE4B49EC6139011A5BFEC6FA87FE46" author="Mamaev, B. M. &amp; Zaitzev, A. I." box="[873,1195,361,388]" pageId="3" pageNumber="264" pagination="5 - 13" refId="ref9578" refString="Mamaev, B. M. &amp; Zaitzev, A. I. (1997) New genus and species of free-developing gall midges of the subfamily Porricondylinae from Somalia (Diptera, Cecidomyiinae). Journal of the Ukrainian Entomological Society, 3 (2), 5 - 13." type="journal article" year="1997">Mamaev &amp; Zaitzev’s (1997)</bibRefCitation>
decision to associate two new
<taxonomicName id="4C5F4D3BEC613901181BFE21FF88FE62" baseAuthorityName="Gagne &amp; Jaschhof" baseAuthorityYear="2021" box="[297,420,397,423]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
from Somalian material with the then monotypic, Nearctic genus
<taxonomicName id="4C5F4D3BEC6139011DB3FE21FB70FE62" authorityName="Parnell" authorityYear="1971" box="[1153,1372,397,423]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011DB3FE21FB70FE62" box="[1153,1372,397,423]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
(
<typeStatus id="54E4881AEC6139011C58FE21FBB0FE62" box="[1386,1436,397,423]" pageId="3" pageNumber="264">type</typeStatus>
species:
<taxonomicName id="4C5F4D3BEC6139011818FE1EFC6AFE0E" authority="Felt" authorityName="Felt" authorityYear="1907" box="[298,582,433,460]" class="Insecta" family="Cecidomyiidae" genus="Asynapta" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="phosphila">
<emphasis id="B92BEAAAEC6139011818FE1EFC27FE0E" box="[298,523,433,460]" italics="true" pageId="3" pageNumber="264">Asynapta phosphila</emphasis>
Felt
</taxonomicName>
), was presumably determined by male terminalia characters. While these authors ascribed one species (
<taxonomicName id="4C5F4D3BEC6139011B12FE79FC53FE2A" authorityName="Mamaev &amp; Zaitzev" authorityYear="1997" box="[544,639,469,495]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B92BEAAAEC6139011B12FE79FC53FE2A" box="[544,639,469,495]" italics="true" pageId="3" pageNumber="264">africana</emphasis>
</taxonomicName>
) to
<emphasis id="B92BEAAAEC6139011B80FE79FDF9FE2A" box="[690,981,469,495]" italics="true" pageId="3" pageNumber="264">
<taxonomicName id="4C5F4D3BEC6139011B80FE79FDA1FE2A" authorityName="Parnell" authorityYear="1971" box="[690,909,469,495]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus" sensu="stricto">Pseudocamptomyia</taxonomicName>
<taxonomicNameLabel id="A21857D1EC6139011AA5FE7AFDF9FE2A" box="[919,981,470,495]" pageId="3" pageNumber="264" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
, the other (
<taxonomicName id="4C5F4D3BEC6139011D6CFE79FA98FE2A" authorityName="Mamaev &amp; Zaitzev" authorityYear="1997" box="[1118,1204,469,495]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="palpata">
<emphasis id="B92BEAAAEC6139011D6CFE79FA98FE2A" box="[1118,1204,469,495]" italics="true" pageId="3" pageNumber="264">palpata</emphasis>
</taxonomicName>
) was classified in a new subgenus,
<taxonomicName id="4C5F4D3BEC6139011846FE56FFC3FDD1" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[372,495,506,532]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011846FE56FFC3FDD1" box="[372,495,506,532]" italics="true" pageId="3" pageNumber="264">Asycampta</emphasis>
</taxonomicName>
, distinguished on three characters: the number of palpal segments, the dentition of the tarsal claws, and the outline of vein
<emphasis id="B92BEAAAEC6139011BB8FDB2FC90FDFD" box="[650,700,542,568]" italics="true" pageId="3" pageNumber="264">CuA</emphasis>
.
<taxonomicName id="4C5F4D3BEC6139011BF9FDB2FD6AFDFD" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[715,838,542,568]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011BF9FDB2FD6AFDFD" box="[715,838,542,568]" italics="true" pageId="3" pageNumber="264">Asycampta</emphasis>
</taxonomicName>
was defined as having a foreshortened, 2-segmented palpus, toothless claws, and distinct vein
<emphasis id="B92BEAAAEC6139011BABFDEEFCE7FD99" box="[665,715,578,604]" italics="true" pageId="3" pageNumber="264">CuA</emphasis>
, whereas
<emphasis id="B92BEAAAEC6139011A0EFDEDFA76FD9E" box="[828,1114,577,603]" italics="true" pageId="3" pageNumber="264">
<taxonomicName id="4C5F4D3BEC6139011A0EFDEDFA3BFD9E" authorityName="Parnell" authorityYear="1971" box="[828,1047,577,603]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus" sensu="stricto">Pseudocamptomyia</taxonomicName>
<taxonomicNameLabel id="A21857D1EC6139011D2DFDEEFA76FD9E" box="[1055,1114,578,603]" pageId="3" pageNumber="264" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
was characterized as having a long, 4-segmented palpus, unidentate claws, and indistinct vein
<emphasis id="B92BEAAAEC6139011A90FDCAFDF8FD45" box="[930,980,614,640]" italics="true" pageId="3" pageNumber="264">CuA</emphasis>
. While
<bibRefCitation id="EFCE4B49EC6139011D1EFDCAFB7AFDBA" author="Mamaev, B. M. &amp; Zaitzev, A. I." box="[1068,1366,613,640]" pageId="3" pageNumber="264" pagination="5 - 13" refId="ref9578" refString="Mamaev, B. M. &amp; Zaitzev, A. I. (1997) New genus and species of free-developing gall midges of the subfamily Porricondylinae from Somalia (Diptera, Cecidomyiinae). Journal of the Ukrainian Entomological Society, 3 (2), 5 - 13." type="journal article" year="1997">Mamaev &amp; Zaitzev (1997)</bibRefCitation>
failed to clarify the meaning of “distinct” and “indistinct” with respect to vein
<emphasis id="B92BEAAAEC6139011D3AFD26FA16FD61" box="[1032,1082,650,676]" italics="true" pageId="3" pageNumber="264">CuA</emphasis>
