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<document id="542CA061A90468B34B3C8AD3F0714D82" ID-DOI="10.11646/zootaxa.5446.1.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="11084732" ID-ZooBank="7EAF7685-8C2E-4C6F-A3A3-350C29B85198" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1714390694158" checkinUser="plazi" docAuthor="Novelo-Gutiérrez, Rodolfo &amp; Gómez-Anaya, José Antonio" docDate="2024" docId="03D387AC451F3A00E3C7FD1746A4E893" docLanguage="en" docName="zootaxa.5446.1.8.pdf" docOrigin="Zootaxa 5446 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5446.1.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epigomphus sulcatistyla Donnelly 1989" docType="treatment" docVersion="1" lastPageNumber="141" masterDocId="FFEAFFD4451E3A08E350FFC04453EA62" masterDocTitle="The rediscovery of Epigomphus sulcatistyla Donnelly, 1989, with a description of its larva and female (Odonata: Gomphidae)" masterLastPageNumber="142" masterPageNumber="133" pageNumber="134" updateTime="1714503160068" updateUser="jonas" zenodo-license-document="CLOSED">
<mods:mods id="810A4DC29E812EFD294FA9711A264BB2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6E4B9A23097BBE1496DCCDB7D339646C">
<mods:title id="8F3C4CDC6F13447178C1AAD5F6E6E7A0">The rediscovery of Epigomphus sulcatistyla Donnelly, 1989, with a description of its larva and female (Odonata: Gomphidae)</mods:title>
</mods:titleInfo>
<mods:name id="CC800E75BC971D291E068E6DCC5081AD" type="personal">
<mods:role id="57F30F34550E2B63FB028B1A920E1DB1">
<mods:roleTerm id="32C73AB9632C615E98A4D628B0B5312B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FF3A53532CE9ED41456364F0B2B305CA">Novelo-Gutiérrez, Rodolfo</mods:namePart>
<mods:affiliation id="C18C2D19DA1DC550838D4757D801898E">Instituto de Ecología, A. C. Red de Biodiversidad y Sistemática. Carretera antigua a Coatepec 351, El Haya 91073 Xalapa, Veracruz, MEXICO</mods:affiliation>
</mods:name>
<mods:name id="460685F91BE2343E26F996639C8B48B9" type="personal">
<mods:role id="FC53A53066AB95BE926F0EF4D05E7826">
<mods:roleTerm id="3A6A01F4FD34AA7D8B2B99F5F0E561F8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="03C4AE40DED4CA33A75A14B14ACF19CD">Gómez-Anaya, José Antonio</mods:namePart>
<mods:nameIdentifier id="A3B527FEC7CFC1747277A5120C7218E0" type="ORCID">0000-0002-4598-9663</mods:nameIdentifier>
<mods:affiliation id="A40F8008D6A8586300EAC6F3D8B145B6">Instituto de Ecología, A. C. Red de Biodiversidad y Sistemática. Carretera antigua a Coatepec 351, El Haya 91073 Xalapa, Veracruz, MEXICO</mods:affiliation>
<mods:nameIdentifier id="EB6E7BBD15985683D24B08D89AC8F659" type="email">antonio.gomez@inecol.mx</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="18C973231BAC573890201FA45382AF31">text</mods:typeOfResource>
<mods:relatedItem id="843B32CD352AE9BE6D0F923158AFBBF8" type="host">
<mods:titleInfo id="A7151604073D469CA6CFBED707F04368">
<mods:title id="0FA3E2E66F185BECF34866443E2A2193">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6F46AF7F70F710EC6FBD3840FD29A809">
<mods:date id="C3C03C0D67BE68B7FEE7183289535E44">2024</mods:date>
<mods:detail id="16DABBEA4A219B25E15FF083F170FFE9" type="pubDate">
<mods:number id="D1C6FBE0C6875216B9BAF53462E741EA">2024-04-29</mods:number>
</mods:detail>
<mods:detail id="2BD731CF2B1CF8C81C3A2751E4F35A98" type="volume">
<mods:number id="CE7C86F58BC371E43FFBC40115AAE417">5446</mods:number>
</mods:detail>
<mods:detail id="62969B385BD8986A65EB0B644BEC73EB" type="issue">
<mods:number id="5D91D54ABDFAA6AF100F748E117F2D6F">1</mods:number>
</mods:detail>
<mods:extent id="0EE650852048FE0131CFF53E40FCA87A" unit="page">
<mods:start id="A0D79ECD57C8D0BD185C7D22334CC086">133</mods:start>
<mods:end id="799E61E251E5DFDAE85425CE50D64412">142</mods:end>
</mods:extent>
</mods:part>
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<mods:location id="2C61F4F19F3AD43D344DE1BA0AA40725">
<mods:url id="F38EFC458F23BB4F5DE6A5DEB8228F17">http://dx.doi.org/10.11646/zootaxa.5446.1.8</mods:url>
</mods:location>
<mods:classification id="808AB6634FAED5B8297422C9162E690E">journal article</mods:classification>
<mods:identifier id="C6CF570989FE362A2A8A23552CAC3FE0" type="DOI">10.11646/zootaxa.5446.1.8</mods:identifier>
<mods:identifier id="4AD789C53F38A5A975D490402FD3F12E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6781771C3E7CE1B95317E9568AD53503" type="Zenodo-Dep">11084732</mods:identifier>
<mods:identifier id="EEC54C8868919053804ABDF7A1B3E641" type="ZooBank">7EAF7685-8C2E-4C6F-A3A3-350C29B85198</mods:identifier>
</mods:mods>
<treatment id="03D387AC451F3A00E3C7FD1746A4E893" LSID="urn:lsid:plazi:treatment:03D387AC451F3A00E3C7FD1746A4E893" httpUri="http://treatment.plazi.org/id/03D387AC451F3A00E3C7FD1746A4E893" lastPageId="8" lastPageNumber="141" pageId="1" pageNumber="134">
<subSubSection id="C3606531451F3A09E3C7FD17476DE893" box="[151,830,727,754]" pageId="1" pageNumber="134" type="nomenclature">
<paragraph id="8BC536BA451F3A09E3C7FD17476DE893" blockId="1.[151,830,727,789]" box="[151,830,727,754]" pageId="1" pageNumber="134">
<heading id="D08D81D6451F3A09E3C7FD17476DE893" bold="true" box="[151,830,727,754]" fontSize="11" level="1" pageId="1" pageNumber="134" reason="1">
<emphasis id="B90EEAA8451F3A09E3C7FD17476DE893" bold="true" box="[151,830,727,754]" pageId="1" pageNumber="134">
Larva description of
<taxonomicName id="4C7A4D39451F3A09E2CEFD17476DE893" ID-CoL="6G5K7" authority="Donnelly" authorityName="Donnelly" authorityYear="1989" box="[414,830,727,754]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="sulcatistyla">
<emphasis id="B90EEAA8451F3A09E2CEFD174695E890" bold="true" box="[414,710,727,754]" italics="true" pageId="1" pageNumber="134">Epigomphus sulcatistyla</emphasis>
Donnelly
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3606531451F3A09E3C7FD3B4541E977" box="[151,274,763,789]" pageId="1" pageNumber="134" type="description">
<paragraph id="8BC536BA451F3A09E3C7FD3B4541E977" blockId="1.[151,830,727,789]" box="[151,274,763,789]" pageId="1" pageNumber="134">
(
<figureCitation id="13412A3F451F3A09E3CFFD3B455AE977" box="[159,265,763,789]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1943,1967]" captionStartId-1="3.[151,250,1943,1967]" captionStartId-2="4.[151,250,1943,1967]" captionTargetBox-0="[194,1394,191,1919]" captionTargetBox-1="[236,1352,181,1920]" captionTargetBox-2="[432,1155,181,1919]" captionTargetId-0="figure-23@2.[194,1394,181,1919]" captionTargetId-1="figure-17@3.[236,1352,181,1920]" captionTargetId-2="figure-23@4.[432,1155,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Epigomphus sulcatistyla: a) last stadium male exuvia, dorsal view; b) Ídem, ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view." captionText-1="FIGURE 2. Epigomphus sulcatistyla: a) antennae of a young larva, dorsal view; b) prementum of the last stadium exuvia, ventral view; c) detail of the palps and ligula, ventral view; d) S7–10 and caudal appendages of male, dorsal view; e) sterna 9–10 of the female showing the vestigial gonapophyses." captionText-2="FIGURE 3. Details of the mouthparts of the last stadium exuvia of Epigomphus sulcatistyla: a–d) mandibles: a–b, right mandible, ventral and mesal view, respectively; c–d, left mandible, ventro-mesal and mesal view, respectively; e–f) left maxilla, ventral and mesal view, respectively." figureDoi-0="http://doi.org/10.5281/zenodo.11084734" figureDoi-1="http://doi.org/10.5281/zenodo.11084736" figureDoi-2="http://doi.org/10.5281/zenodo.11084742" httpUri-0="https://zenodo.org/record/11084734/files/figure.png" httpUri-1="https://zenodo.org/record/11084736/files/figure.png" httpUri-2="https://zenodo.org/record/11084742/files/figure.png" pageId="1" pageNumber="134">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3606531451F3A09E3C7FC83459DEE3B" pageId="1" pageNumber="134" type="materials_examined">
<paragraph id="8BC536BA451F3A09E3C7FC83459DEE3B" blockId="1.[151,1437,835,2014]" pageId="1" pageNumber="134">
<emphasis id="B90EEAA8451F3A09E3C7FC834555E93F" bold="true" box="[151,262,835,861]" pageId="1" pageNumber="134">Material.</emphasis>
<materialsCitation id="3B123CE7451F3A09E25FFC83403BE9C7" collectingDate="2023-04-17" collectingDateMax="2023-05-24" collectingDateMin="2023-04-17" collectorName="J. A. Gomez &amp; R. Novelo" country="Mexico" county="Eight" elevation="102" latitude="18.305584" location="Estacion de Biologia Tropical" longLatPrecision="1" longitude="-94.98755" municipality="Municipality of San Andres Tuxtla" pageId="1" pageNumber="134" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Veracruz">
Eight exuviae (
<specimenCount id="9D7CFD33451F3A09E2EBFC8345ADE93C" box="[443,510,835,862]" count="4" pageId="1" pageNumber="134" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="9D7CFD33451F3A09E15BFC83461FE93C" box="[523,588,835,862]" count="4" pageId="1" pageNumber="134" type="female">4 ♀♀</specimenCount>
, reared), 8 young larvae.
