treatments-xml/data/03/F3/87/03F387C57A23FFB0FF6C072130C0B1B6.xml

<document id="602D9FD910B3E575C95E315DD2A6568E" ID-CLB-Dataset="8143" ID-DOI="10.2478/aemnp-2019-0020" ID-GBIF-Dataset="a03ecce2-b526-4f94-aa65-620016956099" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488918" ID-ZooBank="9F309FCC-A2ED-47B9-BC37-D0C4A3B482E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612219281920" checkinUser="felipe" docAuthor="Fikáček, Martin &amp; Liu, Hsing-Che" docDate="2019" docId="03F387C57A23FFB0FF6C072130C0B1B6" docLanguage="en" docName="ActaEntMusNatPra.59.1.229-252.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 59 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Thysanarthria championi" docType="treatment" docVersion="9" lastPageNumber="243" masterDocId="FFCAFFBD7A2EFFBEFF8901283431B709" masterDocTitle="A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)" masterLastPageNumber="252" masterPageNumber="229" pageNumber="242" updateTime="1698926077238" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods id="1DF141BCA73A53B5D281269950289E9B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9AE21DD299B595907190B096181F3F9D">
<mods:title id="C62F05E0A5AD24E539D54AD91733B120">A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)</mods:title>
</mods:titleInfo>
<mods:name id="F12774E981CD5621EFA868A538F89FD5" type="personal">
<mods:role id="E2AC5C09EFD8E2DE78D2EAC1C4607A79">
<mods:roleTerm id="5BA1770D208CA1E74F7D6B1BD23F1553">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0993F41016B98E8E4F043B4C88556704">Fikáček, Martin</mods:namePart>
<mods:affiliation id="BBC3B25C6595B262A55C1C5EDCA6D0D9">Department of Entomology, National Museum, Cirkusová 1740, CZ- 19100 Praha 9 – Horní Počernice, Czech Republic &amp; Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844 Praha 2, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="65BDE31D4FB8149D9EE86075A21CB91A" type="email">mfikacek@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="DA1CE49ADD462AD515F5D93C049CD9F4" type="personal">
<mods:role id="2F3AC700F93B66CC8A46B040681A8C0D">
<mods:roleTerm id="6ED5080006D665654849088CD3647FD6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8902B8B6A992F31B92FC1333AC8CCD25">Liu, Hsing-Che</mods:namePart>
<mods:affiliation id="81DF6FAFAB439285768E12AF9247FCB8">Department of Environmental Engineering and Management, Chaoyang University of Technology, No. 168, Jifeng E. Rd., Taichung City 413, Taiwan</mods:affiliation>
<mods:nameIdentifier id="78E41F04052971B8C53DA3238AA1B3A3" type="email">td965771@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="77D624C5121AC3A2C30C7FF994D3F4C3">text</mods:typeOfResource>
<mods:relatedItem id="EE9FB24AD91E42A73046E970E92980CB" type="host">
<mods:titleInfo id="609FF517310F3A14B5EABEFDD734B975">
<mods:title id="0F3A42673721D43B32815DA53FCF90EF">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="5CAFE48F3367935008EB53B71D997A92">
<mods:date id="39840FCC8FABCB495E2505756BF3067C">2019</mods:date>
<mods:detail id="2E0CF22FF63DAB64262870A601B4CD94" type="pubDate">
<mods:number id="78D5C8B20789E621FB445722A20C9811">2019-06-20</mods:number>
</mods:detail>
<mods:detail id="4A0561B756954EB3FEB73ABE9B4F349E" type="volume">
<mods:number id="5AF77B6E85BBD3526C4DBF44C8A94A95">59</mods:number>
</mods:detail>
<mods:detail id="0096FAEC6E4213F11E4A3944B0ED8DFD" type="issue">
<mods:number id="5E4C0EEE36A57527FA036AFAC9FD2051">1</mods:number>
</mods:detail>
<mods:extent id="8E87EF8C1CA9F76D20BFB2909D86951E" unit="page">
<mods:start id="6D21125E819A4AF1CBEA3778A9040613">229</mods:start>
<mods:end id="2F3958DA784B909AEEA4F6AB5854DE13">252</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1B3BF4E5ADB1F9111228E7FC673D8750">journal article</mods:classification>
<mods:identifier id="832F7DD90630FAD765B3D8DA4BB8F498" type="CLB-Dataset">8143</mods:identifier>
<mods:identifier id="9A93F690B2FB28C4A5D3032907F01249" type="DOI">10.2478/aemnp-2019-0020</mods:identifier>
<mods:identifier id="E0445321D41CCA80523FD90AE829879D" type="GBIF-Dataset">a03ecce2-b526-4f94-aa65-620016956099</mods:identifier>
<mods:identifier id="9C4C6361F4FE38BAA4BCF8C23E15D53C" type="ISSN">1804-6487</mods:identifier>
<mods:identifier id="6096E562D29BA748DE93BE192047CA52" type="Zenodo-Dep">4488918</mods:identifier>
<mods:identifier id="92D6E979E3937768CDAA70CD91C57D8C" type="ZooBank">9F309FCC-A2ED-47B9-BC37-D0C4A3B482E5</mods:identifier>
</mods:mods>
<treatment id="03F387C57A23FFB0FF6C072130C0B1B6" ID-DOI="http://doi.org/10.5281/zenodo.4548833" ID-GBIF-Taxon="177423424" ID-Zenodo-Dep="4548833" LSID="urn:lsid:plazi:treatment:03F387C57A23FFB0FF6C072130C0B1B6" httpUri="http://treatment.plazi.org/id/03F387C57A23FFB0FF6C072130C0B1B6" lastPageId="14" lastPageNumber="243" pageId="13" pageNumber="242">
<subSubSection id="C34065587A23FFB3FF6C0721369BB129" box="[229,682,1545,1568]" pageId="13" pageNumber="242" type="nomenclature">
<paragraph id="8BE536D37A23FFB3FF6C0721369BB129" blockId="13.[229,682,1545,1568]" box="[229,682,1545,1568]" pageId="13" pageNumber="242">
<heading id="D0AD81BF7A23FFB3FF6C0721369BB129" box="[229,682,1545,1568]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="242" reason="3">
<taxonomicName id="4C5A4D507A23FFB3FF6C0721369BB129" ID-CoL="56TH4" authority="(Knisch, 1924)" baseAuthorityName="Knisch" baseAuthorityYear="1924" box="[229,682,1545,1568]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B92EEAC17A23FFB3FF6C0721369BB129" bold="true" box="[229,682,1545,1568]" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FF6C072135CEB129" bold="true" box="[229,511,1545,1568]" italics="true" pageId="13" pageNumber="242">Thysanarthria championi</emphasis>
(
<bibRefCitation id="EFCB4B227A23FFB3FD8407213692B129" author="KNISCH A." bookContentInfo="306 pp." box="[525,675,1545,1568]" editor="JUNK W. &amp; SCHENKLING S." journalOrPublisher="W. Junk, Berlin" pageId="13" pageNumber="242" refId="ref18391" refString="KNISCH A. 1924 a: Hydrophilidae. In: JUNK W. &amp; SCHENKLING S. (eds): Coleopterorum Catalogus. Vol. 14, part 79. W. Junk, Berlin, 306 pp." title="Hydrophilidae" type="book chapter" volumeTitle="Coleopterorum Catalogus. Vol. 14, part 79." year="1924">Knisch, 1924</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34065587A23FFB3FEBC0700366BB132" box="[309,602,1576,1595]" pageId="13" pageNumber="242" type="description">
<paragraph id="8BE536D37A23FFB3FEBC0700366BB132" blockId="13.[151,761,1576,1869]" box="[309,602,1576,1595]" pageId="13" pageNumber="242">
(
<figureCitation id="13612A567A23FFB3FEB207003543B132" box="[315,370,1576,1595]" captionStart="Fig" captionStartId="3.[151,182,1959,1978]" captionTargetBox="[157,1426,396,1946]" captionTargetId="figure-14@3.[151,1436,395,1949]" captionTargetPageId="3" captionText="Fig. 2. Morphological differences among Thysanarthria and principal groups of Chaetarthria. A–Z – male genitalia and surrounding sclerites; a–e – details of elytral pubescence. A – Chaetarthria similis Wollaston, 1864; B–G, c – Chaetarthria seminulum (Herbst, 1797); H–M, b – Thysanarthria championi (Knisch, 1924); N–S, d – Chaetarthria panda Orchymont, 1939; T–Z, e – Chaetarthria malickyi Hebauer, 1995; a – Thysanarthria atriceps (Régimbart, 1903). A–D, K–M, Q–S, X–Z – aedeagus in dorsal, lateral and ventral view; E, J, P, V – male sternite 9; F, H, N, T – male sternite 8; G, I, O, U – male tergite 8. Figures a–e are all to the same scale." figureDoi="http://doi.org/10.5281/zenodo.4488924" httpUri="https://zenodo.org/record/4488924/files/figure.png" pageId="13" pageNumber="242">Figs 2</figureCitation>
H–M,b, 
