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<mods:title id="87FBDB5BD9126F068099EEFE4DD0F3F9">Procladius Skuse from Lake Winnipeg, Manitoba, Canada, with keys to some females and immature stages of the genus (Diptera: Chironomidae)</mods:title>
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Key to known Nearctic larvae of
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<emphasis id="B903EACAFC67FFA4AE1AF9BAFD06730D" bold="true" box="[562,693,1596,1622]" italics="true" pageId="21" pageNumber="55">Procladius</emphasis>
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<paragraph id="8BC836D8FC67FFA4ACBFF904FA2F73B4" blockId="21.[151,1437,1666,2010]" pageId="21" pageNumber="55">
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1. Two small claws with broad bases and short distal ends, with or without teeth on inner margin, longest claws with pointed apices (
<figureCitation id="134C2A5DFC67FFA4AD24F919FEE373EF" box="[268,336,1695,1716]" captionStart="FIGURE 14" captionStartId="18.[151,250,1880,1903]" captionTargetBox="[277,1316,423,1827]" captionTargetId="figure@18.[252,1335,411,1858]" captionTargetPageId="18" captionText="FIGURE 14. Procladius spp., larvae; maxillae ventral view (A B), dorsal view (C); maxillary palp (D H), claws of posterior parapods (I L). A, D, I, J— Procladius (P.) bellus (Loew); B, F, K— Procladius (H.) culiciformis; C, G, L— Procladius (H.) denticulatus Sublette; E— Procladius (H.) freemani Sublette; H— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200044/files/figure.png" pageId="21" pageNumber="55">
Fig.
<date id="FFC91018FC67FFA4AD10F919FEE373EF" box="[312,336,1695,1716]" pageId="21" pageNumber="55" value="1914-01">14</date>
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I, J); hypopharyngeal pecten (
<figureCitation id="134C2A5DFC67FFA4AE5AF919FD0573EF" box="[626,694,1695,1716]" captionStart="FIGURE 13" captionStartId="17.[151,250,1705,1728]" captionTargetBox="[254,1327,215,1677]" captionTargetId="figure@17.[248,1339,193,1684]" captionTargetPageId="17" captionText="FIGURE 13. Procladius spp., larvae, antennal apex (A E), antenna (F G), labrum (H J), and ligula, paraligula and hypopharyngeal pecten. A, F, H, K— Procladius (P.) bellus (Loew); B, I, L— Procladius (H.) freemani Sublette; C— Procladius (H.) culiciformis (L.); D— Procladius (H.) denticulatus Sublette; E, G, J, M— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200043/files/figure.png" pageId="21" pageNumber="55">Fig. 13</figureCitation>
K) with 69, usually 6 or 7, teeth; blade (
<figureCitation id="134C2A5DFC67FFA4A86BF919FB3273EF" box="[1091,1153,1695,1716]" captionStart="FIGURE 13" captionStartId="17.[151,250,1705,1728]" captionTargetBox="[254,1327,215,1677]" captionTargetId="figure@17.[248,1339,193,1684]" captionTargetPageId="17" captionText="FIGURE 13. Procladius spp., larvae, antennal apex (A E), antenna (F G), labrum (H J), and ligula, paraligula and hypopharyngeal pecten. A, F, H, K— Procladius (P.) bellus (Loew); B, I, L— Procladius (H.) freemani Sublette; C— Procladius (H.) culiciformis (L.); D— Procladius (H.) denticulatus Sublette; E, G, J, M— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200043/files/figure.png" pageId="21" pageNumber="55">Fig.13</figureCitation>
A) as long as segments 2 and 3 combined; lamellae of galea and chaetulae of palpifer (
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A) broad and undivided; head capsule length
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.................................................................................................... 2
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- All claws with narrow bases, without teeth although small spines may be present on inner margins, long claws with attenuated whip-like apices (
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K, L); hypopharyngeal pecten (