<collectingCountry id="F36D762A451F3A09E021FC8347B7E93F" box="[881,996,835,861]" name="Mexico" pageId="1" pageNumber="134">MEXICO</collectingCountry>
:
<collectingRegion id="49BEF858451F3A09E0BFFC844005E93C" box="[1007,1110,836,862]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="1" pageNumber="134">Veracruz</collectingRegion>
;
<collectingMunicipality id="6BA1ACC0451F3A09E732FC8344B1E9E3" pageId="1" pageNumber="134">Municipality of San Andrés Tuxtla</collectingMunicipality>
,
<location id="8EA56061451F3A09E3BDFCA7466FE9E3" LSID="urn:lsid:plazi:treatment:03D387AC451F3A00E3C7FD1746A4E893:8EA56061451F3A09E3BDFCA7466FE9E3" box="[237,572,871,897]" country="Mexico" county="Eight" latitude="18.305584" longLatPrecision="1" longitude="-94.98755" municipality="Municipality of San Andres Tuxtla" name="Estacion de Biologia Tropical" pageId="1" pageNumber="134" stateProvince="Veracruz">Estación de Biología Tropical</location>
“Los Tuxtlas”, UNAM, stream elev.
<quantity id="4C829B5F451F3A09E08EFCA74076E9E0" box="[990,1061,871,898]" metricMagnitude="2" metricUnit="m" metricValue="1.02" pageId="1" pageNumber="134" unit="m" value="102.0">
<elevation id="0057D189451F3A09E08EFCA74076E9E0" box="[990,1061,871,898]" metricMagnitude="2" metricUnit="m" metricValue="1.02" pageId="1" pageNumber="134" unit="m" value="102.0">102 m</elevation>
</quantity>
(
<geoCoordinate id="EE4E507D451F3A09E764FCA7409BE9E0" box="[1076,1224,871,898]" degrees="18" direction="north" minutes="29.193" orientation="latitude" pageId="1" pageNumber="134" precision="1" value="18.48655">18°29.193 N</geoCoordinate>
,
<geoCoordinate id="EE4E507D451F3A09E783FCA7413FE9E3" box="[1235,1388,871,898]" degrees="95" direction="west" minutes="04.896" orientation="longitude" pageId="1" pageNumber="134" precision="1" value="-95.0816">95°04.896 W</geoCoordinate>
),
<date id="FFC4107A451F3A09E6D0FCA74547E9C7" pageId="1" pageNumber="134" value="2023-04-17">
<collectingDate id="EF80E992451F3A09E6D0FCA74547E9C7" pageId="1" pageNumber="134" value="2023-04-17">17 April 2023</collectingDate>
</date>
, 2 F-0 larvae (emerged
<date id="FFC4107A451F3A09E14FFC4B46E6E9C7" box="[543,693,907,934]" pageId="1" pageNumber="134" value="2023-05-23">
<collectingDate id="EF80E992451F3A09E14FFC4B46E6E9C7" box="[543,693,907,934]" pageId="1" pageNumber="134" value="2023-05-23">23 May 2023</collectingDate>
</date>
at 13:15 h (
<specimenCount id="9D7CFD33451F3A09E06EFC4B4703E9C7" box="[830,848,907,933]" count="1" pageId="1" pageNumber="134" type="female">♀</specimenCount>
) and
<date id="FFC4107A451F3A09E0C2FC4B407BE9C7" box="[914,1064,907,934]" pageId="1" pageNumber="134" value="2023-05-24">
<collectingDate id="EF80E992451F3A09E0C2FC4B407BE9C7" box="[914,1064,907,934]" pageId="1" pageNumber="134" value="2023-05-24">24 May 2023</collectingDate>
</date>
(
<specimenCount id="9D7CFD33451F3A09E768FC4B401DE9C7" box="[1080,1102,907,933]" count="1" pageId="1" pageNumber="134" type="male">♂</specimenCount>
));
</materialsCitation>
<materialsCitation id="3B123CE7451F3A09E73FFC4B454AE98F" collectingDate="2023-04-17" collectingDateMax="2023-06-01" collectingDateMin="2022-10-04" collectorName="Dos Amates" country="Mexico" county="Eight" elevation="351" latitude="18.305584" location="Municipality of Catemaco" longLatPrecision="1" longitude="-94.98755" municipality="Municipality of Catemaco" pageId="1" pageNumber="134" specimenCount="1" specimenCount-female="8" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave">
<collectingMunicipality id="6BA1ACC0451F3A09E73FFC4B41CBE9C7" box="[1135,1432,907,933]" pageId="1" pageNumber="134">Municipality of Catemaco</collectingMunicipality>
,
<collectorName id="268F536C451F3A09E3C7FC704572E9AB" box="[151,289,943,970]" pageId="1" pageNumber="134">Dos Amates</collectorName>
, stream elev.