<figureCitation id="13612A567A23FFB3FE30070035F3B132" box="[441,450,1576,1595]" captionStart="Fig" captionStartId="6.[151,182,1970,1989]" captionTargetBox="[178,1407,394,1955]" captionTargetId="figure-13@6.[176,1412,393,1956]" captionTargetPageId="6" captionText="Fig. 3. Morphology of Thysanarthria. A – mentum and maxillary palps; B – meso- and metaventrite and meso- and metafemora; C – head and prothorax in ventral view; D – antenna; E – ultimate maxillary palpomere with basal peg-like setae; F – metatarsus; G – ventral view with gelatinous substrance on the base of abdomen; H – detail of abdomen with gelatinous substance in the cavity on ventrites 1–2. A, C–F – Thysanarthria championi (Knisch, 1924); B, G–H – T. atriceps (Régimbart, 1903)." figureDoi="http://doi.org/10.5281/zenodo.4488928" httpUri="https://zenodo.org/record/4488928/files/figure.png" pageId="13" pageNumber="242">3</figureCitation>
A,C–F, 
<figureCitation id="13612A567A23FFB3FD8F07003621B132" box="[518,528,1576,1595]" captionStart="Fig" captionStartId="14.[151,182,1334,1353]" captionTargetBox="[177,1428,199,1315]" captionTargetId="figure-13@14.[151,1436,186,1319]" captionTargetPageId="14" captionText="Fig. 7. Male genitalia of Thysanarthria championi (Knisch, 1924).A–E – lectotype of Chaetarthria championi Knisch, 1924 (A–C – whole aedeagus in dorsal, lateral and ventral view; D – detail of median lobe and parameres in dorsal view; E – same after KOH treatment, with fully everted internal sac). F–J – lectotype of Chaetarthriomorphus sulcatus Chiesa, 1967 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488938" httpUri="https://zenodo.org/record/4488938/files/figure.png" pageId="13" pageNumber="242">7</figureCitation>
A–J, 
<figureCitation id="13612A567A23FFB3FDB707003662B132" box="[574,595,1576,1595]" captionStart="Fig" captionStartId="20.[151,182,1949,1968]" captionTargetBox="[151,1436,401,1931]" captionTargetId="figure-13@20.[151,1437,401,1931]" captionTargetPageId="20" captionText="Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu." figureDoi="http://doi.org/10.5281/zenodo.4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="13" pageNumber="242">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34065587A23FFB3FF1E076234C9B044" pageId="13" pageNumber="242" type="reference_group">
<paragraph id="8BE536D37A23FFB3FF1E0762363EB154" blockId="13.[151,761,1576,1869]" box="[151,527,1610,1629]" pageId="13" pageNumber="242">
<treatmentCitationGroup id="AB4A11FD7A23FFB3FF1E0762363EB154" box="[151,527,1610,1629]" pageId="13" pageNumber="242">
<treatmentCitation id="0AFB10C27A23FFB3FF1E0762363AB154" author="KNISCH A." box="[151,523,1610,1629]" page="40" pageId="13" pageNumber="242" year="1924">
<taxonomicName id="4C5A4D507A23FFB3FF1E0762363AB154" ID-CoL="5XNRN" authority="Knisch, 1924 b: 40" authorityName="Knisch" authorityPageNumber="40" authorityYear="1924" box="[151,523,1610,1629]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B92EEAC17A23FFB3FF1E07623557B154" box="[151,358,1610,1629]" italics="true" pageId="13" pageNumber="242">Chaetarthria championi</emphasis>
<bibRefCitation id="EFCB4B227A23FFB3FEE20762363AB154" author="Knisch" box="[363,523,1610,1629]" firstAuthor="Knisch" page="40" pageId="13" pageNumber="242" pagination="29 - 41" refId="ref18431" refString="KNISCH A. 1924 b: Neue palpicornier aus dem sudlichen Himalaya (Col. Hydrophilidae - Op. 15). Wiener Entomologische Zeitung 41: 29 - 41." type="journal article" year="1924" yearSuffix="b">Knisch, 1924b: 40</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536D37A23FFB3FF1E074D36C8B1C1" blockId="13.[151,761,1576,1869]" pageId="13" pageNumber="242">
<treatmentCitationGroup id="AB4A11FD7A23FFB3FF1E074D36C8B1C1" pageId="13" pageNumber="242">
<treatmentCitation id="0AFB10C27A23FFB3FF1E074D3673B171" author="ORCHYMONT A. d'" box="[151,578,1637,1656]" page="195" pageId="13" pageNumber="242" year="1926">
<taxonomicName id="4C5A4D507A23FFB3FF1E074D3673B171" ID-CoL="56TH4" authority=": ORCHYMONT (1926 a: 195" authorityName=": ORCHYMONT" authorityYear="1926" box="[151,578,1637,1656]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B92EEAC17A23FFB3FF1E074D355DB171" box="[151,364,1637,1656]" italics="true" pageId="13" pageNumber="242">Thysanarthria championi</emphasis>
: 
<bibRefCitation id="EFCB4B227A23FFB3FEFF074D3673B171" author="ORCHYMONT" box="[374,578,1637,1656]" firstAuthor="ORCHYMONT" page="195" pageId="13" pageNumber="242" pagination="195 - 196" refId="ref18492" refString="ORCHYMONT A. d' 1926 a: Note sur Thysanarthria n. g. Bulletin et Annales de la Societe Entomologique de Belgique 66: 195 - 196." type="journal article" year="1926" yearSuffix="a">ORCHYMONT (1926a: 195</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, transfer to 
<taxonomicName id="4C5A4D507A23FFB3FD2B074D34ECB19B" ID-CoL="8X7X2" authorityName="Fikáček &amp; Liu" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A23FFB3FD2B074D34ECB19B" italics="true" pageId="13" pageNumber="242">Thysanarthria</emphasis>
</taxonomicName>
); 
<treatmentCitation id="0AFB10C27A23FFB3FF670757358BB19B" author="ORCHYMONT A." box="[238,442,1663,1682]" page="242" pageId="13" pageNumber="242" year="1926">
<bibRefCitation id="EFCB4B227A23FFB3FF670757358BB19B" author="ORCHYMONT" box="[238,442,1663,1682]" firstAuthor="ORCHYMONT" page="242" pageId="13" pageNumber="242" pagination="201 - 248" refId="ref18521" refString="ORCHYMONT A. d' 1926 b: Contribution a l'etude des Hydrophilides VI. Bulletin et Annales de la Societe Entomologique de Belgique 66: 201 - 248." type="journal article" year="1926" yearSuffix="b">ORCHYMONT (1926b:242</bibRefCitation>
</treatmentCitation>
, transfer to 
<taxonomicName id="4C5A4D507A23FFB3FD91075736BEB19B" ID-CoL="8X7X2" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[536,655,1663,1682]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A23FFB3FD91075736BEB19B" box="[536,655,1663,1682]" italics="true" pageId="13" pageNumber="242">Thysanarthria</emphasis>
</taxonomicName>
explained in more detail, comparison with 
<taxonomicName id="4C5A4D507A23FFB3FE3D07B2363FB1A4" ID-CoL="56TGX" baseAuthorityName="Regimbart" baseAuthorityYear="1903" box="[436,526,1690,1709]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="atriceps">
<emphasis id="B92EEAC17A23FFB3FE3D07B2363FB1A4" box="[436,526,1690,1709]" italics="true" pageId="13" pageNumber="242">T. atriceps</emphasis>
</taxonomicName>
); 
<treatmentCitation id="0AFB10C27A23FFB3FDA907B236F7B1A4" author="HANSEN M." box="[544,710,1690,1709]" page="105" pageId="13" pageNumber="242" year="1999">
<bibRefCitation id="EFCB4B227A23FFB3FDA907B236F7B1A4" author="HANSEN" box="[544,710,1690,1709]" firstAuthor="HANSEN" page="105" pageId="13" pageNumber="242" pagination="1 - 416" refId="ref18053" refString="HANSEN M. 1999: Hydrophiloidea (s. str.) (Coleoptera). World Catalogue of Insects 2: 1 - 416." type="journal article" year="1999">HANSEN (1999: 105</bibRefCitation>
</treatmentCitation>
, catalogue); 
<treatmentCitation id="0AFB10C27A23FFB3FF7C079D35AEB1C1" author="HEBAUER F." box="[245,415,1717,1736]" page="398" pageId="13" pageNumber="242" year="2001">
<bibRefCitation id="EFCB4B227A23FFB3FF7C079D35AEB1C1" author="HEBAUER" box="[245,415,1717,1736]" firstAuthor="HEBAUER" page="398" pageId="13" pageNumber="242" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">HEBAUER (2001:398</bibRefCitation>
</treatmentCitation>
, redescription and update of distribution).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536D37A23FFB3FF1E07E736A7B1EB" blockId="13.[151,761,1576,1869]" box="[151,662,1743,1762]" pageId="13" pageNumber="242">
