treatments-xml/data/03/E2/87/03E287ED1A68CE41EAFAFB78FE4DD22D.xml

<document id="B8DBD5715CD5520154CA8DB2A80C464E" ID-DOI="10.11646/zootaxa.3931.3.1" ID-GBIF-Dataset="3740223f-a8b0-4fd6-ae57-aec2e7eefde1" ID-ISSN="1175-5326" ID-Zenodo-Dep="244189" ID-ZooBank="5C5A381C-7EB6-4C1F-BAD2-30B45B019E1F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461010551014" checkinUser="plazi" docAuthor="Moraza, María L. &amp; Lindquist, Evert E." docDate="2015" docId="03E287ED1A68CE41EAFAFB78FE4DD22D" docLanguage="en" docName="zt03931p351.pdf" docOrigin="Zootaxa 3931 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Makarovaia ornata Moraza &amp; Lindquist, 2015, new species" docType="treatment" docVersion="8" lastPageNumber="326" masterDocId="FFDBFF951A66CE58EA6DFFB7FFD4D035" masterDocTitle="Systematics and biology of mites associated with neotropical hispine beetles in unfurled leaves of Heliconia, with descriptions of two new genera of the family Melicharidae (Acari: Mesostigmata: Gamasina: Ascoidea)" masterLastPageNumber="351" masterPageNumber="301" pageNumber="315" updateTime="1698566786881" updateUser="plazi">
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<mods:title id="01695D08F7A1C5CCD6DF04A37C71001B">Systematics and biology of mites associated with neotropical hispine beetles in unfurled leaves of Heliconia, with descriptions of two new genera of the family Melicharidae (Acari: Mesostigmata: Gamasina: Ascoidea)</mods:title>
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<mods:namePart id="D69977DC50B8418894AFC750B778E802">Moraza, María L.</mods:namePart>
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<mods:namePart id="E4F0BB4EE41E2B985011CCE3A3546984">Lindquist, Evert E.</mods:namePart>
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<mods:date id="8938FE8F522D34CE7564017CC4F89FB9">2015</mods:date>
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<mods:number id="765F49871670313ACC240E7E1C2DA98D">3931</mods:number>
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<treatment id="03E287ED1A68CE41EAFAFB78FE4DD22D" ID-DOI="http://doi.org/10.5281/zenodo.6115412" ID-GBIF-Taxon="119600159" ID-Zenodo-Dep="6115412" LSID="urn:lsid:plazi:treatment:03E287ED1A68CE41EAFAFB78FE4DD22D" httpUri="http://treatment.plazi.org/id/03E287ED1A68CE41EAFAFB78FE4DD22D" lastPageId="25" lastPageNumber="326" pageId="14" pageNumber="315">
<subSubSection id="C35165701A68CE56EAFAFB78FE4BD53E" pageId="14" pageNumber="315" type="nomenclature">
<paragraph id="8BF436FB1A68CE56EAFAFB78FDC3D4DF" blockId="14.[151,535,1231,1291]" box="[151,535,1231,1258]" pageId="14" pageNumber="315">
<heading id="D0BC81971A68CE56EAFAFB78FDC3D4DF" bold="true" box="[151,535,1231,1258]" fontSize="11" level="1" pageId="14" pageNumber="315" reason="1">
<emphasis id="B93FEAE91A68CE56EAFAFB78FDC3D4DF" bold="true" box="[151,535,1231,1258]" pageId="14" pageNumber="315">
<taxonomicName id="4C4B4D781A68CE56EAFAFB78FE54D4DC" box="[151,384,1231,1257]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B93FEAE91A68CE56EAFAFB78FE54D4DC" bold="true" box="[151,384,1231,1257]" italics="true" pageId="14" pageNumber="315">Makarovaia ornata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C57921A68CE56EBEAFB67FDC3D4DF" box="[391,535,1232,1258]" pageId="14" pageNumber="315" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF436FB1A68CE56EAFAFB45FE4BD53E" blockId="14.[151,535,1231,1291]" box="[151,415,1266,1291]" pageId="14" pageNumber="315">
(
<figureCitation id="13702A7E1A68CE56EAF3FB45FEE1D53E" box="[158,309,1266,1291]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 7" captionStart-2="FIGURES 8 – 15" captionStart-3="FIGURES 16 – 20" captionStart-4="FIGURES 21 – 24" captionStart-5="FIGURES 25 – 28" captionStart-6="FIGURES 29 – 33" captionStart-7="FIGURES 34 – 40" captionStartId-0="15.[151,264,1556,1579]" captionStartId-1="17.[151,264,1946,1969]" captionStartId-2="18.[151,264,1710,1733]" captionStartId-3="19.[151,264,1593,1616]" captionStartId-4="20.[151,264,1968,1991]" captionStartId-5="21.[151,264,1659,1682]" captionStartId-6="22.[151,264,1838,1861]" captionStartId-7="24.[151,264,1946,1969]" captionTargetBox-0="[163,1436,203,1519]" captionTargetBox-1="[175,1415,424,1907]" captionTargetBox-2="[155,1436,196,1664]" captionTargetBox-3="[152,1428,559,1592]" captionTargetBox-4="[160,1426,682,1929]" captionTargetBox-5="[166,1429,207,1619]" captionTargetBox-6="[156,1431,193,1800]" captionTargetBox-7="[151,1431,409,1907]" captionTargetId-0="figure@15.[151,1436,193,1536]" captionTargetId-1="figure@17.[158,1428,409,1925]" captionTargetId-2="figure@18.[151,1436,193,1689]" captionTargetId-3="figure@19.[151,1436,553,1611]" captionTargetId-4="figure@20.[151,1436,661,1947]" captionTargetId-5="figure@21.[151,1436,193,1638]" captionTargetId-6="figure@22.[151,1436,193,1817]" captionTargetId-7="figure@24.[151,1436,409,1925]" captionTargetPageId-0="15" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionTargetPageId-7="24" captionText-0="FIGURES 1 – 4. Makarovaia ornata n. sp., adult female: 1. Idiosoma, dorsal aspect; 2. Gnathotectum; 3. Spermathecal apparatus, sclerotised part; 4. Detail of separate views of dorsal setae denoted." captionText-1="FIGURES 5 – 7. Makarovaia ornata n. sp., adult female: 5. Idiosoma, ventral aspect; 6. Detail of cheliceral dorsal setae; 7. Chelicera, lateral view." captionText-2="FIGURES 8 – 15. Makarovaia ornata n. sp., adult female: 8 a. Palp, dorsal view, palptarsus removed; 8 b. Palptarsus, dorsal view; 8 c. Palp claw; 9. Detail of corniculus and salivary stylus, dorsal view; 10. Subcapitulum; 11. Tritosternum. 12 – 15: right legs I – IV, excepting tarsi, dorsal view: 12. Leg I; 13. Leg II; 14. Leg III; 15. Leg IV (solid black color refers to dorsal setae)." captionText-3="FIGURES 16 – 20. Makarovaia ornata n. sp.: 16. Tarsus II distally and detail of pretarsus, dorsal view, adult female; 17. Tarsus II, ventral view, adult female, right leg; 18. Tarsus II, adult male, right leg; 19. Tarsus I, dorsal view, adult female, right leg (basal dorsal setae in solid black color); 20. Detail of sensorial apical setae of tarsus I, adult female." captionText-4="FIGURES 21 – 24. Makarovaia ornata n. sp., adult male: 21. Idiosoma, dorsal aspect; 22. Gnathotectum; 23. Detail of cheliceral fixed digit, antaxial view; 24. Chelicera, paraxial view (arthrodial process visible)." captionText-5="FIGURES 25 – 28. Makarovaia ornata n. sp.: 25. Idiosoma, ventral aspect, adult male; 26. Detail of tarsus II, ventro – apical process, ventral view, male deutonymph; 27. Detail of hyaline basal process of corniculus, adult male; 28. Subcapitulum, adult male." captionText-6="FIGURES 29 – 33. Makarovaia ornata n. sp.: 29. Deutonymph, idiosoma, dorsal aspect; 30. Deutonymph, idiosoma, ventral aspect; 31. Deutonymph, detail of corniculus with hyaline process; 32. Protonymph, idiosoma, dorsal aspect; 33. Protonymph, idiosoma, ventral aspect .." captionText-7="FIGURES 34 – 40. Makarovaia ornata n. sp.: 34. Larva, idiosoma, dorsal aspect; 35. Larva, idiosoma, ventral aspect; 36. Larva, gnathotectum; 37. Detail of subcapitulum: a, larva (s. s. – indicates salivary stylus); b, protonymph; 38. Chelicera, lateral view, larva; 39. Chelicera, lateral view, protonymph (arrow points to mucro); 40. Chelicera, lateral view, male deutonymph." httpUri-0="https://zenodo.org/record/244190/files/figure.png" httpUri-1="https://ze
, 
<figureCitation id="13702A7E1A68CE56EB2CFB44FE43D53E" box="[321,407,1267,1291]" captionStart="FIGURES 98 – 101" captionStartId="42.[151,264,1744,1767]" captionTargetBox="[153,1431,185,1707]" captionTargetId="figure@42.[151,1436,180,1730]" captionTargetPageId="42" captionText="FIGURES 98 – 101. Adult idiosomas, ventral view. 98, 99. Makarovaia ornata n. sp.: 98. Female, idiosoma, ventral view; 99. Male, idiosoma, ventral view with attached fungi. 100, 101. Detail of sternal shield, female: 100. Makarovaia ornata n. sp; 101. Hispiniphis parva n. sp." httpUri="https://zenodo.org/record/244210/files/figure.png" pageId="14" pageNumber="315">98–100</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35165701A68CE56EAFAFA8DFC2DD68E" pageId="14" pageNumber="315" type="diagnosis">
<paragraph id="8BF436FB1A68CE56EAFAFA8DFC2DD68E" blockId="14.[151,1437,1338,2011]" pageId="14" pageNumber="315">
<emphasis id="B93FEAE91A68CE56EAFAFA8DFDFCD566" bold="true" box="[151,552,1338,1363]" pageId="14" pageNumber="315">
Diagnosis. 
<emphasis id="B93FEAE91A68CE56EB75FA8DFDCBD566" bold="true" box="[280,543,1338,1363]" italics="true" pageId="14" pageNumber="315">Adult female and male</emphasis>
:
</emphasis>
Dorsal shield reticulated marginally, with punctate central region, and with 18 pairs of smooth, similar, moderately short setae, except minute 
<emphasis id="B93FEAE91A68CE56E90CFAE8FCA8D543" box="[865,892,1375,1398]" italics="true" pageId="14" pageNumber="315">J5</emphasis>
barbed basally (
<figureCitation id="13702A7E1A68CE56EE5BFAE9FBA8D542" box="[1078,1148,1374,1399]" captionStart="FIGURES 1 – 4" captionStartId="15.[151,264,1556,1579]" captionTargetBox="[163,1436,203,1519]" captionTargetId="figure@15.[151,1436,193,1536]" captionTargetPageId="15" captionText="FIGURES 1 – 4. Makarovaia ornata n. sp., adult female: 1. Idiosoma, dorsal aspect; 2. Gnathotectum; 3. Spermathecal apparatus, sclerotised part; 4. Detail of separate views of dorsal setae denoted." httpUri="https://zenodo.org/record/244190/files/figure.png" pageId="14" pageNumber="315">Fig. 4</figureCitation>
). Peritremes extending to posterior margin of coxae II in female and anterior border of coxae III in male. 
<emphasis id="B93FEAE91A68CE56EE7CFA35FB7AD5AE" bold="true" box="[1041,1198,1410,1435]" pageId="14" pageNumber="315">
<emphasis id="B93FEAE91A68CE56EE7CFA35FB71D5AE" bold="true" box="[1041,1189,1410,1435]" italics="true" pageId="14" pageNumber="315">Adult female</emphasis>
:
</emphasis>
Sternal shield lateral remnants reticulated, as long as epigynal shield; epigynal shield drop-shaped, posteriorly rounded, with elongate reticula; anal shield wider than long. Peritrematal shield fragmented, with anterior free platelet bearing 
<emphasis id="B93FEAE91A68CE56EF77FA7CFA90D5D6" box="[1306,1348,1483,1507]" italics="true" pageId="14" pageNumber="315">gp1</emphasis>
at level of coxae II. Fixed cheliceral digit with 9–11 teeth along masticatory surface, movable digit with five or six teeth. 
<emphasis id="B93FEAE91A68CE56EAFAF9A5FECBD61E" bold="true" box="[151,287,1554,1579]" pageId="14" pageNumber="315">
<emphasis id="B93FEAE91A68CE56EAFAF9A5FEC3D61E" bold="true" box="[151,279,1554,1579]" italics="true" pageId="14" pageNumber="315">Adult male</emphasis>
:
</emphasis>
Sternitigenital shield fully reticulated, with five pairs of setae; anal shield and opisthogastric setae as in female. Chelicera with paraxial arthrodial process at base of movable digit finely ciliate; fixed digit with eight teeth; movable digit with two minute teeth; spermatodactyl as long as middle article (including fixed digit) of cheliceral shaft. 
<emphasis id="B93FEAE91A68CE56EB37F9C9FE15D6A2" bold="true" box="[346,449,1662,1687]" pageId="14" pageNumber="315">
<emphasis id="B93FEAE91A68CE56EB37F9C9FE6DD6A2" bold="true" box="[346,441,1662,1687]" italics="true" pageId="14" pageNumber="315">Nymphs</emphasis>
:
</emphasis>
Opisthosoma normally developed, not reduced in form and size; legs IV extended posterolaterally, inserted normally, far from posterior margin of opisthosoma.
</paragraph>
</subSubSection>
<subSubSection id="C35165701A68CE4FEAAAF971FB17D731" lastPageId="23" lastPageNumber="324" pageId="14" pageNumber="315" type="description">
<paragraph id="8BF436FB1A68CE56EAAAF971FD27D7EE" blockId="14.[151,1437,1338,2011]" pageId="14" pageNumber="315">
<emphasis id="B93FEAE91A68CE56EAAAF971FDD2D6EA" bold="true" box="[199,518,1734,1759]" pageId="14" pageNumber="315">Description. Adult female.</emphasis>
<emphasis id="B93FEAE91A68CE56E87DF971FD32D6EA" box="[528,742,1734,1759]" italics="true" pageId="14" pageNumber="315">Idiosomatic dorum</emphasis>
. Idiosoma oval, 775–850 long, 450–500 wide; dorsal shield 462–500 long, 309–335 at its greatest width at level of setae 
<emphasis id="B93FEAE91A68CE56E950F95CFC82D736" box="[829,854,1771,1795]" italics="true" pageId="14" pageNumber="315">r3</emphasis>
(n = 7); shield with nearly parallel sides, reticulated laterally, changing to transverse lines anteriorly between setae 
<emphasis id="B93FEAE91A68CE56E93EF8B8FCBED712" box="[851,874,1807,1831]" italics="true" pageId="14" pageNumber="315">j1</emphasis>
and 
<emphasis id="B93FEAE91A68CE56E9F2F8B8FC61D712" box="[927,949,1807,1831]" italics="true" pageId="14" pageNumber="315">j3</emphasis>
, and densely punctate elsewhere; glandular poroids at level of setae 
<emphasis id="B93FEAE91A68CE56EBC8F884FE10D77E" box="[421,452,1843,1867]" italics="true" pageId="14" pageNumber="315">Z3</emphasis>
conspicuously captured by horn-like extensions of shield. Dorsal shield with 18 pairs of smooth, similar, moderately short setae, except basally spiculate, minute 
<emphasis id="B93FEAE91A68CE56E986F8E0FBD1D75B" box="[1003,1029,1879,1902]" italics="true" pageId="14" pageNumber="315">J5</emphasis>
(
<figureCitation id="13702A7E1A68CE56EE75F8E1FBB4D75A" box="[1048,1120,1878,1903]" captionStart="FIGURES 1 – 4" captionStartId="15.[151,264,1556,1579]" captionTargetBox="[163,1436,203,1519]" captionTargetId="figure@15.[151,1436,193,1536]" captionTargetPageId="15" captionText="FIGURES 1 – 4. Makarovaia ornata n. sp., adult female: 1. Idiosoma, dorsal aspect; 2. Gnathotectum; 3. Spermathecal apparatus, sclerotised part; 4. Detail of separate views of dorsal setae denoted." httpUri="https://zenodo.org/record/244190/files/figure.png" pageId="14" pageNumber="315">Fig. 2</figureCitation>
); setal lengths: 
<emphasis id="B93FEAE91A68CE56EF71F8E0FAE6D75A" box="[1308,1330,1879,1903]" italics="true" pageId="14" pageNumber="315">j1</emphasis>
(15–20), 
<emphasis id="B93FEAE91A68CE56EAFAF8CCFF04D7A6" box="[151,208,1914,1939]" italics="true" pageId="14" pageNumber="315">j3–j6</emphasis>
, 
<emphasis id="B93FEAE91A68CE56EA8CF8CCFF2ED7A6" box="[225,250,1915,1939]" italics="true" pageId="14" pageNumber="315">z2</emphasis>
, 
<emphasis id="B93FEAE91A68CE56EB66F8CCFEF0D7A6" box="[267,292,1915,1939]" italics="true" pageId="14" pageNumber="315">z4</emphasis>
(17–23), 
<emphasis id="B93FEAE91A68CE56EBFBF8CCFE7BD7A7" box="[406,431,1915,1938]" italics="true" pageId="14" pageNumber="315">z5</emphasis>
, 
<emphasis id="B93FEAE91A68CE56EBADF8CCFE0CD7A6" box="[448,472,1915,1939]" italics="true" pageId="14" pageNumber="315">s4</emphasis>
(16–18), 
<emphasis id="B93FEAE91A68CE56E821F8CCFDB1D7A6" box="[588,613,1915,1939]" italics="true" pageId="14" pageNumber="315">s1</emphasis>
(10–13), 
<emphasis id="B93FEAE91A68CE56E8BAF8CCFD25D7A6" box="[727,753,1915,1939]" italics="true" pageId="14" pageNumber="315">r2</emphasis>
(12), 
<emphasis id="B93FEAE91A68CE56E954F8CCFC86D7A6" box="[825,850,1915,1939]" italics="true" pageId="14" pageNumber="315">r3</emphasis>
(17), 
<emphasis id="B93FEAE91A68CE56E9F6F8CCFC61D7A6" box="[923,949,1915,1939]" italics="true" pageId="14" pageNumber="315">J4</emphasis>
(18–20), 
<emphasis id="B93FEAE91A68CE56EE45F8CCFB91D7A6" box="[1064,1093,1915,1939]" italics="true" pageId="14" pageNumber="315">Z2</emphasis>
, 
<emphasis id="B93FEAE91A68CE56EE3BF8CCFBA7D7A6" box="[1110,1139,1915,1939]" italics="true" pageId="14" pageNumber="315">Z3</emphasis>
, 
<emphasis id="B93FEAE91A68CE56EEE9F8CCFB76D7A6" box="[1156,1186,1915,1939]" italics="true" pageId="14" pageNumber="315">Z4</emphasis>
(14–16), 
<emphasis id="B93FEAE91A68CE56EF78F8CCFAE7D7A7" box="[1301,1331,1915,1938]" italics="true" pageId="14" pageNumber="315">Z5</emphasis>
(20–25), clunal 
<emphasis id="B93FEAE91A68CE56EA89F828FF2AD783" box="[228,254,1951,1974]" italics="true" pageId="14" pageNumber="315">J5</emphasis>
(4–6) (
<figureCitation id="13702A7E1A68CE56EB39F829FE68D782" box="[340,444,1950,1975]" captionStart="FIGURES 1 – 4" captionStartId="15.[151,264,1556,1579]" captionTargetBox="[163,1436,203,1519]" captionTargetId="figure@15.[151,1436,193,1536]" captionTargetPageId="15" captionText="FIGURES 1 – 4. Makarovaia ornata n. sp., adult female: 1. Idiosoma, dorsal aspect; 2. Gnathotectum; 3. Spermathecal apparatus, sclerotised part; 4. Detail of separate views of dorsal setae denoted." httpUri="https://zenodo.org/record/244190/files/figure.png" pageId="14" pageNumber="315">Figs 1, 4</figureCitation>
). Lateral soft cuticle with setae 
<emphasis id="B93FEAE91A68CE56E947F828FC97D783" box="[810,835,1951,1974]" italics="true" pageId="14" pageNumber="315">s5</emphasis>
, 
<emphasis id="B93FEAE91A68CE56E93EF829FCB8D782" box="[851,876,1950,1975]" italics="true" pageId="14" pageNumber="315">s6</emphasis>
, 
<emphasis id="B93FEAE91A68CE56E916F828FC40D783" box="[891,916,1951,1974]" italics="true" pageId="14" pageNumber="315">r5</emphasis>
, 
<emphasis id="B93FEAE91A68CE56E9CEF828FC3DD783" box="[931,1001,1951,1975]" italics="true" pageId="14" pageNumber="315">S1–S5</emphasis>
, 
<emphasis id="B93FEAE91A68CE56E994F828FB91D782" box="[1017,1093,1950,1975]" italics="true" pageId="14" pageNumber="315">R1–R6</emphasis>
and several 
<emphasis id="B93FEAE91A68CE56EEBBF828FAD8D783" box="[1238,1292,1951,1974]" italics="true" pageId="14" pageNumber="315">UR’s</emphasis>
17–21 long. Peritrematal shields free from dorsal shield anteriorly.