<quantity id="4C829B5F451F3A09E2E5FC6F45A8E9A8" box="[437,507,943,970]" metricMagnitude="2" metricUnit="m" metricValue="3.51" pageId="1" pageNumber="134" unit="m" value="351.0">
<elevation id="0057D189451F3A09E2E5FC6F45A8E9A8" box="[437,507,943,970]" metricMagnitude="2" metricUnit="m" metricValue="3.51" pageId="1" pageNumber="134" unit="m" value="351.0">351 m</elevation>
</quantity>
(
<geoCoordinate id="EE4E507D451F3A09E158FC6F46C8E9A8" box="[520,667,943,970]" degrees="18" direction="north" minutes="25.622" orientation="latitude" pageId="1" pageNumber="134" precision="1" value="18.427032">18°25.622 N</geoCoordinate>
,
<geoCoordinate id="EE4E507D451F3A09E1F5FC6F476FE9AB" box="[677,828,943,970]" degrees="95" direction="west" minutes="04.995" orientation="longitude" pageId="1" pageNumber="134" precision="1" value="-95.08325">95°04.995 W</geoCoordinate>
),
<date id="FFC4107A451F3A09E01FFC6F47B8E9AB" box="[847,1003,943,969]" pageId="1" pageNumber="134" value="2023-04-17">
<collectingDate id="EF80E992451F3A09E01FFC6F47B8E9AB" box="[847,1003,943,969]" pageId="1" pageNumber="134" value="2023-04-17">17 April 2023</collectingDate>
</date>
, one F-0 larva (
<specimenCount id="9D7CFD33451F3A09E7C6FC6F40F8E9AB" box="[1174,1195,943,969]" count="1" pageId="1" pageNumber="134" type="male">♂</specimenCount>
emerged
<date id="FFC4107A451F3A09E647FC6F41CFE9AB" box="[1303,1436,943,970]" pageId="1" pageNumber="134" value="2023-06-01">1 June 2023</date>
at 11:50 h);
</materialsCitation>
<materialsCitation id="3B123CE7451F3A09E270FC1445B9EE73" collectingDate="2022-10-04" collectingDateMax="2023-04-18" collectingDateMin="2022-10-04" collectorName="J. A. Gomez &amp; R. Novelo" country="Mexico" county="Eight" elevation="102" latitude="18.305584" location="road Tebanca-Miguel Hidalgo" longLatPrecision="1" longitude="-94.98755" municipality="Municipality of San Andres Tuxtla" pageId="1" pageNumber="134" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Veracruz de Ignacio de la Llave">
same data but km 7.1
<location id="8EA56061451F3A09E149FC13473EE98F" LSID="urn:lsid:plazi:treatment:03D387AC451F3A00E3C7FD1746A4E893:8EA56061451F3A09E149FC13473EE98F" box="[537,877,979,1005]" country="Mexico" county="Eight" latitude="18.305584" longLatPrecision="1" longitude="-94.98755" municipality="Municipality of San Andres Tuxtla" name="road Tebanca-Miguel Hidalgo" pageId="1" pageNumber="134" stateProvince="Veracruz de Ignacio de la Llave">road Tebanca-Miguel Hidalgo</location>
, stream elev. 402 (
<geoCoordinate id="EE4E507D451F3A09E711FC134086E98C" box="[1089,1237,979,1006]" degrees="18" direction="north" minutes="15.349" orientation="latitude" pageId="1" pageNumber="134" precision="1" value="18.255817">18°15.349 N</geoCoordinate>
,
<geoCoordinate id="EE4E507D451F3A09E7B1FC134129E98F" box="[1249,1402,979,1006]" degrees="94" direction="west" minutes="57.516" orientation="longitude" pageId="1" pageNumber="134" precision="1" value="-94.9586">94°57.516 W</geoCoordinate>
),
<date id="FFC4107A451F3A09E6DEFC134560EE73" pageId="1" pageNumber="134" value="2022-10-04">
<collectingDate id="EF80E992451F3A09E6DEFC134560EE73" pageId="1" pageNumber="134" value="2022-10-04">4 October 2022</collectingDate>
</date>
, 8 young larvae;
</materialsCitation>
<materialsCitation id="3B123CE7451F3A09E2A1FC38459DEE3B" collectingDate="2023-04-18" collectingDateMax="2023-04-18" collectingDateMin="2022-10-04" collectorName="J. A. Gomez &amp; R. Novelo" country="Mexico" county="Eight" elevation="102" latitude="18.305584" location="road Tebanca-Miguel Hidalgo" longLatPrecision="1" longitude="-94.98755" municipality="Municipality of San Andres Tuxtla" pageId="1" pageNumber="134" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Veracruz de Ignacio de la Llave">
same data but km 8.4
<location id="8EA56061451F3A09E1B4FC374065EE73" LSID="urn:lsid:plazi:treatment:03D387AC451F3A00E3C7FD1746A4E893:8EA56061451F3A09E1B4FC374065EE73" box="[740,1078,1015,1041]" country="Mexico" county="Eight" latitude="18.305584" longLatPrecision="1" longitude="-94.98755" municipality="Municipality of San Andres Tuxtla" name="road Tebanca-Miguel Hidalgo" pageId="1" pageNumber="134" stateProvince="Veracruz de Ignacio de la Llave">road Tebanca-Miguel Hidalgo</location>
, stream elev. 507 (
<geoCoordinate id="EE4E507D451F3A09E655FC3741CBEE70" box="[1285,1432,1015,1042]" degrees="18" direction="north" minutes="18.335" orientation="latitude" pageId="1" pageNumber="134" precision="1" value="18.305584">18°18.335 N</geoCoordinate>
,
<geoCoordinate id="EE4E507D451F3A09E3C7FBDB4563EE57" box="[151,304,1051,1078]" degrees="94" direction="west" minutes="59.253" orientation="longitude" pageId="1" pageNumber="134" precision="1" value="-94.98755">94°59.253 W</geoCoordinate>
),
<date id="FFC4107A451F3A09E214FBDB45B0EE57" box="[324,483,1051,1077]" pageId="1" pageNumber="134" value="2023-04-18">
<collectingDate id="EF80E992451F3A09E214FBDB45B0EE57" box="[324,483,1051,1077]" pageId="1" pageNumber="134" value="2023-04-18">18 April 2023</collectingDate>
</date>
, 5 F-0 larvae (
<specimenCount id="9D7CFD33451F3A09E1D5FBDB4694EE54" box="[645,711,1051,1078]" count="2" pageId="1" pageNumber="134" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D7CFD33451F3A09E184FBDB475FEE57" box="[724,780,1051,1077]" count="3" pageId="1" pageNumber="134" type="female">3♀♀</specimenCount>
, all emerged between
<date id="FFC4107A451F3A09E758FBDC40BCEE57" box="[1032,1263,1051,1078]" pageId="1" pageNumber="134">
<collectingDate id="EF80E992451F3A09E758FBDC40BCEE57" box="[1032,1263,1051,1078]" pageId="1" pageNumber="134">May 27 and June 03</collectingDate>
</date>
); all specimens
<collectorName id="268F536C451F3A09E3C7FBFF454CEE3B" box="[151,287,1087,1113]" pageId="1" pageNumber="134">J.A. Gómez</collectorName>
,
<collectorName id="268F536C451F3A09E27AFB8045CEEE3B" box="[298,413,1087,1114]" pageId="1" pageNumber="134">R. Novelo</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3606531451F3A0DE397FBA34487EF53" lastPageId="5" lastPageNumber="138" pageId="1" pageNumber="134" type="description">
<paragraph id="8BC536BA451F3A09E397FBA347A7EEA7" blockId="1.[151,1437,835,2014]" pageId="1" pageNumber="134">
<emphasis id="B90EEAA8451F3A09E397FBA3450AEE1F" bold="true" box="[199,345,1123,1149]" pageId="1" pageNumber="134">Description.</emphasis>
Medium-size exuviae (25.8–30.0 mm total length), body robust, antennae, legs and sides of thorax setose, gently tapering caudad, body light yellow-brown lacking any particular pattern (
<figureCitation id="13412A3F451F3A09E72EFB47409AEEC3" box="[1150,1225,1159,1185]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-23@2.[194,1394,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus sulcatistyla: a) last stadium male exuvia, dorsal view; b) Ídem, ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11084734" httpUri="https://zenodo.org/record/11084734/files/figure.png" pageId="1" pageNumber="134">Figs. 1</figureCitation>
a-b), young larvae light yellow with a dark brown, complex color pattern on abdomen (
<figureCitation id="13412A3F451F3A09E0C5FB6B47B6EEA7" box="[917,997,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-23@2.[194,1394,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus sulcatistyla: a) last stadium male exuvia, dorsal view; b) Ídem, ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11084734" httpUri="https://zenodo.org/record/11084734/files/figure.png" pageId="1" pageNumber="134">Fig. 1c</figureCitation>
).
