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<document id="05945598728D7E7BEB651248F8D62A5E" ID-CLB-Dataset="31254" ID-DOI="10.11646/zootaxa.4362.1.3" ID-GBIF-Dataset="e7983c67-0ce5-4b7f-9373-8782b86e059b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1076176" ID-ZooBank="61A41EC1-2557-429D-B47C-595D12F2D8E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1512393169915" checkinUser="plazi" docAuthor="Polotow, Daniele &amp; Griswold, Charles" docDate="2017" docId="03D38783FF86FF85E6B368A3FEBA617E" docLanguage="en" docName="zootaxa.4362.1.3.pdf" docOrigin="Zootaxa 4362 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Campostichomma Karsch 1892" docType="treatment" docVersion="7" lastPageNumber="54" masterDocId="FFEAFFFBFF84FF86E6246E6DFFC96633" masterDocTitle="Cleaning old cabinets: revealing the taxonomy of Sri Lankan wolf spiders (Araneae, Udubidae and Zoropsidae)" masterLastPageNumber="74" masterPageNumber="51" pageNumber="53" updateTime="1698506327670" updateUser="ExternalLinkService">
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<mods:title id="E1E8881269C4F8D432B500763FB6E10B">Cleaning old cabinets: revealing the taxonomy of Sri Lankan wolf spiders (Araneae, Udubidae and Zoropsidae)</mods:title>
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<mods:namePart id="16E865791E29AE63A9B0C360CDC3C0D9">Polotow, Daniele</mods:namePart>
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<mods:namePart id="A5D380A65871DBDCEED24A63D6E5887F">Griswold, Charles</mods:namePart>
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<treatment id="03D38783FF86FF85E6B368A3FEBA617E" ID-DOI="http://doi.org/10.5281/zenodo.5999768" ID-GBIF-Taxon="137584768" ID-Zenodo-Dep="5999768" LSID="urn:lsid:plazi:treatment:03D38783FF86FF85E6B368A3FEBA617E" httpUri="http://treatment.plazi.org/id/03D38783FF86FF85E6B368A3FEBA617E" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53">
<subSubSection id="C360651EFF86FF84E6B368A3FDD460DB" box="[151,541,1742,1768]" pageId="2" pageNumber="53" type="nomenclature">
<paragraph id="8BC53695FF86FF84E6B368A3FDD460DB" blockId="2.[151,541,1742,1768]" box="[151,541,1742,1768]" pageId="2" pageNumber="53">
<heading id="D08D81F9FF86FF84E6B368A3FDD460DB" bold="true" box="[151,541,1742,1768]" fontSize="11" level="1" pageId="2" pageNumber="53" reason="1">
<taxonomicName id="4C7A4D16FF86FF84E6B368A3FDD460DB" authority="Karsch, 1892" authorityName="Karsch" authorityYear="1892" box="[151,541,1742,1768]" class="Arachnida" family="Zorocratidae" genus="Campostichomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B90EEA87FF86FF84E6B368A3FDD460DB" bold="true" box="[151,541,1742,1768]" pageId="2" pageNumber="53">
<emphasis id="B90EEA87FF86FF84E6B368A3FEB860DB" bold="true" box="[151,369,1742,1768]" italics="true" pageId="2" pageNumber="53">Campostichomma</emphasis>
Karsch, 1892
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C360651EFF86FF84E6B36978FC69619E" pageId="2" pageNumber="53" type="reference_group">
<paragraph id="8BC53695FF86FF84E6B36978FD85615B" blockId="2.[151,1436,1813,1896]" pageId="2" pageNumber="53">
<treatmentCitationGroup id="AB6A11BBFF86FF84E6B36978FD85615B" pageId="2" pageNumber="53">
<taxonomicName id="4C7A4D16FF86FF84E6B36978FE2E6118" authority="KArsch, 1892" authorityName="Karsch" authorityYear="1892" box="[151,487,1813,1835]" class="Arachnida" family="Zorocratidae" genus="Campostichomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B90EEA87FF86FF84E6B36978FE876118" box="[151,334,1813,1835]" italics="true" pageId="2" pageNumber="53">Campostichomma</emphasis>
KArsch, 1892
</taxonomicName>
: 296, pl. 12, FIg. 19 (TypE spEcIEs by OrIgInAl dEsIgnAtIOn:
<taxonomicName id="4C7A4D16FF86FF84E24E6978FE986179" authority="KArsch, 1892" authorityName="Karsch" authorityYear="1892" class="Arachnida" family="Zorocratidae" genus="Campostichomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="manicatum">
<emphasis id="B90EEA87FF86FF84E24E6978FA556118" box="[1130,1436,1813,1835]" italics="true" pageId="2" pageNumber="53">Campostichomma manicatum</emphasis>
KArsch, 1892
</taxonomicName>
); SImOn 1898A: 7;
<treatmentCitation id="0ADB1084FF86FF84E4346959FD106179" author="LEhtInEn" box="[528,729,1844,1866]" page="221" pageId="2" pageNumber="53" year="1967">