, the other two characters were useful for separating the two subgenera. The same system ceases to work, however, when additional species are taken into consideration. In
<taxonomicName id="4C5F4D3BEC6139011B18FD7EFD6DFD2E" authority="Jaschhof &amp; Jaschhof, 2023" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2023" box="[554,833,721,748]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="karkloofensis" status="sp. nov.">
<emphasis id="B92BEAAAEC6139011B18FD7EFD6DFD2E" box="[554,833,721,748]" italics="true" pageId="3" pageNumber="264">Asycampta karkloofensis</emphasis>
</taxonomicName>
<emphasis id="B92BEAAAEC6139011A7BFD7DFD8FFD2E" bold="true" box="[841,931,721,747]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="A21857D1EC6139011A7BFD7DFD8FFD2E" box="[841,931,721,747]" pageId="3" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, as described below, the 1-segmented palpus (supposedly indicative of
<taxonomicName id="4C5F4D3BEC61390118DFFD5AFC44FCD5" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[493,616,758,784]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC61390118DFFD5AFC44FCD5" box="[493,616,758,784]" italics="true" pageId="3" pageNumber="264">Asycampta</emphasis>
</taxonomicName>
) occurs in combination with unidentate claws (supposedly indicative of
<taxonomicName id="4C5F4D3BEC61390119F4FCB5FF8DFCF6" authorityName="Parnell" authorityYear="1971" box="[198,417,793,819]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC61390119F4FCB5FF8DFCF6" box="[198,417,793,819]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
). Also, vein
<emphasis id="B92BEAAAEC6139011B1FFCB6FC73FCF1" box="[557,607,794,820]" italics="true" pageId="3" pageNumber="264">CuA</emphasis>
of
<taxonomicName id="4C5F4D3BEC6139011BB3FCB6FD18FCF6" authority="Jaschhof &amp; Jaschhof, 2023" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2023" box="[641,820,793,820]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="karkloofensis" status="sp. nov.">
<emphasis id="B92BEAAAEC6139011BB3FCB6FCBEFCF1" box="[641,658,794,820]" italics="true" pageId="3" pageNumber="264">A</emphasis>
.
<emphasis id="B92BEAAAEC6139011BADFCB5FD18FCF6" box="[671,820,793,819]" italics="true" pageId="3" pageNumber="264">karkloofensis</emphasis>
</taxonomicName>
<emphasis id="B92BEAAAEC6139011A08FCB5FDBEFCF6" bold="true" box="[826,914,793,819]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="A21857D1EC6139011A08FCB5FDBEFCF6" box="[826,914,793,819]" pageId="3" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
extends to the wing margin, which matches the condition found in
<taxonomicName id="4C5F4D3BEC61390118AFFC92FC06FC92" authorityName="Felt" authorityYear="1907" box="[413,554,829,856]" class="Insecta" family="Cecidomyiidae" genus="Asynapta" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="phosphila">
<emphasis id="B92BEAAAEC61390118AFFC92FF82FC92" box="[413,430,830,855]" italics="true" pageId="3" pageNumber="264">P</emphasis>
.
<emphasis id="B92BEAAAEC613901188EFC91FC06FC92" box="[444,554,829,855]" italics="true" pageId="3" pageNumber="264">phosphila</emphasis>
</taxonomicName>
, but not the situation in
<taxonomicName id="4C5F4D3BEC6139011A0FFC92FD97FC92" authorityName="Mamaev &amp; Zaitzev" authorityYear="1997" box="[829,955,829,856]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B92BEAAAEC6139011A0FFC92FD62FC92" box="[829,846,830,855]" italics="true" pageId="3" pageNumber="264">P</emphasis>
.
<emphasis id="B92BEAAAEC6139011A6EFC91FD97FC92" box="[860,955,829,855]" italics="true" pageId="3" pageNumber="264">africana</emphasis>
</taxonomicName>
, in which vein
<emphasis id="B92BEAAAEC6139011D5BFC92FAB7FC9D" box="[1129,1179,830,856]" italics="true" pageId="3" pageNumber="264">CuA</emphasis>
is evanescent apically. Likewise, in
<taxonomicName id="4C5F4D3BEC613901186EFCCEFC6CFCBE" authority="Jaschhof &amp; Jaschhof, 2023" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2023" box="[348,576,866,892]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="umngeni" status="sp. nov.">
<emphasis id="B92BEAAAEC613901186EFCCEFC6CFCBE" box="[348,576,866,892]" italics="true" pageId="3" pageNumber="264">Asycampta umngeni</emphasis>
</taxonomicName>
<emphasis id="B92BEAAAEC6139011B78FCCDFC8AFCBE" bold="true" box="[586,678,865,891]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="A21857D1EC6139011B78FCCDFC8AFCBE" box="[586,678,865,891]" pageId="3" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the presence of a 4-segmented palpus and unidentate claws (both supposedly indicative of
<taxonomicName id="4C5F4D3BEC61390118D0FC29FC91FC5A" authorityName="Parnell" authorityYear="1971" box="[482,701,901,927]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC61390118D0FC29FC91FC5A" box="[482,701,901,927]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
) are associated with a vein
<emphasis id="B92BEAAAEC6139011AC9FC2AFA01FC65" box="[1019,1069,902,928]" italics="true" pageId="3" pageNumber="264">CuA</emphasis>
that extends to the wing margin and forms a fork with vein
<emphasis id="B92BEAAAEC61390118C9FC06FC36FC0F" box="[507,538,938,970]" italics="true" pageId="3" pageNumber="264">
M
<subScript id="17DB34FDEC6139011B20FC10FC36FC0F" attach="left" box="[530,538,956,970]" fontSize="6" pageId="3" pageNumber="264">4</subScript>
</emphasis>
(whereas
<emphasis id="B92BEAAAEC6139011BBDFC06FC82FC0F" box="[655,686,938,970]" italics="true" pageId="3" pageNumber="264">
M