</paragraph>
<caption id="DF3466731A69CE57EAFAF9A3FC7BD67F" httpUri="https://zenodo.org/record/244190/files/figure.png" pageId="15" pageNumber="316" targetBox="[163,1436,203,1519]" targetPageId="15">
<paragraph id="8BF436FB1A69CE57EAFAF9A3FC7BD67F" blockId="15.[151,1436,1556,1610]" pageId="15" pageNumber="316">
<emphasis id="B93FEAE91A69CE57EAFAF9A3FEEDD61F" bold="true" box="[151,313,1556,1579]" pageId="15" pageNumber="316">FIGURES 1–4</emphasis>
. 
<taxonomicName id="4C4B4D781A69CE57EB26F9A2FDC2D61E" box="[331,534,1557,1579]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B93FEAE91A69CE57EB26F9A2FDC2D61E" box="[331,534,1557,1579]" italics="true" pageId="15" pageNumber="316">Makarovaia ornata</emphasis>
</taxonomicName>
<emphasis id="B93FEAE91A69CE57E84CF9A1FDB4D61E" bold="true" box="[545,608,1558,1579]" pageId="15" pageNumber="316">
<taxonomicNameLabel id="A20C57921A69CE57E84CF9A1FDB4D61E" box="[545,608,1558,1579]" pageId="15" pageNumber="316" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult female: 1. Idiosoma, dorsal aspect; 2. Gnathotectum; 3. Spermathecal apparatus, sclerotised part; 4. Detail of separate views of dorsal setae denoted.
</paragraph>
</caption>
<paragraph id="8BF436FB1A69CE48EAAAF9C6FCFDD26A" blockId="15.[151,1436,1649,2034]" lastBlockId="16.[151,1437,151,2013]" lastPageId="16" lastPageNumber="317" pageId="15" pageNumber="316">
<emphasis id="B93FEAE91A69CE57EAAAF9C6FE4FD6BC" box="[199,411,1649,1674]" italics="true" pageId="15" pageNumber="316">Idiosomatic venter</emphasis>
. Tritosternum with base variably but similarly as long (25–30) as wide (20–28), with laciniae free for about 90 percent of total length (98–112 excluding base), lacineae scarcely pilose (
<figureCitation id="13702A7E1A69CE57EF55F922FA5AD69B" box="[1336,1422,1685,1710]" captionStart="FIGURES 8 – 15" captionStartId="18.[151,264,1710,1733]" captionTargetBox="[155,1436,196,1664]" captionTargetId="figure@18.[151,1436,193,1689]" captionTargetPageId="18" captionText="FIGURES 8 – 15. Makarovaia ornata n. sp., adult female: 8 a. Palp, dorsal view, palptarsus removed; 8 b. Palptarsus, dorsal view; 8 c. Palp claw; 9. Detail of corniculus and salivary stylus, dorsal view; 10. Subcapitulum; 11. Tritosternum. 12 – 15: right legs I – IV, excepting tarsi, dorsal view: 12. Leg I; 13. Leg II; 14. Leg III; 15. Leg IV (solid black color refers to dorsal setae)." httpUri="https://zenodo.org/record/244192/files/figure.png" pageId="15" pageNumber="316">Fig. 11</figureCitation>
). Presternal area weakly transversely lineate (
<figureCitation id="13702A7E1A69CE57E8F4F90EFD35D6E7" box="[665,737,1721,1746]" captionStart="FIGURES 5 – 7" captionStartId="17.[151,264,1946,1969]" captionTargetBox="[175,1415,424,1907]" captionTargetId="figure@17.[158,1428,409,1925]" captionTargetPageId="17" captionText="FIGURES 5 – 7. Makarovaia ornata n. sp., adult female: 5. Idiosoma, ventral aspect; 6. Detail of cheliceral dorsal setae; 7. Chelicera, lateral view." httpUri="https://zenodo.org/record/244191/files/figure.png" pageId="15" pageNumber="316">Fig. 5</figureCitation>
). Sternal shield with lateral elements reticulate and densely punctate; desclerotised region between elements weakly lineate; lateral element with anterior margin lobulate at base of sternal setae 
<emphasis id="B93FEAE91A69CE57EBE4F8B5FE7DD72F" box="[393,425,1794,1818]" italics="true" pageId="15" pageNumber="316">st1</emphasis>
, posterior margin irregularly convex posterior to 
<emphasis id="B93FEAE91A69CE57E988F8B5FBDCD72F" box="[997,1032,1794,1818]" italics="true" pageId="15" pageNumber="316">
<date id="FFF5103B1A69CE57E988F8B5FBDCD72F" box="[997,1032,1794,1818]" pageId="15" pageNumber="316">iv2</date>
</emphasis>
, paraxial lateral margin with small notch between setae 
<emphasis id="B93FEAE91A69CE57EBECF891FE04D70B" box="[385,464,1830,1854]" italics="true" pageId="15" pageNumber="316">st1–st2</emphasis>
, and adaxial lateral margin with elongated, free, narrow endopodal extension between coxae I–II bearing 
<emphasis id="B93FEAE91A69CE57EB07F8FEFE47D757" box="[362,403,1865,1890]" italics="true" pageId="15" pageNumber="316">gvb</emphasis>
, and with well developed, small endopodal extension between coxae II–III; narrowest width of “entire” sternal shield between legs II (87–112), of which surfaces of both lateral sclerites constitute about 40%; single sclerite length 109–128, narrowest width between legs 
<date id="FFF5103B1A69CE57E90FF825FC1DD79F" box="[866,969,1937,1962]" pageId="15" pageNumber="316" value="1915-02-25">II 15–25</date>
; sternal shield setae 
<emphasis id="B93FEAE91A69CE57EED6F825FB0FD79F" box="[1211,1243,1938,1962]" italics="true" pageId="15" pageNumber="316">st1</emphasis>
(33–37) slightly shorter than 
<emphasis id="B93FEAE91A69CE57EB48F801FE92D7FB" box="[293,326,1974,1998]" italics="true" pageId="15" pageNumber="316">st2</emphasis>
(37–43) (
<figureCitation id="13702A7E1A69CE57EBDAF802FDCED7FB" box="[439,538,1973,1998]" captionStart="FIGURES 98 – 101" captionStartId="42.[151,264,1744,1767]" captionTargetBox="[153,1431,185,1707]" captionTargetId="figure@42.[151,1436,180,1730]" captionTargetPageId="42" captionText="FIGURES 98 – 101. Adult idiosomas, ventral view. 98, 99. Makarovaia ornata n. sp.: 98. Female, idiosoma, ventral view; 99. Male, idiosoma, ventral view with attached fungi. 100, 101. Detail of sternal shield, female: 100. Makarovaia ornata n. sp; 101. Hispiniphis parva n. sp." httpUri="https://zenodo.org/record/244210/files/figure.png" pageId="15" pageNumber="316">Fig. 100</figureCitation>
). Setae 
<emphasis id="B93FEAE91A69CE57E81EF801FD40D7FB" box="[627,660,1974,1998]" italics="true" pageId="15" pageNumber="316">st3</emphasis>
(35–39) and 
<emphasis id="B93FEAE91A69CE57E940F801FC9AD7FB" box="[813,846,1974,1998]" italics="true" pageId="15" pageNumber="316">st4</emphasis>
(30–40) on soft cuticle, gland pore 
<emphasis id="B93FEAE91A69CE57EE8AF801FADBD7FB" box="[1255,1295,1974,1998]" italics="true" pageId="15" pageNumber="316">gv1</emphasis>
and poroids 
<emphasis id="B93FEAE91A69CE57EAFAF86DFF6DD7C7" box="[151,185,2010,2034]" italics="true" pageId="15" pageNumber="316">
<date id="FFF5103B1A69CE57EAFAF86DFF6DD7C7" box="[151,185,2010,2034]" pageId="15" pageNumber="316">iv3</date>
</emphasis>
absent. Endopodal strips well developed alongside coxae III and IV. Epigynal shield ornamented with elongate reticula, 130–142 long, 44–52 at widest region at level of setae 
<emphasis id="B93FEAE91A76CE48E9F8FF2FFC62D09A" box="[917,950,152,175]" italics="true" pageId="16" pageNumber="317">st5</emphasis>
, 18–23 at narrowest middle region, with rounded posterior margin; setae 
<emphasis id="B93FEAE91A76CE48E865FF0AFDFDD0E1" box="[520,553,189,212]" italics="true" pageId="16" pageNumber="317">st5</emphasis>
similarly as long as other sternal setae (33–41). Anal shield reticulated, with broadly rounded anterior margin; greatest width (104–122) at level of 
<emphasis id="B93FEAE91A76CE48E9C1FF57FC00D0CD" box="[940,980,224,248]" italics="true" pageId="16" pageNumber="317">gv3</emphasis>
, about 1.3 its mid-length (84–95); gland pores 
<emphasis id="B93FEAE91A76CE48EAB1FEB3FED0D129" box="[220,260,260,284]" italics="true" pageId="16" pageNumber="317">gv3</emphasis>
at level of posterior border of anus; postanal seta (19–22) as long as para-anal setae (19–22); posterior shield margin throughout with thick cribrum (10–15). Soft cuticle with about nine pairs of opisthogastric setae of similar length, 
<emphasis id="B93FEAE91A76CE48EB23FEFBFEADD151" box="[334,377,332,356]" italics="true" pageId="16" pageNumber="317">JV1</emphasis>
(19–25), 
<emphasis id="B93FEAE91A76CE48EB98FEFBFDF5D151" box="[501,545,332,356]" italics="true" pageId="16" pageNumber="317">JV2</emphasis>
(18–20), 
<emphasis id="B93FEAE91A76CE48E8F0FEFBFCD7D151" box="[669,771,332,356]" italics="true" pageId="16" pageNumber="317">JV3–JV5</emphasis>
, 
<emphasis id="B93FEAE91A76CE48E97AFEFBFC92D151" box="[791,838,332,356]" italics="true" pageId="16" pageNumber="317">ZV2</emphasis>
(16–18) slightly thinner than other ventral setae, 
<emphasis id="B93FEAE91A76CE48EAFAFEC7FED7D1BD" box="[151,259,368,392]" italics="true" pageId="16" pageNumber="317">ZV3–ZV4</emphasis>
(18–19); these setae flanked by several pairs of posteriormost setae of 
<emphasis id="B93FEAE91A76CE48EE32FEC6FB50D1BD" box="[1119,1156,369,392]" italics="true" pageId="16" pageNumber="317">UR</emphasis>
- and 
<emphasis id="B93FEAE91A76CE48EEA2FEC6FB34D1BD" box="[1231,1248,369,392]" italics="true" pageId="16" pageNumber="317">R</emphasis>
-series. Anterior fragment of peritrematal shield bearing 
<emphasis id="B93FEAE91A76CE48E839FE23FDAAD199" box="[596,638,404,428]" italics="true" pageId="16" pageNumber="317">gp1</emphasis>
between coxae I and II; main posterior part of shield consolidated with exopodal strip curving behind coxa IV, and bearing poroid 
<emphasis id="B93FEAE91A76CE48E955FE0FFC88D1E5" box="[824,860,440,464]" italics="true" pageId="16" pageNumber="317">ip2</emphasis>
and gland pore 
<emphasis id="B93FEAE91A76CE48EE78FE0FFBEBD1E5" box="[1045,1087,440,464]" italics="true" pageId="16" pageNumber="317">
gp 
<quantity id="4CB39B1E1A76CE48EE5FFE0FFBEBD1E5" box="[1074,1087,440,464]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="317" unit="in" value="2.0">2</quantity>
</emphasis>
in the middle, and poroids 
<emphasis id="B93FEAE91A76CE48EF16FE0EFA48D1E5" box="[1403,1436,441,464]" italics="true" pageId="16" pageNumber="317">ips</emphasis>
and 
<emphasis id="B93FEAE91A76CE48EAA5FE6BFF38D1C1" box="[200,236,476,500]" italics="true" pageId="16" pageNumber="317">ip3</emphasis>
and gland pore 
<emphasis id="B93FEAE91A76CE48EBC4FE6BFE07D1C1" box="[425,467,476,500]" italics="true" pageId="16" pageNumber="317">gp3</emphasis>
behind stigma; peritremes reaching posterior margin of coxae II (121–135) (
<figureCitation id="13702A7E1A76CE48EF2AFE6BFA5AD1C0" box="[1351,1422,476,501]" captionStart="FIGURES 5 – 7" captionStartId="17.[151,264,1946,1969]" captionTargetBox="[175,1415,424,1907]" captionTargetId="figure@17.[158,1428,409,1925]" captionTargetPageId="17" captionText="FIGURES 5 – 7. Makarovaia ornata n. sp., adult female: 5. Idiosoma, ventral aspect; 6. Detail of cheliceral dorsal setae; 7. Chelicera, lateral view." httpUri="https://zenodo.org/record/244191/files/figure.png" pageId="16" pageNumber="317">Fig. 5</figureCitation>
). Spermathecal apparatus with small, curved tubular proximal section (curved length 
<emphasis id="B93FEAE91A76CE48EE15FDB6FB47D22D" box="[1144,1171,513,536]" italics="true" pageId="16" pageNumber="317">ca</emphasis>
. 25, tubular width 2), widened distally (width nearly 10), discernible within body between coxae III and IV (
<figureCitation id="13702A7E1A76CE48EE32FD93FB76D209" box="[1119,1186,548,573]" captionStart="FIGURES 1 – 4" captionStartId="15.[151,264,1556,1579]" captionTargetBox="[163,1436,203,1519]" captionTargetId="figure@15.[151,1436,193,1536]" captionTargetPageId="15" captionText="FIGURES 1 – 4. Makarovaia ornata n. sp., adult female: 1. Idiosoma, dorsal aspect; 2. Gnathotectum; 3. Spermathecal apparatus, sclerotised part; 4. Detail of separate views of dorsal setae denoted." httpUri="https://zenodo.org/record/244190/files/figure.png" pageId="16" pageNumber="317">Fig. 3</figureCitation>
), solenostome located slightly paraxially from between bases of coxae III and IV.