<bibRefCitation id="EFEB4B64FF86FF84E4346959FD6F6179" author="LEhtInEn" box="[528,678,1844,1866]" pageId="2" pageNumber="53" refString="LEhtInEn, P. T. (1967) ClAssIFIcAtIOn OF thE crIbEllAtE spIdErs And sOmE AllIEd FAmIlIEs, wIth nOtEs On thE EVOlutIOn OF thE subOrdEr ArAnEOmOrphA. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">LEhtInEn 1967</bibRefCitation>
: 221
</treatmentCitation>
;
<treatmentCitation id="0ADB1084FF86FF84E4C76959FC5F6179" author="GrIswOld" box="[739,918,1844,1866]" page="7" pageId="2" pageNumber="53" year="1993">
<bibRefCitation id="EFEB4B64FF86FF84E4C76959FCB46179" author="GrIswOld" box="[739,893,1844,1866]" pageId="2" pageNumber="53" refString="GrIswOld, C. E. (1993) InVEstIgAtIOns IntO thE phylOgEny OF thE lycOsOId spIdErs And thEIr kIn (ArAchnIdA: ArAnEAE: LycOsOIdEA). Smithsonian Contributions to Zoology, 539, 1 - 39. https: // dOI. Org / 10.5479 / sI. 00810282.539" type="journal article" year="1993">GrIswOld 1993</bibRefCitation>
: 7
</treatmentCitation>
;
<treatmentCitation id="0ADB1084FF86FF84E5856959FB506179" author="GrIswOld" box="[929,1177,1844,1866]" page="59" pageId="2" pageNumber="53" year="1999">
<bibRefCitation id="EFEB4B64FF86FF84E5856959FBBA6179" author="GrIswOld" box="[929,1139,1844,1866]" pageId="2" pageNumber="53" refString="GrIswOld, C. E., COddIngtOn, J. A., PlAtnIck, N. I. &amp; FOrstEr, R. R. (1999) TOwArds A phylOgEny OF EntElEgynE spIdErs (ArAnEAE, ArAnEOmOrphAE, EntElEgynAE). Journal of Arachnology, 27, 53 - 63." type="journal article" year="1999">
GrIswOld
<emphasis id="B90EEA87FF86FF84E2216958FBF16179" box="[1029,1080,1844,1866]" italics="true" pageId="2" pageNumber="53">et al.</emphasis>
1999
</bibRefCitation>
: 59
</treatmentCitation>
;
<treatmentCitation id="0ADB1084FF86FF84E2876959FA556179" author="POlOtOw" box="[1187,1436,1844,1866]" page="151" pageId="2" pageNumber="53" year="2015">
<bibRefCitation id="EFEB4B64FF86FF84E2876959FAA46179" author="POlOtOw" box="[1187,1389,1844,1866]" pageId="2" pageNumber="53" refString="POlOtOw, D., CArmIchEAl, A. &amp; GrIswOld, C. E. (2015) TOtAl EVIdEncE AnAlysIs OF thE phylOgEnEtIc rElAtIOnshIps OF LycOsOIdEA spIdErs (ArAnEAE, EntElEgynAE). Invertebrate Systematics, 29, 124 - 163. https: // dOI. Org / 10.1071 / IS 14041" type="journal article" year="2015">
POlOtOw
<emphasis id="B90EEA87FF86FF84E2DA6958FAF86179" box="[1278,1329,1844,1866]" italics="true" pageId="2" pageNumber="53">et al.</emphasis>
2015
</bibRefCitation>
: 151
</treatmentCitation>
(trAnsFErrEd tO
<taxonomicName id="4C7A4D16FF86FF84E779693FFE0C615B" ID-CoL="HN5" authorityName="Polotow &amp; Griswold" authorityYear="2015" box="[349,453,1874,1896]" class="Arachnida" family="Udubidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="53" phylum="Arthropoda" rank="family">UdubIdAE</taxonomicName>
); WSC 2017.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC53695FF86FF84E6B369F8FC69619E" blockId="2.[151,1436,1941,2038]" box="[151,928,1941,1966]" pageId="2" pageNumber="53">
<emphasis id="B90EEA87FF86FF84E6B369F8FF1F619D" bold="true" box="[151,214,1941,1966]" pageId="2" pageNumber="53">Note.</emphasis>
Name is neuter in gender, hence
<taxonomicName id="4C7A4D16FF86FF84E46F69F8FC51619E" authorityName="Karsch" authorityYear="1892" box="[587,920,1941,1965]" class="Arachnida" family="Zorocratidae" genus="Campostichomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="manicatum">
<emphasis id="B90EEA87FF86FF84E46F69F8FC51619E" box="[587,920,1941,1965]" italics="true" pageId="2" pageNumber="53">Campostichomma manicatum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C360651EFF86FF85E6E369D4FD9C603B" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53" type="diagnosis">
<paragraph id="8BC53695FF86FF85E6E369D4FE2E672F" blockId="2.[151,1436,1941,2038]" lastBlockId="3.[151,1437,151,1869]" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53">
<emphasis id="B90EEA87FF86FF84E6E369D4FE8861E1" bold="true" box="[199,321,1977,2002]" pageId="2" pageNumber="53">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C7A4D16FF86FF84E79169D4FDB661E2" authorityName="Karsch" authorityYear="1892" box="[437,639,1977,2001]" class="Arachnida" family="Zorocratidae" genus="Campostichomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B90EEA87FF86FF84E79169D4FDB661E2" box="[437,639,1977,2001]" italics="true" pageId="2" pageNumber="53">Campostichomma</emphasis>