</paragraph>
<paragraph id="8BC536BA451F3A0DE397FB0F4727EA9B" blockId="1.[151,1437,835,2014]" lastBlockId="5.[151,1437,151,1401]" lastPageId="5" lastPageNumber="138" pageId="1" pageNumber="134">
<emphasis id="B90EEAA8451F3A09E397FB0F4557EE8B" box="[199,260,1231,1257]" italics="true" pageId="1" pageNumber="134">Head</emphasis>
: Wider than long, wider than prothorax but narrower than pterothorax and abdomen (
<figureCitation id="13412A3F451F3A09E7B5FB0F417AEE8B" box="[1253,1321,1231,1257]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-23@2.[194,1394,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus sulcatistyla: a) last stadium male exuvia, dorsal view; b) Ídem, ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11084734" httpUri="https://zenodo.org/record/11084734/files/figure.png" pageId="1" pageNumber="134">Fig. 1</figureCitation>
). Labrum mostly bare, anterior border with dense brush of golden setae, flattened ventrally; anteclypeus bare, postclypeus, frons, vertex and occiput with minute, moderately abundant scale-like setae, a tuft of long, upturned, golden setae on anterolateral margins of frons (between antenna and anterior margin of compound eye), occiput mostly bare, each occipital lobe with two large, circular, bare areas and with long, golden setae on lateral margins. Anterior margin of frons shallowly concave. Antennae 4-segmented (
<figureCitation id="13412A3F451F3A09E191FA434741EFFF" box="[705,786,1411,1437]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[236,1352,181,1920]" captionTargetId="figure-17@3.[236,1352,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Epigomphus sulcatistyla: a) antennae of a young larva, dorsal view; b) prementum of the last stadium exuvia, ventral view; c) detail of the palps and ligula, ventral view; d) S7–10 and caudal appendages of male, dorsal view; e) sterna 9–10 of the female showing the vestigial gonapophyses." figureDoi="http://doi.org/10.5281/zenodo.11084736" httpUri="https://zenodo.org/record/11084736/files/figure.png" pageId="1" pageNumber="134">Fig. 2a</figureCitation>
), covered with minute, scale-like setae dorsally, and dense rows of long, yellowish setae on lateral margins, those on external margins longest, those on internal margins stiff, scape globose, pedicel subglobose, 3
<superScript id="7C0F9BF2451F3A09E167FA0B4616EFBB" attach="left" box="[567,581,1483,1497]" fontSize="6" pageId="1" pageNumber="134">rd</superScript>
antennomere largest, spindle-shaped, flattened dorsoventrally, almost twice as long as widest part, 4
<superScript id="7C0F9BF2451F3A09E2FDFA2F45E9EF9F" attach="left" box="[429,442,1519,1533]" fontSize="6" pageId="1" pageNumber="134">th</superScript>
antennomere a small blunt cone; scape, pedicel and 3
<superScript id="7C0F9BF2451F3A09E775FA2F4060EF9F" attach="left" box="[1061,1075,1519,1533]" fontSize="6" pageId="1" pageNumber="134">rd</superScript>
antennomere yellow-brown, 4
<superScript id="7C0F9BF2451F3A09E6C0FA2F41CEEF9F" attach="none" box="[1424,1437,1519,1533]" fontSize="6" pageId="1" pageNumber="134">th</superScript>
antennomere reddish brown, size proportions: 0.30, 0.20, 1.0, 0.10. Compound eyes moderately developed; ocelli pale. Occipital lobes rounded, bulging; a well-developed longitudinal carina beset with small, stiff setae to each side of ventral surface of head. Mandibles (
<figureCitation id="13412A3F451F3A09E16FF99B46DAEC17" box="[575,649,1627,1653]" captionStart="FIGURE 3" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[432,1155,181,1919]" captionTargetId="figure-23@4.[432,1155,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Details of the mouthparts of the last stadium exuvia of Epigomphus sulcatistyla: a–d) mandibles: a–b, right mandible, ventral and mesal view, respectively; c–d, left mandible, ventro-mesal and mesal view, respectively; e–f) left maxilla, ventral and mesal view, respectively." figureDoi="http://doi.org/10.5281/zenodo.11084742" httpUri="https://zenodo.org/record/11084742/files/figure.png" pageId="1" pageNumber="134">Figs. 3</figureCitation>
a-d) with molar crest, formula: L 1234 -y a(m
<superScript id="7C0F9BF2451F3A09E72EF99B40CCEC0B" attach="left" box="[1150,1183,1627,1641]" fontSize="6" pageId="1" pageNumber="134">1,2,3</superScript>
)b / R 1234 y a(m
<superScript id="7C0F9BF2451F3A09E60DF99B412DEC0B" attach="left" box="[1373,1406,1627,1641]" fontSize="6" pageId="1" pageNumber="134">1,2,3</superScript>
)b, in both mandibles tooth a&gt;b. Maxillae: galeolacinia (
<figureCitation id="13412A3F451F3A09E1B4F9BF477CECFB" box="[740,815,1663,1689]" captionStart="FIGURE 3" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[432,1155,181,1919]" captionTargetId="figure-23@4.[432,1155,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Details of the mouthparts of the last stadium exuvia of Epigomphus sulcatistyla: a–d) mandibles: a–b, right mandible, ventral and mesal view, respectively; c–d, left mandible, ventro-mesal and mesal view, respectively; e–f) left maxilla, ventral and mesal view, respectively." figureDoi="http://doi.org/10.5281/zenodo.11084742" httpUri="https://zenodo.org/record/11084742/files/figure.png" pageId="1" pageNumber="134">Figs. 3</figureCitation>
e-f) with 7 moderately incurved teeth, three dorsal teeth more or less of same length and robustness, four ventral teeth of different size, apical one largest; maxillary palp thick and robust. Ventral pad of hypopharynx pentagonal, whitish, soft, with a row of ventral, subapical, long, stiff setae, basal margin concave, sclerotized. Labium: prementum-postmentum articulation reaching posterior margin of procoxae (
<figureCitation id="13412A3F451F3A09E25EF8CF4530ED4B" box="[270,355,1807,1833]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[236,1352,181,1920]" captionTargetId="figure-17@3.[236,1352,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Epigomphus sulcatistyla: a) antennae of a young larva, dorsal view; b) prementum of the last stadium exuvia, ventral view; c) detail of the palps and ligula, ventral view; d) S7–10 and caudal appendages of male, dorsal view; e) sterna 9–10 of the female showing the vestigial gonapophyses." figureDoi="http://doi.org/10.5281/zenodo.11084736" httpUri="https://zenodo.org/record/11084736/files/figure.png" pageId="1" pageNumber="134">Fig. 2b</figureCitation>
). Prementum reddish brown, subrectangular, slightly longer than its widest part, lateral margins smooth, slightly sinuate, convex at apical 0.50, convergent on basal 0.37, basal margin shallowly concave (
<figureCitation id="13412A3F451F3A09E66BF8F341DDED2F" box="[1339,1422,1843,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[236,1352,181,1920]" captionTargetId="figure-17@3.[236,1352,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Epigomphus sulcatistyla: a) antennae of a young larva, dorsal view; b) prementum of the last stadium exuvia, ventral view; c) detail of the palps and ligula, ventral view; d) S7–10 and caudal appendages of male, dorsal view; e) sterna 9–10 of the female showing the vestigial gonapophyses." figureDoi="http://doi.org/10.5281/zenodo.11084736" httpUri="https://zenodo.org/record/11084736/files/figure.png" pageId="1" pageNumber="134">Fig. 2b</figureCitation>
); without a longitudinal, central sulcus on ventral surface, a small, shallow, concavity just below ligula beset with some long setae; ligula poorly developed (
<figureCitation id="13412A3F451F3A09E125F8BB4694EDF7" box="[629,711,1915,1941]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[236,1352,181,1920]" captionTargetId="figure-17@3.[236,1352,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Epigomphus sulcatistyla: a) antennae of a young larva, dorsal view; b) prementum of the last stadium exuvia, ventral view; c) detail of the palps and ligula, ventral view; d) S7–10 and caudal appendages of male, dorsal view; e) sterna 9–10 of the female showing the vestigial gonapophyses." figureDoi="http://doi.org/10.5281/zenodo.11084736" httpUri="https://zenodo.org/record/11084736/files/figure.png" pageId="1" pageNumber="134">Fig. 2c</figureCitation>
), W/L ratio 1:0.20, apical margin slightly convex, with a ventral row of 8 (88 %) to 10 (12 %) reddish brown, short, truncate teeth on middle, a dorsal row of long, whitish piliform setae, and a middle row of short, stout, brown setae; labial palp stout (
<figureCitation id="13412A3F451F3A09E0C9F80347BAEDBF" box="[921,1001,1987,2013]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[236,1352,181,1920]" captionTargetId="figure-17@3.[236,1352,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Epigomphus sulcatistyla: a) antennae of a young larva, dorsal view; b) prementum of the last stadium exuvia, ventral view; c) detail of the palps and ligula, ventral view; d) S7–10 and caudal appendages of male, dorsal view; e) sterna 9–10 of the female showing the vestigial gonapophyses." figureDoi="http://doi.org/10.5281/zenodo.11084736" httpUri="https://zenodo.org/record/11084736/files/figure.png" pageId="1" pageNumber="134">Fig. 2c</figureCitation>
), reddish brown, mostly glabrous, with some sparse, long, delicate setae on external surface; apical lobe stout, tip rounded and finely scalloped, internal margin almost straight with 10–11 small, truncate teeth, external margin gently convex and smooth; movable hook reddish-brown, shorter than palp, sharp and moderately incurved.