<treatmentCitationGroup id="AB4A11FD7A23FFB3FF1E07E736A7B1EB" box="[151,662,1743,1762]" pageId="13" pageNumber="242">
<treatmentCitation id="0AFB10C27A23FFB3FF1E07E7360FB1EB" author="CHIESA A." box="[151,574,1743,1762]" page="276" pageId="13" pageNumber="242" year="1967">
<taxonomicName id="4C5A4D507A23FFB3FF1E07E7360FB1EB" ID-CoL="5XNTC" authority="Chiesa, 1967: 276" authorityName="Chiesa" authorityPageNumber="276" authorityYear="1967" box="[151,574,1743,1762]" class="Insecta" family="Hydrophilidae" genus="Chaetarthriomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B92EEAC17A23FFB3FF1E07E735ACB1EB" box="[151,413,1743,1762]" italics="true" pageId="13" pageNumber="242">Chaetarthriomorphus sulcatus</emphasis>
<bibRefCitation id="EFCB4B227A23FFB3FE2B07E7360FB1EB" author="Chiesa" box="[418,574,1743,1762]" firstAuthor="Chiesa" page="276" pageId="13" pageNumber="242" pagination="275 - 277" refId="ref17771" refString="CHIESA A. 1967: Compimento di una revisione di Hydrophilidae del Afghanistan (Coleoptera: Hydrophilidae). Annales Historico-Naturales Musei Nationalis Hungarici 59: 275 - 277." type="journal article" year="1967">Chiesa, 1967: 276</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 
<emphasis id="B92EEAC17A23FFB3FDCE07E736A7B1EB" bold="true" box="[583,662,1743,1762]" pageId="13" pageNumber="242">
<taxonomicNameLabel id="A21D57BA7A23FFB3FDCE07E736A7B1EB" box="[583,662,1743,1762]" pageId="13" pageNumber="242">syn. nov.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536D37A23FFB3FF1E07C234C9B044" blockId="13.[151,761,1576,1869]" pageId="13" pageNumber="242">
<treatmentCitationGroup id="AB4A11FD7A23FFB3FF1E07C234C9B044" pageId="13" pageNumber="242">
<treatmentCitation id="0AFB10C27A23FFB3FF1E07C23638B1F4" author="HANSEN M." box="[151,521,1770,1789]" page="126" pageId="13" pageNumber="242" year="1991">
<taxonomicName id="4C5A4D507A23FFB3FF1E07C23638B1F4" authority=": HANSEN (1991: 126" authorityName=": HANSEN" authorityYear="1991" box="[151,521,1770,1789]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="sulcata">
<emphasis id="B92EEAC17A23FFB3FF1E07C23564B1F4" box="[151,341,1770,1789]" italics="true" pageId="13" pageNumber="242">Thysanarthria sulcata</emphasis>
: 
<bibRefCitation id="EFCB4B227A23FFB3FEE807C23638B1F4" author="HANSEN" box="[353,521,1770,1789]" firstAuthor="HANSEN" page="126" pageId="13" pageNumber="242" pagination="1 - 368" refId="ref18021" refString="HANSEN M. 1991: The hydrophiloid beetles. Phylogeny, classification and a revision of the genera (Coleoptera, Hydrophiloidea). Biologiske Skrifter 40: 1 - 368." type="journal article" year="1991">HANSEN (1991: 126</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, transfer to 
<taxonomicName id="4C5A4D507A23FFB3FDF907C236DBB1F4" ID-CoL="8X7X2" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[624,746,1770,1789]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A23FFB3FDF907C236DBB1F4" box="[624,746,1770,1789]" italics="true" pageId="13" pageNumber="242">Thysanarthria</emphasis>
</taxonomicName>
); 
<treatmentCitation id="0AFB10C27A23FFB3FF3C062D3568B011" author="HANSEN M." box="[181,345,1797,1816]" page="105" pageId="13" pageNumber="242" year="1999">
<bibRefCitation id="EFCB4B227A23FFB3FF3C062D3568B011" author="HANSEN" box="[181,345,1797,1816]" firstAuthor="HANSEN" page="105" pageId="13" pageNumber="242" pagination="1 - 416" refId="ref18053" refString="HANSEN M. 1999: Hydrophiloidea (s. str.) (Coleoptera). World Catalogue of Insects 2: 1 - 416." type="journal article" year="1999">HANSEN (1999: 105</bibRefCitation>
</treatmentCitation>
, catalogue).All other records of this species refer to different species, see under 
<taxonomicName id="4C5A4D507A23FFB3FE4A06373628B03B" ID-CoL="8YBVV" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[451,537,1823,1842]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="persica" status="sp. nov.">
<emphasis id="B92EEAC17A23FFB3FE4A06373628B03B" box="[451,537,1823,1842]" italics="true" pageId="13" pageNumber="242">T. persica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A23FFB3FDA806373657B03B" box="[545,614,1823,1842]" pageId="13" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C5A4D507A23FFB3FD1B063736C6B03B" ID-CoL="8YBVY" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[658,759,1823,1842]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="wadicola" status="sp. nov.">
<emphasis id="B92EEAC17A23FFB3FD1B063736C6B03B" box="[658,759,1823,1842]" italics="true" pageId="13" pageNumber="242">T. wadicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A23FFB3FF3C061234C9B044" box="[181,248,1850,1869]" pageId="13" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34065587A23FFB3FF1E064C31AFB408" pageId="13" pageNumber="242" type="materials_examined">
<paragraph id="8BE536D37A23FFB3FF1E064C31B0B60F" blockId="13.[151,761,1892,2071]" lastBlockId="13.[828,1438,190,769]" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FF1E064C35B1B07E" bold="true" box="[151,384,1892,1911]" pageId="13" pageNumber="242">Type material examined.</emphasis>
<materialsCitation id="3B323C8E7A23FFB3FE01064C3653B0CE" ID-GBIF-Occurrence="3756360309" collectionCode="BMNH" collectorName="H. G. C. G. C. Champion" country="India" determinerName="Knisch" location="'Ranikhet" municipality="Kumaon" pageId="13" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="UTTARKHAND" typeStatus="lectotype">
<emphasis id="B92EEAC17A23FFB3FE01064C3652B07E" box="[392,611,1892,1911]" italics="true" pageId="13" pageNumber="242">
<taxonomicName id="4C5A4D507A23FFB3FE01064C366EB07E" authorityName="Knisch" authorityYear="1924" box="[392,607,1892,1911]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="championi">
<originalDetermination id="A8448E2F7A23FFB3FE01064C366EB07E" box="[392,607,1892,1911]" pageId="13" pageNumber="242">Chaetarthria championi</originalDetermination>
</taxonomicName>
.
</emphasis>
<typeStatus id="54E188717A23FFB3FDE0064C36F2B07C" box="[617,707,1892,1911]" pageId="13" pageNumber="242" type="lectotype">LECTOTYPE</typeStatus>
(here designated): 
<specimenCount id="9D5CFD5A7A23FFB3FE8C06503524B098" box="[261,277,1912,1937]" pageId="13" pageNumber="242" type="male">♂</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FEA806563554B098" box="[289,357,1918,1937]" pageId="13" pageNumber="242">BMNH</collectionCode>
), 
<emphasis id="B92EEAC17A23FFB3FEFD06563607B098" bold="true" box="[372,566,1918,1937]" pageId="13" pageNumber="242">
<collectingCountry id="F34D76437A23FFB3FEFD06563587B098" box="[372,438,1918,1937]" name="India" pageId="13" pageNumber="242">INDIA</collectingCountry>
: 
<collectingRegion id="499EF8317A23FFB3FE490656361EB099" box="[448,559,1918,1937]" pageId="13" pageNumber="242">UTTARKHAND</collectingRegion>
:
</emphasis>
<location id="8E8560087A23FFB3FDB5065636A0B098" LSID="urn:lsid:plazi:treatment:03F387C57A23FFB0FF6C072130C0B1B6:8E8560087A23FFB3FDB5065636A0B098" box="[572,657,1918,1937]" country="India" municipality="Kumaon" name="'Ranikhet" pageId="13" pageNumber="242" stateProvince="UTTARKHAND">ʻRanikhet</location>
/ 
<collectingMunicipality id="6B81ACA97A23FFB3FD2B065636DAB098" box="[674,747,1918,1937]" pageId="13" pageNumber="242">Kumaon</collectingMunicipality>
/ India, 
<collectorName id="26AF53057A23FFB3FF4406B13584B0A5" box="[205,437,1945,1964]" pageId="13" pageNumber="242">H. G. C. // G. C. Champion</collectorName>
/ Brit. Mus. / 1924-42. // 
<taxonomicName id="4C5A4D507A23FFB3FD0706B136C6B0A5" authorityName="Stephens" authorityYear="1835" box="[654,759,1945,1964]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="251" phylum="Arthropoda" rank="genus">Chaetarthria</taxonomicName>
/ championi / Knisch // det. 