</taxonomicName>
can be distinguished from those of other
<taxonomicName id="4C7A4D16FF86FF84E27E69D4FB0E61E1" authorityName="Polotow &amp; Griswold" authorityYear="2015" box="[1114,1223,1977,2002]" class="Arachnida" family="Udubidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="53" phylum="Arthropoda" rank="family">Udubidae</taxonomicName>
by the palpal tibia length less than ½ length of cymbium (
<figureCitation id="13412A10FF86FF84E47369B0FD7B61C5" box="[599,690,2013,2038]" captionStart="FIGURE 3" captionStartId="7.[151,250,1732,1754]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@7.[151,1436,193,1711]" captionTargetPageId="7" captionText="FIGURE 3. Campostichomma manicatum KArsch, 1892. (A–D) mAlE lEFt pAlp; (A) VEntrAl VIEw, (B) rEtrOlAtErAl VIEw, (C) prOlAtErAl VIEw; (D) rIght pAlp, rEtrOlAtErAl VIEw, ExpAndEd; (E) mAlE lEg IV; (F–G) EpIgynum; (F) VEntrAl VIEw; (G) dOrsAl VIEw.AbbrEVIAtIOns:LO, lOckIng lObEs;TC, tIbIAl crAck." httpUri="https://zenodo.org/record/1076188/files/figure.png" pageId="2" pageNumber="53">Fig. 3A</figureCitation>
), the cymbium with a deep groove retrolaterally (
<figureCitation id="13412A10FF86FF84E2CC69B0FAAA61C5" box="[1256,1379,2013,2038]" captionStart="FIGURE 4" captionStartId="8.[151,250,1910,1932]" captionTargetBox="[151,1419,600,1845]" captionTargetId="figure@8.[149,1434,579,1940]" captionTargetPageId="8" captionText="FIGURE 4. Campostichomma manicatum KArsch, 1892. (A–C) mAlE lEFt pAlp; (A) VEntrAl VIEw; (B) prOlAtErAl VIEw; (C) rEtrOlAtErAl VIEw; (D–E) EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw. AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RG, rEtrOlAtErAl grOOVE OF cymbIum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs;S, spErmAthEcAE;ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076192/files/figure.png" pageId="2" pageNumber="53">Fig. 4A, C</figureCitation>
) and the embolus elongate, threadlike, extending 360
<taxonomicName id="4C7A4D16FF87FF85E4EF6EFAFCE26683" box="[715,811,151,176]" pageId="3" pageNumber="54" rank="subSpecies" subSpecies="around">˚ around</taxonomicName>
tegulum (
<figureCitation id="13412A10FF87FF85E5826EFAFBC16683" box="[934,1032,151,176]" captionStart="FIGURE 3" captionStartId="7.[151,250,1732,1754]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@7.[151,1436,193,1711]" captionTargetPageId="7" captionText="FIGURE 3. Campostichomma manicatum KArsch, 1892. (A–D) mAlE lEFt pAlp; (A) VEntrAl VIEw, (B) rEtrOlAtErAl VIEw, (C) prOlAtErAl VIEw; (D) rIght pAlp, rEtrOlAtErAl VIEw, ExpAndEd; (E) mAlE lEg IV; (F–G) EpIgynum; (F) VEntrAl VIEw; (G) dOrsAl VIEw.AbbrEVIAtIOns:LO, lOckIng lObEs;TC, tIbIAl crAck." httpUri="https://zenodo.org/record/1076188/files/figure.png" pageId="3" pageNumber="54">Figs 3D</figureCitation>
,
<figureCitation id="13412A10FF87FF85E23C6EF5FBF76683" box="[1048,1086,152,176]" captionStart="FIGURE 4" captionStartId="8.[151,250,1910,1932]" captionTargetBox="[151,1419,600,1845]" captionTargetId="figure@8.[149,1434,579,1940]" captionTargetPageId="8" captionText="FIGURE 4. Campostichomma manicatum KArsch, 1892. (A–C) mAlE lEFt pAlp; (A) VEntrAl VIEw; (B) prOlAtErAl VIEw; (C) rEtrOlAtErAl VIEw; (D–E) EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw. AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RG, rEtrOlAtErAl grOOVE OF cymbIum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs;S, spErmAthEcAE;ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076192/files/figure.png" pageId="3" pageNumber="54">4A</figureCitation>
). Females have a long, broad epigynal plate divided into medial and lateral lobes (
<figureCitation id="13412A10FF87FF85E4CD6ED1FC8E66E7" box="[745,839,188,213]" captionStart="FIGURE 4" captionStartId="8.[151,250,1910,1932]" captionTargetBox="[151,1419,600,1845]" captionTargetId="figure@8.[149,1434,579,1940]" captionTargetPageId="8" captionText="FIGURE 4. Campostichomma manicatum KArsch, 1892. (A–C) mAlE lEFt pAlp; (A) VEntrAl VIEw; (B) prOlAtErAl VIEw; (C) rEtrOlAtErAl VIEw; (D–E) EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw. AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RG, rEtrOlAtErAl grOOVE OF cymbIum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs;S, spErmAthEcAE;ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076192/files/figure.png" pageId="3" pageNumber="54">Figs 4D</figureCitation>