<figureCitation id="134C2A5DFC67FFA4AED2F893FC8D7271" box="[762,830,1813,1834]" captionStart="FIGURE 13" captionStartId="17.[151,250,1705,1728]" captionTargetBox="[254,1327,215,1677]" captionTargetId="figure@17.[248,1339,193,1684]" captionTargetPageId="17" captionText="FIGURE 13. Procladius spp., larvae, antennal apex (A E), antenna (F G), labrum (H J), and ligula, paraligula and hypopharyngeal pecten. A, F, H, K— Procladius (P.) bellus (Loew); B, I, L— Procladius (H.) freemani Sublette; C— Procladius (H.) culiciformis (L.); D— Procladius (H.) denticulatus Sublette; E, G, J, M— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200043/files/figure.png" pageId="21" pageNumber="55">Fig. 13</figureCitation>
L, M) with 1014, usually 1113, teeth; usually at least as long as segments 24 combined; at least some lamellae of galea and chaetulae of palpifer (
<figureCitation id="134C2A5DFC67FFA4A807F8B4FBDE721C" box="[1071,1133,1842,1863]" captionStart="FIGURE 14" captionStartId="18.[151,250,1880,1903]" captionTargetBox="[277,1316,423,1827]" captionTargetId="figure@18.[252,1335,411,1858]" captionTargetPageId="18" captionText="FIGURE 14. Procladius spp., larvae; maxillae ventral view (A B), dorsal view (C); maxillary palp (D H), claws of posterior parapods (I L). A, D, I, J— Procladius (P.) bellus (Loew); B, F, K— Procladius (H.) culiciformis; C, G, L— Procladius (H.) denticulatus Sublette; E— Procladius (H.) freemani Sublette; H— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200044/files/figure.png" pageId="21" pageNumber="55">Fig.14</figureCitation>
B, C) divided at apical half to third; head length more than
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..................................................................... 3
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2. Two of the smaller posterior parapod claws sharply hooked, no claws with strong points on inner side..............................................................................................
<taxonomicName id="4C774D5BFC67FFA4A84CF80CFA2672C4" ID-CoL="8DGVG" authority="Roback" authorityName="Roback" box="[1124,1429,1930,1951]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="55" phylum="Arthropoda" rank="species" species="nietus" subGenus="Psilotanypus">
<emphasis id="B903EACAFC67FFA4A84CF80CFAF572C4" box="[1124,1350,1930,1951]" italics="true" pageId="21" pageNumber="55">P. (Psilotanypus) nietus</emphasis>
Roback
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</paragraph>
<paragraph id="8BC836D8FC67FFA4ACBFF821FA2F7281" blockId="21.[151,1437,1666,2010]" pageId="21" pageNumber="55">
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- The smaller posterior parapod claws not sharply hooked, 0, 1, or 2 claws with strong points on inner side.............................................................................................
<taxonomicName id="4C774D5BFC67FFA4A854F843FA2F7281" ID-CoL="8DGV2" authority="Loew" authorityName="Loew" box="[1148,1436,1989,2010]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="55" phylum="Arthropoda" rank="species" species="bellus" subGenus="Psilotanypus">
<emphasis id="B903EACAFC67FFA4A854F843FAEC7281" box="[1148,1375,1989,2010]" italics="true" pageId="21" pageNumber="55">P. (Psilotanypus) bellus</emphasis>
Loew
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<keyStep id="30832E7DFC64FFA7ACBFFF11FA2F7479" pageId="22" pageNumber="56">
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3. Blade shorter than segments 24 combined (