</paragraph>
<caption id="DF056632451C3A0AE3C7F85746C7EDB6" ID-DOI="http://doi.org/10.5281/zenodo.11084734" ID-Zenodo-Dep="11084734" httpUri="https://zenodo.org/record/11084734/files/figure.png" pageId="2" pageNumber="135" startId="2.[151,250,1943,1967]" targetBox="[194,1394,191,1919]" targetPageId="2" targetType="figure">
<paragraph id="8BC536BA451C3A0AE3C7F85746C7EDB6" blockId="2.[151,1436,1943,2004]" pageId="2" pageNumber="135">
<emphasis id="B90EEAA8451C3A0AE3C7F8574540EDCD" bold="true" box="[151,275,1943,1967]" pageId="2" pageNumber="135">FIGURE 1.</emphasis>
<taxonomicName id="4C7A4D39451C3A0AE24AF8584641EDD2" authorityName="Donnelly" authorityYear="1989" box="[282,530,1944,1968]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="135" phylum="Arthropoda" rank="species" species="sulcatistyla">
<emphasis id="B90EEAA8451C3A0AE24AF8584641EDD2" box="[282,530,1944,1968]" italics="true" pageId="2" pageNumber="135">Epigomphus sulcatistyla</emphasis>
</taxonomicName>
: a) last stadium male exuvia, dorsal view; b)
<emphasis id="B90EEAA8451C3A0AE0B0F8594049EDD0" box="[992,1050,1945,1970]" italics="true" pageId="2" pageNumber="135">Ídem,</emphasis>
ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view.
</paragraph>
</caption>
<caption id="DF056632451D3A0BE3C7F85746CBED9A" ID-DOI="http://doi.org/10.5281/zenodo.11084736" ID-Zenodo-Dep="11084736" httpUri="https://zenodo.org/record/11084736/files/figure.png" pageId="3" pageNumber="136" startId="3.[151,250,1943,1967]" targetBox="[236,1352,181,1920]" targetPageId="3" targetType="figure">
<paragraph id="8BC536BA451D3A0BE3C7F85746CBED9A" blockId="3.[151,1436,1943,2040]" pageId="3" pageNumber="136">
<emphasis id="B90EEAA8451D3A0BE3C7F8574546EDCD" bold="true" box="[151,277,1943,1967]" pageId="3" pageNumber="136">FIGURE 2.</emphasis>
<emphasis id="B90EEAA8451D3A0BE24EF8584671EDD2" box="[286,546,1944,1968]" italics="true" pageId="3" pageNumber="136">
<taxonomicName id="4C7A4D39451D3A0BE24EF858464FEDD2" authorityName="Donnelly" authorityYear="1989" box="[286,540,1944,1968]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="sulcatistyla">Epigomphus sulcatistyla</taxonomicName>
:
</emphasis>
a) antennae of a young larva, dorsal view; b) prementum of the last stadium exuvia, ventral view; c) detail of the palps and ligula, ventral view; d) S7–10 and caudal appendages of male, dorsal view; e) sterna 9–10 of the female showing the vestigial gonapophyses.
</paragraph>
</caption>
<caption id="DF056632451A3A0CE3C7F857465EED95" ID-DOI="http://doi.org/10.5281/zenodo.11084742" ID-Zenodo-Dep="11084742" httpUri="https://zenodo.org/record/11084742/files/figure.png" pageId="4" pageNumber="137" startId="4.[151,250,1943,1967]" targetBox="[432,1155,181,1919]" targetPageId="4" targetType="figure">
<paragraph id="8BC536BA451A3A0CE3C7F857465EED95" blockId="4.[151,1436,1943,2039]" pageId="4" pageNumber="137">
<emphasis id="B90EEAA8451A3A0CE3C7F8574544EDCD" bold="true" box="[151,279,1943,1967]" pageId="4" pageNumber="137">FIGURE 3.</emphasis>
Details of the mouthparts of the last stadium exuvia of
<taxonomicName id="4C7A4D39451A3A0CE028F8574027EDCD" authorityName="Donnelly" authorityYear="1989" box="[888,1140,1943,1967]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="sulcatistyla">
<emphasis id="B90EEAA8451A3A0CE028F8574027EDCD" box="[888,1140,1943,1967]" italics="true" pageId="4" pageNumber="137">Epigomphus sulcatistyla</emphasis>
</taxonomicName>
: a–d) mandibles: a–b, right mandible, ventral and mesal view, respectively; c–d, left mandible, ventro-mesal and mesal view, respectively; e–f) left maxilla, ventral and mesal view, respectively.
</paragraph>
</caption>
<paragraph id="8BC536BA451B3A0DE397FEC34633E8E7" blockId="5.[151,1437,151,1401]" pageId="5" pageNumber="138">
<emphasis id="B90EEAA8451B3A0DE397FEC3454BEB7F" box="[199,280,259,285]" italics="true" pageId="5" pageNumber="138">Thorax</emphasis>
: Prothorax slightly narrower than head (
<figureCitation id="13412A3F451B3A0DE18AFEC3477BEB7F" box="[730,808,259,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-23@2.[194,1394,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus sulcatistyla: a) last stadium male exuvia, dorsal view; b) Ídem, ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11084734" httpUri="https://zenodo.org/record/11084734/files/figure.png" pageId="5" pageNumber="138">Fig. 1a</figureCitation>
), setose on inferior border. Anterior margin of pronotum slightly concave, lateral margins convex and bulging, posterior margin convex; a large, subquadrate glabrous area on each side of midline, remainder of pronotum with sparse, minute, scale-like setae. Pterothorax mostly bare, with minute, sparse, scale-like setae and some tufts of long, curled setae on inferior margins, mesospiracles well-developed, metaspiracles reduced. Legs short (
<figureCitation id="13412A3F451B3A0DE1F9FE5346B9EBCF" box="[681,746,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-23@2.[194,1394,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus sulcatistyla: a) last stadium male exuvia, dorsal view; b) Ídem, ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11084734" httpUri="https://zenodo.org/record/11084734/files/figure.png" pageId="5" pageNumber="138">Fig. 1</figureCitation>
) (when fully extended, hind legs scarcely reaching posterior margin of S7), strongly setose, with long, yellow-brown, delicate setae mainly on sides and shorter, stiff, reddish setae mainly on anterior surfaces of tibiae and tarsi; burrowing hooks moderately developed (
<figureCitation id="13412A3F451B3A0DE793FE1B4141EB97" box="[1219,1298,475,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-23@2.[194,1394,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus sulcatistyla: a) last stadium male exuvia, dorsal view; b) Ídem, ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11084734" httpUri="https://zenodo.org/record/11084734/files/figure.png" pageId="5" pageNumber="138">Figs. 1</figureCitation>
,
<figureCitation id="13412A3F451B3A0DE672FE1B4112EB97" box="[1314,1345,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[236,1352,181,1920]" captionTargetId="figure-17@3.[236,1352,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Epigomphus sulcatistyla: a) antennae of a young larva, dorsal view; b) prementum of the last stadium exuvia, ventral view; c) detail of the palps and ligula, ventral view; d) S7–10 and caudal appendages of male, dorsal view; e) sterna 9–10 of the female showing the vestigial gonapophyses." figureDoi="http://doi.org/10.5281/zenodo.11084736" httpUri="https://zenodo.org/record/11084736/files/figure.png" pageId="5" pageNumber="138">2b</figureCitation>
); dorsal margins of metafemora same length as metatibiae (
<figureCitation id="13412A3F451B3A0DE183FE3F4770E87B" box="[723,803,511,537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-23@2.[194,1394,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus sulcatistyla: a) last stadium male exuvia, dorsal view; b) Ídem, ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11084734" httpUri="https://zenodo.org/record/11084734/files/figure.png" pageId="5" pageNumber="138">Fig. 1a</figureCitation>
); tarsal formula 2-2-3, claws simple, with a pulvilliform empodium, metatarsal claws yellow-brown on basal 0.75, distal 0.25 reddish-brown. Wing sheaths scarcely surpassing anterior margin of S4, strongly divergent (
<figureCitation id="13412A3F451B3A0DE1B9FD87476BE803" box="[745,824,583,609]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-23@2.[194,1394,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus sulcatistyla: a) last stadium male exuvia, dorsal view; b) Ídem, ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11084734" httpUri="https://zenodo.org/record/11084734/files/figure.png" pageId="5" pageNumber="138">Fig. 1a</figureCitation>
), mostly bare, with sparse, minute scale-like setae, and abundant long, delicate setae on borders.