</paragraph>
<paragraph id="8BF436FB1A76CE48EAAAFDDBFE31D401" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="317">
<taxonomicName id="4C4B4D781A76CE48EAAAFDDBFE82D2B0" box="[199,342,620,645]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="16" pageNumber="317" phylum="Nematoda" rank="genus">
<emphasis id="B93FEAE91A76CE48EAAAFDDBFE82D2B0" box="[199,342,620,645]" italics="true" pageId="16" pageNumber="317">Gnathosoma</emphasis>
</taxonomicName>
. Gnathotectum (
<figureCitation id="13702A7E1A76CE48E87CFDDBFD80D2B1" box="[529,596,620,645]" captionStart="FIGURES 1 – 4" captionStartId="15.[151,264,1556,1579]" captionTargetBox="[163,1436,203,1519]" captionTargetId="figure@15.[151,1436,193,1536]" captionTargetPageId="15" captionText="FIGURES 1 – 4. Makarovaia ornata n. sp., adult female: 1. Idiosoma, dorsal aspect; 2. Gnathotectum; 3. Spermathecal apparatus, sclerotised part; 4. Detail of separate views of dorsal setae denoted." httpUri="https://zenodo.org/record/244190/files/figure.png" pageId="16" pageNumber="317">Fig. 2</figureCitation>
) with dorsal face convex, sparsely lineate, with prominent convex, serrated anterior margin. Cheliceral shaft, excluding basal section 116–130 long, with moderate sized digits (
<figureCitation id="13702A7E1A76CE48EE90FD38FA95D29D" box="[1277,1345,655,680]" captionStart="FIGURES 5 – 7" captionStartId="17.[151,264,1946,1969]" captionTargetBox="[175,1415,424,1907]" captionTargetId="figure@17.[158,1428,409,1925]" captionTargetPageId="17" captionText="FIGURES 5 – 7. Makarovaia ornata n. sp., adult female: 5. Idiosoma, ventral aspect; 6. Detail of cheliceral dorsal setae; 7. Chelicera, lateral view." httpUri="https://zenodo.org/record/244191/files/figure.png" pageId="16" pageNumber="317">Fig. 7</figureCitation>
); dorsal face of fixed digit with basally widened dorsal seta (
<figureCitation id="13702A7E1A76CE48E88BFD03FCFED2F8" box="[742,810,692,717]" captionStart="FIGURES 5 – 7" captionStartId="17.[151,264,1946,1969]" captionTargetBox="[175,1415,424,1907]" captionTargetId="figure@17.[158,1428,409,1925]" captionTargetPageId="17" captionText="FIGURES 5 – 7. Makarovaia ornata n. sp., adult female: 5. Idiosoma, ventral aspect; 6. Detail of cheliceral dorsal setae; 7. Chelicera, lateral view." httpUri="https://zenodo.org/record/244191/files/figure.png" pageId="16" pageNumber="317">Fig. 6</figureCitation>
); fixed digit with offset subapical tooth by apical hook, with 9–11 teeth along masticatory surface; movable digit (45–50) with five or six teeth. Corniculus moderately short (21–27), stout (width 9–11 where inserted), with acuminate paraxial process (17–22), extending beyond main tip of corniculus (
<figureCitation id="13702A7E1A76CE48EB0BFCA8FE68D30D" box="[358,444,799,824]" captionStart="FIGURES 8 – 15" captionStartId="18.[151,264,1710,1733]" captionTargetBox="[155,1436,196,1664]" captionTargetId="figure@18.[151,1436,193,1689]" captionTargetPageId="18" captionText="FIGURES 8 – 15. Makarovaia ornata n. sp., adult female: 8 a. Palp, dorsal view, palptarsus removed; 8 b. Palptarsus, dorsal view; 8 c. Palp claw; 9. Detail of corniculus and salivary stylus, dorsal view; 10. Subcapitulum; 11. Tritosternum. 12 – 15: right legs I – IV, excepting tarsi, dorsal view: 12. Leg I; 13. Leg II; 14. Leg III; 15. Leg IV (solid black color refers to dorsal setae)." httpUri="https://zenodo.org/record/244192/files/figure.png" pageId="16" pageNumber="317">Fig. 10</figureCitation>
); corniculi inserted at level of 
<emphasis id="B93FEAE91A76CE48E94EFC96FCEAD30D" box="[803,830,801,824]" italics="true" pageId="16" pageNumber="317">ca</emphasis>
. 20 µm posterior to insertions of setae 
<emphasis id="B93FEAE91A76CE48EF67FCA8FAE0D30D" box="[1290,1332,799,824]" italics="true" pageId="16" pageNumber="317">hp1</emphasis>
. Internal malae wide basally, bifid, tapered, long (50–64 from base of 
<emphasis id="B93FEAE91A76CE48E90EFCF3FC59D368" box="[867,909,836,861]" italics="true" pageId="16" pageNumber="317">hp1</emphasis>
) extending beyond tip of setae 
<emphasis id="B93FEAE91A76CE48EF6EFCF3FAF8D368" box="[1283,1324,836,861]" italics="true" pageId="16" pageNumber="317">hp1</emphasis>
and with surfaces finely pilose; salivary styli tapered (
<figureCitation id="13702A7E1A76CE48E8FAFCD0FD0AD3B5" box="[663,734,871,896]" captionStart="FIGURES 8 – 15" captionStartId="18.[151,264,1710,1733]" captionTargetBox="[155,1436,196,1664]" captionTargetId="figure@18.[151,1436,193,1689]" captionTargetPageId="18" captionText="FIGURES 8 – 15. Makarovaia ornata n. sp., adult female: 8 a. Palp, dorsal view, palptarsus removed; 8 b. Palptarsus, dorsal view; 8 c. Palp claw; 9. Detail of corniculus and salivary stylus, dorsal view; 10. Subcapitulum; 11. Tritosternum. 12 – 15: right legs I – IV, excepting tarsi, dorsal view: 12. Leg I; 13. Leg II; 14. Leg III; 15. Leg IV (solid black color refers to dorsal setae)." httpUri="https://zenodo.org/record/244192/files/figure.png" pageId="16" pageNumber="317">Fig. 9</figureCitation>
). Labrum long, (60–68 from base of setae 
<emphasis id="B93FEAE91A76CE48EEA1FCD0FB22D3B5" box="[1228,1270,871,896]" italics="true" pageId="16" pageNumber="317">hp1</emphasis>
), extending to palptibia. Subcapitulum with eight multidenticulate (6–8 denticles each) rows of deutosternal denticles. Subcapitulum with setae 
<emphasis id="B93FEAE91A76CE48EBD2FC18FE3ED3FD" box="[447,490,943,968]" italics="true" pageId="16" pageNumber="317">hp1</emphasis>
conspicuously thicker and longer (44–52) than other setae; 
<emphasis id="B93FEAE91A76CE48EEDEFC18FB09D3FD" box="[1203,1245,943,968]" italics="true" pageId="16" pageNumber="317">hp2</emphasis>
(17–22) and 
<emphasis id="B93FEAE91A76CE48EFECFC06FA4FD3FD" box="[1409,1435,945,968]" italics="true" pageId="16" pageNumber="317">pc</emphasis>
(23–30) shorter and thinner than 
<emphasis id="B93FEAE91A76CE48E873FC63FD93D3D8" box="[542,583,980,1005]" italics="true" pageId="16" pageNumber="317">hp3</emphasis>
(30–40). Palpus length (145–155); palptrochanter with inner seta longer (28–32) than outer seta (12–20); palpfemoral seta 
<emphasis id="B93FEAE91A76CE48E8BBFC40FD38D425" box="[726,748,1015,1040]" italics="true" pageId="16" pageNumber="317">al</emphasis>
and palpgenual setae 
<emphasis id="B93FEAE91A76CE48E987FC40FBC8D425" box="[1002,1052,1015,1040]" italics="true" pageId="16" pageNumber="317">al–1</emphasis>
and 
<emphasis id="B93FEAE91A76CE48EE3BFC40FB5CD425" box="[1110,1160,1015,1040]" italics="true" pageId="16" pageNumber="317">al–2</emphasis>
relatively short (11–12) with tapered tips (
<figureCitation id="13702A7E1A76CE48EB0DFBABFE73D401" box="[352,423,1052,1077]" captionStart="FIGURES 8 – 15" captionStartId="18.[151,264,1710,1733]" captionTargetBox="[155,1436,196,1664]" captionTargetId="figure@18.[151,1436,193,1689]" captionTargetPageId="18" captionText="FIGURES 8 – 15. Makarovaia ornata n. sp., adult female: 8 a. Palp, dorsal view, palptarsus removed; 8 b. Palptarsus, dorsal view; 8 c. Palp claw; 9. Detail of corniculus and salivary stylus, dorsal view; 10. Subcapitulum; 11. Tritosternum. 12 – 15: right legs I – IV, excepting tarsi, dorsal view: 12. Leg I; 13. Leg II; 14. Leg III; 15. Leg IV (solid black color refers to dorsal setae)." httpUri="https://zenodo.org/record/244192/files/figure.png" pageId="16" pageNumber="317">Figs 8</figureCitation>
a–c).
</paragraph>
<paragraph id="8BF436FB1A76CE48EAAAFBF6FC2AD688" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="317">
<emphasis id="B93FEAE91A76CE48EAAAFBF6FF28D46D" box="[199,252,1089,1112]" italics="true" pageId="16" pageNumber="317">Legs</emphasis>
. At level from femur to tibia, respectively, legs I as wide as legs IV; widths of femur and tibia: leg 
<date id="FFF5103B1A76CE48EFFEFBF7FF67D448" pageId="16" pageNumber="317" value="1955-01">I 55</date>
–76, 46–60; leg 
<date id="FFF5103B1A76CE48EB02FBD2FE7CD448" box="[367,424,1124,1149]" pageId="16" pageNumber="317" value="1965-02">II 65</date>
–90, 50–69: leg 
<date id="FFF5103B1A76CE48E809FBD2FD71D448" box="[612,677,1124,1149]" pageId="16" pageNumber="317" value="1965-03">III 65</date>
–85, 55–64; leg 
<date id="FFF5103B1A76CE48E90FFBD2FC72D448" box="[866,934,1124,1149]" pageId="16" pageNumber="317" value="1960-04">IV 60</date>
–75, 48–60. Legs I to IV (
<figureCitation id="13702A7E1A76CE48EEB8FBD3FA83D448" box="[1237,1367,1124,1149]" captionStart="FIGURES 8 – 15" captionStartId="18.[151,264,1710,1733]" captionTargetBox="[155,1436,196,1664]" captionTargetId="figure@18.[151,1436,193,1689]" captionTargetPageId="18" captionText="FIGURES 8 – 15. Makarovaia ornata n. sp., adult female: 8 a. Palp, dorsal view, palptarsus removed; 8 b. Palptarsus, dorsal view; 8 c. Palp claw; 9. Detail of corniculus and salivary stylus, dorsal view; 10. Subcapitulum; 11. Tritosternum. 12 – 15: right legs I – IV, excepting tarsi, dorsal view: 12. Leg I; 13. Leg II; 14. Leg III; 15. Leg IV (solid black color refers to dorsal setae)." httpUri="https://zenodo.org/record/244192/files/figure.png" pageId="16" pageNumber="317">Figs 12–15</figureCitation>
) each clearly shorter than dorsal shield length; legs I slightly shorter than legs II–IV, lengths, excluding pretarsi: I 360–380, II 372–405, III 374–410, IV 379–430. Leg I tarsus (91–102) about 1.7–1.8 longer than each of the femur (57–63), genu (52–60), and tibia (53–58). Tarsus I with nearly truncate distal margin, pretarsus (15–23 to base of claws) with large claws (19–23) and pulvillus; tarsus I without markedly elongated setae apically (
<figureCitation id="13702A7E1A76CE48EE88FB43FAECD539" box="[1253,1336,1268,1293]" captionStart="FIGURES 16 – 20" captionStartId="19.[151,264,1593,1616]" captionTargetBox="[152,1428,559,1592]" captionTargetId="figure@19.[151,1436,553,1611]" captionTargetPageId="19" captionText="FIGURES 16 – 20. Makarovaia ornata n. sp.: 16. Tarsus II distally and detail of pretarsus, dorsal view, adult female; 17. Tarsus II, ventral view, adult female, right leg; 18. Tarsus II, adult male, right leg; 19. Tarsus I, dorsal view, adult female, right leg (basal dorsal setae in solid black color); 20. Detail of sensorial apical setae of tarsus I, adult female." httpUri="https://zenodo.org/record/244193/files/figure.png" pageId="16" pageNumber="317">Fig. 19</figureCitation>
). Legs II to IV with tarsus about 1.8–2.1 times as long as tibia. Tarsi II–IV with pair of apical setal processes 
<emphasis id="B93FEAE91A76CE48EF48FAA0FA8CD505" box="[1317,1368,1303,1328]" italics="true" pageId="16" pageNumber="317">ad -1</emphasis>
, 
<emphasis id="B93FEAE91A76CE48EF04FAA0FA48D505" box="[1385,1436,1303,1328]" italics="true" pageId="16" pageNumber="317">pd -1</emphasis>
(10–14) inconspicuous, shorter than length of pretarsus to base of claws (18–27); tarsal ventral projection truncate apically, with small central point (11–13) (
<figureCitation id="13702A7E1A76CE48E8F3FAE8FCFBD54D" box="[670,815,1375,1400]" captionStart="FIGURES 16 – 20" captionStartId="19.[151,264,1593,1616]" captionTargetBox="[152,1428,559,1592]" captionTargetId="figure@19.[151,1436,553,1611]" captionTargetPageId="19" captionText="FIGURES 16 – 20. Makarovaia ornata n. sp.: 16. Tarsus II distally and detail of pretarsus, dorsal view, adult female; 17. Tarsus II, ventral view, adult female, right leg; 18. Tarsus II, adult male, right leg; 19. Tarsus I, dorsal view, adult female, right leg (basal dorsal setae in solid black color); 20. Detail of sensorial apical setae of tarsus I, adult female." httpUri="https://zenodo.org/record/244193/files/figure.png" pageId="16" pageNumber="317">Figs 16, 17</figureCitation>
); pretarsi with paradactyli inconspicuous as narrow sclerotised shafts flanking, and about as long as claws (19–23). Tarsi II–IV with ventral setae 
<emphasis id="B93FEAE91A76CE48EEDAFA32FB3DD5A9" box="[1207,1257,1412,1436]" italics="true" pageId="16" pageNumber="317">av -1</emphasis>
, 
<emphasis id="B93FEAE91A76CE48EE95FA32FAFED5A9" box="[1272,1322,1412,1436]" italics="true" pageId="16" pageNumber="317">pv -1</emphasis>
similar in length to attenuated setae 
<emphasis id="B93FEAE91A76CE48EBDAFA10FE30D5F5" box="[439,484,1447,1472]" italics="true" pageId="16" pageNumber="317">al -2</emphasis>
, 
<emphasis id="B93FEAE91A76CE48EB9CFA10FDCAD5F5" box="[497,542,1447,1472]" italics="true" pageId="16" pageNumber="317">pl -1</emphasis>
, and at least twice as long as other ventral setae (as in 
<figureCitation id="13702A7E1A76CE48EE17FA10FB18D5F5" box="[1146,1228,1447,1472]" captionStart="FIGURES 16 – 20" captionStartId="19.[151,264,1593,1616]" captionTargetBox="[152,1428,559,1592]" captionTargetId="figure@19.[151,1436,553,1611]" captionTargetPageId="19" captionText="FIGURES 16 – 20. Makarovaia ornata n. sp.: 16. Tarsus II distally and detail of pretarsus, dorsal view, adult female; 17. Tarsus II, ventral view, adult female, right leg; 18. Tarsus II, adult male, right leg; 19. Tarsus I, dorsal view, adult female, right leg (basal dorsal setae in solid black color); 20. Detail of sensorial apical setae of tarsus I, adult female." httpUri="https://zenodo.org/record/244193/files/figure.png" pageId="16" pageNumber="317">Fig. 17</figureCitation>
); 
<emphasis id="B93FEAE91A76CE48EE8DFA1EFAC6D5F5" box="[1248,1298,1448,1472]" italics="true" pageId="16" pageNumber="317">av -2</emphasis>
, 
<emphasis id="B93FEAE91A76CE48EF72FA1FFA9CD5F5" box="[1311,1352,1448,1472]" italics="true" pageId="16" pageNumber="317">pv2</emphasis>
similar in shorter length and thicker shape to 
<emphasis id="B93FEAE91A76CE48E80FFA7BFD5BD5D1" box="[610,655,1484,1509]" italics="true" pageId="16" pageNumber="317">al -1</emphasis>
(18), and 
<emphasis id="B93FEAE91A76CE48E97FFA7AFCE7D5D1" box="[786,819,1485,1508]" italics="true" pageId="16" pageNumber="317">mv</emphasis>
with bulbose base and slightly shorter. Coxae I–IV reticulate on ventral surfaces; coxal setae simple, attenuate (
<figureCitation id="13702A7E1A76CE48E95EFA58FCA2D63D" box="[819,886,1519,1544]" captionStart="FIGURES 5 – 7" captionStartId="17.[151,264,1946,1969]" captionTargetBox="[175,1415,424,1907]" captionTargetId="figure@17.[158,1428,409,1925]" captionTargetPageId="17" captionText="FIGURES 5 – 7. Makarovaia ornata n. sp., adult female: 5. Idiosoma, ventral aspect; 6. Detail of cheliceral dorsal setae; 7. Chelicera, lateral view." httpUri="https://zenodo.org/record/244191/files/figure.png" pageId="16" pageNumber="317">Fig. 5</figureCitation>
), coxa I seta 
<emphasis id="B93FEAE91A76CE48EE64FA46FBF0D63D" box="[1033,1060,1521,1544]" italics="true" pageId="16" pageNumber="317">av</emphasis>
(20–22), 
<emphasis id="B93FEAE91A76CE48EEFDFA46FB7FD63D" box="[1168,1195,1521,1544]" italics="true" pageId="16" pageNumber="317">pv</emphasis>
(16–18); coxae II–III 
<emphasis id="B93FEAE91A76CE48EAFAF9A2FF66D619" box="[151,178,1557,1580]" italics="true" pageId="16" pageNumber="317">av</emphasis>
(
<emphasis id="B93FEAE91A76CE48EAAFF9A2FF08D619" box="[194,220,1557,1580]" italics="true" pageId="16" pageNumber="317">ca</emphasis>
. 25), 
<emphasis id="B93FEAE91A76CE48EB72F9A2FEEED619" box="[287,314,1557,1580]" italics="true" pageId="16" pageNumber="317">pv</emphasis>
(
<emphasis id="B93FEAE91A76CE48EB27F9A2FEB0D619" box="[330,356,1557,1580]" italics="true" pageId="16" pageNumber="317">ca</emphasis>
. 22); coxae I–IV with serrate posterior distal margin, and a second ventral serrate ridge parallel to that on coxa I; trochanter I with a dorsal serrate ridge; trochanters I–IV with few denticles on basal margin. Legs I to IV with chaetotactic formulae of segments as described for genus. Leg segments with several setae on small tubercles and thickened, spine-like setae as described for genus (
<figureCitation id="13702A7E1A76CE48E908F9C8FC33D6AD" box="[869,999,1663,1688]" captionStart="FIGURES 8 – 15" captionStartId="18.[151,264,1710,1733]" captionTargetBox="[155,1436,196,1664]" captionTargetId="figure@18.[151,1436,193,1689]" captionTargetPageId="18" captionText="FIGURES 8 – 15. Makarovaia ornata n. sp., adult female: 8 a. Palp, dorsal view, palptarsus removed; 8 b. Palptarsus, dorsal view; 8 c. Palp claw; 9. Detail of corniculus and salivary stylus, dorsal view; 10. Subcapitulum; 11. Tritosternum. 12 – 15: right legs I – IV, excepting tarsi, dorsal view: 12. Leg I; 13. Leg II; 14. Leg III; 15. Leg IV (solid black color refers to dorsal setae)." httpUri="https://zenodo.org/record/244192/files/figure.png" pageId="16" pageNumber="317">Figs 12–15</figureCitation>
). Seta 
<emphasis id="B93FEAE91A76CE48EE5DF936FBB6D6AD" box="[1072,1122,1664,1688]" italics="true" pageId="16" pageNumber="317">pv -2</emphasis>
on trochanters I–IV and 
<emphasis id="B93FEAE91A76CE48EF15F936FF71D689" italics="true" pageId="16" pageNumber="317">av - 1</emphasis>
on trochanters II and IV (
<figureCitation id="13702A7E1A76CE48EBA5F913FD9CD688" box="[456,584,1700,1725]" captionStart="FIGURES 8 – 15" captionStartId="18.[151,264,1710,1733]" captionTargetBox="[155,1436,196,1664]" captionTargetId="figure@18.[151,1436,193,1689]" captionTargetPageId="18" captionText="FIGURES 8 – 15. Makarovaia ornata n. sp., adult female: 8 a. Palp, dorsal view, palptarsus removed; 8 b. Palptarsus, dorsal view; 8 c. Palp claw; 9. Detail of corniculus and salivary stylus, dorsal view; 10. Subcapitulum; 11. Tritosternum. 12 – 15: right legs I – IV, excepting tarsi, dorsal view: 12. Leg I; 13. Leg II; 14. Leg III; 15. Leg IV (solid black color refers to dorsal setae)." httpUri="https://zenodo.org/record/244192/files/figure.png" pageId="16" pageNumber="317">Figs 12–15</figureCitation>
) attenuate but not markedly elongated.