<subScript id="17DB34FDEC6139011B94FC10FC82FC0F" attach="left" box="[678,686,956,970]" fontSize="6" pageId="3" pageNumber="264">4</subScript>
</emphasis>
is purportedly absent in both
<taxonomicName id="4C5F4D3BEC6139011ACCFC06FA55FC01" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[1022,1145,938,964]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011ACCFC06FA55FC01" box="[1022,1145,938,964]" italics="true" pageId="3" pageNumber="264">Asycampta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D3BEC6139011D83FC05FBA0FC06" authorityName="Parnell" authorityYear="1971" box="[1201,1420,937,963]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011D83FC05FBA0FC06" box="[1201,1420,937,963]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
). There are basically two different options to resolve the conflict between the subgeneric definitions proposed by
<bibRefCitation id="EFCE4B49EC61390119DBFC5EFC34FBCE" author="Mamaev, B. M. &amp; Zaitzev, A. I." box="[233,536,1009,1036]" pageId="3" pageNumber="264" pagination="5 - 13" refId="ref9578" refString="Mamaev, B. M. &amp; Zaitzev, A. I. (1997) New genus and species of free-developing gall midges of the subfamily Porricondylinae from Somalia (Diptera, Cecidomyiinae). Journal of the Ukrainian Entomological Society, 3 (2), 5 - 13." type="journal article" year="1997">Mamaev &amp; Zaitzev (1997)</bibRefCitation>
: on the one hand,
<taxonomicName id="4C5F4D3BEC6139011BD0FC5EFD71FBC9" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[738,861,1010,1036]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011BD0FC5EFD71FBC9" box="[738,861,1010,1036]" italics="true" pageId="3" pageNumber="264">Asycampta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D3BEC6139011AA6FC5DFA43FBCE" authorityName="Parnell" authorityYear="1971" box="[916,1135,1009,1035]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011AA6FC5DFA43FBCE" box="[916,1135,1009,1035]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
are maintained as separate subgenera/genera and then need to be defined more precisely, on the other, both names are considered as synonyms.
</paragraph>
<paragraph id="8BE036B8EC61390119F4FBF1FCD1F989" blockId="3.[151,1437,325,1756]" pageId="3" pageNumber="264">
The situation becomes more complex with the inclusion of
<taxonomicName id="4C5F4D3BEC6139011A52FBF1FAE6FBB2" authority="Jaschhof &amp; Jaschhof" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[864,1226,1117,1143]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011A52FBF1FDF4FBB2" box="[864,984,1117,1143]" italics="true" pageId="3" pageNumber="264">Zadbimyia</emphasis>
Jaschhof &amp; Jaschhof
</taxonomicName>
, a genus described from
<collectingCountry id="F3487628EC6139011831FB2DFF51FB5E" box="[259,381,1153,1179]" name="Costa Rica" pageId="3" pageNumber="264">Costa Rica</collectingCountry>
several years after the publication of
<bibRefCitation id="EFCE4B49EC6139011A10FB2EFA73FB5E" author="Mamaev, B. M. &amp; Zaitzev, A. I." box="[802,1119,1153,1180]" pageId="3" pageNumber="264" pagination="5 - 13" refId="ref9578" refString="Mamaev, B. M. &amp; Zaitzev, A. I. (1997) New genus and species of free-developing gall midges of the subfamily Porricondylinae from Somalia (Diptera, Cecidomyiinae). Journal of the Ukrainian Entomological Society, 3 (2), 5 - 13." type="journal article" year="1997">Mamaev &amp; Zaitzev’s (1997)</bibRefCitation>
paper (
<bibRefCitation id="EFCE4B49EC6139011D81FB2DFF2FFB7A" author="Jaschhof, M. &amp; Jaschhof, C." pageId="3" pageNumber="264" pagination="1 - 29" refId="ref8736" refString="Jaschhof, M. &amp; Jaschhof, C. (2014) Zadbimyia, a new genus of asynaptine Porricondylinae (Diptera: Cecidomyiidae) with twenty-two new species from the cloud forest of Costa Rica. Zootax a, 3866 (1), 1 - 29. https: // doi. org / 10.11646 / zootaxa. 3866.1.1" type="journal article" year="2014">Jaschhof &amp; Jaschhof 2014</bibRefCitation>
). While
<taxonomicName id="4C5F4D3BEC613901185BFB09FFCDFB7A" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[361,481,1189,1215]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC613901185BFB09FFCDFB7A" box="[361,481,1189,1215]" italics="true" pageId="3" pageNumber="264">Zadbimyia</emphasis>
</taxonomicName>
is variable regarding the characters discussed above (
<bibRefCitation id="EFCE4B49EC6139011D6DFB09FBB4FB7A" author="Jaschhof, M. &amp; Jaschhof, C." box="[1119,1432,1189,1215]" pageId="3" pageNumber="264" pagination="1 - 29" refId="ref8736" refString="Jaschhof, M. &amp; Jaschhof, C. (2014) Zadbimyia, a new genus of asynaptine Porricondylinae (Diptera: Cecidomyiidae) with twenty-two new species from the cloud forest of Costa Rica. Zootax a, 3866 (1), 1 - 29. https: // doi. org / 10.11646 / zootaxa. 3866.1.1" type="journal article" year="2014">Jaschhof &amp; Jaschhof 2014</bibRefCitation>
, fig. 2), the basic construction of male terminalia structures is stable (
<bibRefCitation id="EFCE4B49EC6139011AD0FB65FB39FB26" author="Jaschhof, M. &amp; Jaschhof, C." box="[994,1301,1225,1251]" pageId="3" pageNumber="264" pagination="1 - 29" refId="ref8736" refString="Jaschhof, M. &amp; Jaschhof, C. (2014) Zadbimyia, a new genus of asynaptine Porricondylinae (Diptera: Cecidomyiidae) with twenty-two new species from the cloud forest of Costa Rica. Zootax a, 3866 (1), 1 - 29. https: // doi. org / 10.11646 / zootaxa. 3866.1.1" type="journal article" year="2014">Jaschhof &amp; Jaschhof 2014</bibRefCitation>
, fig. 4) and resembles that described here for three new species of