</paragraph>
<paragraph id="8BC536BA451B3A0DE397FD4F4509EEC3" blockId="5.[151,1437,151,1401]" pageId="5" pageNumber="138">
<emphasis id="B90EEAA8451B3A0DE397FD4F4562E8CB" box="[199,305,655,681]" italics="true" pageId="5" pageNumber="138">Abdomen</emphasis>
: Yellow-brown, with a complex pattern of dark and pale areas only visible in larvae not exuviae (
<figureCitation id="13412A3F451B3A0DE3CFFD7344A1E8AF" box="[159,242,691,717]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-23@2.[194,1394,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus sulcatistyla: a) last stadium male exuvia, dorsal view; b) Ídem, ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11084734" httpUri="https://zenodo.org/record/11084734/files/figure.png" pageId="5" pageNumber="138">Fig. 1c</figureCitation>
), more or less spindle-shaped, ventral surface flat, dorsal surface convex, lacking dorsal protuberances, widest on S5; lateral margins of S1–9 with long, stiff yellowish setae, with scale-like setae on S10, lateral margins of S8–9 serrate, serrations stout and sharp, S7–9 ending in slightly upturned, sharply pointed spines increasing in size and robustness posteriorly, those of S7–8 slightly divergent (
<figureCitation id="13412A3F451B3A0DE0F6FCDF47AEE95B" box="[934,1021,799,825]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[236,1352,181,1920]" captionTargetId="figure-17@3.[236,1352,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Epigomphus sulcatistyla: a) antennae of a young larva, dorsal view; b) prementum of the last stadium exuvia, ventral view; c) detail of the palps and ligula, ventral view; d) S7–10 and caudal appendages of male, dorsal view; e) sterna 9–10 of the female showing the vestigial gonapophyses." figureDoi="http://doi.org/10.5281/zenodo.11084736" httpUri="https://zenodo.org/record/11084736/files/figure.png" pageId="5" pageNumber="138">Fig. 2d</figureCitation>
); tergites 2–10 with combined short and long, delicate setae mainly on dorsolateral thirds and also with minute scale-like setae, all of them sparse except on posterior margins where scale-like setae are abundant; tergites 3–8 with a thin, longitudinal, middle suture. Sternites following same color pattern as tergites; sternites 3–8 divided into five plates, sternites 2 and 9 divided into three plates, ventral sutures parallel on 2–3, diverging gradually on 4–8, strongly divergent on 9. Male gonapophyses lacking, with evident circular gonopore (
<figureCitation id="13412A3F451B3A0DE05CFC13470CE98F" box="[780,863,979,1005]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-23@2.[194,1394,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus sulcatistyla: a) last stadium male exuvia, dorsal view; b) Ídem, ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11084734" httpUri="https://zenodo.org/record/11084734/files/figure.png" pageId="5" pageNumber="138">Fig. 1b</figureCitation>
); female gonapophyses vestigial, roundly-pointed, slightly divergent distally (
<figureCitation id="13412A3F451B3A0DE282FC37467BEE73" box="[466,552,1015,1041]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[236,1352,181,1920]" captionTargetId="figure-17@3.[236,1352,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Epigomphus sulcatistyla: a) antennae of a young larva, dorsal view; b) prementum of the last stadium exuvia, ventral view; c) detail of the palps and ligula, ventral view; d) S7–10 and caudal appendages of male, dorsal view; e) sterna 9–10 of the female showing the vestigial gonapophyses." figureDoi="http://doi.org/10.5281/zenodo.11084736" httpUri="https://zenodo.org/record/11084736/files/figure.png" pageId="5" pageNumber="138">Fig. 2e</figureCitation>
). Caudal appendages (
<figureCitation id="13412A3F451B3A0DE062FC3747D7EE73" box="[818,900,1015,1041]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-23@2.[194,1394,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Epigomphus sulcatistyla: a) last stadium male exuvia, dorsal view; b) Ídem, ventral view; c) Young larva showing the complex abdominal color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11084734" httpUri="https://zenodo.org/record/11084734/files/figure.png" pageId="5" pageNumber="138">Figs. 1</figureCitation>
,
<figureCitation id="13412A3F451B3A0DE0C6FC3747E7EE73" box="[918,948,1015,1041]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[236,1352,181,1920]" captionTargetId="figure-17@3.[236,1352,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Epigomphus sulcatistyla: a) antennae of a young larva, dorsal view; b) prementum of the last stadium exuvia, ventral view; c) detail of the palps and ligula, ventral view; d) S7–10 and caudal appendages of male, dorsal view; e) sterna 9–10 of the female showing the vestigial gonapophyses." figureDoi="http://doi.org/10.5281/zenodo.11084736" httpUri="https://zenodo.org/record/11084736/files/figure.png" pageId="5" pageNumber="138">2d</figureCitation>
) dark reddish brown, pyramidal, sharplypointed, with abundant setae on internal margins, with scale-like setae on external margins of cerci and paraprocts, external margins concave, tips of cerci and paraprocts more or less parallel, paraprocts and cerci subequal, epiproct slightly longer; male epiproct with a pair of bare dorsal tubercles along basal 0.50, rounded apically and divergent distally (
<figureCitation id="13412A3F451B3A0DE3A9FB47451EEEC3" box="[249,333,1159,1185]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[236,1352,181,1920]" captionTargetId="figure-17@3.[236,1352,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Epigomphus sulcatistyla: a) antennae of a young larva, dorsal view; b) prementum of the last stadium exuvia, ventral view; c) detail of the palps and ligula, ventral view; d) S7–10 and caudal appendages of male, dorsal view; e) sterna 9–10 of the female showing the vestigial gonapophyses." figureDoi="http://doi.org/10.5281/zenodo.11084736" httpUri="https://zenodo.org/record/11084736/files/figure.png" pageId="5" pageNumber="138">Fig. 2d</figureCitation>
).
</paragraph>
<paragraph id="8BC536BA451B3A0DE397FB6B4487EF53" blockId="5.[151,1437,151,1401]" pageId="5" pageNumber="138">
<emphasis id="B90EEAA8451B3A0DE397FB6B46AAEEA7" bold="true" box="[199,761,1194,1221]" pageId="5" pageNumber="138">Measurements [in mm, averages in brackets]:</emphasis>
Exuviae (N = 7): TL 25.8–30.0 [27.8]; AL (ventral, excl. caudal app.) 15–17 [16.2]; MWh 5.8–6.5 [6.1]; HfL (lateral) 5.0–5.2 [5.1]; MWa (ventral) 6.8–8.0 [7.3]; Ep 1.1–1.5 [1.4], Ce 1.1–1.4 [1.3], Pp 1.1–1.4 [1.3]; spine on S7 0.2-0.3 [0.2], spine on S8 0.4-0.5 [0.4], spine on S9 0.6-0.8 [0.7].