,
<figureCitation id="13412A10FF87FF85E5776ED1FC5E66E7" box="[851,919,188,212]" captionStart="FIGURE8" captionStartId="12.[151,250,1887,1909]" captionTargetBox="[167,1428,218,1840]" captionTargetId="figure@12.[151,1436,193,1866]" captionTargetPageId="12" captionText="FIGURE8. Campostichomma spp., FEmAlE gEnItAlIA.(A–B) C. alawala sp. nov., EpIgynum;(A) VEntrAl VIEw;(B) dOrsAlVIEw; (C–D) C. harasbedda sp. nov., EpIgynum; (C) VEntrAl VIEw; (D) dOrsAl VIEw;(E–F) C. mudduk sp. nov., EpIgynum; (E)VEntrAl VIEw; (F) dOrsAl VIEw. AbbrEVIAtIOns: CD, cOpulAtOry ducts; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAn lObEOF EpIgynum;S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076206/files/figure.png" pageId="3" pageNumber="54">8A–F</figureCitation>
), lateral lobes with (
<figureCitation id="13412A10FF87FF85E25A6ED1FAD166E7" box="[1150,1304,188,213]" captionStart="FIGURE8" captionStartId="12.[151,250,1887,1909]" captionTargetBox="[167,1428,218,1840]" captionTargetId="figure@12.[151,1436,193,1866]" captionTargetPageId="12" captionText="FIGURE8. Campostichomma spp., FEmAlE gEnItAlIA.(A–B) C. alawala sp. nov., EpIgynum;(A) VEntrAl VIEw;(B) dOrsAlVIEw; (C–D) C. harasbedda sp. nov., EpIgynum; (C) VEntrAl VIEw; (D) dOrsAl VIEw;(E–F) C. mudduk sp. nov., EpIgynum; (E)VEntrAl VIEw; (F) dOrsAl VIEw. AbbrEVIAtIOns: CD, cOpulAtOry ducts; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAn lObEOF EpIgynum;S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076206/files/figure.png" pageId="3" pageNumber="54">Fig. 8A, C, E</figureCitation>
) or without teeth (
<figureCitation id="13412A10FF87FF85E6F96EB2FEFF66CB" box="[221,310,223,248]" captionStart="FIGURE 4" captionStartId="8.[151,250,1910,1932]" captionTargetBox="[151,1419,600,1845]" captionTargetId="figure@8.[149,1434,579,1940]" captionTargetPageId="8" captionText="FIGURE 4. Campostichomma manicatum KArsch, 1892. (A–C) mAlE lEFt pAlp; (A) VEntrAl VIEw; (B) prOlAtErAl VIEw; (C) rEtrOlAtErAl VIEw; (D–E) EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw. AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RG, rEtrOlAtErAl grOOVE OF cymbIum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs;S, spErmAthEcAE;ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076192/files/figure.png" pageId="3" pageNumber="54">Fig. 4D</figureCitation>
), and internally the copulatory ducts have several loops, which can be used to identify the species of the genus (
<figureCitation id="13412A10FF87FF85E7346F69FEA2672F" box="[272,363,260,285]" captionStart="FIGURE 4" captionStartId="8.[151,250,1910,1932]" captionTargetBox="[151,1419,600,1845]" captionTargetId="figure@8.[149,1434,579,1940]" captionTargetPageId="8" captionText="FIGURE 4. Campostichomma manicatum KArsch, 1892. (A–C) mAlE lEFt pAlp; (A) VEntrAl VIEw; (B) prOlAtErAl VIEw; (C) rEtrOlAtErAl VIEw; (D–E) EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw. AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RG, rEtrOlAtErAl grOOVE OF cymbIum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs;S, spErmAthEcAE;ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076192/files/figure.png" pageId="3" pageNumber="54">Figs 4E</figureCitation>
,
<figureCitation id="13412A10FF87FF85E7526F69FE13672F" box="[374,474,260,284]" captionStart="FIGURE8" captionStartId="12.[151,250,1887,1909]" captionTargetBox="[167,1428,218,1840]" captionTargetId="figure@12.[151,1436,193,1866]" captionTargetPageId="12" captionText="FIGURE8. Campostichomma spp., FEmAlE gEnItAlIA.(A–B) C. alawala sp. nov., EpIgynum;(A) VEntrAl VIEw;(B) dOrsAlVIEw; (C–D) C. harasbedda sp. nov., EpIgynum; (C) VEntrAl VIEw; (D) dOrsAl VIEw;(E–F) C. mudduk sp. nov., EpIgynum; (E)VEntrAl VIEw; (F) dOrsAl VIEw. AbbrEVIAtIOns: CD, cOpulAtOry ducts; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAn lObEOF EpIgynum;S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076206/files/figure.png" pageId="3" pageNumber="54">8B, D, F</figureCitation>
).