<taxonomicName id="4C5F4D3BEC6139011A7BFB42FDE8FACD" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[841,964,1262,1288]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011A7BFB42FDE8FACD" box="[841,964,1262,1288]" italics="true" pageId="3" pageNumber="264">Asycampta</emphasis>
</taxonomicName>
. This finding challenges the distinctness of
<taxonomicName id="4C5F4D3BEC61390119D8FABDFF4EFAEE" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[234,354,1297,1323]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC61390119D8FABDFF4EFAEE" box="[234,354,1297,1323]" italics="true" pageId="3" pageNumber="264">Zadbimyia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D3BEC6139011896FABDFC53FAEE" authorityName="Parnell" authorityYear="1971" box="[420,639,1297,1323]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011896FABDFC53FAEE" box="[420,639,1297,1323]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
(now including
<taxonomicName id="4C5F4D3BEC6139011A78FABEFDE9FAE9" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[842,965,1298,1324]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011A78FABEFDE9FAE9" box="[842,965,1298,1324]" italics="true" pageId="3" pageNumber="264">Asycampta</emphasis>
</taxonomicName>
) as postulated by
<bibRefCitation id="EFCE4B49EC6139011D9BFABDFF38FA8A" author="Jaschhof, M. &amp; Jaschhof, C." pageId="3" pageNumber="264" pagination="1 - 29" refId="ref8736" refString="Jaschhof, M. &amp; Jaschhof, C. (2014) Zadbimyia, a new genus of asynaptine Porricondylinae (Diptera: Cecidomyiidae) with twenty-two new species from the cloud forest of Costa Rica. Zootax a, 3866 (1), 1 - 29. https: // doi. org / 10.11646 / zootaxa. 3866.1.1" type="journal article" year="2014">Jaschhof &amp; Jaschhof (2014)</bibRefCitation>
. The decision here to maintain all three genera as discrete, rather than to subsume both
<taxonomicName id="4C5F4D3BEC6139011DC3FA9AFB40FA95" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[1265,1388,1334,1360]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011DC3FA9AFB40FA95" box="[1265,1388,1334,1360]" italics="true" pageId="3" pageNumber="264">Asycampta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D3BEC61390119F4FAF5FF12FAB6" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[198,318,1369,1395]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC61390119F4FAF5FF12FAB6" box="[198,318,1369,1395]" italics="true" pageId="3" pageNumber="264">Zadbimyia</emphasis>
</taxonomicName>
under a broad concept of
<taxonomicName id="4C5F4D3BEC6139011B48FAF5FD79FAB6" authorityName="Parnell" authorityYear="1971" box="[634,853,1369,1395]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011B48FAF5FD79FAB6" box="[634,853,1369,1395]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
, is backed by male morphological indicators (see below under B). As a further argument, at least
<taxonomicName id="4C5F4D3BEC6139011BCAFAD2FD5FFA5D" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[760,883,1406,1432]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011BCAFAD2FD5FFA5D" box="[760,883,1406,1432]" italics="true" pageId="3" pageNumber="264">Asycampta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D3BEC6139011A82FAD1FAA7FA52" authorityName="Parnell" authorityYear="1971" box="[944,1163,1405,1431]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011A82FAD1FAA7FA52" box="[944,1163,1405,1431]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
+
<taxonomicName id="4C5F4D3BEC6139011DA9FAD1FB3FFA52" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[1179,1299,1405,1431]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011DA9FAD1FB3FFA52" box="[1179,1299,1405,1431]" italics="true" pageId="3" pageNumber="264">Zadbimyia</emphasis>
</taxonomicName>
likely have long, independent evolutionary histories that began with the separation of South America and Africa some 130 MYA. The sister group relationship postulated here for
<taxonomicName id="4C5F4D3BEC6139011A0BFA69FA38FA1A" authorityName="Parnell" authorityYear="1971" box="[825,1044,1477,1503]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011A0BFA69FA38FA1A" box="[825,1044,1477,1503]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D3BEC6139011D7EFA69FAE8FA1A" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[1100,1220,1477,1503]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011D7EFA69FAE8FA1A" box="[1100,1220,1477,1503]" italics="true" pageId="3" pageNumber="264">Zadbimyia</emphasis>
</taxonomicName>
cannot be justified by synapomorphies, also because the states of several characters in
<taxonomicName id="4C5F4D3BEC6139011AF2FA45FAB7F9C6" authorityName="Parnell" authorityYear="1971" box="[960,1179,1513,1539]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011AF2FA45FAB7F9C6" box="[960,1179,1513,1539]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
are beyond judgement (see below). The morphology of
<taxonomicName id="4C5F4D3BEC6139011B7BF9A1FCEDF9E2" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[585,705,1549,1575]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011B7BF9A1FCEDF9E2" box="[585,705,1549,1575]" italics="true" pageId="3" pageNumber="264">Zadbimyia</emphasis>
</taxonomicName>
is, compared with
<taxonomicName id="4C5F4D3BEC6139011A95F9A1FAAEF9E2" authorityName="Parnell" authorityYear="1971" box="[935,1154,1549,1575]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011A95F9A1FAAEF9E2" box="[935,1154,1549,1575]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