</paragraph>
</subSubSection>
<subSubSection id="C3606531451B3A0DE397FAFA46EDEF1B" pageId="5" pageNumber="138" type="biology_ecology">
<paragraph id="8BC536BA451B3A0DE397FAFA46EDEF1B" blockId="5.[151,1437,151,1401]" pageId="5" pageNumber="138">
<emphasis id="B90EEAA8451B3A0DE397FAFA4579EF36" bold="true" box="[199,298,1338,1364]" pageId="5" pageNumber="138">Habitat.</emphasis>
Larvae were found in small, shallow streams of 2nd and 3rd order, at sluggish margins in shady places, inhabiting banks of fine sand, mud and leaf litter.
</paragraph>
</subSubSection>
<subSubSection id="C3606531451B3A00E3C7FA0B46A4E893" lastPageId="8" lastPageNumber="141" pageId="5" pageNumber="138" type="description">
<paragraph id="8BC536BA451B3A0DE3C7FA0B45D5EF86" blockId="5.[151,390,1482,1545]" box="[151,390,1482,1509]" pageId="5" pageNumber="138">
<heading id="D08D81D6451B3A0DE3C7FA0B45D5EF86" bold="true" box="[151,390,1482,1509]" fontSize="11" level="1" pageId="5" pageNumber="138" reason="1">
<emphasis id="B90EEAA8451B3A0DE3C7FA0B45D5EF86" bold="true" box="[151,390,1482,1509]" pageId="5" pageNumber="138">Female description</emphasis>
</heading>
</paragraph>
<paragraph id="8BC536BA451B3A0DE3C7FA2F44B8EC6B" blockId="5.[151,390,1482,1545]" box="[151,235,1519,1545]" pageId="5" pageNumber="138">
(
<figureCitation id="13412A3F451B3A0DE3CFFA2F44B0EC6B" box="[159,227,1519,1545]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="5" pageNumber="138">Fig. 4</figureCitation>
)
</paragraph>
<paragraph id="8BC536BA451B3A0DE3C7F9F7401FEC33" blockId="5.[151,1437,1591,2013]" box="[151,1100,1591,1617]" pageId="5" pageNumber="138">The following description is based upon four teneral females reared from F-0 larvae.</paragraph>
<paragraph id="8BC536BA451B3A0DE397F99B4178EDBF" blockId="5.[151,1437,1591,2013]" pageId="5" pageNumber="138">
<emphasis id="B90EEAA8451B3A0DE397F99B4557EC17" box="[199,260,1627,1653]" italics="true" pageId="5" pageNumber="138">Head</emphasis>
: Labrum grayish, ventrolateral margins black, distal margin fringed with dark brown, bristle-like setae, some scattered bristle-like setae on surface to each side of midline (
<figureCitation id="13412A3F451B3A0DE0C1F9BF47B7ECFB" box="[913,996,1663,1689]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="5" pageNumber="138">Fig. 4a</figureCitation>
); sides of mandibles and genae creamy pale; anteclypeus distally grayish, yellow basally, postclypeus light brown, wider than anteclypeus with the anterior corners, rounded, bulging, and with marginal, black, bristle-like setae (
<figureCitation id="13412A3F451B3A0DE093F907404BEC83" box="[963,1048,1735,1761]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="5" pageNumber="138">Fig. 4a</figureCitation>
); antefrons light brown, postfrons shallowly concave at middle, with a large, pale spot to each side of midline and with scattered, black, bristle-like setae (
<figureCitation id="13412A3F451B3A0DE38EF8CF4504ED4B" box="[222,343,1807,1833]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="5" pageNumber="138">Figs. 4a–b</figureCitation>
); vertex brown, distally striated, ocelli yellow, with a low, mound-like tubercle behind each lateral ocellus (
<figureCitation id="13412A3F451B3A0DE3AAF8F34501ED2F" box="[250,338,1843,1869]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="5" pageNumber="138">Fig. 4b</figureCitation>
). Compound eyes grayish-yellow (
<figureCitation id="13412A3F451B3A0DE1BAF8F34736ED2F" box="[746,869,1843,1869]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="5" pageNumber="138">Figs. 4a–b</figureCitation>
) (pale grayish-blue when alive). Occiput mostly yellowish-brown, brown at sides, striated on anterior half (
<figureCitation id="13412A3F451B3A0DE07FF89747D0ED13" box="[815,899,1879,1905]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="5" pageNumber="138">Fig. 4b</figureCitation>
), with a large concavity to each side of midline (
<figureCitation id="13412A3F451B3A0DE3CFF8BB454AEDF7" box="[159,281,1915,1941]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="5" pageNumber="138">Figs. 4b–c</figureCitation>
), posterior margin sinuose, concave at middle (
<figureCitation id="13412A3F451B3A0DE07CF8BB47D3EDF7" box="[812,896,1915,1941]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="5" pageNumber="138">Fig. 4b</figureCitation>
), with a small, subglobose, tubercle at each side (
<figureCitation id="13412A3F451B3A0DE3CFF85F44A7EDDB" box="[159,244,1951,1977]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="5" pageNumber="138">Fig. 4b</figureCitation>
). Rear of head light brown, a large ear-like pit on postgenae (
<emphasis id="B90EEAA8451B3A0DE0F3F85F47B2EDDA" box="[931,993,1951,1976]" italics="true" pageId="5" pageNumber="138">sensu</emphasis>
<bibRefCitation id="EFEB4B4B451B3A0DE0B8F85F4029EDDB" author="Calvert, P. P." box="[1000,1146,1951,1977]" pageId="5" pageNumber="138" pagination="323 - 354" refId="ref4417" refString="Calvert, P. P. (1920) The Costa Rican species of Epigomphus and their mutual mating adaptations (Odonata). Transactions of the American Entomological Society, 46, 323 - 354." type="journal article" year="1920">Calvert 1920</bibRefCitation>
) (
<figureCitation id="13412A3F451B3A0DE7C2F85F40B0EDDB" box="[1170,1251,1951,1977]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="5" pageNumber="138">Fig. 4c</figureCitation>
), each pit with a mesal and lateral ridge, the mesal ridge followed by vertical, well-marked, impressed striae (
<figureCitation id="13412A3F451B3A0DE7F5F8034173EDBF" box="[1189,1312,1987,2013]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="5" pageNumber="138">Figs. 4b–c</figureCitation>
).
</paragraph>
<caption id="DF05663245183A0EE3C7F857474BED9A" ID-DOI="http://doi.org/10.5281/zenodo.11084744" ID-Zenodo-Dep="11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="6" pageNumber="139" startId="6.[151,250,1943,1967]" targetBox="[170,1417,203,1893]" targetPageId="6" targetType="figure">
<paragraph id="8BC536BA45183A0EE3C7F857474BED9A" blockId="6.[151,1436,1943,2040]" pageId="6" pageNumber="139">
<emphasis id="B90EEAA845183A0EE3C7F8574541EDCD" bold="true" box="[151,274,1943,1967]" pageId="6" pageNumber="139">FIGURE 4.</emphasis>
Details of the morphology of the female imago of
<taxonomicName id="4C7A4D3945183A0EE042F8584050EDD2" authorityName="Donnelly" authorityYear="1989" box="[786,1027,1944,1968]" class="Insecta" family="Gomphidae" genus="Epgomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="139" phylum="Arthropoda" rank="species" species="sulcatistyla">
<emphasis id="B90EEAA845183A0EE042F8584050EDD2" box="[786,1027,1944,1968]" italics="true" pageId="6" pageNumber="139">Epgomphus sulcatistyla</emphasis>
</taxonomicName>
: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina.