</paragraph>
<paragraph id="8BF436FB1A76CE48EAAAF970FA98D7A1" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="317">
<emphasis id="B93FEAE91A76CE48EAAAF970FE82D6D5" bold="true" box="[199,342,1735,1760]" pageId="16" pageNumber="317">Adult male.</emphasis>
<emphasis id="B93FEAE91A76CE48EB0CF970FD90D6D5" box="[353,580,1735,1760]" italics="true" pageId="16" pageNumber="317">Idiosomatic dorsum</emphasis>
. Idiosoma oval, 
<emphasis id="B93FEAE91A76CE48E966F97EFCF1D6D5" box="[779,805,1737,1760]" italics="true" pageId="16" pageNumber="317">ca</emphasis>
. 600–675 long, 450–500 wide; dorsal shield 465–600 long, 350–437 wide at level of setae 
<emphasis id="B93FEAE91A76CE48E85AF95BFD84D731" box="[567,592,1772,1796]" italics="true" pageId="16" pageNumber="317">r3</emphasis>
(n = 4); shield with ornamentation and complement of pore-like structures similar to that of female, but slightly more oval in shape, and slightly more widened anterolaterally to include anterior fragment of peritrematal shield bearing 
<emphasis id="B93FEAE91A76CE48E8C2F883FD0DD779" box="[687,729,1844,1868]" italics="true" pageId="16" pageNumber="317">gp1</emphasis>
above coxae I and II. Dorsal shield with 18 pairs of setae; form and lengths of setae as in female, including reduced size and form of 
<emphasis id="B93FEAE91A76CE48E9C4F8EFFC10D75A" box="[937,964,1880,1903]" italics="true" pageId="16" pageNumber="317">J5</emphasis>
(4–6). Lateral soft cuticle thickened as in female, usually with four pairs of setae 
<emphasis id="B93FEAE91A76CE48E83FF8CAFDA3D7A1" box="[594,631,1917,1940]" italics="true" pageId="16" pageNumber="317">UR</emphasis>
. Peritrematal shields free from dorsal shield anteriorly (
<figureCitation id="13702A7E1A76CE48EE86F8CBFAEBD7A1" box="[1259,1343,1916,1941]" captionStart="FIGURES 21 – 24" captionStartId="20.[151,264,1968,1991]" captionTargetBox="[160,1426,682,1929]" captionTargetId="figure@20.[151,1436,661,1947]" captionTargetPageId="20" captionText="FIGURES 21 – 24. Makarovaia ornata n. sp., adult male: 21. Idiosoma, dorsal aspect; 22. Gnathotectum; 23. Detail of cheliceral fixed digit, antaxial view; 24. Chelicera, paraxial view (arthrodial process visible)." httpUri="https://zenodo.org/record/244194/files/figure.png" pageId="16" pageNumber="317">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8BF436FB1A76CE49EAAAF817FB0BD150" blockId="16.[151,1437,151,2013]" lastBlockId="17.[151,1437,151,357]" lastPageId="17" lastPageNumber="318" pageId="16" pageNumber="317">
<emphasis id="B93FEAE91A76CE48EAAAF817FE48D78D" box="[199,412,1952,1977]" italics="true" pageId="16" pageNumber="317">Idiosomatic venter</emphasis>
. Tritosternum much as in female, with shorter base (length 14–20, width 18–24) and laciniae free for about 90% of total length (82–96). Presternal ornamentation as in female. Sternitigenital shield conspicuously reticulate (
<figureCitation id="13702A7E1A77CE49EBDAFF20FDDFD085" box="[439,523,151,176]" captionStart="FIGURES 25 – 28" captionStartId="21.[151,264,1659,1682]" captionTargetBox="[166,1429,207,1619]" captionTargetId="figure@21.[151,1436,193,1638]" captionTargetPageId="21" captionText="FIGURES 25 – 28. Makarovaia ornata n. sp.: 25. Idiosoma, ventral aspect, adult male; 26. Detail of tarsus II, ventro – apical process, ventral view, male deutonymph; 27. Detail of hyaline basal process of corniculus, adult male; 28. Subcapitulum, adult male." httpUri="https://zenodo.org/record/244195/files/figure.png" pageId="17" pageNumber="318">Fig. 25</figureCitation>
). Anterior margin of sternitigenital shield lobed on either side of genital opening (
<figureCitation id="13702A7E1A77CE49EAF2FF0BFF27D0E0" box="[159,243,188,213]" captionStart="FIGURES 25 – 28" captionStartId="21.[151,264,1659,1682]" captionTargetBox="[166,1429,207,1619]" captionTargetId="figure@21.[151,1436,193,1638]" captionTargetPageId="21" captionText="FIGURES 25 – 28. Makarovaia ornata n. sp.: 25. Idiosoma, ventral aspect, adult male; 26. Detail of tarsus II, ventro – apical process, ventral view, male deutonymph; 27. Detail of hyaline basal process of corniculus, adult male; 28. Subcapitulum, adult male." httpUri="https://zenodo.org/record/244195/files/figure.png" pageId="17" pageNumber="318">Fig. 25</figureCitation>
); total shield length 318–337, narrowest widths 99–112 and 90–109 at levels between coxae II and coxae IV respectively; lengths setae 
<emphasis id="B93FEAE91A77CE49EB87FF57FDDFD0CD" box="[490,523,224,248]" italics="true" pageId="17" pageNumber="318">st1</emphasis>
(28–34), 
<emphasis id="B93FEAE91A77CE49E817FF57FD1DD0CD" box="[634,713,224,248]" italics="true" pageId="17" pageNumber="318">st2–st3</emphasis>
(35–37), 
<emphasis id="B93FEAE91A77CE49E955FF57FC8CD0CD" box="[824,856,224,248]" italics="true" pageId="17" pageNumber="318">st4</emphasis>
(34), and 
<emphasis id="B93FEAE91A77CE49E9A0FF57FC3AD0C2" box="[973,1006,224,247]" italics="true" pageId="17" pageNumber="318">st5</emphasis>
shorter (26–28). Anal shield 123–130 wide at level of 
<emphasis id="B93FEAE91A77CE49EB26FEB3FEA7D129" box="[331,371,260,284]" italics="true" pageId="17" pageNumber="318">gv3</emphasis>
, and 104–112 long, ornamented as in female; postanal seta (14–18) nearly as long as para-anal setae (16–19). Soft cuticle with nine pairs of opisthogastric setae similar in length (13–19) and shape as in female. Form of peritrematal and exopodal shields posteriorly as in female, peritremes 149 long (
<figureCitation id="13702A7E1A77CE49EE13FEFBFB06D150" box="[1150,1234,332,357]" captionStart="FIGURES 25 – 28" captionStartId="21.[151,264,1659,1682]" captionTargetBox="[166,1429,207,1619]" captionTargetId="figure@21.[151,1436,193,1638]" captionTargetPageId="21" captionText="FIGURES 25 – 28. Makarovaia ornata n. sp.: 25. Idiosoma, ventral aspect, adult male; 26. Detail of tarsus II, ventro – apical process, ventral view, male deutonymph; 27. Detail of hyaline basal process of corniculus, adult male; 28. Subcapitulum, adult male." httpUri="https://zenodo.org/record/244195/files/figure.png" pageId="17" pageNumber="318">Fig. 25</figureCitation>
).
</paragraph>
<caption id="DF3466731A77CE49EAFAF82DFE55D7E5" httpUri="https://zenodo.org/record/244191/files/figure.png" pageId="17" pageNumber="318" targetBox="[175,1415,424,1907]" targetPageId="17">
<paragraph id="8BF436FB1A77CE49EAFAF82DFE55D7E5" blockId="17.[151,1436,1946,2000]" pageId="17" pageNumber="318">
<emphasis id="B93FEAE91A77CE49EAFAF82DFEE8D785" bold="true" box="[151,316,1946,1969]" pageId="17" pageNumber="318">FIGURES 5–7.</emphasis>
<taxonomicName id="4C4B4D781A77CE49EB28F82CFDD8D784" box="[325,524,1947,1969]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B93FEAE91A77CE49EB28F82CFDD8D784" box="[325,524,1947,1969]" italics="true" pageId="17" pageNumber="318">Makarovaia ornata</emphasis>
</taxonomicName>
<emphasis id="B93FEAE91A77CE49E879F82CFD84D785" bold="true" box="[532,592,1947,1968]" pageId="17" pageNumber="318">
<taxonomicNameLabel id="A20C57921A77CE49E879F82CFD84D785" box="[532,592,1947,1968]" pageId="17" pageNumber="318" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult female: 5. Idiosoma, ventral aspect; 6. Detail of cheliceral dorsal setae; 7. Chelicera, lateral view.
</paragraph>
</caption>
<caption id="DF3466731A74CE4AEAFAF919FA51D737" httpUri="https://zenodo.org/record/244192/files/figure.png" pageId="18" pageNumber="319" targetBox="[155,1436,196,1664]" targetPageId="18">
<paragraph id="8BF436FB1A74CE4AEAFAF919FA51D737" blockId="18.[151,1436,1710,1794]" pageId="18" pageNumber="319">
<emphasis id="B93FEAE91A74CE4AEAFAF919FE9DD6F1" bold="true" box="[151,329,1710,1733]" pageId="18" pageNumber="319">FIGURES 8–15.</emphasis>
<taxonomicName id="4C4B4D781A74CE4AEB3FF919FDCED6F0" box="[338,538,1710,1733]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B93FEAE91A74CE4AEB3FF919FDCED6F0" box="[338,538,1710,1733]" italics="true" pageId="18" pageNumber="319">Makarovaia ornata</emphasis>
</taxonomicName>
<emphasis id="B93FEAE91A74CE4AE84FF918FD8AD6F1" bold="true" box="[546,606,1711,1732]" pageId="18" pageNumber="319">
<taxonomicNameLabel id="A20C57921A74CE4AE84FF918FD8AD6F1" box="[546,606,1711,1732]" pageId="18" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult female: 8a. Palp, dorsal view, palptarsus removed; 8b. Palptarsus, dorsal view; 8c. Palp claw; 9. Detail of corniculus and salivary stylus, dorsal view; 10. Subcapitulum; 11. Tritosternum. 12–15: right legs I–IV, excepting tarsi, dorsal view: 12. Leg I; 13. Leg II; 14. Leg III; 15. Leg IV (solid black color refers to dorsal setae).
</paragraph>
</caption>
<paragraph id="8BF436FB1A74CE4BEAAAF89EFE78D0CD" blockId="18.[151,1436,1833,2038]" lastBlockId="19.[151,1436,151,501]" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319">
<taxonomicName id="4C4B4D781A74CE4AEAAAF89EFE82D777" box="[199,342,1833,1858]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="18" pageNumber="319" phylum="Nematoda" rank="genus">
<emphasis id="B93FEAE91A74CE4AEAAAF89EFE82D777" box="[199,342,1833,1858]" italics="true" pageId="18" pageNumber="319">Gnathosoma</emphasis>
</taxonomicName>
. Gnathotectum similar in form to female (
<figureCitation id="13702A7E1A74CE4AE95CF89EFC50D777" box="[817,900,1833,1858]" captionStart="FIGURES 21 – 24" captionStartId="20.[151,264,1968,1991]" captionTargetBox="[160,1426,682,1929]" captionTargetId="figure@20.[151,1436,661,1947]" captionTargetPageId="20" captionText="FIGURES 21 – 24. Makarovaia ornata n. sp., adult male: 21. Idiosoma, dorsal aspect; 22. Gnathotectum; 23. Detail of cheliceral fixed digit, antaxial view; 24. Chelicera, paraxial view (arthrodial process visible)." httpUri="https://zenodo.org/record/244194/files/figure.png" pageId="18" pageNumber="319">Fig. 22</figureCitation>
), 90 long, 118 wide. Cheliceral shaft, excluding basal section, 117–125 long; fixed digit with offset subapical tooth by apical hook followed by eight or nine moderate sized teeth, pilus dentilis indicated only by alveolar remnant; movable digit (39–40) with apical hook and two small teeth; spermatodactyl long (112–125), 
<emphasis id="B93FEAE91A74CE4AE8A1F821FD33D798" box="[716,743,1942,1965]" italics="true" pageId="18" pageNumber="319">ca</emphasis>
. 7 wide at its base and 
<emphasis id="B93FEAE91A74CE4AE991F821FBC3D798" box="[1020,1047,1942,1965]" italics="true" pageId="18" pageNumber="319">ca</emphasis>
. 4 at its apical portion, extending posteroventrally from base of cheliceral shaft (
<figureCitation id="13702A7E1A74CE4AE8F2F80EFCF5D7E7" box="[671,801,1977,2002]" captionStart="FIGURES 21 – 24" captionStartId="20.[151,264,1968,1991]" captionTargetBox="[160,1426,682,1929]" captionTargetId="figure@20.[151,1436,661,1947]" captionTargetPageId="20" captionText="FIGURES 21 – 24. Makarovaia ornata n. sp., adult male: 21. Idiosoma, dorsal aspect; 22. Gnathotectum; 23. Detail of cheliceral fixed digit, antaxial view; 24. Chelicera, paraxial view (arthrodial process visible)." httpUri="https://zenodo.org/record/244194/files/figure.png" pageId="18" pageNumber="319">Figs 23, 24</figureCitation>
); paraxial arthrodial process finely ciliate (20). Corniculi (25–27) more widely spaced basally, with inner acuminate process (22–23) thinner basally than in female (
<figureCitation id="13702A7E1A74CE4AEF56F86AFA5BD7C3" box="[1339,1423,2013,2038]" captionStart="FIGURES 25 – 28" captionStartId="21.[151,264,1659,1682]" captionTargetBox="[166,1429,207,1619]" captionTargetId="figure@21.[151,1436,193,1638]" captionTargetPageId="21" captionText="FIGURES 25 – 28. Makarovaia ornata n. sp.: 25. Idiosoma, ventral aspect, adult male; 26. Detail of tarsus II, ventro – apical process, ventral view, male deutonymph; 27. Detail of hyaline basal process of corniculus, adult male; 28. Subcapitulum, adult male." httpUri="https://zenodo.org/record/244195/files/figure.png" pageId="18" pageNumber="319">Fig. 27</figureCitation>
); internal malae as in female. Subcapitulum (
<figureCitation id="13702A7E1A75CE4BE8E7FF20FD09D085" box="[650,733,151,176]" captionStart="FIGURES 25 – 28" captionStartId="21.[151,264,1659,1682]" captionTargetBox="[166,1429,207,1619]" captionTargetId="figure@21.[151,1436,193,1638]" captionTargetPageId="21" captionText="FIGURES 25 – 28. Makarovaia ornata n. sp.: 25. Idiosoma, ventral aspect, adult male; 26. Detail of tarsus II, ventro – apical process, ventral view, male deutonymph; 27. Detail of hyaline basal process of corniculus, adult male; 28. Subcapitulum, adult male." httpUri="https://zenodo.org/record/244195/files/figure.png" pageId="19" pageNumber="320">Fig. 28</figureCitation>
) with seven rows of deutosternal denticles, form and relative lengths of setae much as in female, e.g. 