, in many respects more advanced and largely similar to that of
<taxonomicName id="4C5F4D3BEC6139011B49F99EFCDAF989" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[635,758,1586,1612]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011B49F99EFCDAF989" box="[635,758,1586,1612]" italics="true" pageId="3" pageNumber="264">Asycampta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE036B8EC61390119F4F9FAFA15F919" blockId="3.[151,1437,325,1756]" pageId="3" pageNumber="264">
At this point
<taxonomicName id="4C5F4D3BEC6139011869F9FAFCE5F9AA" authority="Fedotova &amp; Sidorenko" authorityName="Fedotova &amp; Sidorenko" authorityYear="2007" box="[347,713,1621,1647]" class="Insecta" family="Cecidomyiidae" genus="Larimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011869F9FAFFEEF9AA" box="[347,450,1622,1647]" italics="true" pageId="3" pageNumber="264">Larimyia</emphasis>
Fedotova &amp; Sidorenko
</taxonomicName>
must be mentioned, a monotypic genus of
<taxonomicName id="4C5F4D3BEC6139011D9DF9F9FB06F9AA" baseAuthorityName="Gagne &amp; Jaschhof" baseAuthorityYear="2021" box="[1199,1322,1621,1647]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
occurring in the eastern Palearctic Region, the wing and terminalia of which (
<bibRefCitation id="EFCE4B49EC6139011A9EF9D5FAC0F956" author="Fedotova, Z. A. &amp; Sidorenko, V. S." box="[940,1260,1657,1683]" pageId="3" pageNumber="264" pagination="69 - 103" refId="ref8442" refString="Fedotova, Z. A. &amp; Sidorenko, V. S. (2007) New taxa of gall midges from tribes Porricondylini, Bryocryptini, Asynaptini and Winnertziini (Diptera, Cecidomyiidae, Porricondylinae) from the Russian Far East. International Journal of Dipterological Research, 18, 69 - 103." type="journal article" year="2007">Fedotova &amp; Sidorenko 2007</bibRefCitation>
, figs 84, 93) are reminiscent of the genera under discussion here. The original description does not, however, provide sufficient detail to permit any conclusions regarding the systematic position of
<taxonomicName id="4C5F4D3BEC6139011AF9F96EFA1EF91E" authorityName="Fedotova &amp; Sidorenko" authorityYear="2007" box="[971,1074,1730,1755]" class="Insecta" family="Cecidomyiidae" genus="Larimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011AF9F96EFA1EF91E" box="[971,1074,1730,1755]" italics="true" pageId="3" pageNumber="264">Larimyia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE036B8EC61390619A5F8A8FF84FE4C" blockId="3.[151,1437,1796,2039]" lastBlockId="4.[198,1437,151,753]" lastPageId="4" lastPageNumber="265" pageId="3" pageNumber="264">
(B) Definitions.The
<taxonomicName id="4C5F4D3BEC6139011845F8A8FC7CF8DB" authorityName="Parnell" authorityYear="1971" box="[375,592,1796,1822]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011845F8A8FC7CF8DB" box="[375,592,1796,1822]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
groupofgenera(synapomorphiesunderlined):
<taxonomicName id="4C5F4D3BEC6139011D73F8A8FA97F8DB" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[1089,1211,1796,1822]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011D73F8A8FA97F8DB" box="[1089,1211,1796,1822]" italics="true" pageId="3" pageNumber="264">Asycampta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5F4D3BEC6139011DF1F8A8FBB0F8DB" authorityName="Parnell" authorityYear="1971" box="[1219,1436,1796,1822]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6139011DF1F8A8FBB0F8DB" box="[1219,1436,1796,1822]" italics="true" pageId="3" pageNumber="264">Pseudocamptomyia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D3BEC61390119C5F884FF42F887" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[247,366,1832,1858]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC61390119C5F884FF42F887" box="[247,366,1832,1858]" italics="true" pageId="3" pageNumber="264">Zadbimyia</emphasis>
</taxonomicName>
form a generic group that differs from other
<taxonomicName id="4C5F4D3BEC6139011A40F884FDC0F887" baseAuthorityName="Gagne &amp; Jaschhof" baseAuthorityYear="2021" box="[882,1004,1832,1858]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
in the following combination of male character states (females and larvae are unknown): the scape has a medial cluster of dense setae; the scutum lacks transparent windows; vein
<emphasis id="B92BEAAAEC61390118C3F8DDFC0FF855" box="[497,547,1905,1936]" italics="true" pageId="3" pageNumber="264">
M
<subScript id="17DB34FDEC6139011B3AF82EFC0FF855" attach="left" box="[520,547,1922,1936]" fontSize="6" pageId="3" pageNumber="264">1+2</subScript>
</emphasis>
is always and vein
<emphasis id="B92BEAAAEC6139011BC8F8DDFD35F855" box="[762,793,1905,1936]" italics="true" pageId="3" pageNumber="264">
M
<subScript id="17DB34FDEC6139011A23F82EFD35F855" attach="left" box="[785,793,1922,1936]" fontSize="6" pageId="3" pageNumber="264">4</subScript>
</emphasis>
usually absent; the gonostylus lacks a pectinate tooth; the parameres are merged medially to form the tegmen that is closely tied to the apex of the phallapodeme (only two species of
<taxonomicName id="4C5F4D3BEC613901180DF814FF9AF817" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[319,438,1976,2002]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC613901180DF814FF9AF817" box="[319,438,1976,2002]" italics="true" pageId="3" pageNumber="264">Zadbimyia</emphasis>
</taxonomicName>
, the affiliation of which is disputable, have separate parameres); the structural complex made of tegmen and phallapodeme is equipped with processes and lobes of varying size, shape and orientation.