</paragraph>
<paragraph id="8BC53695FF87FF85E6E36F4AFD9C603B" blockId="3.[151,1437,151,1869]" pageId="3" pageNumber="54">
<emphasis id="B90EEA87FF87FF85E6E36F4AFE906773" bold="true" box="[199,345,295,320]" pageId="3" pageNumber="54">DesCription.</emphasis>
Medium sized-spiders, total length 4.30–7.40; sexual dimorphism slight (
<figureCitation id="13412A10FF87FF85E2E56F4AFA8B6773" box="[1217,1346,295,320]" captionStart="FIGURE2" captionStartId="6.[151,250,1960,1982]" captionTargetBox="[235,1353,193,1941]" captionTargetId="figure@6.[235,1353,193,1941]" captionTargetPageId="6" captionText="FIGURE2. Campostichomma manicatumKArsch, 1892. (A) dOrsAl And (B) VEntrAl hAbItus OF mAlE;(C) dOrsAl And (D) FEmAlE hAbItus,VEntrAl VIEw; (E) mAlE And (F) FEmAlE OculAr ArEA, AntErIOrVIEw." httpUri="https://zenodo.org/record/1076184/files/figure.png" pageId="3" pageNumber="54">Fig. 2A–D</figureCitation>
), males smaller with relatively longer legs. Carapace pear-shaped in dorsal view, domed; fovea linear, deep (
<figureCitation id="13412A10FF87FF85E3366F21FA466757" box="[1298,1423,332,357]" captionStart="FIGURE2" captionStartId="6.[151,250,1960,1982]" captionTargetBox="[235,1353,193,1941]" captionTargetId="figure@6.[235,1353,193,1941]" captionTargetPageId="6" captionText="FIGURE2. Campostichomma manicatumKArsch, 1892. (A) dOrsAl And (B) VEntrAl hAbItus OF mAlE;(C) dOrsAl And (D) FEmAlE hAbItus,VEntrAl VIEw; (E) mAlE And (F) FEmAlE OculAr ArEA, AntErIOrVIEw." httpUri="https://zenodo.org/record/1076184/files/figure.png" pageId="3" pageNumber="54">Fig. 2A, C</figureCitation>
). Ocular area broad, ocular quadrangle trapezoidal, widest behind; eight eyes in 2 rows, anterior row slightly recurved, posterior row nearly straight (
<figureCitation id="13412A10FF87FF85E4726FF9FD04679F" box="[598,717,404,429]" captionStart="FIGURE2" captionStartId="6.[151,250,1960,1982]" captionTargetBox="[235,1353,193,1941]" captionTargetId="figure@6.[235,1353,193,1941]" captionTargetPageId="6" captionText="FIGURE2. Campostichomma manicatumKArsch, 1892. (A) dOrsAl And (B) VEntrAl hAbItus OF mAlE;(C) dOrsAl And (D) FEmAlE hAbItus,VEntrAl VIEw; (E) mAlE And (F) FEmAlE OculAr ArEA, AntErIOrVIEw." httpUri="https://zenodo.org/record/1076184/files/figure.png" pageId="3" pageNumber="54">Figs 2E–F</figureCitation>
,
<figureCitation id="13412A10FF87FF85E4FC6FF9FD32679E" box="[728,763,404,429]" captionStart="FIGURE5" captionStartId="9.[151,250,1741,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@9.[151,1436,193,1719]" captionTargetPageId="9" captionText="FIGURE5. Campostichomma alawalasp. nov., FEmAlE.(A) hAbItus,dOrsAl VIEw;(B) VEntrAl VIEw; (C)AntErIOr VIEw OF OculAr ArEA;(D–F) EpIgynum; (D)VEntrAl VIEw;(E) dOrsAl VIEw; (F) lAtErAl VIEw." httpUri="https://zenodo.org/record/1076194/files/figure.png" pageId="3" pageNumber="54">5C</figureCitation>
,
<figureCitation id="13412A10FF87FF85E5236FF9FCE3679E" box="[775,810,404,429]" captionStart="FIGURE 6" captionStartId="10.[151,250,1707,1729]" captionTargetBox="[151,1436,337,1685]" captionTargetId="figure@10.[151,1436,337,1685]" captionTargetPageId="10" captionText="FIGURE 6. Campostichomma harasbedda sp. nov., FEmAlE. (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA, nOtE dIVIdEd chIlum; (C) pEdIcEl,dOrsAl VIEw;(D–E) EpIgynum; (D) VEntrAl VIEw; (E)dOrsAl VIEw." httpUri="https://zenodo.org/record/1076198/files/figure.png" pageId="3" pageNumber="54">6B</figureCitation>
,
<figureCitation id="13412A10FF87FF85E5126FF9FC92679F" box="[822,859,404,428]" captionStart="FIGURE7" captionStartId="11.[151,250,1967,1989]" captionTargetBox="[151,1436,983,1945]" captionTargetId="figure@11.[151,1436,983,1945]" captionTargetPageId="11" captionText="FIGURE7. Campostichomma mudduk sp. nov., FEmAlE.(A) dOrsAl hAbItus;(B) AntErIOr VIEw OF OculAr ArEA;(C–D) EpIgynum; (C) VEntrAl VIEw;(D) dOrsAl VIEw." httpUri="https://zenodo.org/record/1076202/files/figure.png" pageId="3" pageNumber="54">7B</figureCitation>
). Clypeus low (