<emphasis id="B93FEAE91A75CE4BE83FFF0BFDA9D0E0" box="[594,637,188,213]" italics="true" pageId="19" pageNumber="320">hp1</emphasis>
(41–45) prominent, 
<emphasis id="B93FEAE91A75CE4BE90FFF0BFC58D0E0" box="[866,908,188,213]" italics="true" pageId="19" pageNumber="320">hp2</emphasis>
(13–15) shortest. Palpi (length 
<emphasis id="B93FEAE91A75CE4BEE81FF0AFAD2D0E1" box="[1260,1286,189,212]" italics="true" pageId="19" pageNumber="320">ca</emphasis>
. 125) similar in structure as in female.
</paragraph>
<paragraph id="8BF436FB1A75CE4BEAAAFEB2FC6DD1C1" blockId="19.[151,1436,151,501]" pageId="19" pageNumber="320">
<emphasis id="B93FEAE91A75CE4BEAAAFEB2FF28D129" box="[199,252,261,284]" italics="true" pageId="19" pageNumber="320">Legs</emphasis>
. Relative leg thicknesses and lengths, excluding pretarsi, similar to those of female: I 367–395, II 401–425, III 420–480, IV 442–480. Legs with segment length ratios much as in female, tibia and genu 
<date id="FFF5103B1A75CE4BEF7AFE9FFA96D175" box="[1303,1346,295,320]" pageId="19" pageNumber="320" value="1955-01">I 55</date>
–62 and 57–62 respectively, but femur I longer (65–73). Tarsus I (77–90) bearing pretarsus (13–17 to base of claws), claws 20–25. Tarsi II–IV with pair of apical setal processes 
<emphasis id="B93FEAE91A75CE4BE969FED8FCE3D1BD" box="[772,823,367,392]" italics="true" pageId="19" pageNumber="320">ad -1</emphasis>
, 
<emphasis id="B93FEAE91A75CE4BE92AFED8FCAFD1BD" box="[839,891,367,392]" italics="true" pageId="19" pageNumber="320">pd -1</emphasis>
(14–15), apical ventral process under pretarsus truncate (13–15), apex without central point present in female (
<figureCitation id="13702A7E1A75CE4BE90CFE23FC60D199" box="[865,948,404,429]" captionStart="FIGURES 16 – 20" captionStartId="19.[151,264,1593,1616]" captionTargetBox="[152,1428,559,1592]" captionTargetId="figure@19.[151,1436,553,1611]" captionTargetPageId="19" captionText="FIGURES 16 – 20. Makarovaia ornata n. sp.: 16. Tarsus II distally and detail of pretarsus, dorsal view, adult female; 17. Tarsus II, ventral view, adult female, right leg; 18. Tarsus II, adult male, right leg; 19. Tarsus I, dorsal view, adult female, right leg (basal dorsal setae in solid black color); 20. Detail of sensorial apical setae of tarsus I, adult female." httpUri="https://zenodo.org/record/244193/files/figure.png" pageId="19" pageNumber="320">Fig. 18</figureCitation>
), pretarsus (20–27) and pretarsal structures formed as in female, claws 20–25. Scant male dimorphism discerned among leg setae, except for ventral setae and 
<emphasis id="B93FEAE91A75CE4BEAFAFE6BFF10D1C1" box="[151,196,476,501]" italics="true" pageId="19" pageNumber="320">al -1</emphasis>
on tarsi II–IV more thickly spinelike with a bulbous base (
<figureCitation id="13702A7E1A75CE4BE934FE6BFC78D1C1" box="[857,940,476,501]" captionStart="FIGURES 16 – 20" captionStartId="19.[151,264,1593,1616]" captionTargetBox="[152,1428,559,1592]" captionTargetId="figure@19.[151,1436,553,1611]" captionTargetPageId="19" captionText="FIGURES 16 – 20. Makarovaia ornata n. sp.: 16. Tarsus II distally and detail of pretarsus, dorsal view, adult female; 17. Tarsus II, ventral view, adult female, right leg; 18. Tarsus II, adult male, right leg; 19. Tarsus I, dorsal view, adult female, right leg (basal dorsal setae in solid black color); 20. Detail of sensorial apical setae of tarsus I, adult female." httpUri="https://zenodo.org/record/244193/files/figure.png" pageId="19" pageNumber="320">Fig. 18</figureCitation>
).
</paragraph>
<caption id="DF3466731A75CE4BEAFAF98EFB5CD6B9" httpUri="https://zenodo.org/record/244193/files/figure.png" pageId="19" pageNumber="320" targetBox="[152,1428,559,1592]" targetPageId="19">
<paragraph id="8BF436FB1A75CE4BEAFAF98EFB5CD6B9" blockId="19.[151,1436,1593,1676]" pageId="19" pageNumber="320">
<emphasis id="B93FEAE91A75CE4BEAFAF98EFE87D67B" bold="true" box="[151,339,1593,1616]" pageId="19" pageNumber="320">FIGURES 16–20.</emphasis>
<taxonomicName id="4C4B4D781A75CE4BEB34F98EFDCAD67A" box="[345,542,1593,1615]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B93FEAE91A75CE4BEB34F98EFDCAD67A" box="[345,542,1593,1615]" italics="true" pageId="19" pageNumber="320">Makarovaia ornata</emphasis>
</taxonomicName>
<emphasis id="B93FEAE91A75CE4BE84EF98DFD88D67A" bold="true" box="[547,604,1594,1615]" pageId="19" pageNumber="320">
<taxonomicNameLabel id="A20C57921A75CE4BE84EF98DFD88D67A" box="[547,604,1594,1615]" pageId="19" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: 16. Tarsus II distally and detail of pretarsus, dorsal view, adult female; 17. Tarsus II, ventral view, adult female, right leg; 18. Tarsus II, adult male, right leg; 19. Tarsus I, dorsal view, adult female, right leg (basal dorsal setae in solid black color); 20. Detail of sensorial apical setae of tarsus I, adult female.
</paragraph>
</caption>
<paragraph id="8BF436FB1A75CE4BEAAAF903FCA9D7B4" blockId="19.[151,1436,1716,2029]" pageId="19" pageNumber="320">
<emphasis id="B93FEAE91A75CE4BEAAAF903FEBDD6F8" bold="true" box="[199,361,1716,1741]" pageId="19" pageNumber="320">Deutonymph.</emphasis>
<emphasis id="B93FEAE91A75CE4BEB1CF903FD84D6F8" box="[369,592,1716,1741]" italics="true" pageId="19" pageNumber="320">Idiosomatic dorsum</emphasis>
. Idiosoma broadly oval, length 458– 
<specimenCount id="9D4DFD721A75CE4BE99DF903FBCED6F8" box="[1008,1050,1716,1741]" pageId="19" pageNumber="320" type="female">512</specimenCount>
female, 
<specimenCount id="9D4DFD721A75CE4BEE11F903FB33D6F8" box="[1148,1255,1716,1741]" pageId="19" pageNumber="320" type="male">539 male</specimenCount>
, width 320– 
<specimenCount id="9D4DFD721A75CE4BEF1EF903FA48D6F8" box="[1395,1436,1716,1741]" pageId="19" pageNumber="320" type="female">365</specimenCount>
female, 
<specimenCount id="9D4DFD721A75CE4BEA9BF96FFEB1D6C4" box="[246,357,1752,1777]" pageId="19" pageNumber="320" type="male">445 male</specimenCount>
, dorsal shielding not clearly delineated, without lateral incisions (
<figureCitation id="13702A7E1A75CE4BEE1FF96FFB18D6C5" box="[1138,1228,1752,1777]" captionStart="FIGURES 29 – 33" captionStartId="22.[151,264,1838,1861]" captionTargetBox="[156,1431,193,1800]" captionTargetId="figure@22.[151,1436,193,1817]" captionTargetPageId="22" captionText="FIGURES 29 – 33. Makarovaia ornata n. sp.: 29. Deutonymph, idiosoma, dorsal aspect; 30. Deutonymph, idiosoma, ventral aspect; 31. Deutonymph, detail of corniculus with hyaline process; 32. Protonymph, idiosoma, dorsal aspect; 33. Protonymph, idiosoma, ventral aspect .." httpUri="https://zenodo.org/record/244196/files/figure.png" pageId="19" pageNumber="320">Fig. 29</figureCitation>
); dorsal shielding similarly ornamented as in female in matured specimens, but with fainter rugose cuticle when young. Most dorsal setae short (7–20), clunals 
<emphasis id="B93FEAE91A75CE4BEBBAF896FE25D70D" box="[471,497,1825,1848]" italics="true" pageId="19" pageNumber="320">J5</emphasis>
barbed basally and minute (3–5); central setae 
<emphasis id="B93FEAE91A75CE4BEE44F897FBEBD70D" box="[1065,1087,1824,1848]" italics="true" pageId="19" pageNumber="320">j1</emphasis>
(9–14), 
<emphasis id="B93FEAE91A75CE4BEEC5F897FB6AD70D" box="[1192,1214,1824,1848]" italics="true" pageId="19" pageNumber="320">j3</emphasis>
(12–17), 
<emphasis id="B93FEAE91A75CE4BEF58F897FA9FD70D" box="[1333,1355,1824,1848]" italics="true" pageId="19" pageNumber="320">j4</emphasis>
, 
<emphasis id="B93FEAE91A75CE4BEF30F896FAA7D70D" box="[1373,1395,1825,1848]" italics="true" pageId="19" pageNumber="320">j5</emphasis>
, 
<emphasis id="B93FEAE91A75CE4BEFEBF897FA48D70C" box="[1414,1436,1824,1849]" italics="true" pageId="19" pageNumber="320">j6</emphasis>
(15–17), 
<emphasis id="B93FEAE91A75CE4BEB6DF8F2FECFD768" box="[256,283,1861,1885]" italics="true" pageId="19" pageNumber="320">J4</emphasis>
(11–15); 
<emphasis id="B93FEAE91A75CE4BEBE0F8F2FE72D768" box="[397,422,1861,1885]" italics="true" pageId="19" pageNumber="320">z2</emphasis>
(13–14), 
<emphasis id="B93FEAE91A75CE4BE874F8F2FDE6D768" box="[537,562,1861,1885]" italics="true" pageId="19" pageNumber="320">z4</emphasis>
, 
<emphasis id="B93FEAE91A75CE4BE82EF8F2FD88D769" box="[579,604,1861,1884]" italics="true" pageId="19" pageNumber="320">z5</emphasis>
, 
<emphasis id="B93FEAE91A75CE4BE801F8F2FD50D768" box="[620,644,1861,1885]" italics="true" pageId="19" pageNumber="320">s4</emphasis>
(12–20), 
<emphasis id="B93FEAE91A75CE4BE895F8F2FCECD768" box="[760,824,1860,1885]" italics="true" pageId="19" pageNumber="320">s5–s6</emphasis>
(16), 
<emphasis id="B93FEAE91A75CE4BE9ECF8F2FC4ED768" box="[897,922,1861,1885]" italics="true" pageId="19" pageNumber="320">s1</emphasis>
, 
<emphasis id="B93FEAE91A75CE4BE9C7F8F2FC3ED768" box="[938,1002,1861,1885]" italics="true" pageId="19" pageNumber="320">r2–r3</emphasis>
(10–11), 
<emphasis id="B93FEAE91A75CE4BEE31F8F2FBADD768" box="[1116,1145,1861,1885]" italics="true" pageId="19" pageNumber="320">Z2</emphasis>
(16–19), 
<emphasis id="B93FEAE91A75CE4BEE81F8F2FAE2D768" box="[1260,1334,1861,1885]" italics="true" pageId="19" pageNumber="320">Z3–Z4</emphasis>
(16), 
<emphasis id="B93FEAE91A75CE4BEF13F8F2FA48D769" box="[1406,1436,1861,1884]" italics="true" pageId="19" pageNumber="320">Z5</emphasis>
(15–20); other peripheral setae on soft cuticle 
<emphasis id="B93FEAE91A75CE4BE8F1F8DEFD7ED7B5" box="[668,682,1897,1920]" italics="true" pageId="19" pageNumber="320">S</emphasis>
, 
<emphasis id="B93FEAE91A75CE4BE8D5F8DEFD1DD7B5" box="[696,713,1897,1920]" italics="true" pageId="19" pageNumber="320">R</emphasis>
and 
<emphasis id="B93FEAE91A75CE4BE892F8DEFCF0D7B5" box="[767,804,1897,1920]" italics="true" pageId="19" pageNumber="320">UR</emphasis>
(5–14).
</paragraph>
<paragraph id="8BF436FB1A75CE4CEAAAF83BFD4FD198" blockId="19.[151,1436,1716,2029]" lastBlockId="20.[151,1436,151,608]" lastPageId="20" lastPageNumber="321" pageId="19" pageNumber="320">
<emphasis id="B93FEAE91A75CE4BEAAAF83BFE4DD791" box="[199,409,1932,1957]" italics="true" pageId="19" pageNumber="320">Idiosomatic venter</emphasis>
. Tritosternum similar in form and extent of laciniae fusion to that of adult female, base length (29–30), lacinia with 13–14% of length (68–74) fused. Presternal area bare. Sternal shield with four pairs sternal setae 
<emphasis id="B93FEAE91A75CE4BEB5FF862FE55D7D8" box="[306,385,2005,2029]" italics="true" pageId="19" pageNumber="320">st1–st4</emphasis>
and two pairs of poroids (
<figureCitation id="13702A7E1A75CE4BE8ABF863FCCAD7D8" box="[710,798,2004,2029]" captionStart="FIGURES 29 – 33" captionStartId="22.[151,264,1838,1861]" captionTargetBox="[156,1431,193,1800]" captionTargetId="figure@22.[151,1436,193,1817]" captionTargetPageId="22" captionText="FIGURES 29 – 33. Makarovaia ornata n. sp.: 29. Deutonymph, idiosoma, dorsal aspect; 30. Deutonymph, idiosoma, ventral aspect; 31. Deutonymph, detail of corniculus with hyaline process; 32. Protonymph, idiosoma, dorsal aspect; 33. Protonymph, idiosoma, ventral aspect .." httpUri="https://zenodo.org/record/244196/files/figure.png" pageId="19" pageNumber="320">Fig. 30</figureCitation>
), length 
<emphasis id="B93FEAE91A75CE4BE9E7F862FC70D7D9" box="[906,932,2005,2028]" italics="true" pageId="19" pageNumber="320">ca</emphasis>
. 275, width 123–140 at level of coxae II; endopodal extensions distinct; sternal setae 
<emphasis id="B93FEAE91A72CE4CE8E5FF2FFD0CD085" box="[648,728,152,176]" italics="true" pageId="20" pageNumber="321">st1–st3</emphasis>
similar in length (16–30), 
<emphasis id="B93FEAE91A72CE4CEE67FF2FFBFFD085" box="[1034,1067,152,176]" italics="true" pageId="20" pageNumber="321">st4</emphasis>
shorter (8–13); setae 
<emphasis id="B93FEAE91A72CE4CEF49FF2FFA91D09A" box="[1316,1349,152,175]" italics="true" pageId="20" pageNumber="321">st5</emphasis>
on soft cuticle, minute (5–8). Opisthogastric region length, from insertion of 
<emphasis id="B93FEAE91A72CE4CE9C3FF0AFC1BD0E1" box="[942,975,189,212]" italics="true" pageId="20" pageNumber="321">st5</emphasis>
to posterior margin of idiosoma, 
<emphasis id="B93FEAE91A72CE4CEF22FF0AFABDD0E1" box="[1359,1385,189,212]" italics="true" pageId="20" pageNumber="321">ca</emphasis>
. 0.3 idiosomatic length. Anal shield poorly sclerotised, with thick cribrum as in adults; greatest width (88–100) of shield at level of postanal seta similar to midline length 90–99; postanal seta (6–8) slightly shorter than para-anal setae (9–10). Opisthogastric soft cuticle with at least nine pairs of setae (
<emphasis id="B93FEAE91A72CE4CE9BBFE9FFBEFD10A" box="[982,1083,296,320]" italics="true" pageId="20" pageNumber="321">JV1–JV5</emphasis>
, 
<emphasis id="B93FEAE91A72CE4CEE27FE9FFB61D10A" box="[1098,1205,296,320]" italics="true" pageId="20" pageNumber="321">ZV2–ZV5</emphasis>
), 
<emphasis id="B93FEAE91A72CE4CEEA3FE9FFB2ED175" box="[1230,1274,296,320]" italics="true" pageId="20" pageNumber="321">JV1</emphasis>
(9–11) longer than 
<emphasis id="B93FEAE91A72CE4CEAA2FEFBFF2ED151" box="[207,250,332,356]" italics="true" pageId="20" pageNumber="321">JV2</emphasis>
, and 
<emphasis id="B93FEAE91A72CE4CEB54FEFAFEB1D151" box="[313,357,333,356]" italics="true" pageId="20" pageNumber="321">JV5</emphasis>
, 
<emphasis id="B93FEAE91A72CE4CEB1EFEFBFE0AD151" box="[371,478,332,356]" italics="true" pageId="20" pageNumber="321">ZV2–ZV5</emphasis>
minute, spine-like (5–7) (
<figureCitation id="13702A7E1A72CE4CE96BFEFBFC8FD151" box="[774,859,332,357]" captionStart="FIGURES 29 – 33" captionStartId="22.[151,264,1838,1861]" captionTargetBox="[156,1431,193,1800]" captionTargetId="figure@22.[151,1436,193,1817]" captionTargetPageId="22" captionText="FIGURES 29 – 33. Makarovaia ornata n. sp.: 29. Deutonymph, idiosoma, dorsal aspect; 30. Deutonymph, idiosoma, ventral aspect; 31. Deutonymph, detail of corniculus with hyaline process; 32. Protonymph, idiosoma, dorsal aspect; 33. Protonymph, idiosoma, ventral aspect .." httpUri="https://zenodo.org/record/244196/files/figure.png" pageId="20" pageNumber="321">Fig. 30</figureCitation>
). Metapodal plates absent. Peritremes short (35–50 including stigma), reaching to mid-level of coxae III. Peritrematal shield poorly developed and not including peritrematal poroids which are on soft cuticle.