<taxonomicName id="4C5F4D3BEC66390619F4FF3BFF6DFF74" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[198,321,151,177]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC66390619F4FF3BFF6DFF74" box="[198,321,151,177]" italics="true" pageId="4" pageNumber="265">Asycampta</emphasis>
</taxonomicName>
. Flagellomeral necks lack microtrichia; circumfila are irregularly, strongly looped. Both labellum and palpus are atrophied; the palpus, which is markedly shorter than the head height, has one to four setaebearing segments. Basitarsal spines are present. The gonostylar apex is either unmodified, equipped with a bunch of large microtrichia (spines), or reinforced by strong sclerotization and then without microtrichia. The geographical distribution is exclusively Afrotropical. The definition given here differs from that by
<bibRefCitation id="EFCE4B49EC66390619F4FEE7FFD9FEA0" author="Mamaev, B. M. &amp; Zaitzev, A. I." box="[198,501,331,357]" pageId="4" pageNumber="265" pagination="5 - 13" refId="ref9578" refString="Mamaev, B. M. &amp; Zaitzev, A. I. (1997) New genus and species of free-developing gall midges of the subfamily Porricondylinae from Somalia (Diptera, Cecidomyiinae). Journal of the Ukrainian Entomological Society, 3 (2), 5 - 13." type="journal article" year="1997">Mamaev &amp; Zaitzev (1997)</bibRefCitation>
for it emphasizes the distinctions separating
<taxonomicName id="4C5F4D3BEC6639061ACAFEE7FA5FFEA0" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[1016,1139,331,357]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061ACAFEE7FA5FFEA0" box="[1016,1139,331,357]" italics="true" pageId="4" pageNumber="265">Asycampta</emphasis>
</taxonomicName>
from both
<taxonomicName id="4C5F4D3BEC6639061DC6FEE7FB40FEA0" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[1268,1388,331,357]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061DC6FEE7FB40FEA0" box="[1268,1388,331,357]" italics="true" pageId="4" pageNumber="265">Zadbimyia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D3BEC66390619F4FEC3FF8DFE4C" authorityName="Parnell" authorityYear="1971" box="[198,417,367,393]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC66390619F4FEC3FF8DFE4C" box="[198,417,367,393]" italics="true" pageId="4" pageNumber="265">Pseudocamptomyia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE036B8EC66390619F4FE3FFC56FDF8" blockId="4.[198,1437,151,753]" pageId="4" pageNumber="265">
<taxonomicName id="4C5F4D3BEC66390619F4FE3FFF12FE68" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[198,318,403,429]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC66390619F4FE3FFF12FE68" box="[198,318,403,429]" italics="true" pageId="4" pageNumber="265">Zadbimyia</emphasis>
</taxonomicName>
. Flagellomeral necks are, in most of the species, partly or entirely microtrichose; circumfila are irregularly, strongly looped. Both labellum and palpus are atrophied; the palpus, which is markedly shorter than the head height, has one to four setae-bearing segments. Basitarsal spines are present. The gonostylar apex is variously, densely microtrichose, with conspicuously large microtrichae (spines) being absent. The geographical distribution is exclusively Neotropical.
</paragraph>
<paragraph id="8BE036B8EC66390619F4FDEBFCC0FD34" blockId="4.[198,1437,151,753]" pageId="4" pageNumber="265">
<taxonomicName id="4C5F4D3BEC66390619F4FDEBFF8DFDA4" authorityName="Parnell" authorityYear="1971" box="[198,417,583,609]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC66390619F4FDEBFF8DFDA4" box="[198,417,583,609]" italics="true" pageId="4" pageNumber="265">Pseudocamptomyia</emphasis>
</taxonomicName>
. Flagellomeral necks lack microtrichia; circumfila are slightly sinuous. The palpus has four setae-bearing segments; its length, as well as the structure of the labella, are unknown, due to the poor condition of available specimens. Basitarsi have a microtrichose apical projection rather than a spine. The gonostylar apex is possibly equipped with one or several small spines, which cannot be decided from existing specimens. The geographical distribution is exclusively Nearctic.
</paragraph>
</subSubSection>
<subSubSection id="C3456533EC66390619A5FCB3FFA5FA04" pageId="4" pageNumber="265" type="discussion">
<paragraph id="8BE036B8EC66390619A5FCB3FFA5FA04" blockId="4.[151,1437,799,1473]" pageId="4" pageNumber="265">
(C) Discussion. Besides
<taxonomicName id="4C5F4D3BEC6639061883FCB3FC35FCFC" authorityName="Loew" authorityYear="1850" box="[433,537,799,825]" class="Insecta" family="Cecidomyiidae" genus="Asynapta" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061883FCB3FC35FCFC" box="[433,537,799,825]" italics="true" pageId="4" pageNumber="265">Asynapta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D3BEC6639061B61FCB3FCCEFCFC" authorityName="Kieffer" authorityYear="1894" box="[595,738,799,825]" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061B61FCB3FCCEFCFC" box="[595,738,799,825]" italics="true" pageId="4" pageNumber="265">Camptomyia</emphasis>
</taxonomicName>
, the
<taxonomicName id="4C5F4D3BEC6639061A2CFCB3FDD5FCFC" authorityName="Parnell" authorityYear="1971" box="[798,1017,799,825]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061A2CFCB3FDD5FCFC" box="[798,1017,799,825]" italics="true" pageId="4" pageNumber="265">Pseudocamptomyia</emphasis>
</taxonomicName>
group of genera represents a further extraordinarily speciose and complex subset of
<taxonomicName id="4C5F4D3BEC6639061BFEFCEFFD67FC98" baseAuthorityName="Gagne &amp; Jaschhof" baseAuthorityYear="2021" box="[716,843,835,861]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