</paragraph>
</caption>
<caption id="DF05663245193A0FE3C7F8B44544ED9A" ID-DOI="http://doi.org/10.5281/zenodo.11084746" ID-Zenodo-Dep="11084746" httpUri="https://zenodo.org/record/11084746/files/figure.png" pageId="7" pageNumber="140" startId="7.[151,250,1908,1932]" targetBox="[192,1395,196,1860]" targetPageId="7" targetType="figure">
<paragraph id="8BC536BA45193A0FE3C7F8B44544ED9A" blockId="7.[151,1436,1908,2040]" pageId="7" pageNumber="140">
<emphasis id="B90EEAA845193A0FE3C7F8B44540EDEE" bold="true" box="[151,275,1908,1932]" pageId="7" pageNumber="140">FIGURE 5.</emphasis>
Details of the morphology of the female imagoes of
<taxonomicName id="4C7A4D3945193A0FE062F8B447B6EDEE" box="[818,997,1908,1932]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="140" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90EEAA845193A0FE062F8B447E2EDEE" box="[818,945,1908,1932]" italics="true" pageId="7" pageNumber="140">Epigomphus</emphasis>
spp.
</taxonomicName>
: a, c, e, g, and h)
<taxonomicName id="4C7A4D3945193A0FE7C9F8B4412EEDEE" baseAuthorityName="Gonzalez-Soriano &amp; Cook" baseAuthorityYear="1988" box="[1177,1405,1908,1932]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="140" phylum="Arthropoda" rank="species" species="donnellyi">
<emphasis id="B90EEAA845193A0FE7C9F8B4412EEDEE" box="[1177,1405,1908,1932]" italics="true" pageId="7" pageNumber="140">Epigomphus donnellyi</emphasis>
</taxonomicName>
: a, c) head, dorsoposterior and dorsal view, respectively; b, d, and f)
<taxonomicName id="4C7A4D3945193A0FE07AF858404CEDD2" baseAuthorityName="Calvert" baseAuthorityYear="1903" box="[810,1055,1944,1968]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="140" phylum="Arthropoda" rank="species" species="subobtusus">
<emphasis id="B90EEAA845193A0FE07AF858404CEDD2" box="[810,1055,1944,1968]" italics="true" pageId="7" pageNumber="140">Epigomphus subobtusus</emphasis>
</taxonomicName>
: b, d) head, dorsal and dorsoposterior view, respectively; e) vulvar lamina, ventral view; f)
<emphasis id="B90EEAA845193A0FE1E5F87E46BBEDB5" box="[693,744,1982,2007]" italics="true" pageId="7" pageNumber="140">Ídem</emphasis>
; g-h) S9–10 and caudal appendages in dorsal and left lateral views, respectively.
</paragraph>
</caption>
<paragraph id="8BC536BA45163A00E397FF574516EBB3" blockId="8.[151,1437,151,753]" pageId="8" pageNumber="141">
<emphasis id="B90EEAA845163A00E397FF57454BEAD3" box="[199,280,151,177]" italics="true" pageId="8" pageNumber="141">Thorax</emphasis>
: Prothorax pale brown (
<figureCitation id="13412A3F45163A00E171FF574620EAD3" box="[545,627,151,177]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="8" pageNumber="141">Fig. 4d</figureCitation>
), middle lobe pale at sides with paired yellow mesal spots on the posterior margin; posterior lobe brown. Pterothorax pale brown above, pale below (
<figureCitation id="13412A3F45163A00E0B8FF7B4037EAB7" box="[1000,1124,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="8" pageNumber="141">Figs. 4d–e</figureCitation>
), yellow stripes as follows: 1
<superScript id="7C0F9BF245163A00E3F5FF1F44E3EA8F" attach="left" box="[165,176,223,237]" fontSize="6" pageId="8" pageNumber="141">st</superScript>
antehumeral stripe not connected to pale anterior collar (
<figureCitation id="13412A3F45163A00E06DFF1F47C3EA9B" box="[829,912,223,249]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="8" pageNumber="141">Fig. 4e</figureCitation>
), 2
<superScript id="7C0F9BF245163A00E0E4FF1E4797EA8E" attach="left" box="[948,964,222,236]" fontSize="6" pageId="8" pageNumber="141">nd</superScript>
antehumeral stripe thin, wider at anterior end reaching the yellow mesokatepisternum (
<figureCitation id="13412A3F45163A00E1C9FEC346BEEB7F" box="[665,749,259,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="8" pageNumber="141">Fig. 4e</figureCitation>
); mesepimeral stripe subequal in width throughout, reaching the subalar carina; metepisternal stripe wide, narrowing gradually posteriorly not reaching the subalar carina and connected anteriorly with the yellow metakatepisternum; metepisternum mostly light yellow; metasternum pale (
<figureCitation id="13412A3F45163A00E3CFFEAF44A0EBEB" box="[159,243,367,393]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="8" pageNumber="141">Fig. 4e</figureCitation>
); antealar crest and sinuses pale brown, pale laterally. Legs (
<figureCitation id="13412A3F45163A00E0F9FEAF47ADEBEB" box="[937,1022,367,393]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="8" pageNumber="141">Fig. 4e</figureCitation>
): coxae yellow, trochanter and basal half of femora grayish, tibiae brownish-black, tarsi and spines armature black. Wing venation black, pterostigmata grayish-yellow.
</paragraph>
<paragraph id="8BC536BA45163A00E397FE1B46C8E8AF" blockId="8.[151,1437,151,753]" pageId="8" pageNumber="141">
<emphasis id="B90EEAA845163A00E397FE1B4562EB97" box="[199,305,475,501]" italics="true" pageId="8" pageNumber="141">Abdomen</emphasis>
: Dorsally brown, laterally pale on S1, S2 with a full-length pale stripe, S3 with a pale stripe on basal 0.80, S4–6 with a pale ring on basal 0.33, S7 with basal half pale, S8–10 light brown with some irregular yellow spots (
<figureCitation id="13412A3F45163A00E267FDE345DBE85F" box="[311,392,547,573]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="8" pageNumber="141">Fig. 4f</figureCitation>
), S10 yellow laterally (
<figureCitation id="13412A3F45163A00E1C4FDE346BAE85F" box="[660,745,547,573]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="8" pageNumber="141">
Fig.
<quantity id="4C829B5F45163A00E19AFDE346BAE85F" box="[714,745,547,573]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="8" pageNumber="141" unit="g" value="4.0">4g</quantity>
</figureCitation>
). Caudal appendages (
<figureCitation id="13412A3F45163A00E0BBFDE34036E85F" box="[1003,1125,547,573]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="8" pageNumber="141">Figs. 4f–g</figureCitation>
): epiproct light brown with posterior margin black, distally roundly-pointed; cerci reddish-brown, acutely-pointed, the longest, 4.8x as long as its maximum width; paraprocts yellowish-brown, with dorsal borders creamy pale, roundly-pointed, the shortest. Vulvar lamina (
<figureCitation id="13412A3F45163A00E213FD4F45C7E8CB" box="[323,404,655,681]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[170,1417,203,1893]" captionTargetId="figure-23@6.[170,1417,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Details of the morphology of the female imago of Epgomphus sulcatistyla: a–c) head: frontal, dorsal, and posterior views, respectively; d) thorax, dorsal view; e) head and thorax, lateral view; f–h) S9–10 and caudal appendages: f) dorsal view; g) left lateral view; h) ventral view, showing the vulvar lamina." figureDoi="http://doi.org/10.5281/zenodo.11084744" httpUri="https://zenodo.org/record/11084744/files/figure.png" pageId="8" pageNumber="141">Fig. 4h</figureCitation>
) widely U-shaped, reaching basal half of sternum 9, with tips strongly divergent, each branch comprising 0.75 of vulvar lamina total length.
</paragraph>
<paragraph id="8BC536BA45163A00E397FD1746A4E893" blockId="8.[151,1437,151,753]" box="[199,759,726,753]" pageId="8" pageNumber="141">
<emphasis id="B90EEAA845163A00E397FD17458EE892" bold="true" box="[199,477,726,753]" pageId="8" pageNumber="141">Measurements (in mm)</emphasis>
: AL 38–39; HwL 41–42.
</paragraph>
</subSubSection>
</treatment>
</document>
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