</paragraph>
<paragraph id="8BF436FB1A72CE4CEAAAFE00FD0FD255" blockId="20.[151,1436,151,608]" pageId="20" pageNumber="321">
<taxonomicName id="4C4B4D781A72CE4CEAAAFE00FE82D1E5" box="[199,342,439,464]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="20" pageNumber="321" phylum="Nematoda" rank="genus">
<emphasis id="B93FEAE91A72CE4CEAAAFE00FE82D1E5" box="[199,342,439,464]" italics="true" pageId="20" pageNumber="321">Gnathosoma</emphasis>
</taxonomicName>
. Gnathotectum (100 long, 110 wide), chelicerae (in female deutonymph), ventral gnathosomatic structures and palpi (107–123 long) as in adult female. In male deutonymph, cheliceral shaft, excluding basal section, 120 long, with movable digit (37–39) flattened (12–14 wide) and with sperm duct primordium, one minute tooth and minute distal hook discernible (
<figureCitation id="13702A7E1A72CE4CE81DFD93FD12D209" box="[624,710,548,573]" captionStart="FIGURES 34 – 40" captionStartId="24.[151,264,1946,1969]" captionTargetBox="[151,1431,409,1907]" captionTargetId="figure@24.[151,1436,409,1925]" captionTargetPageId="24" captionText="FIGURES 34 – 40. Makarovaia ornata n. sp.: 34. Larva, idiosoma, dorsal aspect; 35. Larva, idiosoma, ventral aspect; 36. Larva, gnathotectum; 37. Detail of subcapitulum: a, larva (s. s. – indicates salivary stylus); b, protonymph; 38. Chelicera, lateral view, larva; 39. Chelicera, lateral view, protonymph (arrow points to mucro); 40. Chelicera, lateral view, male deutonymph." httpUri="https://zenodo.org/record/244197/files/figure.png" pageId="20" pageNumber="321">Fig. 40</figureCitation>
); cheliceral dorsal seta short (9). Female deutonymph cheliceral shaft 106, movable digit as in adult female (37–42).
</paragraph>
<caption id="DF3466731A72CE4CEAFAF807FB97D7D0" httpUri="https://zenodo.org/record/244194/files/figure.png" pageId="20" pageNumber="321" targetBox="[160,1426,682,1929]" targetPageId="20">
<paragraph id="8BF436FB1A72CE4CEAFAF807FB97D7D0" blockId="20.[151,1436,1968,2021]" pageId="20" pageNumber="321">
<emphasis id="B93FEAE91A72CE4CEAFAF807FE8DD7F3" bold="true" box="[151,345,1968,1991]" pageId="20" pageNumber="321">FIGURES 21–24.</emphasis>
<taxonomicName id="4C4B4D781A72CE4CEB09F806FDFBD7F2" box="[356,559,1969,1991]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B93FEAE91A72CE4CEB09F806FDFBD7F2" box="[356,559,1969,1991]" italics="true" pageId="20" pageNumber="321">Makarovaia ornata</emphasis>
</taxonomicName>
<emphasis id="B93FEAE91A72CE4CE857F806FDADD7F3" bold="true" box="[570,633,1969,1990]" pageId="20" pageNumber="321">
<taxonomicNameLabel id="A20C57921A72CE4CE857F806FDADD7F3" box="[570,633,1969,1990]" pageId="20" pageNumber="321" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult male: 21. Idiosoma, dorsal aspect; 22. Gnathotectum; 23. Detail of cheliceral fixed digit, antaxial view; 24. Chelicera, paraxial view (arthrodial process visible).
</paragraph>
</caption>
<caption id="DF3466731A73CE4DEAFAF9CCFF04D6FB" httpUri="https://zenodo.org/record/244195/files/figure.png" pageId="21" pageNumber="322" targetBox="[166,1429,207,1619]" targetPageId="21">
<paragraph id="8BF436FB1A73CE4DEAFAF9CCFF04D6FB" blockId="21.[151,1436,1659,1742]" pageId="21" pageNumber="322">
<emphasis id="B93FEAE91A73CE4DEAFAF9CCFE82D6A5" bold="true" box="[151,342,1659,1682]" pageId="21" pageNumber="322">FIGURES 25–28.</emphasis>
<taxonomicName id="4C4B4D781A73CE4DEB30F9CCFDF1D6A4" box="[349,549,1659,1681]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B93FEAE91A73CE4DEB30F9CCFDF1D6A4" box="[349,549,1659,1681]" italics="true" pageId="21" pageNumber="322">Makarovaia ornata</emphasis>
</taxonomicName>
<emphasis id="B93FEAE91A73CE4DE841F9CBFDBCD6A4" bold="true" box="[556,616,1660,1681]" pageId="21" pageNumber="322">
<taxonomicNameLabel id="A20C57921A73CE4DE841F9CBFDBCD6A4" box="[556,616,1660,1681]" pageId="21" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: 25. Idiosoma, ventral aspect, adult male; 26. Detail of tarsus II, ventro–apical process, ventral view, male deutonymph; 27. Detail of hyaline basal process of corniculus, adult male; 28. Subcapitulum, adult male.
</paragraph>
</caption>
<paragraph id="8BF436FB1A73CE4DEAAAF940FD0CD7F6" blockId="21.[151,1437,1782,1987]" pageId="21" pageNumber="322">
<emphasis id="B93FEAE91A73CE4DEAAAF940FF28D73B" box="[199,252,1783,1806]" italics="true" pageId="21" pageNumber="322">Legs</emphasis>
. Relative leg lengths, excluding pretarsi, similar to those of female: female DN, I 336–350, II 341–350, III 354–360, IV 361–362; male DN, I 357–362, II 377–393, III 371–395, IV 361–388. Tarsus I with pretarsus 
<emphasis id="B93FEAE91A73CE4DEF16F8ACFA41D707" box="[1403,1429,1819,1842]" italics="true" pageId="21" pageNumber="322">ca</emphasis>
. 10–15 to base of claws (
<emphasis id="B93FEAE91A73CE4DEBC5F888FE17D763" box="[424,451,1855,1878]" italics="true" pageId="21" pageNumber="322">ca</emphasis>
. 18). Tarsi II–IV with pair of apical setal processes 
<emphasis id="B93FEAE91A73CE4DEE64F889FBE8D762" box="[1033,1084,1854,1879]" italics="true" pageId="21" pageNumber="322">ad -1</emphasis>
, 
<emphasis id="B93FEAE91A73CE4DEE27F889FBA9D762" box="[1098,1149,1854,1879]" italics="true" pageId="21" pageNumber="322">pd -1</emphasis>
(12–16) similar to length of pretarsus to base of claws (14–15); with apical ventral projection short (8–12), spade-shaped (
<figureCitation id="13702A7E1A73CE4DEE9CF8D5FA9CD74E" box="[1265,1352,1890,1915]" captionStart="FIGURES 25 – 28" captionStartId="21.[151,264,1659,1682]" captionTargetBox="[166,1429,207,1619]" captionTargetId="figure@21.[151,1436,193,1638]" captionTargetPageId="21" captionText="FIGURES 25 – 28. Makarovaia ornata n. sp.: 25. Idiosoma, ventral aspect, adult male; 26. Detail of tarsus II, ventro – apical process, ventral view, male deutonymph; 27. Detail of hyaline basal process of corniculus, adult male; 28. Subcapitulum, adult male." httpUri="https://zenodo.org/record/244195/files/figure.png" pageId="21" pageNumber="322">Fig. 26</figureCitation>
). Leg I segment length ratios and legs I–IV coxal structures and relative lengths and form of leg setae as in adult female. Trochanters I–IV with seta 
<emphasis id="B93FEAE91A73CE4DEBA7F81CFE2FD7F7" box="[458,507,1962,1986]" italics="true" pageId="21" pageNumber="322">pv -2</emphasis>
slightly attenuated.
</paragraph>
<caption id="DF3466731A70CE4EEAFAF899FE4DD7B7" httpUri="https://zenodo.org/record/244196/files/figure.png" pageId="22" pageNumber="323" targetBox="[156,1431,193,1800]" targetPageId="22">
<paragraph id="8BF436FB1A70CE4EEAFAF899FE4DD7B7" blockId="22.[151,1436,1838,1922]" pageId="22" pageNumber="323">
<emphasis id="B93FEAE91A70CE4EEAFAF899FE9BD771" bold="true" box="[151,335,1838,1861]" pageId="22" pageNumber="323">FIGURES 29–33</emphasis>
. 
<taxonomicName id="4C4B4D781A70CE4EEB30F899FDF0D770" box="[349,548,1838,1861]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B93FEAE91A70CE4EEB30F899FDF0D770" box="[349,548,1838,1861]" italics="true" pageId="22" pageNumber="323">Makarovaia ornata</emphasis>
</taxonomicName>
<emphasis id="B93FEAE91A70CE4EE841F898FDB3D771" bold="true" box="[556,615,1839,1860]" pageId="22" pageNumber="323">
<taxonomicNameLabel id="A20C57921A70CE4EE841F898FDB3D771" box="[556,615,1839,1860]" pageId="22" pageNumber="323" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: 29. Deutonymph, idiosoma, dorsal aspect; 30. Deutonymph, idiosoma, ventral aspect; 31. Deutonymph, detail of corniculus with hyaline process; 32. Protonymph, idiosoma, dorsal aspect; 33. Protonymph, idiosoma, ventral aspect..
</paragraph>
</caption>
<paragraph id="8BF436FB1A70CE4FEAAAF81EFD3CD0CD" blockId="22.[151,1436,1961,2022]" lastBlockId="23.[151,1437,151,2013]" lastPageId="23" lastPageNumber="324" pageId="22" pageNumber="323">
<emphasis id="B93FEAE91A70CE4EEAAAF81EFEB0D7F7" bold="true" box="[199,356,1961,1986]" pageId="22" pageNumber="323">Protonymph.</emphasis>
<emphasis id="B93FEAE91A70CE4EEB06F81EFD9ED7F7" box="[363,586,1961,1986]" italics="true" pageId="22" pageNumber="323">Idiosomatic dorsum</emphasis>
. Idiosoma ovoid in shape, 500–575 long, 
<emphasis id="B93FEAE91A70CE4EEE72F81DFBEDD7F4" box="[1055,1081,1962,1985]" italics="true" pageId="22" pageNumber="323">ca</emphasis>
. 400–500 wide (n = 3); dorsum poorly sclerotised (
<figureCitation id="13702A7E1A70CE4EEB1DF87AFE11D7D3" box="[368,453,1997,2022]" captionStart="FIGURES 29 – 33" captionStartId="22.[151,264,1838,1861]" captionTargetBox="[156,1431,193,1800]" captionTargetId="figure@22.[151,1436,193,1817]" captionTargetPageId="22" captionText="FIGURES 29 – 33. Makarovaia ornata n. sp.: 29. Deutonymph, idiosoma, dorsal aspect; 30. Deutonymph, idiosoma, ventral aspect; 31. Deutonymph, detail of corniculus with hyaline process; 32. Protonymph, idiosoma, dorsal aspect; 33. Protonymph, idiosoma, ventral aspect .." httpUri="https://zenodo.org/record/244196/files/figure.png" pageId="22" pageNumber="323">Fig. 32</figureCitation>
), without clearly delineated shields and with rugose, faint reticulation laterally, similar to that of larva and young deutonymph. Dorsal podonotal setae slightly longer than opisthonotal setae: 
<emphasis id="B93FEAE91A71CE4FEF41FF2FFA96D085" box="[1324,1346,152,176]" italics="true" pageId="23" pageNumber="324">j1</emphasis>
(9–10), 
<emphasis id="B93FEAE91A71CE4FEAFAFF0BFF05D0E1" box="[151,209,188,212]" italics="true" pageId="23" pageNumber="324">j3–j5</emphasis>
(8–13), 
<emphasis id="B93FEAE91A71CE4FEB59FF0BFE9DD0E0" box="[308,329,188,213]" italics="true" pageId="23" pageNumber="324">j6</emphasis>
(20), 
<emphasis id="B93FEAE91A71CE4FEBFDFF0BFE7DD0E1" box="[400,425,188,212]" italics="true" pageId="23" pageNumber="324">z2</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEBD4FF0BFE06D0E1" box="[441,466,188,212]" italics="true" pageId="23" pageNumber="324">z4</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEB8FFF0AFE2FD0E1" box="[482,507,189,212]" italics="true" pageId="23" pageNumber="324">z5</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FE867FF0BFDF7D0E1" box="[522,547,188,212]" italics="true" pageId="23" pageNumber="324">r2</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FE85EFF0BFD98D0E1" box="[563,588,188,212]" italics="true" pageId="23" pageNumber="324">r3</emphasis>
(17–18), 
<emphasis id="B93FEAE91A71CE4FE8D0FF0BFD03D0E1" box="[701,727,188,212]" italics="true" pageId="23" pageNumber="324">J4</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FE88AFF0BFCE4D0E1" box="[743,816,188,212]" italics="true" pageId="23" pageNumber="324">Z2–Z4</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FE92DFF0BFC52D0E1" box="[832,902,188,212]" italics="true" pageId="23" pageNumber="324">S3–S5</emphasis>
(10–11), clunals 
<emphasis id="B93FEAE91A71CE4FEE23FF0AFBBCD0E1" box="[1102,1128,189,212]" italics="true" pageId="23" pageNumber="324">J5</emphasis>
minute, thorn-like, basally spiculate (3–4), and other marginal setae short (5–6).
</paragraph>
<paragraph id="8BF436FB1A71CE4FEAAAFEB3FB41D2B1" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="324">
<emphasis id="B93FEAE91A71CE4FEAAAFEB3FE4CD129" box="[199,408,260,285]" italics="true" pageId="23" pageNumber="324">Idiosomatic venter</emphasis>
. Tritosternum similar in form to that in deutonymph and adult female, but with length of base (14–16) less than width (22–26), and laciniae free for about 75% of total length (110–116). Presternal area bare. Sternal shield (
<emphasis id="B93FEAE91A71CE4FEBEFFEFAFE48D151" box="[386,412,333,356]" italics="true" pageId="23" pageNumber="324">ca</emphasis>
. 
<quantity id="4CB39B1E1A71CE4FEBC1FEFBFDF8D150" box="[428,556,332,357]" metricMagnitude="0" metricUnit="m" metricValue="7.112" metricValueMax="7.62" metricValueMin="6.603999999999999" pageId="23" pageNumber="324" unit="in" value="280.0" valueMax="300.0" valueMin="260.0">260–300 in</quantity>
length), faintly sclerotised, reticulate, with setae 
<emphasis id="B93FEAE91A71CE4FEE33FEFBFB7AD151" box="[1118,1198,332,356]" italics="true" pageId="23" pageNumber="324">st1–st3</emphasis>
(15–17) and poroids 
<emphasis id="B93FEAE91A71CE4FEAFAFEC7FF6DD1BD" box="[151,185,368,392]" italics="true" pageId="23" pageNumber="324">
<date id="FFF5103B1A71CE4FEAFAFEC7FF6DD1BD" box="[151,185,368,392]" pageId="23" pageNumber="324">iv1</date>
</emphasis>
, 
<date id="FFF5103B1A71CE4FEAADFEC7FF3DD1B2" box="[192,233,368,392]" pageId="23" pageNumber="324">
<emphasis id="B93FEAE91A71CE4FEAADFEC7FF37D1BD" box="[192,227,368,392]" italics="true" pageId="23" pageNumber="324">iv2</emphasis>
.