. One of the included genera,
<taxonomicName id="4C5F4D3BEC6639061DB6FCEFFAD0FC98" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[1156,1276,835,861]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061DB6FCEFFAD0FC98" box="[1156,1276,835,861]" italics="true" pageId="4" pageNumber="265">Zadbimyia</emphasis>
</taxonomicName>
, is remarkable for its regional and local diversity, with 47 species, both named and unnamed, known to be present in
<collectingCountry id="F3487628EC6639061C2FFCCBFBB4FC44" box="[1309,1432,871,897]" name="Costa Rica" pageId="4" pageNumber="265">Costa Rica</collectingCountry>
, including 35 species known to occur at a single site of cloud forest (
<bibRefCitation id="EFCE4B49EC6639061AE7FC27FB2BFC60" author="Jaschhof, M. &amp; Jaschhof, C." box="[981,1287,907,933]" pageId="4" pageNumber="265" pagination="1 - 29" refId="ref8736" refString="Jaschhof, M. &amp; Jaschhof, C. (2014) Zadbimyia, a new genus of asynaptine Porricondylinae (Diptera: Cecidomyiidae) with twenty-two new species from the cloud forest of Costa Rica. Zootax a, 3866 (1), 1 - 29. https: // doi. org / 10.11646 / zootaxa. 3866.1.1" type="journal article" year="2014">Jaschhof &amp; Jaschhof 2014</bibRefCitation>
; unpublished data). The fact that the genus has not yet been found outside
<collectingCountry id="F3487628EC6639061ABDFC03FA20FC0C" box="[911,1036,943,969]" name="Costa Rica" pageId="4" pageNumber="265">Costa Rica</collectingCountry>
is certainly due to the shortage of surveys targeting
<taxonomicName id="4C5F4D3BEC66390618BDFC7FFC6EFC28" authorityName="Kieffer" authorityYear="1913" box="[399,578,979,1005]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="subFamily" subFamily="Porricondylinae">Porricondylinae</taxonomicName>
in the Neotropics. There is every indication that
<taxonomicName id="4C5F4D3BEC6639061D41FC7FFAC2FC28" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[1139,1262,979,1005]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061D41FC7FFAC2FC28" box="[1139,1262,979,1005]" italics="true" pageId="4" pageNumber="265">Asycampta</emphasis>
</taxonomicName>
plays a similar role in the Afrotropics. The three species described here as new were found co-occurring at a single site, and
<emphasis id="B92BEAAAEC66390619F4FBB7FF6CFBF1" box="[198,320,1051,1076]" italics="true" pageId="4" pageNumber="265">en passant</emphasis>
rather than through targeted search. The same is true for the two species (including
<taxonomicName id="4C5F4D3BEC6639061C29FBB7FF9DFB9C" authority="(Mamaev &amp; Zaitzev)" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2023" baseAuthorityName="Jaschhof &amp; Jaschhof" baseAuthorityYear="2023" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="africana" status="comb. nov.">
<emphasis id="B92BEAAAEC6639061C29FBB7FB00FBF0" box="[1307,1324,1051,1077]" italics="true" pageId="4" pageNumber="265">A</emphasis>
.
<emphasis id="B92BEAAAEC6639061C0FFBB7FBB0FBF0" box="[1341,1436,1051,1077]" italics="true" pageId="4" pageNumber="265">africana</emphasis>
(Mamaev &amp; Zaitzev)
</taxonomicName>
<emphasis id="B92BEAAAEC6639061885FB93FC1AFB9C" bold="true" box="[439,566,1087,1113]" pageId="4" pageNumber="265">
<taxonomicNameLabel id="A21857D1EC6639061885FB93FC1AFB9C" box="[439,566,1087,1113]" pageId="4" pageNumber="265" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
) described from
<collectingCountry id="F3487628EC6639061BC6FB93FD7FFB9C" box="[756,851,1087,1113]" name="Somalia" pageId="4" pageNumber="265">Somalia</collectingCountry>
. As a further indication of
<taxonomicName id="4C5F4D3BEC6639061D4BFB93FAD8FB9C" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[1145,1268,1087,1113]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061D4BFB93FAD8FB9C" box="[1145,1268,1087,1113]" italics="true" pageId="4" pageNumber="265">Asycampta</emphasis>
</taxonomicName>
’s diversity, the five species now known of the genus exhibit great morphological complexity, which is comparable to that found in
<taxonomicName id="4C5F4D3BEC66390619D3FB2BFF75FB64" authorityName="Jaschhof &amp; Jaschhof" authorityYear="2014" box="[225,345,1159,1185]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC66390619D3FB2BFF75FB64" box="[225,345,1159,1185]" italics="true" pageId="4" pageNumber="265">Zadbimyia</emphasis>
</taxonomicName>
. A number of issues regarding the
<taxonomicName id="4C5F4D3BEC6639061BE4FB2BFD9DFB64" authorityName="Parnell" authorityYear="1971" box="[726,945,1159,1185]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061BE4FB2BFD9DFB64" box="[726,945,1159,1185]" italics="true" pageId="4" pageNumber="265">Pseudocamptomyia</emphasis>
</taxonomicName>
group remain unresolved for the time being. It remains to be shown, for example, whether
<taxonomicName id="4C5F4D3BEC6639061BFAFB07FD8FFB00" authorityName="Parnell" authorityYear="1971" box="[712,931,1195,1221]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061BFAFB07FD8FFB00" box="[712,931,1195,1221]" italics="true" pageId="4" pageNumber="265">Pseudocamptomyia</emphasis>
</taxonomicName>
is exclusively Nearctic in distribution and as poor in species as present data suggest; whether
<taxonomicName id="4C5F4D3BEC6639061BDCFB63FD79FB2D" authorityName="Fedotova &amp; Sidorenko" authorityYear="2007" box="[750,853,1231,1256]" class="Insecta" family="Cecidomyiidae" genus="Larimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061BDCFB63FD79FB2D" box="[750,853,1231,1256]" italics="true" pageId="4" pageNumber="265">Larimyia</emphasis>
</taxonomicName>
is a representative and if so the