<determinerName id="7E39DA177A23FFB3FE05069C35F9B0CE" box="[396,456,1972,1991]" pageId="13" pageNumber="242">Knisch</determinerName>
/ W. E. Z. 1924ʼ.
</materialsCitation>
<materialsCitation id="3B323C8E7A23FFB3FDE0069C31B0B60F" ID-GBIF-Occurrence="3756360347" pageId="13" pageNumber="242" typeStatus="paralectotype">
<typeStatus id="54E188717A23FFB3FDE0069C36C3B0CC" box="[617,754,1972,1991]" pageId="13" pageNumber="242" type="paralectotype">PARALECTOTYPES</typeStatus>
: 
<specimenCount id="9D5CFD5A7A23FFB3FF1E06E634E7B0E8" box="[151,214,1998,2017]" pageId="13" pageNumber="242" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FF6B06E63517B0E8" box="[226,294,1998,2017]" pageId="13" pageNumber="242">BMNH</collectionCode>
), same label data as the 
<typeStatus id="54E188717A23FFB3FD8A06E63665B0E8" box="[515,596,1998,2017]" pageId="13" pageNumber="242" type="lectotype">lectotype</typeStatus>
; 
<specimenCount id="9D5CFD5A7A23FFB3FDD706E636ACB0E8" box="[606,669,1998,2017]" pageId="13" pageNumber="242" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FD2306E636DEB0E8" box="[682,751,1998,2017]" pageId="13" pageNumber="242">BMNH</collectionCode>
): ʻW. Almora / Kumaon U.P. / 
<collectingCountry id="F34D76437A23FFB3FE1F06C135F5B0F5" box="[406,452,2025,2044]" name="India" pageId="13" pageNumber="242">India</collectingCountry>
, H. G. C. // G. C. Champion / Brit. Mus. / 1924-42. // 
<taxonomicName id="4C5A4D507A23FFB3FEBD092C35ACBF1E" authorityName="Stephens" authorityYear="1835" box="[308,413,2052,2071]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="251" phylum="Arthropoda" rank="genus">Chaetarthria</taxonomicName>
/ championi / Knisch // Knisch det. 1922 / 
<taxonomicName id="4C5A4D507A23FFB3FCC101963780B7D8" authorityName="Stephens" authorityYear="1835" box="[840,945,190,209]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="251" phylum="Arthropoda" rank="genus">Chaetarthria</taxonomicName>
/ championiʼ; 
<specimenCount id="9D5CFD5A7A23FFB3FBBB0196305EB7D8" box="[1074,1135,190,209]" pageId="13" pageNumber="242" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FBF30196308EB7D8" box="[1146,1215,190,209]" pageId="13" pageNumber="242">BMNH</collectionCode>
): ʻW. Almora / Kumaon / 
<collectingCountry id="F34D76437A23FFB3FCB501F1375BB7E5" box="[828,874,217,236]" name="India" pageId="13" pageNumber="242">India</collectingCountry>
, H. G. C. // G. C. Champion / Brit. Mus. / 1924-42. // 
<taxonomicName id="4C5A4D507A23FFB3FABA01F131ADB7E5" authorityName="Stephens" authorityYear="1835" box="[1331,1436,217,236]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="251" phylum="Arthropoda" rank="genus">Chaetarthria</taxonomicName>
/ championi / Knisch // Knisch det. 1922 / 
<taxonomicName id="4C5A4D507A23FFB3FB2F01DB313EB60F" authorityName="Stephens" authorityYear="1835" box="[1190,1295,243,262]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="251" phylum="Arthropoda" rank="genus">Chaetarthria</taxonomicName>
/ championiʼ
</materialsCitation>
</paragraph>
<paragraph id="8BE536D37A23FFB3FCD300263144B685" blockId="13.[828,1438,190,769]" pageId="13" pageNumber="242">
<materialsCitation id="3B323C8E7A23FFB3FCD300263144B685" ID-GBIF-Occurrence="3756360316" collectionCode="HNHM" country="Afghanistan" elevation="1200" location="Bashgultal" municipality="Bashgultal" pageId="13" pageNumber="242" specimenCount="1" specimenCount-male="1" stateProvince="Nuristan" typeStatus="lectotype">
<taxonomicName id="4C5A4D507A23FFB3FCD30026305CB628" authorityName="Chiesa" authorityYear="1967" box="[858,1133,270,289]" class="Insecta" family="Hydrophilidae" genus="Chaetarthriomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B92EEAC17A23FFB3FCD30026305CB628" box="[858,1133,270,289]" italics="true" pageId="13" pageNumber="242">
<originalDetermination id="A8448E2F7A23FFB3FCD30026305CB628" box="[858,1133,270,289]" pageId="13" pageNumber="242">Chaetarthriomorphus sulcatus</originalDetermination>
</emphasis>
</taxonomicName>
: 
<typeStatus id="54E188717A23FFB3FBF5002630E9B616" box="[1148,1240,270,289]" pageId="13" pageNumber="242" type="lectotype">LECTOTYPE</typeStatus>
(here designated): 
<specimenCount id="9D5CFD5A7A23FFB3FA04002F31ACB629" box="[1421,1437,263,288]" pageId="13" pageNumber="242" type="male">♂</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FCCB000137B9B635" box="[834,904,297,316]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="13" pageNumber="242">HNHM</collectionCode>
): 
<emphasis id="B92EEAC17A23FFB3FC1D00013004B635" bold="true" box="[916,1077,297,316]" pageId="13" pageNumber="242">
<collectingCountry id="F34D76437A23FFB3FC1D00013003B635" box="[916,1074,297,316]" name="Afghanistan" pageId="13" pageNumber="242">AFGHANISTAN</collectingCountry>
:
</emphasis>
ʻNO.Afghan.1953 / J. Klapperich // 
<collectingRegion id="499EF8317A23FFB3FAEF00013769B65F" country="Afghanistan" name="Nuristan" pageId="13" pageNumber="242">Nuristan</collectingRegion>
, 
<quantity id="4CA29B367A23FFB3FCD6006B37AFB65F" box="[863,926,323,342]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="242" unit="m" value="1200.0">
<elevation id="0077D1E07A23FFB3FCD6006B37AFB65F" box="[863,926,323,342]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="242" unit="m" value="1200.0">1200 m</elevation>
</quantity>
/ 
<collectingMunicipality id="6B81ACA97A23FFB3FC25006B3038B65F" box="[940,1033,323,342]" pageId="13" pageNumber="242">Bashgultal</collectingMunicipality>
, 20.IV. // Paratypus 1964 / 
<taxonomicName id="4C5A4D507A23FFB3FB6E006B31ADB65F" authorityName="Chiesa" authorityYear="1967" box="[1255,1436,323,342]" class="Insecta" family="Hydrophilidae" genus="Chaetarthriomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="243" phylum="Arthropoda" rank="genus">Chaetarthriomorphus</taxonomicName>
/ sulcatus / Chiesa // 
<taxonomicName id="4C5A4D507A23FFB3FC6F007630C1B678" authorityName="Chiesa" authorityYear="1967" box="[998,1264,350,369]" class="Insecta" family="Hydrophilidae" genus="Chaetarthriomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="243" phylum="Arthropoda" rank="genus">CHAETARTHRIOMORPHUS</taxonomicName>
/ sulcatus / CHIESA / CHIESA DET. // AEDEAGUS / DRAWN BY / P. D. PERKINS’.