<figureCitation id="13412A10FF87FF85E22F6FF9FB4B679F" box="[1035,1154,404,429]" captionStart="FIGURE2" captionStartId="6.[151,250,1960,1982]" captionTargetBox="[235,1353,193,1941]" captionTargetId="figure@6.[235,1353,193,1941]" captionTargetPageId="6" captionText="FIGURE2. Campostichomma manicatumKArsch, 1892. (A) dOrsAl And (B) VEntrAl hAbItus OF mAlE;(C) dOrsAl And (D) FEmAlE hAbItus,VEntrAl VIEw; (E) mAlE And (F) FEmAlE OculAr ArEA, AntErIOrVIEw." httpUri="https://zenodo.org/record/1076184/files/figure.png" pageId="3" pageNumber="54">Figs 2E–F</figureCitation>
,
<figureCitation id="13412A10FF87FF85E2A96FF9FB79679E" box="[1165,1200,404,429]" captionStart="FIGURE5" captionStartId="9.[151,250,1741,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@9.[151,1436,193,1719]" captionTargetPageId="9" captionText="FIGURE5. Campostichomma alawalasp. nov., FEmAlE.(A) hAbItus,dOrsAl VIEw;(B) VEntrAl VIEw; (C)AntErIOr VIEw OF OculAr ArEA;(D–F) EpIgynum; (D)VEntrAl VIEw;(E) dOrsAl VIEw; (F) lAtErAl VIEw." httpUri="https://zenodo.org/record/1076194/files/figure.png" pageId="3" pageNumber="54">5C</figureCitation>
,
<figureCitation id="13412A10FF87FF85E2996FF9FB29679E" box="[1213,1248,404,429]" captionStart="FIGURE 6" captionStartId="10.[151,250,1707,1729]" captionTargetBox="[151,1436,337,1685]" captionTargetId="figure@10.[151,1436,337,1685]" captionTargetPageId="10" captionText="FIGURE 6. Campostichomma harasbedda sp. nov., FEmAlE. (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA, nOtE dIVIdEd chIlum; (C) pEdIcEl,dOrsAl VIEw;(D–E) EpIgynum; (D) VEntrAl VIEw; (E)dOrsAl VIEw." httpUri="https://zenodo.org/record/1076198/files/figure.png" pageId="3" pageNumber="54">6B</figureCitation>
,
<figureCitation id="13412A10FF87FF85E2C86FF9FAD9679F" box="[1260,1296,404,428]" captionStart="FIGURE7" captionStartId="11.[151,250,1967,1989]" captionTargetBox="[151,1436,983,1945]" captionTargetId="figure@11.[151,1436,983,1945]" captionTargetPageId="11" captionText="FIGURE7. Campostichomma mudduk sp. nov., FEmAlE.(A) dOrsAl hAbItus;(B) AntErIOr VIEw OF OculAr ArEA;(C–D) EpIgynum; (C) VEntrAl VIEw;(D) dOrsAl VIEw." httpUri="https://zenodo.org/record/1076202/files/figure.png" pageId="3" pageNumber="54">7B</figureCitation>
). Chelicerae stout with large boss, especially in female, pro- and retromargins of fang furrow with three and four stout teeth, respectively. Sternum broadly oval, bluntly curved or with small point posteriorly; labium with shallow basal notch; palpal coxae converging slightly anteriorly (
<figureCitation id="13412A10FF87FF85E4F76F92FC9B642B" box="[723,850,511,536]" captionStart="FIGURE2" captionStartId="6.[151,250,1960,1982]" captionTargetBox="[235,1353,193,1941]" captionTargetId="figure@6.[235,1353,193,1941]" captionTargetPageId="6" captionText="FIGURE2. Campostichomma manicatumKArsch, 1892. (A) dOrsAl And (B) VEntrAl hAbItus OF mAlE;(C) dOrsAl And (D) FEmAlE hAbItus,VEntrAl VIEw; (E) mAlE And (F) FEmAlE OculAr ArEA, AntErIOrVIEw." httpUri="https://zenodo.org/record/1076184/files/figure.png" pageId="3" pageNumber="54">Figs 2B, D</figureCitation>
), serrula in a single row along outer margin. Pedicel elongated and not fused (
<figureCitation id="13412A10FF87FF85E7966C49FDC3640E" box="[434,522,548,573]" captionStart="FIGURE 6" captionStartId="10.[151,250,1707,1729]" captionTargetBox="[151,1436,337,1685]" captionTargetId="figure@10.[151,1436,337,1685]" captionTargetPageId="10" captionText="FIGURE 6. Campostichomma harasbedda sp. nov., FEmAlE. (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA, nOtE dIVIdEd chIlum; (C) pEdIcEl,dOrsAl VIEw;(D–E) EpIgynum; (D) VEntrAl VIEw; (E)dOrsAl VIEw." httpUri="https://zenodo.org/record/1076198/files/figure.png" pageId="3" pageNumber="54">Fig. 6C</figureCitation>
). Legs moderately long; leg formula 4123; integument finely wrinkled (
<bibRefCitation id="EFEB4B64FF87FF85E3166C49FEE06453" author="Griswold" pageId="3" pageNumber="54" refString="GrIswOld, C. E. (1993) InVEstIgAtIOns IntO thE phylOgEny OF thE lycOsOId spIdErs And thEIr kIn (ArAchnIdA: ArAnEAE: LycOsOIdEA). Smithsonian Contributions to Zoology, 539, 1 - 39. https: // dOI. Org / 10.5479 / sI. 00810282.539" type="journal article" year="1993">Griswold 1993: fig. 77</bibRefCitation>