</date>
Setae 
<emphasis id="B93FEAE91A71CE4FEB51FEC7FE88D1B2" box="[316,348,368,391]" italics="true" pageId="23" pageNumber="324">st5</emphasis>
minute (3–4), on soft cuticle between bases of legs IV (
<figureCitation id="13702A7E1A71CE4FE991FED8FB87D1BD" box="[1020,1107,367,392]" captionStart="FIGURES 29 – 33" captionStartId="22.[151,264,1838,1861]" captionTargetBox="[156,1431,193,1800]" captionTargetId="figure@22.[151,1436,193,1817]" captionTargetPageId="22" captionText="FIGURES 29 – 33. Makarovaia ornata n. sp.: 29. Deutonymph, idiosoma, dorsal aspect; 30. Deutonymph, idiosoma, ventral aspect; 31. Deutonymph, detail of corniculus with hyaline process; 32. Protonymph, idiosoma, dorsal aspect; 33. Protonymph, idiosoma, ventral aspect .." httpUri="https://zenodo.org/record/244196/files/figure.png" pageId="23" pageNumber="324">Fig. 33</figureCitation>
). Anal shield faintly ornate, slightly elongate in form as on deutonymph (length 85–110, width 
<emphasis id="B93FEAE91A71CE4FE9E3FE22FC7DD199" box="[910,937,405,428]" italics="true" pageId="23" pageNumber="324">ca</emphasis>
. 80–100); gland pores 
<emphasis id="B93FEAE91A71CE4FEEC2FE23FB03D199" box="[1199,1239,404,428]" italics="true" pageId="23" pageNumber="324">gv3</emphasis>
on posterolateral corners of shield, poroids 
<emphasis id="B93FEAE91A71CE4FEBABFE0EFE3CD1E5" box="[454,488,441,464]" italics="true" pageId="23" pageNumber="324">ivp</emphasis>
on adjacent soft cuticle; postanal seta (9–10) slightly shorter than para-anal setae (10–13); cribum not only similar in width (
<emphasis id="B93FEAE91A71CE4FE8E1FE6AFD73D1C1" box="[652,679,477,500]" italics="true" pageId="23" pageNumber="324">ca</emphasis>
. 90) to anal shield as in subsequent instars but thicker, extending from posterior shield margin to insertion of postanal seta. Opisthogastric setae on soft cuticle short (3–4). Opisthogaster length, from setae 
<emphasis id="B93FEAE91A71CE4FE863FD92FDFBD209" box="[526,559,549,572]" italics="true" pageId="23" pageNumber="324">st5</emphasis>
to posterior margin of idiosoma 160–185, 
<emphasis id="B93FEAE91A71CE4FEE7BFD92FBE4D209" box="[1046,1072,549,572]" italics="true" pageId="23" pageNumber="324">ca</emphasis>
. 0.3 of total idiosomatic length. Peritremes short (
<emphasis id="B93FEAE91A71CE4FEB32FDFEFEADD255" box="[351,377,585,608]" italics="true" pageId="23" pageNumber="324">ca</emphasis>
. 32–35, including stigma) extending anteriorly to posterior margins of coxae III; peritrematal plate hardly discernible, with peritrematal poroids and gland pores on soft cuticle (
<figureCitation id="13702A7E1A71CE4FEE59FDDBFB53D2B1" box="[1076,1159,620,645]" captionStart="FIGURES 29 – 33" captionStartId="22.[151,264,1838,1861]" captionTargetBox="[156,1431,193,1800]" captionTargetId="figure@22.[151,1436,193,1817]" captionTargetPageId="22" captionText="FIGURES 29 – 33. Makarovaia ornata n. sp.: 29. Deutonymph, idiosoma, dorsal aspect; 30. Deutonymph, idiosoma, ventral aspect; 31. Deutonymph, detail of corniculus with hyaline process; 32. Protonymph, idiosoma, dorsal aspect; 33. Protonymph, idiosoma, ventral aspect .." httpUri="https://zenodo.org/record/244196/files/figure.png" pageId="23" pageNumber="324">Fig. 33</figureCitation>
).
</paragraph>
<paragraph id="8BF436FB1A71CE4FEAAAFD38FB1AD369" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="324">
<taxonomicName id="4C4B4D781A71CE4FEAAAFD38FE82D29D" box="[199,342,655,680]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="23" pageNumber="324" phylum="Nematoda" rank="genus">
<emphasis id="B93FEAE91A71CE4FEAAAFD38FE82D29D" box="[199,342,655,680]" italics="true" pageId="23" pageNumber="324">Gnathosoma</emphasis>
</taxonomicName>
. Gnathotectum similar in form to subsequent instars, but dorsal face unornamented. Cheliceral shaft, excluding basal section, 94–105 long; fixed digit with offset subapical tooth followed by row of seven teeth; movable digit (30–35) with four teeth and a ventral proximal protuberance (
<figureCitation id="13702A7E1A71CE4FEE5BFD60FB45D2C5" box="[1078,1169,727,752]" captionStart="FIGURES 34 – 40" captionStartId="24.[151,264,1946,1969]" captionTargetBox="[151,1431,409,1907]" captionTargetId="figure@24.[151,1436,409,1925]" captionTargetPageId="24" captionText="FIGURES 34 – 40. Makarovaia ornata n. sp.: 34. Larva, idiosoma, dorsal aspect; 35. Larva, idiosoma, ventral aspect; 36. Larva, gnathotectum; 37. Detail of subcapitulum: a, larva (s. s. – indicates salivary stylus); b, protonymph; 38. Chelicera, lateral view, larva; 39. Chelicera, lateral view, protonymph (arrow points to mucro); 40. Chelicera, lateral view, male deutonymph." httpUri="https://zenodo.org/record/244197/files/figure.png" pageId="23" pageNumber="324">Fig. 39</figureCitation>
). Other gnathosomatic structures, including inner acuminate process of corniculus and seven rows of deutosternal denticles (four to six denticles each), similar to those in subsequent instars; corniculi inserted at level of 
<emphasis id="B93FEAE91A71CE4FEE5FFC96FB99D30D" box="[1074,1101,801,824]" italics="true" pageId="23" pageNumber="324">ca</emphasis>
. 10 µm posterior to insertions of setae 
<emphasis id="B93FEAE91A71CE4FEA99FCF3FECBD368" box="[244,287,836,861]" italics="true" pageId="23" pageNumber="324">hp1</emphasis>
(
<figureCitation id="13702A7E1A71CE4FEB40FCF3FE54D368" box="[301,384,836,861]" captionStart="FIGURES 34 – 40" captionStartId="24.[151,264,1946,1969]" captionTargetBox="[151,1431,409,1907]" captionTargetId="figure@24.[151,1436,409,1925]" captionTargetPageId="24" captionText="FIGURES 34 – 40. Makarovaia ornata n. sp.: 34. Larva, idiosoma, dorsal aspect; 35. Larva, idiosoma, ventral aspect; 36. Larva, gnathotectum; 37. Detail of subcapitulum: a, larva (s. s. – indicates salivary stylus); b, protonymph; 38. Chelicera, lateral view, larva; 39. Chelicera, lateral view, protonymph (arrow points to mucro); 40. Chelicera, lateral view, male deutonymph." httpUri="https://zenodo.org/record/244197/files/figure.png" pageId="23" pageNumber="324">Fig. 37</figureCitation>
b). Palpi (length 105–115) with normal protonymphal complement of setae.
</paragraph>
<paragraph id="8BF436FB1A71CE4FEAAAFCDEFAB8D425" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="324">
<emphasis id="B93FEAE91A71CE4FEAAAFCDEFF28D3B5" box="[199,252,873,896]" italics="true" pageId="23" pageNumber="324">Legs</emphasis>
. Leg lengths, excluding pretarsi: I 306–325, II 
<emphasis id="B93FEAE91A71CE4FE978FCDEFCE4D3B5" box="[789,816,873,896]" italics="true" pageId="23" pageNumber="324">ca</emphasis>
. 300–350, III 323–350, IV 320–350. Form of legs and their setae as in deutonymph except with normal protonymphal complement of setae (see genus description); tarsus I apically bearing pretarsus (10–16 to base of claws). Tarsi II–IV with apical setal processes 
<emphasis id="B93FEAE91A71CE4FEEAEFC18FB22D3FD" box="[1219,1270,943,968]" italics="true" pageId="23" pageNumber="324">ad -1</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEF6AFC18FAEED3FD" box="[1287,1338,943,968]" italics="true" pageId="23" pageNumber="324">pd -1</emphasis>
(13–15) similar in length to pretarsi (11–15 to base of claws); with ventroapical process under pretarsus as in deutonymph (15–18), bluntly triangular. Form of leg setae much as in deutonymph, but with spinelike setae less developed.
</paragraph>
<paragraph id="8BF436FB1A71CE4FEAAAFBABFCCBD495" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="324">
<emphasis id="B93FEAE91A71CE4FEAAAFBABFEC3D400" bold="true" box="[199,279,1052,1077]" pageId="23" pageNumber="324">Larva.</emphasis>
<emphasis id="B93FEAE91A71CE4FEB72FBABFE2BD400" box="[287,511,1052,1077]" italics="true" pageId="23" pageNumber="324">Idiosomatic dorsum</emphasis>
. Idiosoma oval, 475–580 long, 375–428 wide at level of setae 
<emphasis id="B93FEAE91A71CE4FEEA1FBABFB36D400" box="[1228,1250,1052,1077]" italics="true" pageId="23" pageNumber="324">j6</emphasis>
(n = 2); dorsum weakly sclerotised, without clearly delineated shielding but faintly lineated posterolaterally (
<figureCitation id="13702A7E1A71CE4FEEB3FB88FAECD46D" box="[1246,1336,1087,1112]" captionStart="FIGURES 34 – 40" captionStartId="24.[151,264,1946,1969]" captionTargetBox="[151,1431,409,1907]" captionTargetId="figure@24.[151,1436,409,1925]" captionTargetPageId="24" captionText="FIGURES 34 – 40. Makarovaia ornata n. sp.: 34. Larva, idiosoma, dorsal aspect; 35. Larva, idiosoma, ventral aspect; 36. Larva, gnathotectum; 37. Detail of subcapitulum: a, larva (s. s. – indicates salivary stylus); b, protonymph; 38. Chelicera, lateral view, larva; 39. Chelicera, lateral view, protonymph (arrow points to mucro); 40. Chelicera, lateral view, male deutonymph." httpUri="https://zenodo.org/record/244197/files/figure.png" pageId="23" pageNumber="324">Fig. 34</figureCitation>
). Dorsal podonotal setae slightly longer than opisthonotal setae; 
<emphasis id="B93FEAE91A71CE4FE973FBD3FCE0D449" box="[798,820,1124,1148]" italics="true" pageId="23" pageNumber="324">j1</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FE928FBD3FC88D449" box="[837,860,1124,1148]" italics="true" pageId="23" pageNumber="324">j3</emphasis>
(12–13), 
<emphasis id="B93FEAE91A71CE4FE9BDFBD3FC32D449" box="[976,998,1124,1148]" italics="true" pageId="23" pageNumber="324">j4</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FE99AFBD2FBD9D449" box="[1015,1037,1125,1148]" italics="true" pageId="23" pageNumber="324">j5</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEE73FBD3FBE0D448" box="[1054,1076,1124,1149]" italics="true" pageId="23" pageNumber="324">j6</emphasis>
longest (23–28), 
<emphasis id="B93FEAE91A71CE4FEF69FBD3FAC8D449" box="[1284,1308,1124,1148]" italics="true" pageId="23" pageNumber="324">z2</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEF43FBD3FA93D449" box="[1326,1351,1124,1148]" italics="true" pageId="23" pageNumber="324">z4</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEF34FBD3FAA5D449" box="[1369,1393,1124,1148]" italics="true" pageId="23" pageNumber="324">s4</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEFEEFBD3FA48D448" box="[1411,1436,1124,1149]" italics="true" pageId="23" pageNumber="324">s6</emphasis>
(18–22); 
<emphasis id="B93FEAE91A71CE4FEA93FB3FFECCD495" box="[254,280,1160,1184]" italics="true" pageId="23" pageNumber="324">J2</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEB4AFB3FFE94D495" box="[295,320,1160,1184]" italics="true" pageId="23" pageNumber="324">J3</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEB22FB3FFEBDD495" box="[335,361,1160,1184]" italics="true" pageId="23" pageNumber="324">J4</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEB1AFB3FFE14D4AA" box="[375,448,1160,1184]" italics="true" pageId="23" pageNumber="324">Z3–Z5</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEBA3FB3FFDC1D4AA" box="[462,533,1160,1184]" italics="true" pageId="23" pageNumber="324">S3–S5</emphasis>
(11–13), 
<emphasis id="B93FEAE91A71CE4FE8ECFB3FFD48D4AA" box="[641,668,1160,1183]" italics="true" pageId="23" pageNumber="324">J5</emphasis>
minute (3).
</paragraph>
<paragraph id="8BF436FB1A71CE4FEAAAFB1BFA91D505" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="324">
<emphasis id="B93FEAE91A71CE4FEAAAFB1BFE4ED4F1" box="[199,410,1196,1221]" italics="true" pageId="23" pageNumber="324">Idiosomatic venter</emphasis>
. Tritosternum with base as long (26) as wide (25); laciniae free for about 80% of total length (86 excluding base). Sternal region smooth, with few lines behind setae 
<emphasis id="B93FEAE91A71CE4FEE77FB67FBEED4DD" box="[1050,1082,1232,1256]" italics="true" pageId="23" pageNumber="324">st1</emphasis>
; setae 
<emphasis id="B93FEAE91A71CE4FEEE7FB67FB0ED4DD" box="[1162,1242,1232,1256]" italics="true" pageId="23" pageNumber="324">st1–st3</emphasis>
similar in length (15–17) (
<figureCitation id="13702A7E1A71CE4FEA92FB43FE87D538" box="[255,339,1268,1293]" captionStart="FIGURES 34 – 40" captionStartId="24.[151,264,1946,1969]" captionTargetBox="[151,1431,409,1907]" captionTargetId="figure@24.[151,1436,409,1925]" captionTargetPageId="24" captionText="FIGURES 34 – 40. Makarovaia ornata n. sp.: 34. Larva, idiosoma, dorsal aspect; 35. Larva, idiosoma, ventral aspect; 36. Larva, gnathotectum; 37. Detail of subcapitulum: a, larva (s. s. – indicates salivary stylus); b, protonymph; 38. Chelicera, lateral view, larva; 39. Chelicera, lateral view, protonymph (arrow points to mucro); 40. Chelicera, lateral view, male deutonymph." httpUri="https://zenodo.org/record/244197/files/figure.png" pageId="23" pageNumber="324">Fig. 35</figureCitation>
). Anal shield mostly smooth, as long as wide (
<emphasis id="B93FEAE91A71CE4FE932FB42FCAED539" box="[863,890,1269,1292]" italics="true" pageId="23" pageNumber="324">ca</emphasis>
. 70); para-anal setae (19–22) clearly longer than postanal seta (12–14). Four pairs of opisthogastric setae, 
<emphasis id="B93FEAE91A71CE4FE97BFAAFFC95D505" box="[790,833,1304,1328]" italics="true" pageId="23" pageNumber="324">JV1</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FE922FAAFFCABD505" box="[847,895,1304,1328]" italics="true" pageId="23" pageNumber="324">ZV2</emphasis>
(8–10), 
<emphasis id="B93FEAE91A71CE4FE9B3FAAFFBDED51A" box="[990,1034,1304,1327]" italics="true" pageId="23" pageNumber="324">JV5</emphasis>
(7–8), 
<emphasis id="B93FEAE91A71CE4FEE36FAAFFB52D505" box="[1115,1158,1304,1328]" italics="true" pageId="23" pageNumber="324">JV2</emphasis>
longest (12–15).
</paragraph>
<paragraph id="8BF436FB1A71CE4FEAAAFA8BFB6DD640" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="324">
<taxonomicName id="4C4B4D781A71CE4FEAAAFA8BFE82D560" box="[199,342,1340,1365]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="23" pageNumber="324" phylum="Nematoda" rank="genus">
<emphasis id="B93FEAE91A71CE4FEAAAFA8BFE82D560" box="[199,342,1340,1365]" italics="true" pageId="23" pageNumber="324">Gnathosoma</emphasis>
</taxonomicName>
. Gnathotectum with anterior margin unevenly triangular and smooth (
<figureCitation id="13702A7E1A71CE4FEEE1FA8BFB31D560" box="[1164,1253,1340,1365]" captionStart="FIGURES 34 – 40" captionStartId="24.[151,264,1946,1969]" captionTargetBox="[151,1431,409,1907]" captionTargetId="figure@24.[151,1436,409,1925]" captionTargetPageId="24" captionText="FIGURES 34 – 40. Makarovaia ornata n. sp.: 34. Larva, idiosoma, dorsal aspect; 35. Larva, idiosoma, ventral aspect; 36. Larva, gnathotectum; 37. Detail of subcapitulum: a, larva (s. s. – indicates salivary stylus); b, protonymph; 38. Chelicera, lateral view, larva; 39. Chelicera, lateral view, protonymph (arrow points to mucro); 40. Chelicera, lateral view, male deutonymph." httpUri="https://zenodo.org/record/244197/files/figure.png" pageId="23" pageNumber="324">Fig. 36</figureCitation>
). Fixed digit of chelicera with offset tooth near apical hook, alveolar vestige of pilus dentilis, and a small second tooth 5 µm distant from tip; movable chela (25) bidentate, with small ventral process (
<figureCitation id="13702A7E1A71CE4FE9C8FA33FC2FD5A9" box="[933,1019,1412,1437]" captionStart="FIGURES 34 – 40" captionStartId="24.[151,264,1946,1969]" captionTargetBox="[151,1431,409,1907]" captionTargetId="figure@24.[151,1436,409,1925]" captionTargetPageId="24" captionText="FIGURES 34 – 40. Makarovaia ornata n. sp.: 34. Larva, idiosoma, dorsal aspect; 35. Larva, idiosoma, ventral aspect; 36. Larva, gnathotectum; 37. Detail of subcapitulum: a, larva (s. s. – indicates salivary stylus); b, protonymph; 38. Chelicera, lateral view, larva; 39. Chelicera, lateral view, protonymph (arrow points to mucro); 40. Chelicera, lateral view, male deutonymph." httpUri="https://zenodo.org/record/244197/files/figure.png" pageId="23" pageNumber="324">Fig. 38</figureCitation>
). Palps 98–125 long. Internal malae with external branch thick, pilose, shorter than internal branch (
<figureCitation id="13702A7E1A71CE4FE9E3FA10FC33D5F5" box="[910,999,1447,1472]" captionStart="FIGURES 34 – 40" captionStartId="24.[151,264,1946,1969]" captionTargetBox="[151,1431,409,1907]" captionTargetId="figure@24.[151,1436,409,1925]" captionTargetPageId="24" captionText="FIGURES 34 – 40. Makarovaia ornata n. sp.: 34. Larva, idiosoma, dorsal aspect; 35. Larva, idiosoma, ventral aspect; 36. Larva, gnathotectum; 37. Detail of subcapitulum: a, larva (s. s. – indicates salivary stylus); b, protonymph; 38. Chelicera, lateral view, larva; 39. Chelicera, lateral view, protonymph (arrow points to mucro); 40. Chelicera, lateral view, male deutonymph." httpUri="https://zenodo.org/record/244197/files/figure.png" pageId="23" pageNumber="324">Fig. 37</figureCitation>
a). Deutosternum with seven rows of denticles, basal rows with one to three teeth, distal rows with seven to nine teeth. Hypostomatic setae 
<emphasis id="B93FEAE91A71CE4FEF7FFA7BFAEFD5D0" box="[1298,1339,1484,1509]" italics="true" pageId="23" pageNumber="324">hp1</emphasis>
(36–37) thick, longer than 
<emphasis id="B93FEAE91A71CE4FEB04FA46FE57D63D" box="[361,387,1521,1544]" italics="true" pageId="23" pageNumber="324">pc</emphasis>
(18–20); corniculi (18) inserted at same transverse level as alveoli of setae 
<emphasis id="B93FEAE91A71CE4FEE80FA58FAC3D63D" box="[1261,1303,1519,1544]" italics="true" pageId="23" pageNumber="324">hp1</emphasis>
, with inner acuminate process shorter (8) relative to subsequent instars, its tip not reaching that of main corniculus tip; salivary stylets (19) blunter than in subsequent instars (
<figureCitation id="13702A7E1A71CE4FE8AEF980FCC8D665" box="[707,796,1591,1616]" captionStart="FIGURES 34 – 40" captionStartId="24.[151,264,1946,1969]" captionTargetBox="[151,1431,409,1907]" captionTargetId="figure@24.[151,1436,409,1925]" captionTargetPageId="24" captionText="FIGURES 34 – 40. Makarovaia ornata n. sp.: 34. Larva, idiosoma, dorsal aspect; 35. Larva, idiosoma, ventral aspect; 36. Larva, gnathotectum; 37. Detail of subcapitulum: a, larva (s. s. – indicates salivary stylus); b, protonymph; 38. Chelicera, lateral view, larva; 39. Chelicera, lateral view, protonymph (arrow points to mucro); 40. Chelicera, lateral view, male deutonymph." httpUri="https://zenodo.org/record/244197/files/figure.png" pageId="23" pageNumber="324">Fig. 37</figureCitation>
a). Lateral seta 
<emphasis id="B93FEAE91A71CE4FE9B9F980FBD5D665" box="[980,1025,1591,1616]" italics="true" pageId="23" pageNumber="324">al -1</emphasis>
of palpal femur and genu slender, tapered, as in protonymph; palptarsal claw with dissimilar tines (20 the longer, 15 the shorter).