</materialsCitation>
</paragraph>
<paragraph id="8BE536D37A23FFB3FCB500BB31AFB408" blockId="13.[828,1438,190,769]" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FCB500BB307BB6AF" bold="true" box="[828,1098,403,422]" pageId="13" pageNumber="242">Additional material examined.</emphasis>
<materialsCitation id="3B323C8E7A23FFB3FBC700BB37B9B6D5" ID-GBIF-Occurrence="3756360318" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FBC700BB30AAB6AF" bold="true" box="[1102,1179,403,422]" pageId="13" pageNumber="242">
<collectingCountry id="F34D76437A23FFB3FBC700BB30A6B6AF" box="[1102,1175,403,422]" name="Nepal" pageId="13" pageNumber="242">NEPAL</collectingCountry>
:
</emphasis>
1 ♂ (
<collectionCode id="ED4BAE167A23FFB3FB4100BB3139B6AF" box="[1224,1288,403,422]" country="Czech Republic" name="National Museum Prague" pageId="13" pageNumber="242">NMPC</collectionCode>
), S Ganesh Himal village,near Kali Sundhara Bazar, 
<quantity id="4CA29B367A23FFB3FBD8008630B7B6C8" box="[1105,1158,430,449]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="242" unit="m" value="700.0">700 m</quantity>
, 
<date id="FFE410137A23FFB3FB0400863132B6C8" box="[1165,1283,430,449]" pageId="13" pageNumber="242" value="1996-05-24" valueMax="1996-05-25" valueMin="1996-05-24">24.–25.v.1996</date>
,lgt.Ahrens, Kulbe &amp; Rulik;
</materialsCitation>
<materialsCitation id="3B323C8E7A23FFB3FC0700E13147B6FF" ID-GBIF-Occurrence="3756360305" latitude="84.49695" longitude="27.56667" pageId="13" pageNumber="242">
1 ♂, 
<specimenCount id="9D5CFD5A7A23FFB3FC3200E13015B6D5" box="[955,1060,457,476]" pageId="13" pageNumber="242" type="generic">3 specimens</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FBB800E1305EB6D5" box="[1073,1135,457,476]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="13" pageNumber="242">SMNS</collectionCode>
): 
<collectingRegion id="499EF8317A23FFB3FBF600E130FFB6D5" box="[1151,1230,457,476]" country="Nepal" name="Narayani" pageId="13" pageNumber="242">Narayani</collectingRegion>
, Sauraha, bank of 
<collectingRegion id="499EF8317A23FFB3FAE600E131ACB6D5" box="[1391,1437,457,476]" country="Nepal" name="Rapti" pageId="13" pageNumber="242">Rapti</collectingRegion>
River, light trap, 180, 
<geoCoordinate id="EE6E50147A23FFB3FC7000CB307BB6FF" box="[1017,1098,483,502]" degrees="84.49695" direction="north" orientation="latitude" pageId="13" pageNumber="242" precision="1" value="84.49695">84.49695</geoCoordinate>
, 
<geoCoordinate id="EE6E50147A23FFB3FBDA00CB3095B6FF" box="[1107,1188,483,502]" degrees="27.56667" direction="east" orientation="longitude" pageId="13" pageNumber="242" precision="1" value="27.56667">27.56667</geoCoordinate>
, 
<date id="FFE410137A23FFB3FB2700CB3120B6FF" box="[1198,1297,483,502]" pageId="13" pageNumber="242" value="2000-04-18">2000-04-18</date>
, A. Weigel;
</materialsCitation>
<materialsCitation id="3B323C8E7A23FFB3FAF200CB30A7B518" ID-GBIF-Occurrence="3756360341" pageId="13" pageNumber="242">
1 ♂ (
<collectionCode id="ED4BAE167A23FFB3FCCB00D637B1B518" box="[834,896,510,529]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="13" pageNumber="242">SMNS</collectionCode>
):same label data but lgt.A. Skale.
</materialsCitation>
<materialsCitation id="3B323C8E7A23FFB3FB1100D631AFB54F" ID-GBIF-Occurrence="3756360311" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FB1100D6316CB518" bold="true" box="[1176,1373,510,529]" pageId="13" pageNumber="242">
<collectingCountry id="F34D76437A23FFB3FB1100D630E8B518" box="[1176,1241,510,529]" name="India" pageId="13" pageNumber="242">INDIA</collectingCountry>
: UTTARANCHAL:
</emphasis>
1 ♂, 
<specimenCount id="9D5CFD5A7A23FFB3FA0E00D637A2B525" pageId="13" pageNumber="242" type="generic">22 specimens</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FC15033137ECB525" box="[924,989,537,556]" country="Czech Republic" name="National Museum Prague" pageId="13" pageNumber="242">NMPC</collectionCode>
): ca. 
<quantity id="4CA29B367A23FFB3FB8D03313006B525" box="[1028,1079,537,556]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="242" unit="km" value="13.0">13 km</quantity>
NW of Nainital, Khaira [= Khairna] bridge, near river, at light, 
<quantity id="4CA29B367A23FFB3FC56031B3026B54F" box="[991,1047,563,582]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="242" unit="m" value="900.0">900 m</quantity>
, 
<date id="FFE410137A23FFB3FBA8031B3097B54F" box="[1057,1190,563,582]" pageId="13" pageNumber="242" value="2003-07-13" valueMax="2003-07-17" valueMin="2003-07-13">13.–17.vii.2003</date>
, lgt. Z. Kejval &amp; M. Trýzna.
</materialsCitation>
<materialsCitation id="3B323C8E7A23FFB3FCB5036630FBB575" ID-GBIF-Occurrence="3756360342" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FCB5036637C9B568" bold="true" box="[828,1016,590,609]" pageId="13" pageNumber="242">
<collectingRegion id="499EF8317A23FFB3FCB5036637C0B556" box="[828,1009,590,609]" country="India" name="Arunachal Pradesh" pageId="13" pageNumber="242">ARUNACHAL PRADESH</collectingRegion>
:
</emphasis>
1 ♂, 
<specimenCount id="9D5CFD5A7A23FFB3FBA4036630A7B568" box="[1069,1174,590,609]" pageId="13" pageNumber="242" type="generic">8 specimens</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FB2A036630D5B568" box="[1187,1252,590,609]" country="Czech Republic" name="National Museum Prague" pageId="13" pageNumber="242">NMPC</collectionCode>
): 
<quantity id="4CA29B367A23FFB3FB7D03663111B568" box="[1268,1312,590,609]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="242" unit="km" value="8.0">8 km</quantity>
S Jamiri-Sesa vicinity, 
<quantity id="4CA29B367A23FFB3FC0A03413789B575" box="[899,952,617,636]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="242" unit="m" value="350.0">350 m</quantity>
, 
<date id="FFE410137A23FFB3FC490341301FB575" box="[960,1070,617,636]" pageId="13" pageNumber="242" value="2006-05-04" valueMax="2006-05-26" valueMin="2006-05-04">4.–26.v.2006</date>
, lgt. P.Pacholatko.
</materialsCitation>
<materialsCitation id="3B323C8E7A23FFB3FB4503413069B5B8" ID-GBIF-Occurrence="3756360306" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FB4503413156B575" bold="true" box="[1228,1383,617,636]" pageId="13" pageNumber="242">
<collectingCountry id="F34D76437A23FFB3FB4503413124B575" box="[1228,1301,617,636]" name="China" pageId="13" pageNumber="242">CHINA</collectingCountry>
: 
<collectingRegion id="499EF8317A23FFB3FA9503413151B573" box="[1308,1376,617,636]" country="China" name="Yunnan" pageId="13" pageNumber="242">YUNNAN</collectingRegion>
:
</emphasis>
1 ♂, 
<specimenCount id="9D5CFD5A7A23FFB3FA1803413756B59F" pageId="13" pageNumber="242" type="generic">2 spec.</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FCF903AB378BB59F" box="[880,954,643,662]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="242" type="Museum">NHMW</collectionCode>
): 
<quantity id="4CA29B367A23FFB3FC4E03AB3034B59F" box="[967,1029,643,662]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="242" unit="km" value="100.0">100 km</quantity>
W of Kunming, Diaolin Nature Reserve, 
<date id="FFE410137A23FFB3FAD303AB374CB5B8" pageId="13" pageNumber="242" value="1993-05-22" valueMax="1993-06-02" valueMin="1993-05-22">22.v.–2. vi.1993</date>
, lgt.E. Jendek &amp; O.Šauša;
</materialsCitation>
<materialsCitation id="3B323C8E7A23FFB3FBD203B637EEB5EF" ID-GBIF-Occurrence="3756360350" latitude="24.611668" longitude="97.65667" pageId="13" pageNumber="242">
1♂, 
<specimenCount id="9D5CFD5A7A23FFB3FB0803B6308AB5B8" box="[1153,1211,670,689]" pageId="13" pageNumber="242" type="generic">2 spec.</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FB4A03B63135B5B8" box="[1219,1284,670,689]" country="Czech Republic" name="National Museum Prague" pageId="13" pageNumber="242">NMPC</collectionCode>
):Tongbiguan vill., near river, at light, 
<quantity id="4CA29B367A23FFB3FC5703913010B5C5" box="[990,1057,697,716]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="13" pageNumber="242" unit="m" value="1340.0">1340 m</quantity>
, 
<geoCoordinate id="EE6E50147A23FFB3FBA30391304EB5C5" box="[1066,1151,697,716]" degrees="24" direction="north" minutes="36.7" orientation="latitude" pageId="13" pageNumber="242" precision="92" value="24.611668">24°36.7ʹN</geoCoordinate>
<geoCoordinate id="EE6E50147A23FFB3FB0F039130EAB5C5" box="[1158,1243,697,716]" degrees="97" direction="east" minutes="39.4" orientation="longitude" pageId="13" pageNumber="242" precision="92" value="97.65667">97°39.4ʹE</geoCoordinate>
, 
<date id="FFE410137A23FFB3FB6D03913155B5C5" box="[1252,1380,697,716]" pageId="13" pageNumber="242" value="2018-06-24" valueMax="2018-06-26" valueMin="2018-06-24">24.–26.vi.2018</date>
, lgt. J. Hájek &amp;J.Růžička.