), with plumose setae (
<emphasis id="B90EEA87FF87FF85E4076C24FDA86453" box="[547,609,585,608]" italics="true" pageId="3" pageNumber="54">sensu</emphasis>
<bibRefCitation id="EFEB4B64FF87FF85E4436C2AFCC26453" author="Lehtinen" box="[615,779,583,608]" pageId="3" pageNumber="54" refString="LEhtInEn, P. T. (1975) NOtEs On thE phylOgEnEtIc clAssIFIcAtIOn OF ArAnEAE. Proceedings of the 6 th Arachnological Congress, 1975, 26 - 29." type="journal article" year="1975">Lehtinen 1975</bibRefCitation>
) and feathery scales (
<figureCitation id="13412A10FF87FF85E5D96C2AFB9D6453" box="[1021,1108,583,608]" captionStart="FIGURE 1" captionStartId="4.[151,250,1671,1693]" captionTargetBox="[151,1436,195,1649]" captionTargetId="figure@4.[151,1436,195,1649]" captionTargetPageId="4" captionText="FIGURE 1. Campostichomma alawala sp. nov. (CASENT9067599), FEmAlE lEFt lEg I: (A) clAws; (B) tArsus (ArrOw pOInts tO tArsAl OrgAn); (C) mEtAtArsus, sEtAE And FEAthEry scAlEs; (D) tArsus-mEtAtArsus jOInt, shOwIng mEtAtArsAl stOppEr; (E) tArsAl OrgAn;(F) trIchObOthrIAl bAsE,tArsus." httpUri="https://zenodo.org/record/1076180/files/figure.png" pageId="3" pageNumber="54">Fig. 1C</figureCitation>
;
<bibRefCitation id="EFEB4B64FF87FF85E2446C2AFAC36453" author="Griswold" box="[1120,1290,583,608]" pageId="3" pageNumber="54" refString="GrIswOld, C. E. (1993) InVEstIgAtIOns IntO thE phylOgEny OF thE lycOsOId spIdErs And thEIr kIn (ArAchnIdA: ArAnEAE: LycOsOIdEA). Smithsonian Contributions to Zoology, 539, 1 - 39. https: // dOI. Org / 10.5479 / sI. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: figs 75, 77); trochanters notched. Femora of legs and palpi with dorsal and lateral spines, patellae III and IV with lateral spines, tibiae I and II with 4 pairs of ventral spines and an additional proventral spine just behind apical pair, metatarsi I and II with 3 pairs of ventral spines, males usually with, females without, lateral spines on tibiae and metatarsi I and II; tibiae and metatarsi III and IV with dorsal, lateral, and ventral spines; palpal femora with dorsal and lateral spines, tibiae and tarsi of females with spines, tibiae and tarsi of males with or without spines, cymbium usually lacking spines. Males with tibial crack on legs I–IV (
<figureCitation id="13412A10FF87FF85E4C26D72FCF4650B" box="[742,829,799,824]" captionStart="FIGURE 3" captionStartId="7.[151,250,1732,1754]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@7.[151,1436,193,1711]" captionTargetPageId="7" captionText="FIGURE 3. Campostichomma manicatum KArsch, 1892. (A–D) mAlE lEFt pAlp; (A) VEntrAl VIEw, (B) rEtrOlAtErAl VIEw, (C) prOlAtErAl VIEw; (D) rIght pAlp, rEtrOlAtErAl VIEw, ExpAndEd; (E) mAlE lEg IV; (F–G) EpIgynum; (F) VEntrAl VIEw; (G) dOrsAl VIEw.AbbrEVIAtIOns:LO, lOckIng lObEs;TC, tIbIAl crAck." httpUri="https://zenodo.org/record/1076188/files/figure.png" pageId="3" pageNumber="54">Fig. 3E</figureCitation>
). Preening combs and scopulae absent. Superior tarsal claws pectinate, with inferior tarsal claws, I–II usually medium to large (
<figureCitation id="13412A10FF87FF85E5E66D29FBD2656F" box="[962,1051,836,861]" captionStart="FIGURE 1" captionStartId="4.[151,250,1671,1693]" captionTargetBox="[151,1436,195,1649]" captionTargetId="figure@4.[151,1436,195,1649]" captionTargetPageId="4" captionText="FIGURE 1. Campostichomma alawala sp. nov. (CASENT9067599), FEmAlE lEFt lEg I: (A) clAws; (B) tArsus (ArrOw pOInts tO tArsAl OrgAn); (C) mEtAtArsus, sEtAE And FEAthEry scAlEs; (D) tArsus-mEtAtArsus jOInt, shOwIng mEtAtArsAl stOppEr; (E) tArsAl OrgAn;(F) trIchObOthrIAl bAsE,tArsus." httpUri="https://zenodo.org/record/1076180/files/figure.png" pageId="3" pageNumber="54">Fig. 1A</figureCitation>
;
<bibRefCitation id="EFEB4B64FF87FF85E2036D29FB1B656F" author="Griswold" box="[1063,1234,836,861]" pageId="3" pageNumber="54" refString="GrIswOld, C. E. (1993) InVEstIgAtIOns IntO thE phylOgEny OF thE lycOsOId spIdErs And thEIr kIn (ArAchnIdA: ArAnEAE: LycOsOIdEA). Smithsonian Contributions to Zoology, 539, 1 - 39. https: // dOI. Org / 10.5479 / sI. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: fig. 5), claw tufts absent (