</paragraph>
<paragraph id="8BF436FB1A71CE4FEAAAF936FB17D731" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="324">
<emphasis id="B93FEAE91A71CE4FEAAAF936FF28D6AD" box="[199,252,1665,1688]" italics="true" pageId="23" pageNumber="324">Legs</emphasis>
. Leg lengths similar, excluding pretarsi: I (338–340), II (325), III (325–342). Form of legs and their setae generally as in protonymph. Tarsus I (88–91) twice as long as tibia I (45). Tarsi II and III apically with well defined ventral process (7–9) shorter than setal processes 
<emphasis id="B93FEAE91A71CE4FE8D2F970FD26D6D5" box="[703,754,1735,1760]" italics="true" pageId="23" pageNumber="324">ad -1</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FE96DF970FCE7D6D5" box="[768,819,1735,1760]" italics="true" pageId="23" pageNumber="324">pd -1</emphasis>
(15) and pretarsus (14–16) to base of claws; tarsi II–III with setae a 
<emphasis id="B93FEAE91A71CE4FEB76F95AFEEBD731" box="[283,319,1772,1796]" italics="true" pageId="23" pageNumber="324">
<date id="FFF5103B1A71CE4FEB76F95AFEEBD731" box="[283,319,1772,1796]" bridgedPair="-" pageId="23" pageNumber="324">v -1</date>
</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEB20F95AFEAAD731" box="[333,382,1772,1796]" italics="true" pageId="23" pageNumber="324">pv -1</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEBE1F95BFE6ED731" box="[396,442,1772,1797]" italics="true" pageId="23" pageNumber="324">al -2</emphasis>
, 
<emphasis id="B93FEAE91A71CE4FEBAAF95BFE21D731" box="[455,501,1772,1797]" italics="true" pageId="23" pageNumber="324">pl -1</emphasis>
similarly longer (14–15), more attenuate than other tarsal setae.
</paragraph>
</subSubSection>
<subSubSection id="C35165701A71CE41EAAAF8B8FE02D129" lastPageId="25" lastPageNumber="326" pageId="23" pageNumber="324" type="materials_examined">
<paragraph id="8BF436FB1A71CE40EAAAF8B8FBE1D151" blockId="23.[151,1437,151,2013]" lastBlockId="24.[151,1437,151,357]" lastPageId="24" lastPageNumber="325" pageId="23" pageNumber="324">
<emphasis id="B93FEAE91A71CE4FEAAAF8B8FEA1D71D" bold="true" box="[199,373,1807,1832]" pageId="23" pageNumber="324">
<typeStatus id="54F088591A71CE4FEAAAF8B8FED6D71D" box="[199,258,1807,1832]" pageId="23" pageNumber="324">Type</typeStatus>
material.
</emphasis>
All material collected from one locality, with initials of collectors indicating Arthropods of La Selva parataxonomists (
<collectingCountry id="F35C766B1A71CE4FEBC3F883FE2FD779" box="[430,507,1844,1868]" name="Kazakhstan" pageId="23" pageNumber="324">ALAS</collectingCountry>
, see acknowledgments), Derek M. Johnson (DMJ), Evert E. Lindquist (EEL). 
<collectingCountry id="F35C766B1A71CE4FEAFAF8EFFE96D745" box="[151,322,1880,1904]" name="Costa Rica" pageId="23" pageNumber="324">COSTA RICA</collectingCountry>
. Heredia Province, La Selva Biological station (
<geoCoordinate id="EE7F503C1A71CE4FE910F8E0FBD2D75A" box="[893,1030,1879,1904]" direction="north" orientation="latitude" pageId="23" pageNumber="324" precision="15" value="10.433611">10°26’1” N</geoCoordinate>
, 
<geoCoordinate id="EE7F503C1A71CE4FEE7BF8EFFB41D75A" box="[1046,1173,1880,1904]" direction="west" orientation="longitude" pageId="23" pageNumber="324" precision="15" value="-84.01723">84°1’2” W</geoCoordinate>
, elevation 
<quantity id="4CB39B1E1A71CE4FEF7AF8E0FA44D745" box="[1303,1424,1879,1904]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="23" pageNumber="324" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50–150 m</quantity>
). 
<typeStatus id="54F088591A71CE4FEAFAF8CBFEFBD7A1" box="[151,303,1916,1940]" pageId="23" pageNumber="324" type="holotype">HOLOTYPE</typeStatus>
: adult female, ex adult 
<taxonomicName id="4C4B4D781A71CE4FE85CF8CBFD7AD7A0" box="[561,686,1916,1941]" class="Insecta" family="Chrysomelidae" genus="Chelobasis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEAE91A71CE4FE85CF8CBFD7AD7A0" box="[561,686,1916,1941]" italics="true" pageId="23" pageNumber="324">Chelobasis</emphasis>
</taxonomicName>
sp. on leaf of 
<taxonomicName id="4C4B4D781A71CE4FE921F8CBFB81D7A0" box="[844,1109,1916,1941]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="23" pageNumber="324" phylum="Tracheophyta" rank="species" species="pogognantha">
<emphasis id="B93FEAE91A71CE4FE921F8CBFB81D7A0" box="[844,1109,1916,1941]" italics="true" pageId="23" pageNumber="324">Heliconia pogognantha</emphasis>
</taxonomicName>
, 
<date id="FFF5103B1A71CE4FEE0EF8CBFADED7A1" box="[1123,1290,1916,1941]" pageId="23" pageNumber="324" value="1994-02">February 1994</date>
, coll. EEL &amp; 
<collectingCountry id="F35C766B1A71CE4FEAFAF817FF31D78D" box="[151,229,1952,1976]" name="Kazakhstan" pageId="23" pageNumber="324">ALAS</collectingCountry>
: deposited in Instituto Nacional de Biodiversidad (
<collectionCode id="ED5AAE3E1A71CE4FE92DF817FC5DD78C" box="[832,905,1952,1977]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="23" pageNumber="324">INBio</collectionCode>
) of 
<collectingCountry id="F35C766B1A71CE4FE9ACF817FB97D78C" box="[961,1091,1952,1977]" name="Costa Rica" pageId="23" pageNumber="324">Costa Rica</collectingCountry>
, Santo 
<collectingCountry id="F35C766B1A71CE4FEECDF817FADED78C" box="[1184,1290,1952,1977]" name="Dominican Republic" pageId="23" pageNumber="324">Domingo</collectingCountry>
de Heredia. 
<typeStatus id="54F088591A71CE4FEAFAF873FEE2D7E9" box="[151,310,1988,2012]" pageId="23" pageNumber="324" type="paratype">PARATYPES</typeStatus>
: two adult females, seven adult males, one deutonymph, two protonymphs (one pharate deutonymph), one larva, with same data as 
<typeStatus id="54F088591A7ECE40E8E2FF20FD20D085" box="[655,756,151,176]" pageId="24" pageNumber="325" type="holotype">holotype</typeStatus>
; one adult female, one adult male, ex 
<taxonomicName id="4C4B4D781A7ECE40EEDFFF20FAFBD085" box="[1202,1327,151,176]" class="Insecta" family="Chrysomelidae" genus="Chelobasis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEAE91A7ECE40EEDFFF20FAFBD085" box="[1202,1327,151,176]" italics="true" pageId="24" pageNumber="325">Chelobasis</emphasis>
</taxonomicName>
sp., 
<date id="FFF5103B1A7ECE40EF04FF2FFF06D0E1" pageId="24" pageNumber="325" value="1997-06">June 1997</date>
, coll. DMJ; five adult males, four deutonymphal males, one deutonymphal female, one protonymph, under elytra of 
<taxonomicName id="4C4B4D781A7ECE40EB6AFF68FDE7D0CD" authority="Baly" authorityName="Baly" box="[263,563,223,248]" class="Insecta" family="Chrysomelidae" genus="Chelobasis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="B93FEAE91A7ECE40EB6AFF68FE26D0CD" box="[263,498,223,248]" italics="true" pageId="24" pageNumber="325">Chelobasis perplexa</emphasis>
Baly
</taxonomicName>
from rolled 
<taxonomicName id="4C4B4D781A7ECE40E8BCFF68FCEBD0CD" box="[721,831,223,248]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="24" pageNumber="325" phylum="Tracheophyta" rank="genus">
<emphasis id="B93FEAE91A7ECE40E8BCFF68FCEBD0CD" box="[721,831,223,248]" italics="true" pageId="24" pageNumber="325">Heliconia</emphasis>
</taxonomicName>
leaf, 
<date id="FFF5103B1A7ECE40E9E7FF68FBD4D0CD" box="[906,1024,223,248]" pageId="24" pageNumber="325" value="1997-07">July 1997</date>
, #739025a, coll. 
<collectingCountry id="F35C766B1A7ECE40EEBDFF57FAC8D0CD" box="[1232,1308,224,248]" name="Kazakhstan" pageId="24" pageNumber="325">ALAS</collectingCountry>
; two adult females, two adult males, one deutonymph (pharate adult female), one larva (pharate protonymph), same data as previous entry, #739025b, coll. 
<collectingCountry id="F35C766B1A7ECE40E86EFE9FFD84D175" box="[515,592,296,320]" name="Kazakhstan" pageId="24" pageNumber="325">ALAS</collectingCountry>
; one adult female, same data as previous entry, #739025c, coll. 
<collectingCountry id="F35C766B1A7ECE40EF77FE9FFAB3D175" box="[1306,1383,296,320]" name="Kazakhstan" pageId="24" pageNumber="325">ALAS</collectingCountry>
; two adult females, two adult males, same data as previous entry, #739026, coll. 
<collectingCountry id="F35C766B1A7ECE40E989FEFBFBE5D151" box="[996,1073,332,356]" name="Kazakhstan" pageId="24" pageNumber="325">ALAS</collectingCountry>
.
</paragraph>
<caption id="DF3466731A7ECE40EAFAF82DFAA8D7DB" httpUri="https://zenodo.org/record/244197/files/figure.png" pageId="24" pageNumber="325" targetBox="[151,1431,409,1907]" targetPageId="24">
<paragraph id="8BF436FB1A7ECE40EAFAF82DFAA8D7DB" blockId="24.[151,1436,1946,2030]" pageId="24" pageNumber="325">
<emphasis id="B93FEAE91A7ECE40EAFAF82DFE83D785" bold="true" box="[151,343,1946,1969]" pageId="24" pageNumber="325">FIGURES 34–40.</emphasis>
<taxonomicName id="4C4B4D781A7ECE40EB0CF82CFDFDD784" box="[353,553,1947,1969]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B93FEAE91A7ECE40EB0CF82CFDFDD784" box="[353,553,1947,1969]" italics="true" pageId="24" pageNumber="325">Makarovaia ornata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C57921A7ECE40E85EF82CFDA4D785" box="[563,624,1947,1968]" pageId="24" pageNumber="325" rank="species">
<emphasis id="B93FEAE91A7ECE40E85EF82CFDA4D785" bold="true" box="[563,624,1947,1968]" pageId="24" pageNumber="325">n. sp.</emphasis>
</taxonomicNameLabel>
: 34. Larva, idiosoma, dorsal aspect; 35. Larva, idiosoma, ventral aspect; 36. Larva, gnathotectum; 37. Detail of subcapitulum: a, larva (s.s. – indicates salivary stylus); b, protonymph; 38. Chelicera, lateral view, larva; 39. Chelicera, lateral view, protonymph (arrow points to mucro); 40. Chelicera, lateral view, male deutonymph.
</paragraph>
</caption>
<paragraph id="8BF436FB1A7FCE41EAAAFF20FE02D129" blockId="25.[151,1436,151,536]" pageId="25" pageNumber="326">
<emphasis id="B93FEAE91A7FCE41EAAAFF20FE6ED085" bold="true" box="[199,442,151,176]" pageId="25" pageNumber="326">Host identifications.</emphasis>
The identities of the host beetles and background plant substrate given above are in accord with current molecular studies, which indicate that one haplotype of 
<taxonomicName id="4C4B4D781A7FCE41EE6FFF0BFB3DD0E0" box="[1026,1257,188,213]" class="Insecta" family="Chrysomelidae" genus="Chelobasis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" sensu="stricto" species="perplexa">
<emphasis id="B93FEAE91A7FCE41EE6FFF0BFB3DD0E0" box="[1026,1257,188,213]" italics="true" pageId="25" pageNumber="326">Chelobasis perplexa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C57921A7FCE41EE9EFF0AFA54D0E0" box="[1267,1408,188,213]" pageId="25" pageNumber="326" sensu="stricto">sensu stricto</taxonomicNameLabel>
is found on 
<taxonomicName id="4C4B4D781A7FCE41EB75FF68FDCAD0CD" box="[280,542,223,248]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="25" pageNumber="326" phylum="Tracheophyta" rank="species" species="pogonantha">
<emphasis id="B93FEAE91A7FCE41EB75FF68FDCAD0CD" box="[280,542,223,248]" italics="true" pageId="25" pageNumber="326">Heliconia pogonantha</emphasis>
</taxonomicName>
in the La Selva area of 
<collectingCountry id="F35C766B1A7FCE41E91BFF57FC2FD0CD" box="[886,1019,223,248]" name="Costa Rica" pageId="25" pageNumber="326">Costa Rica</collectingCountry>
(Carlos Garcia-Robledo, personal communication, 
<date id="FFF5103B1A7FCE41EB3EFEB2FE1FD129" box="[339,459,260,284]" pageId="25" pageNumber="326" value="2014-06">June 2014</date>
).
</paragraph>
</subSubSection>
<subSubSection id="C35165701A7FCE41EAAAFE90FA8DD150" pageId="25" pageNumber="326" type="etymology">
<paragraph id="8BF436FB1A7FCE41EAAAFE90FA8DD150" blockId="25.[151,1436,151,536]" pageId="25" pageNumber="326">
<emphasis id="B93FEAE91A7FCE41EAAAFE90FE9AD175" bold="true" box="[199,334,295,320]" pageId="25" pageNumber="326">Etymology.</emphasis>
The specific name indicates the conspicuously reticulate ornamentation of the intercoxal shielding of adults, and also the adults being commonly ornamented with remarkable laboulbeniaceous fungal growth.
</paragraph>
</subSubSection>
<subSubSection id="C35165701A7FCE41EAAAFED8FE4DD22D" pageId="25" pageNumber="326" type="discussion">
<paragraph id="8BF436FB1A7FCE41EAAAFED8FE4DD22D" blockId="25.[151,1436,151,536]" pageId="25" pageNumber="326">
<emphasis id="B93FEAE91A7FCE41EAAAFED8FEE0D1BD" bold="true" box="[199,308,367,392]" pageId="25" pageNumber="326">Remarks</emphasis>
. As noted in the description and confirmed by views of specimens with pharate adults, deutonymphs of 
<taxonomicName id="4C4B4D781A7FCE41EAD8FE23FE44D199" box="[181,400,404,429]" class="Arachnida" family="Ascidae" genus="Makarovaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="B93FEAE91A7FCE41EAD8FE23FE44D199" box="[181,400,404,429]" italics="true" pageId="25" pageNumber="326">Makarovaia ornata</emphasis>
</taxonomicName>
are dimorphic in the form of their movable cheliceral digits. While the movable digit of the female deutonymph is identical in form with that of the female adult, that of the male deutonymph displays anlage of the adult sperm duct, and has a straighter masticatory surface bearing just one minute tooth, instead of a series of five well-formed teeth.
</paragraph>
</subSubSection>
</treatment>
</document>