</materialsCitation>
<materialsCitation id="3B323C8E7A23FFB3FC6803FB31AFB408" ID-GBIF-Occurrence="3756360328" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FC6803FB306BB5EF" bold="true" box="[993,1114,723,742]" pageId="13" pageNumber="242">
<collectingCountry id="F34D76437A23FFB3FC6803FB3064B5EF" box="[993,1109,723,742]" name="Myanmar" pageId="13" pageNumber="242">MYANMAR</collectingCountry>
:
</emphasis>
1♂, 
<specimenCount id="9D5CFD5A7A23FFB3FB0A03FB30D6B5EF" box="[1155,1255,723,742]" pageId="13" pageNumber="242" type="generic">6 specimens</specimenCount>
(
<collectionCode id="ED4BAE167A23FFB3FB7903FB310BB5EF" box="[1264,1338,723,742]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="242" type="Museum">NHMW</collectionCode>
): 
<collectingRegion id="499EF8317A23FFB3FACF03FB31ABB5EF" box="[1350,1434,723,742]" country="Myanmar" name="Mandalay" pageId="13" pageNumber="242">Mandalay</collectingRegion>
, ca. 
<quantity id="4CA29B367A23FFB3FCDE03C637BBB408" box="[855,906,750,769]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="242" unit="km" value="50.0">50 km</quantity>
NW Kalaw,Myitsona river, 
<quantity id="4CA29B367A23FFB3FBF903C63094B408" box="[1136,1189,750,769]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="242" unit="m" value="450.0">450 m</quantity>
, 
<date id="FFE410137A23FFB3FB2503C63131B408" box="[1196,1280,750,769]" pageId="13" pageNumber="242" value="1998-10-25">25.x.1998</date>
,lgt.Schillhammer.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34065587A23FFB3FCB302093107B1D1" pageId="13" pageNumber="242" type="description">
<paragraph id="8BE536D37A23FFB3FCB302093107B1D1" blockId="13.[826,1437,801,2072]" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FCB302093015B431" bold="true" box="[826,1060,801,824]" pageId="13" pageNumber="242">
Redescription. 
<emphasis id="B92EEAC17A23FFB3FC6502093015B431" bold="true" box="[1004,1060,801,824]" italics="true" pageId="13" pageNumber="242">Body</emphasis>
</emphasis>
length 1.5–2.0 mm (
<typeStatus id="54E188717A23FFB3FA9802093140B431" box="[1297,1393,801,824]" pageId="13" pageNumber="242" type="holotype">holotype</typeStatus>
<quantity id="4CA29B367A23FFB3FAF202093758B451" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="242" unit="mm" value="1.9">1.9 mm</quantity>
), maximum body width 1.0– 
<quantity id="4CA29B367A23FFB3FB2402693131B451" box="[1197,1280,833,856]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="242" unit="mm" value="1.1">1.1 mm</quantity>
(
<typeStatus id="54E188717A23FFB3FA9802693141B451" box="[1297,1392,833,856]" pageId="13" pageNumber="242" type="holotype">holotype</typeStatus>
1.0 mm). Head and labrum black; pronotum yellowish with vaguely delimited darker central spot of variable extent or in some specimens (incl. 
<typeStatus id="54E188717A23FFB3FBB4028930ABB4B1" box="[1085,1178,929,952]" pageId="13" pageNumber="242" type="holotype">holotype</typeStatus>
) completely dark brown; elytra yellowish with darkened elytral striae, or with darker lateral parts, or uniformly brown (incl. in 
<typeStatus id="54E188717A23FFB3FB7702C9316FB4F1" box="[1278,1374,993,1016]" pageId="13" pageNumber="242" type="holotype">holotype</typeStatus>
); legs yellowish to brown. 
<emphasis id="B92EEAC17A23FFB3FB9405293069B311" bold="true" box="[1053,1112,1025,1048]" italics="true" pageId="13" pageNumber="242">Head</emphasis>
with weak microsculpture on interstices; punctation sparse, each puncture bearing pointed seta. Eyes separated by 3.1× the width of one eye in dorsal view. 
<emphasis id="B92EEAC17A23FFB3FCF2054937D9B371" bold="true" box="[891,1000,1121,1144]" italics="true" pageId="13" pageNumber="242">Pronotum</emphasis>
with sparse setiferous punctation similar to that on head; interstices without microsculpture. 
<emphasis id="B92EEAC17A23FFB3FAD005A931ADB391" bold="true" box="[1369,1436,1153,1176]" italics="true" pageId="13" pageNumber="242">Elytra</emphasis>
with 10 striae sharply impressed except anteromedially (near scutellar shield) where neither striae nor serial punctures are visible (but darker spots may be present, resembling real punctures); intervals weakly convex at midlength and near apex; interval punctation sparse, setiferous; interstices without microsculpture. 
<emphasis id="B92EEAC17A23FFB3FBB404693097B251" bold="true" box="[1085,1190,1345,1368]" italics="true" pageId="13" pageNumber="242">Aedeagus</emphasis>
(
<figureCitation id="13612A567A23FFB3FB3D046930C7B251" box="[1204,1270,1345,1368]" captionStart="Fig" captionStartId="14.[151,182,1334,1353]" captionTargetBox="[177,1428,199,1315]" captionTargetId="figure-13@14.[151,1436,186,1319]" captionTargetPageId="14" captionText="Fig. 7. Male genitalia of Thysanarthria championi (Knisch, 1924).A–E – lectotype of Chaetarthria championi Knisch, 1924 (A–C – whole aedeagus in dorsal, lateral and ventral view; D – detail of median lobe and parameres in dorsal view; E – same after KOH treatment, with fully everted internal sac). F–J – lectotype of Chaetarthriomorphus sulcatus Chiesa, 1967 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488938" httpUri="https://zenodo.org/record/4488938/files/figure.png" pageId="13" pageNumber="242">Figs 7</figureCitation>
A–J) c. 
<quantity id="4CA29B367A23FFB3FAC2046931ADB251" box="[1355,1436,1345,1368]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="242" unit="mm" value="0.5">0.5 mm</quantity>
long. Phallobase not much wider at base of parameres than more basally, only indistinctly narrowed at midlength; arcuate in lateral view. Paremere wide basally, gradually narrowing towards apex, outer face nearly continuously arcuate, slightly bisinuate in apical fourth, apex bluntly pointed; mesal face with short cuticular asperities (visible in cleaned aedeagus only, 
<figureCitation id="13612A567A23FFB3FBD20709309EB131" box="[1115,1199,1569,1592]" captionStart="Fig" captionStartId="14.[151,182,1334,1353]" captionTargetBox="[177,1428,199,1315]" captionTargetId="figure-13@14.[151,1436,186,1319]" captionTargetPageId="14" captionText="Fig. 7. Male genitalia of Thysanarthria championi (Knisch, 1924).A–E – lectotype of Chaetarthria championi Knisch, 1924 (A–C – whole aedeagus in dorsal, lateral and ventral view; D – detail of median lobe and parameres in dorsal view; E – same after KOH treatment, with fully everted internal sac). F–J – lectotype of Chaetarthriomorphus sulcatus Chiesa, 1967 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488938" httpUri="https://zenodo.org/record/4488938/files/figure.png" pageId="13" pageNumber="242">Fig. 7E</figureCitation>
). Median lobe widely bottle-shaped, without paired subapical projections; apex nearly reaching level of parameral apices, membranous, rounded in relaxed position (
<figureCitation id="13612A567A23FFB3FBE707A930F6B191" box="[1134,1223,1665,1688]" captionStart="Fig" captionStartId="14.[151,182,1334,1353]" captionTargetBox="[177,1428,199,1315]" captionTargetId="figure-13@14.[151,1436,186,1319]" captionTargetPageId="14" captionText="Fig. 7. Male genitalia of Thysanarthria championi (Knisch, 1924).A–E – lectotype of Chaetarthria championi Knisch, 1924 (A–C – whole aedeagus in dorsal, lateral and ventral view; D – detail of median lobe and parameres in dorsal view; E – same after KOH treatment, with fully everted internal sac). F–J – lectotype of Chaetarthriomorphus sulcatus Chiesa, 1967 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488938" httpUri="https://zenodo.org/record/4488938/files/figure.png" pageId="13" pageNumber="242">Figs 7D</figureCitation>
, I–J), with a pair of backwards directed lobes when fully everted (not usually visible); gonopore rounded, situated subapically.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A23FFB0FCB507C936A0B12D" lastPageId="14" lastPageNumber="243" pageId="13" pageNumber="242" type="diagnosis">
<paragraph id="8BE536D37A23FFB0FCB507C936A0B12D" blockId="13.[826,1437,801,2072]" lastBlockId="14.[151,760,1485,2084]" lastPageId="14" lastPageNumber="243" pageId="13" pageNumber="242">
<emphasis id="B92EEAC17A23FFB3FCB507C93018B1F1" bold="true" box="[828,1065,1761,1784]" pageId="13" pageNumber="242">Differential diagnosis.</emphasis>