<figureCitation id="134C2A5DFC64FFA7AEA1FF11FD7B75F7" box="[649,712,151,172]" captionStart="FIGURE 13" captionStartId="17.[151,250,1705,1728]" captionTargetBox="[254,1327,215,1677]" captionTargetId="figure@17.[248,1339,193,1684]" captionTargetPageId="17" captionText="FIGURE 13. Procladius spp., larvae, antennal apex (A E), antenna (F G), labrum (H J), and ligula, paraligula and hypopharyngeal pecten. A, F, H, K— Procladius (P.) bellus (Loew); B, I, L— Procladius (H.) freemani Sublette; C— Procladius (H.) culiciformis (L.); D— Procladius (H.) denticulatus Sublette; E, G, J, M— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200043/files/figure.png" pageId="22" pageNumber="56">Fig.13</figureCitation>
B); two-segmented sensilla basiconica (SBP in
<figureCitation id="134C2A5DFC64FFA7A8BBFF11FB6575F7" box="[1171,1238,151,172]" captionStart="FIGURE 14" captionStartId="18.[151,250,1880,1903]" captionTargetBox="[277,1316,423,1827]" captionTargetId="figure@18.[252,1335,411,1858]" captionTargetPageId="18" captionText="FIGURE 14. Procladius spp., larvae; maxillae ventral view (A B), dorsal view (C); maxillary palp (D H), claws of posterior parapods (I L). A, D, I, J— Procladius (P.) bellus (Loew); B, F, K— Procladius (H.) culiciformis; C, G, L— Procladius (H.) denticulatus Sublette; E— Procladius (H.) freemani Sublette; H— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200044/files/figure.png" pageId="22" pageNumber="56">Fig. 14</figureCitation>
E) of maxillary palp shorter than bisensillum (Bs in
<figureCitation id="134C2A5DFC64FFA7AE25FF33FDF97591" box="[525,586,181,202]" captionStart="FIGURE 14" captionStartId="18.[151,250,1880,1903]" captionTargetBox="[277,1316,423,1827]" captionTargetId="figure@18.[252,1335,411,1858]" captionTargetPageId="18" captionText="FIGURE 14. Procladius spp., larvae; maxillae ventral view (A B), dorsal view (C); maxillary palp (D H), claws of posterior parapods (I L). A, D, I, J— Procladius (P.) bellus (Loew); B, F, K— Procladius (H.) culiciformis; C, G, L— Procladius (H.) denticulatus Sublette; E— Procladius (H.) freemani Sublette; H— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200044/files/figure.png" pageId="22" pageNumber="56">Fig.14</figureCitation>
E), its basal segment shorter than its apical segment; head capsule length 0.620.78....................................................................................
<taxonomicName id="4C774D5BFC64FFA7A886FF54FA2875BC" ID-CoL="8DGSH" authority="Sublette" authorityName="Sublette" box="[1198,1435,210,231]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="56" phylum="Arthropoda" rank="species" species="freemani" subGenus="Holotanypus">
<emphasis id="B903EACAFC64FFA7A886FF54FAF475BC" box="[1198,1351,210,231]" italics="true" pageId="22" pageNumber="56">P. (H.) freemani</emphasis>
Sublette
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</paragraph>
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- Blade at least as long as segments 24 combined or usually longer (
<figureCitation id="134C2A5DFC64FFA7AF58FF69FC07745F" box="[880,948,239,260]" captionStart="FIGURE 13" captionStartId="17.[151,250,1705,1728]" captionTargetBox="[254,1327,215,1677]" captionTargetId="figure@17.[248,1339,193,1684]" captionTargetPageId="17" captionText="FIGURE 13. Procladius spp., larvae, antennal apex (A E), antenna (F G), labrum (H J), and ligula, paraligula and hypopharyngeal pecten. A, F, H, K— Procladius (P.) bellus (Loew); B, I, L— Procladius (H.) freemani Sublette; C— Procladius (H.) culiciformis (L.); D— Procladius (H.) denticulatus Sublette; E, G, J, M— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200043/files/figure.png" pageId="22" pageNumber="56">Fig. 13</figureCitation>
CE); SBP (
<figureCitation id="134C2A5DFC64FFA7A805FF69FBC0745F" box="[1069,1139,239,260]" captionStart="FIGURE 14" captionStartId="18.[151,250,1880,1903]" captionTargetBox="[277,1316,423,1827]" captionTargetId="figure@18.[252,1335,411,1858]" captionTargetPageId="18" captionText="FIGURE 14. Procladius spp., larvae; maxillae ventral view (A B), dorsal view (C); maxillary palp (D H), claws of posterior parapods (I L). A, D, I, J— Procladius (P.) bellus (Loew); B, F, K— Procladius (H.) culiciformis; C, G, L— Procladius (H.) denticulatus Sublette; E— Procladius (H.) freemani Sublette; H— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200044/files/figure.png" pageId="22" pageNumber="56">Fig. 14</figureCitation>