<emphasis id="B92BEAAAEC6639061DF5FB63FADBFB2C" box="[1223,1271,1231,1257]" italics="true" pageId="4" pageNumber="265">only</emphasis>
representative of this genus-group in the Palearctic Region; whether
<taxonomicName id="4C5F4D3BEC6639061A14FB5FFD8DFAC8" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[806,929,1267,1293]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061A14FB5FFD8DFAC8" box="[806,929,1267,1293]" italics="true" pageId="4" pageNumber="265">Asycampta</emphasis>
</taxonomicName>
expands into the Oriental Region or whether the
<taxonomicName id="4C5F4D3BEC66390619C2FABBFFE7FAF4" authorityName="Parnell" authorityYear="1971" box="[240,459,1303,1329]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC66390619C2FABBFFE7FAF4" box="[240,459,1303,1329]" italics="true" pageId="4" pageNumber="265">Pseudocamptomyia</emphasis>
</taxonomicName>
group has a different, yet unknown branch in that part of the world. It is obvious that the phylogenetic and biogeographic relations within the
<taxonomicName id="4C5F4D3BEC6639061A7FFA97FA04FA90" authorityName="Parnell" authorityYear="1971" box="[845,1064,1339,1365]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC6639061A7FFA97FA04FA90" box="[845,1064,1339,1365]" italics="true" pageId="4" pageNumber="265">Pseudocamptomyia</emphasis>
</taxonomicName>
group remain as vague as those within the
<taxonomicName id="4C5F4D3BEC6639061808FAF3FF99FABC" baseAuthorityName="Gagne &amp; Jaschhof" baseAuthorityYear="2021" box="[314,437,1375,1401]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
as a whole (
<bibRefCitation id="EFCE4B49EC6639061B0FFAF3FD44FABC" author="Jaschhof, M. &amp; Jaschhof, C." box="[573,872,1375,1401]" pageId="4" pageNumber="265" pagination="1 - 392" refId="ref8698" refString="Jaschhof, M. &amp; Jaschhof, C. (2013) The Porricondylinae (Diptera: Cecidomyiidae) of Sweden, with notes on extralimital species. Studia dipterologica Supplement, 20, 1 - 392." type="journal article" year="2013">Jaschhof &amp; Jaschhof 2013</bibRefCitation>
). The single most important impediment to a better understanding of asynaptine inter-relationships is the shortage of basic taxonomic information regarding nonEuropean faunas.
</paragraph>
</subSubSection>
<subSubSection id="C3456533EC66390619A5FA45FDB1F91E" pageId="4" pageNumber="265" type="diagnosis">
<paragraph id="8BE036B8EC66390619A5FA45FDB1F91E" blockId="4.[151,1437,1513,1756]" pageId="4" pageNumber="265">
(D) Identification of
<taxonomicName id="4C5F4D3BEC66390618B1FA46FFD2F9C1" authorityName="Mamaev &amp; Zaitzev" authorityYear="2023" box="[387,510,1514,1540]" class="Insecta" family="Cecidomyiidae" genus="Asycampta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEAAAEC66390618B1FA46FFD2F9C1" box="[387,510,1514,1540]" italics="true" pageId="4" pageNumber="265">Asycampta</emphasis>
</taxonomicName>
using male morphology. With only five species known at the present, certain nonterminalia characters suffice for their differentiation (
<bibRefCitation id="EFCE4B49EC6639061A23F9A1FA07F9E2" author="Mamaev, B. M. &amp; Zaitzev, A. I." box="[785,1067,1549,1575]" pageId="4" pageNumber="265" pagination="5 - 13" refId="ref9578" refString="Mamaev, B. M. &amp; Zaitzev, A. I. (1997) New genus and species of free-developing gall midges of the subfamily Porricondylinae from Somalia (Diptera, Cecidomyiinae). Journal of the Ukrainian Entomological Society, 3 (2), 5 - 13." type="journal article" year="1997">Mamaev &amp; Zaitzev 1997</bibRefCitation>
). Those characters may, to a large extent, be observed even in specimens stored in ethanol, meaning they are more readily accessible compared to terminalia characters, which require investigation by light microscopy. Considering the expectation that dozens of
<taxonomicName id="4C5F4D3BEC66390619D4F9D6FF97F956" box="[230,443,1657,1684]" pageId="4" pageNumber="265">
<emphasis id="B92BEAAAEC66390619D4F9D6FF4DF951" box="[230,353,1658,1684]" italics="true" pageId="4" pageNumber="265">Asycampta</emphasis>
species
</taxonomicName>
remain to be discovered and described, the specific diagnoses given below give major weight to terminalia structures, which in speciose genera of
<taxonomicName id="4C5F4D3BEC6639061A55F931FDCEF972" baseAuthorityName="Gagne &amp; Jaschhof" baseAuthorityYear="2021" box="[871,994,1693,1719]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
are proven to provide the only positive interspecific distinctions (
<emphasis id="B92BEAAAEC66390618DBF96EFC3DF91E" box="[489,529,1730,1755]" italics="true" pageId="4" pageNumber="265">e.g.</emphasis>
,
<bibRefCitation id="EFCE4B49EC6639061B2DF96DFD65F91E" author="Jaschhof, M. &amp; Jaschhof, C." box="[543,841,1729,1755]" pageId="4" pageNumber="265" pagination="1 - 392" refId="ref8698" refString="Jaschhof, M. &amp; Jaschhof, C. (2013) The Porricondylinae (Diptera: Cecidomyiidae) of Sweden, with notes on extralimital species. Studia dipterologica Supplement, 20, 1 - 392." type="journal article" year="2013">Jaschhof &amp; Jaschhof 2013</bibRefCitation>
,
<bibRefCitation id="EFCE4B49EC6639061A66F96DFDBDF91E" author="Jaschhof, M. &amp; Jaschhof, C." box="[852,913,1729,1755]" pageId="4" pageNumber="265" pagination="1 - 29" refId="ref8736" refString="Jaschhof, M. &amp; Jaschhof, C. (2014) Zadbimyia, a new genus of asynaptine Porricondylinae (Diptera: Cecidomyiidae) with twenty-two new species from the cloud forest of Costa Rica. Zootax a, 3866 (1), 1 - 29. https: // doi. org / 10.11646 / zootaxa. 3866.1.1" type="journal article" year="2014">2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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