<taxonomicName id="4C5A4D507A23FFB3FBA407C93100B1F1" authorityName=": ORCHYMONT" authorityYear="1926" box="[1069,1329,1761,1784]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B92EEAC17A23FFB3FBA407C93100B1F1" box="[1069,1329,1761,1784]" italics="true" pageId="13" pageNumber="242">Thysanarthria championi</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="4C5A4D507A23FFB3FCB5062937A0B011" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[828,913,1793,1816]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
<emphasis id="B92EEAC17A23FFB3FCB5062937A0B011" box="[828,913,1793,1816]" italics="true" pageId="13" pageNumber="242">T. bifida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A23FFB3FC11062937DDB011" box="[920,1004,1793,1816]" pageId="13" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C5A4D507A23FFB3FC7F06293061B011" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[1014,1104,1793,1816]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="trifida" status="sp. nov.">
<emphasis id="B92EEAC17A23FFB3FC7F06293061B011" box="[1014,1104,1793,1816]" italics="true" pageId="13" pageNumber="242">T. trifida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A23FFB3FBDE0629309AB011" box="[1111,1195,1793,1816]" pageId="13" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
, and 
<taxonomicName id="4C5A4D507A23FFB3FB6B06293118B011" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[1250,1321,1793,1816]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="chui" status="sp. nov.">
<emphasis id="B92EEAC17A23FFB3FB6B06293118B011" box="[1250,1321,1793,1816]" italics="true" pageId="13" pageNumber="242">T. chui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A23FFB3FAB9062931B3B011" box="[1328,1410,1793,1816]" pageId="13" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
in the general morphology of the aedeagus and the basally wide parameres arcuately narrowing into widely to narrowly rounded apex; of these 
<taxonomicName id="4C5A4D507A23FFB3FBA5064930B0B071" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[1068,1153,1889,1912]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="B92EEAC17A23FFB3FBA5064930B0B071" box="[1068,1153,1889,1912]" italics="true" pageId="13" pageNumber="242">T. bifida</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5A4D507A23FFB3FB3B0649313AB071" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[1202,1291,1889,1912]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="trifida">
<emphasis id="B92EEAC17A23FFB3FB3B0649313AB071" box="[1202,1291,1889,1912]" italics="true" pageId="13" pageNumber="242">T. trifida</emphasis>
</taxonomicName>
can be distinguished by the presence of a pair of subapical projections on the median lobe (absent in 
<taxonomicName id="4C5A4D507A23FFB3FBF006893130B0B1" authorityName=": ORCHYMONT" authorityYear="1926" box="[1145,1281,1953,1976]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B92EEAC17A23FFB3FBF006893130B0B1" box="[1145,1281,1953,1976]" italics="true" pageId="13" pageNumber="242">T. championi</emphasis>
</taxonomicName>
); 
<taxonomicName id="4C5A4D507A23FFB3FA910689316EB0B1" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[1304,1375,1953,1976]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="chui">
<emphasis id="B92EEAC17A23FFB3FA910689316EB0B1" box="[1304,1375,1953,1976]" italics="true" pageId="13" pageNumber="242">T. chui</emphasis>
</taxonomicName>
lacks these lobes, but its parameres are relatively shorter and more abruptly narrowed apically, projecting into rounded lobes. 
<taxonomicName id="4C5A4D507A23FFB3FCB509293071BF11" authorityName=": ORCHYMONT" authorityYear="1926" box="[828,1088,2049,2072]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="242" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B92EEAC17A23FFB3FCB509293071BF11" box="[828,1088,2049,2072]" italics="true" pageId="13" pageNumber="242">Thysanarthria championi</emphasis>
</taxonomicName>
is one of four species co-occurring in Himalaya, along with 
<taxonomicName id="4C5A4D507A20FFB0FE2A04E53607B2ED" authorityName="Hebauer" authorityYear="2001" box="[419,566,1485,1508]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="madurensis">
<emphasis id="B92EEAC17A20FFB0FE2A04E53607B2ED" box="[419,566,1485,1508]" italics="true" pageId="14" pageNumber="243">T. madurensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5A4D507A20FFB0FDCC04E536E0B2ED" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[581,721,1485,1508]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="saurahana" status="sp. nov.">
<emphasis id="B92EEAC17A20FFB0FDCC04E536E0B2ED" box="[581,721,1485,1508]" italics="true" pageId="14" pageNumber="243">T. saurahana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A20FFB0FD5304E534F7B10D" pageId="14" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
, and 
<taxonomicName id="4C5A4D507A20FFB0FF7704C5354FB10D" authorityName="Hebauer" authorityYear="2001" box="[254,382,1517,1540]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B92EEAC17A20FFB0FF7704C5354FB10D" box="[254,382,1517,1540]" italics="true" pageId="14" pageNumber="243">T. siamensis</emphasis>
</taxonomicName>
; it can be easily distinguished from all of them by the morphology of male genitalia.
</paragraph>
</subSubSection>
<caption id="DF25665B7A20FFB0FF1E041E3624B290" ID-DOI="http://doi.org/10.5281/zenodo.4488938" ID-Zenodo-Dep="4488938" httpUri="https://zenodo.org/record/4488938/files/figure.png" pageId="14" pageNumber="243" startId="14.[151,182,1334,1353]" targetBox="[177,1428,199,1315]" targetPageId="14">
<paragraph id="8BE536D37A20FFB0FF1E041E3624B290" blockId="14.[151,1437,1334,1433]" pageId="14" pageNumber="243">
Fig. 7. Male genitalia of 
<taxonomicName id="4C5A4D507A20FFB0FEE1041E36F6B240" authority="(Knisch, 1924)" baseAuthorityName="Knisch" baseAuthorityYear="1924" box="[360,711,1334,1353]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B92EEAC17A20FFB0FEE1041E3670B240" box="[360,577,1334,1353]" italics="true" pageId="14" pageNumber="243">Thysanarthria championi</emphasis>
(
<bibRefCitation id="EFCB4B227A20FFB0FDC5041E36F0B240" author="KNISCH A." bookContentInfo="306 pp." box="[588,705,1334,1353]" editor="JUNK W. &amp; SCHENKLING S." journalOrPublisher="W. Junk, Berlin" pageId="14" pageNumber="243" refId="ref18391" refString="KNISCH A. 1924 a: Hydrophilidae. In: JUNK W. &amp; SCHENKLING S. (eds): Coleopterorum Catalogus. Vol. 14, part 79. W. Junk, Berlin, 306 pp." title="Hydrophilidae" type="book chapter" volumeTitle="Coleopterorum Catalogus. Vol. 14, part 79." year="1924">Knisch, 1924</bibRefCitation>
)
</taxonomicName>
.A–E – lectotype of 
<taxonomicName id="4C5A4D507A20FFB0FCFA041E3089B240" authority="Knisch, 1924" authorityName="Knisch" authorityYear="1924" box="[883,1208,1334,1353]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B92EEAC17A20FFB0FCFA041E3070B240" box="[883,1089,1334,1353]" italics="true" pageId="14" pageNumber="243">Chaetarthria championi</emphasis>
<bibRefCitation id="EFCB4B227A20FFB0FBCF041E3089B240" author="KNISCH A." bookContentInfo="306 pp." box="[1094,1208,1334,1353]" editor="JUNK W. &amp; SCHENKLING S." journalOrPublisher="W. Junk, Berlin" pageId="14" pageNumber="243" refId="ref18391" refString="KNISCH A. 1924 a: Hydrophilidae. In: JUNK W. &amp; SCHENKLING S. (eds): Coleopterorum Catalogus. Vol. 14, part 79. W. Junk, Berlin, 306 pp." title="Hydrophilidae" type="book chapter" volumeTitle="Coleopterorum Catalogus. Vol. 14, part 79." year="1924">Knisch, 1924</bibRefCitation>
</taxonomicName>
(A–C – whole aedeagus in dorsal, lateral and ventral view; D – detail of median lobe and parameres in dorsal view; E – same after KOH treatment, with fully everted internal sac). F–J – lectotype of 
<taxonomicName id="4C5A4D507A20FFB0FEB004433686B277" authority="Chiesa, 1967" authorityName="Chiesa" authorityYear="1967" box="[313,695,1387,1406]" class="Insecta" family="Hydrophilidae" genus="Chaetarthriomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B92EEAC17A20FFB0FEB004433671B277" box="[313,576,1387,1406]" italics="true" pageId="14" pageNumber="243">Chaetarthriomorphus sulcatus</emphasis>
<bibRefCitation id="EFCB4B227A20FFB0FDCF04433686B277" author="Chiesa" box="[582,695,1387,1406]" firstAuthor="Chiesa" pageId="14" pageNumber="243" pagination="275 - 277" refId="ref17771" refString="CHIESA A. 1967: Compimento di una revisione di Hydrophilidae del Afghanistan (Coleoptera: Hydrophilidae). Annales Historico-Naturales Musei Nationalis Hungarici 59: 275 - 277." type="journal article" year="1967">Chiesa, 1967</bibRefCitation>
</taxonomicName>
(F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view).