FH) of maxillary palp longer than Bs, its basal segment longer than its apical segment; head capsule length
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.......................... 4
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4. Blade at least as long as segments 24 combined, accessory blade usually nearly as long as blade (
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C); head capsule length
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................................................................................... 5
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- Blade as long as segments 24 combined or, if as long as segments 25 combined, head capsule length over
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; accessory blade distinctly shorter than blade (
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D, E)........................................................................................................ 6
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<paragraph id="8BC836D8FC64FFA7ACBFFE24FA2F74EC" blockId="22.[151,1436,151,703]" box="[151,1436,418,439]" pageId="22" pageNumber="56">
<keyLead id="308695EDFC64FFA7ACBFFE24FA2F74EC" box="[151,1436,418,439]" pageId="22" pageNumber="56">
5. Sclerotized part of basal segment of maxillary palp (
<figureCitation id="134C2A5DFC64FFA7AEFFFE24FCAF74EC" box="[727,796,418,439]" captionStart="FIGURE 14" captionStartId="18.[151,250,1880,1903]" captionTargetBox="[277,1316,423,1827]" captionTargetId="figure@18.[252,1335,411,1858]" captionTargetPageId="18" captionText="FIGURE 14. Procladius spp., larvae; maxillae ventral view (A B), dorsal view (C); maxillary palp (D H), claws of posterior parapods (I L). A, D, I, J— Procladius (P.) bellus (Loew); B, F, K— Procladius (H.) culiciformis; C, G, L— Procladius (H.) denticulatus Sublette; E— Procladius (H.) freemani Sublette; H— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200044/files/figure.png" pageId="22" pageNumber="56">Fig. 14</figureCitation>
F) 1.92.6 times as long as wide.......
<taxonomicName id="4C774D5BFC64FFA7A891FE24FAC274EC" ID-CoL="8DGS7" box="[1209,1393,418,439]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="56" phylum="Arthropoda" rank="species" species="culiciformis" subGenus="Holotanypus">
<emphasis id="B903EACAFC64FFA7A891FE24FAC274EC" box="[1209,1393,418,439]" italics="true" pageId="22" pageNumber="56">P. (H.) culiciformis</emphasis>
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(L.)
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</paragraph>
<paragraph id="8BC836D8FC64FFA7ACBFFE39FA2F748F" blockId="22.[151,1436,151,703]" box="[151,1436,447,468]" pageId="22" pageNumber="56">
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- Sclerotized part of basal segment of maxillary palp about 3.2 times as long as wide................
<taxonomicName id="4C774D5BFC64FFA7A8EEFE39FA2F748F" ID-CoL="8DGT9" authority="Roback" authorityName="Roback" box="[1222,1436,447,468]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="56" phylum="Arthropoda" rank="species" species="wilhmi" subGenus="Holotanypus">
<emphasis id="B903EACAFC64FFA7A8EEFE39FAFF748F" box="[1222,1356,447,468]" italics="true" pageId="22" pageNumber="56">P. (H.) wilhmi</emphasis>
Roback
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<paragraph id="8BC836D8FC64FFA7ACBFFE5BFA2F74A9" blockId="22.[151,1436,151,703]" box="[151,1436,477,498]" pageId="22" pageNumber="56">
<keyLead id="308695EDFC64FFA7ACBFFE5BFA2F74A9" box="[151,1436,477,498]" pageId="22" pageNumber="56">