</paragraph>
</caption>
<subSubSection id="C34065587A20FFB0FF1E070535E6BF2D" pageId="14" pageNumber="243" type="discussion">
<paragraph id="8BE536D37A20FFB0FF1E070535E6BF2D" blockId="14.[151,760,1485,2084]" pageId="14" pageNumber="243">
<emphasis id="B92EEAC17A20FFB0FF1E07053588B14D" bold="true" box="[151,441,1581,1604]" pageId="14" pageNumber="243">Comments on synonymy.</emphasis>
The above type specimen of 
<taxonomicName id="4C5A4D507A20FFB0FF1E076535D5B16D" authorityName="Chiesa" authorityYear="1967" box="[151,484,1613,1636]" class="Insecta" family="Hydrophilidae" genus="Chaetarthriomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B92EEAC17A20FFB0FF1E076535D5B16D" box="[151,484,1613,1636]" italics="true" pageId="14" pageNumber="243">Chaetarthriomorphus sulcatus</emphasis>
</taxonomicName>
is the only found in the Klapperich collection in HNHM. It is largely damaged, with prothorax and one elytron completely missing. However, the abdomen and male genitalia are present, and the morphology of the aedeagus corresponds completely with that of the 
<typeStatus id="54E188717A20FFB0FEC707C53581B00D" box="[334,432,1773,1796]" pageId="14" pageNumber="243" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="4C5A4D507A20FFB0FE5307C53659B00D" authorityName="Knisch" authorityYear="1924" box="[474,616,1773,1796]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B92EEAC17A20FFB0FE5307C53659B00D" box="[474,616,1773,1796]" italics="true" pageId="14" pageNumber="243">T. championi</emphasis>
</taxonomicName>
. The elytron is paler, not dark brown, which further supports the fact that 
<taxonomicName id="4C5A4D507A20FFB0FF4C0605350AB04D" authorityName="Chiesa" authorityYear="1967" box="[197,315,1837,1860]" class="Insecta" family="Hydrophilidae" genus="Chaetarthriomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B92EEAC17A20FFB0FF4C0605350AB04D" box="[197,315,1837,1860]" italics="true" pageId="14" pageNumber="243">C. sulcatus</emphasis>
</taxonomicName>
cannot be conspecific with the specimens from southern 
<collectingCountry id="F34D76437A20FFB0FEBD0665356EB06D" box="[308,351,1869,1892]" name="Iran" pageId="14" pageNumber="243">Iran</collectingCountry>
(described here as 
<taxonomicName id="4C5A4D507A20FFB0FDA4066536A7B06D" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[557,662,1869,1892]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="persica" status="sp. nov.">
<emphasis id="B92EEAC17A20FFB0FDA4066536A7B06D" box="[557,662,1869,1892]" italics="true" pageId="14" pageNumber="243">T. persica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A20FFB0FD14066536C0B06D" box="[669,753,1869,1892]" pageId="14" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
) and the northern Arabian Peninsula (described here as 
<taxonomicName id="4C5A4D507A20FFB0FF1E06A53522B0AD" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[151,275,1933,1956]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="wadicola" status="sp. nov.">
<emphasis id="B92EEAC17A20FFB0FF1E06A53522B0AD" box="[151,275,1933,1956]" italics="true" pageId="14" pageNumber="243">T. wadicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A20FFB0FE9006A5355DB0AD" box="[281,364,1933,1956]" pageId="14" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
) as erroneously reported by 
<bibRefCitation id="EFCB4B227A20FFB0FD1306A534EFB0CD" author="HEBAUER" firstAuthor="HEBAUER" pageId="14" pageNumber="243" pagination="255 - 276" refId="ref18079" refString="HEBAUER F. 1997: Annotated checklist of the Hydrophilidae and Helophoridae (Insecta: Coleoptera) of the Arabian Peninsula with a description of a new genus and species. Fauna of Saudi Arabia 16: 255 - 276." type="journal article" year="1997">HEBAUER (1997)</bibRefCitation>
and 
<bibRefCitation id="EFCB4B227A20FFB0FE9F068535CCB0CD" author="FIKACEK" box="[278,509,1965,1988]" etAl="et al." firstAuthor="FIKACEK" pageId="14" pageNumber="243" pagination="135 - 165" refId="ref17868" refString="FIKACEK M., GENTILI E. &amp; SHORT A. E. Z. 2010: Order Coleoptera, family Hydrophilidae. Pp. 135 - 165. In: HARTEN A. VAN (ed.): Arthropod Fauna of the United Arab Emirates. Volume 3. Al Ummah Printing, Publishing, Distribution and Advertising, Abu Dhabi, 700 pp." type="book chapter" year="2010">FIKÁČEK et al. (2010)</bibRefCitation>
. Following these facts, the examined specimen is designated as the 
<typeStatus id="54E188717A20FFB0FDED06E536F8B0ED" box="[612,713,1997,2020]" pageId="14" pageNumber="243" type="lectotype">lectotype</typeStatus>
, and 
<taxonomicName id="4C5A4D507A20FFB0FF1E06C535E8BF0D" authorityName="Chiesa" authorityYear="1967" box="[151,473,2029,2052]" class="Insecta" family="Hydrophilidae" genus="Chaetarthriomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B92EEAC17A20FFB0FF1E06C535E8BF0D" box="[151,473,2029,2052]" italics="true" pageId="14" pageNumber="243">Chaetarthriomorphus sulcatus</emphasis>
</taxonomicName>
is here placed in synonymy with 
<taxonomicName id="4C5A4D507A20FFB0FF44092535E1BF2D" authorityName="Knisch" authorityYear="1924" box="[205,464,2061,2084]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="243" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B92EEAC17A20FFB0FF44092535E1BF2D" box="[205,464,2061,2084]" italics="true" pageId="14" pageNumber="243">Chaetarthria championi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A20FFB0FCB504E031A6B2F6" pageId="14" pageNumber="243" type="biology_ecology">
<paragraph id="8BE536D37A20FFB0FCB504E031A6B2F6" blockId="14.[828,1437,1480,1727]" pageId="14" pageNumber="243">
<emphasis id="B92EEAC17A20FFB0FCB504E037A6B2D6" bold="true" box="[828,919,1480,1503]" pageId="14" pageNumber="243">Biology.</emphasis>
Part of the examined specimens was collected at light on river banks, no more information is available.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A20FFB0FCB5072030C0B1B6" pageId="14" pageNumber="243" type="distribution">
<paragraph id="8BE536D37A20FFB0FCB5072030C0B1B6" blockId="14.[828,1437,1480,1727]" pageId="14" pageNumber="243">
<emphasis id="B92EEAC17A20FFB0FCB5072037E3B116" bold="true" box="[828,978,1544,1567]" pageId="14" pageNumber="243">Distribution.</emphasis>
The species is widely distributed in the foothills of the Himalaya Mts. and the adjacent mountain systems in eastern 
<collectingCountry id="F34D76437A20FFB0FBCA076030F6B156" box="[1091,1223,1608,1631]" name="Afghanistan" pageId="14" pageNumber="243">Afghanistan</collectingCountry>
(Hindukush Mts.), southwestern 
<collectingCountry id="F34D76437A20FFB0FC5907403021B176" box="[976,1040,1640,1663]" name="China" pageId="14" pageNumber="243">China</collectingCountry>
(
<collectingRegion id="499EF8317A20FFB0FB9607403047B176" box="[1055,1142,1640,1663]" country="China" name="Yunnan" pageId="14" pageNumber="243">Yunnan</collectingRegion>
), and 
<collectingCountry id="F34D76437A20FFB0FB3E07403110B176" box="[1207,1313,1640,1663]" name="Myanmar" pageId="14" pageNumber="243">Myanmar</collectingCountry>
. The specimens listed by 
<bibRefCitation id="EFCB4B227A20FFB0FC5E07A030B3B196" author="HEBAUER" box="[983,1154,1672,1695]" firstAuthor="HEBAUER" pageId="14" pageNumber="243" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">HEBAUER (2001)</bibRefCitation>
from 
<collectingCountry id="F34D76437A20FFB0FB3607A030C2B196" box="[1215,1267,1672,1695]" name="Laos" pageId="14" pageNumber="243">Laos</collectingCountry>
are females and hence their identity cannot be confirmed.
</paragraph>
</subSubSection>
</treatment>
</document>