6. Procercus with 1819 setae; both mandible and basal antennal segment longer than 210 µm............
<taxonomicName id="4C774D5BFC64FFA7A8FDFE5BFA2F74A9" ID-CoL="8DGT2" authority="Roback" authorityName="Roback" box="[1237,1436,477,498]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="56" phylum="Arthropoda" rank="species" species="ruris" subGenus="Holotanypus">
<emphasis id="B903EACAFC64FFA7A8FDFE5BFAFE74A9" box="[1237,1357,477,498]" italics="true" pageId="22" pageNumber="56">P. (H.) ruris</emphasis>
Roback
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</paragraph>
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<keyLead id="308695EDFC64FFA7ACBFFE7CFA2F7754" box="[151,1436,506,527]" pageId="22" pageNumber="56">- Procercus with 914 setae; mandible and basal antennal segment often shorter than 210 µm.......................... 7</keyLead>
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<paragraph id="8BC836D8FC64FFA7ACBFFD91FA26773C" blockId="22.[151,1436,151,703]" pageId="22" pageNumber="56">
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7. Sclerotized part of basal segment of maxillary palp (
<figureCitation id="134C2A5DFC64FFA7AEF5FD91FC907777" box="[733,803,535,556]" captionStart="FIGURE 14" captionStartId="18.[151,250,1880,1903]" captionTargetBox="[277,1316,423,1827]" captionTargetId="figure@18.[252,1335,411,1858]" captionTargetPageId="18" captionText="FIGURE 14. Procladius spp., larvae; maxillae ventral view (A B), dorsal view (C); maxillary palp (D H), claws of posterior parapods (I L). A, D, I, J— Procladius (P.) bellus (Loew); B, F, K— Procladius (H.) culiciformis; C, G, L— Procladius (H.) denticulatus Sublette; E— Procladius (H.) freemani Sublette; H— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200044/files/figure.png" pageId="22" pageNumber="56">Fig. 14</figureCitation>
E) 4558 µm long and 2.42.9, mean 2.5, times as long as wide; basal antennal segment 4.55.6, 5.0 times as long as wide and 3.44.5, mean 4.0, times as long as basal palpal segment; head capsule length
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.........................................................
<taxonomicName id="4C774D5BFC64FFA7A898FDD4FA26773C" ID-CoL="8DGT6" authority="Roback" authorityName="Roback" box="[1200,1429,594,615]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="56" phylum="Arthropoda" rank="species" species="sublettei" subGenus="Holotanypus">
<emphasis id="B903EACAFC64FFA7A898FDD4FAF5773C" box="[1200,1350,594,615]" italics="true" pageId="22" pageNumber="56">P. (H.) sublettei</emphasis>
Roback
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</paragraph>
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<keyLead id="308695EDFC64FFA7ACBFFDE9FA2877E4" pageId="22" pageNumber="56">
- Sclerotized part of basal segment of maxillary palp (
<figureCitation id="134C2A5DFC64FFA7AEF4FDE9FC9177DF" box="[732,802,623,644]" captionStart="FIGURE 14" captionStartId="18.[151,250,1880,1903]" captionTargetBox="[277,1316,423,1827]" captionTargetId="figure@18.[252,1335,411,1858]" captionTargetPageId="18" captionText="FIGURE 14. Procladius spp., larvae; maxillae ventral view (A B), dorsal view (C); maxillary palp (D H), claws of posterior parapods (I L). A, D, I, J— Procladius (P.) bellus (Loew); B, F, K— Procladius (H.) culiciformis; C, G, L— Procladius (H.) denticulatus Sublette; E— Procladius (H.) freemani Sublette; H— Procladius (H.) sublettei Roback." httpUri="https://zenodo.org/record/200044/files/figure.png" pageId="22" pageNumber="56">Fig. 14</figureCitation>
G) 5574 µm long and 2.73.5, mean 3.0, times as long as wide; basal antennal segment 5.16.4, mean 5.5 times as long as wide and 2.93.9, mean 3.4, times as long as basal palpal segment; head capsule length
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long..............................................
<taxonomicName id="4C774D5BFC64FFA7A8A5FD2CFA2877E4" ID-CoL="8DGS9" authority="Sublette" authorityName="Sublette" box="[1165,1435,682,703]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="56" phylum="Arthropoda" rank="species" species="denticulatus" subGenus="Holotanypus">
<emphasis id="B903EACAFC64FFA7A8A5FD2CFAF477E4" box="[1165,1351,682,703]" italics="true" pageId="22" pageNumber="56">P. (H.) denticulatus</emphasis>
Sublette
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