<figureCitation id="13412A10FF87FF85E6CB6D0AFE8065B3" box="[239,329,871,896]" captionStart="FIGURE 1" captionStartId="4.[151,250,1671,1693]" captionTargetBox="[151,1436,195,1649]" captionTargetId="figure@4.[151,1436,195,1649]" captionTargetPageId="4" captionText="FIGURE 1. Campostichomma alawala sp. nov. (CASENT9067599), FEmAlE lEFt lEg I: (A) clAws; (B) tArsus (ArrOw pOInts tO tArsAl OrgAn); (C) mEtAtArsus, sEtAE And FEAthEry scAlEs; (D) tArsus-mEtAtArsus jOInt, shOwIng mEtAtArsAl stOppEr; (E) tArsAl OrgAn;(F) trIchObOthrIAl bAsE,tArsus." httpUri="https://zenodo.org/record/1076180/files/figure.png" pageId="3" pageNumber="54">Fig. 1A</figureCitation>
). Trichobothria on tibiae with basal dorsolateral group and pro- and retrolateral rows, retrolateral row extending to apex, metatarsi with dorsal, irregular row or dorsal row plus dorsobasal group, tarsi with dorsobasal row that divides into 2–3 rows apically; palpal tibiae with pro- and retrolateral rows, apparently absent from tarsi; trichobothrial base with transversely-ridged hood (
<figureCitation id="13412A10FF87FF85E5576DB9FC0465DF" box="[883,973,980,1005]" captionStart="FIGURE 1" captionStartId="4.[151,250,1671,1693]" captionTargetBox="[151,1436,195,1649]" captionTargetId="figure@4.[151,1436,195,1649]" captionTargetPageId="4" captionText="FIGURE 1. Campostichomma alawala sp. nov. (CASENT9067599), FEmAlE lEFt lEg I: (A) clAws; (B) tArsus (ArrOw pOInts tO tArsAl OrgAn); (C) mEtAtArsus, sEtAE And FEAthEry scAlEs; (D) tArsus-mEtAtArsus jOInt, shOwIng mEtAtArsAl stOppEr; (E) tArsAl OrgAn;(F) trIchObOthrIAl bAsE,tArsus." httpUri="https://zenodo.org/record/1076180/files/figure.png" pageId="3" pageNumber="54">Fig. 1F</figureCitation>
;
<bibRefCitation id="EFEB4B64FF87FF85E5FB6DB9FB5965DF" author="Griswold" box="[991,1168,980,1005]" pageId="3" pageNumber="54" refString="GrIswOld, C. E. (1993) InVEstIgAtIOns IntO thE phylOgEny OF thE lycOsOId spIdErs And thEIr kIn (ArAchnIdA: ArAnEAE: LycOsOIdEA). Smithsonian Contributions to Zoology, 539, 1 - 39. https: // dOI. Org / 10.5479 / sI. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: fig. 77). Tarsal organ median, capsulate, orifice oval to round or keyhole-shaped (
<figureCitation id="13412A10FF87FF85E56D6D9AFC696223" box="[841,928,1015,1040]" captionStart="FIGURE 1" captionStartId="4.[151,250,1671,1693]" captionTargetBox="[151,1436,195,1649]" captionTargetId="figure@4.[151,1436,195,1649]" captionTargetPageId="4" captionText="FIGURE 1. Campostichomma alawala sp. nov. (CASENT9067599), FEmAlE lEFt lEg I: (A) clAws; (B) tArsus (ArrOw pOInts tO tArsAl OrgAn); (C) mEtAtArsus, sEtAE And FEAthEry scAlEs; (D) tArsus-mEtAtArsus jOInt, shOwIng mEtAtArsAl stOppEr; (E) tArsAl OrgAn;(F) trIchObOthrIAl bAsE,tArsus." httpUri="https://zenodo.org/record/1076180/files/figure.png" pageId="3" pageNumber="54">Fig. 1E</figureCitation>
;
<bibRefCitation id="EFEB4B64FF87FF85E58B6D9AFB946223" author="Griswold" box="[943,1117,1015,1040]" pageId="3" pageNumber="54" refString="GrIswOld, C. E. (1993) InVEstIgAtIOns IntO thE phylOgEny OF thE lycOsOId spIdErs And thEIr kIn (ArAchnIdA: ArAnEAE: LycOsOIdEA). Smithsonian Contributions to Zoology, 539, 1 - 39. https: // dOI. Org / 10.5479 / sI. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: figs 76–77), with proximal seam. Male palpal tibia short. Cymbium with large, triangular petiole attached to alveolus; subtegulum cup-shaped, with anelli, prolateral margin with lobe that articulates with corresponding lobe on tegulum in unexpanded bulb (
<figureCitation id="13412A10FF87FF85E6BB6A09FF3E624F" box="[159,247,1124,1149]" captionStart="FIGURE 3" captionStartId="7.[151,250,1732,1754]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@7.[151,1436,193,1711]" captionTargetPageId="7" captionText="FIGURE 3. Campostichomma manicatum KArsch, 1892. (A–D) mAlE lEFt pAlp; (A) VEntrAl VIEw, (B) rEtrOlAtErAl VIEw, (C) prOlAtErAl VIEw; (D) rIght pAlp, rEtrOlAtErAl VIEw, ExpAndEd; (E) mAlE lEg IV; (F–G) EpIgynum; (F) VEntrAl VIEw; (G) dOrsAl VIEw.AbbrEVIAtIOns:LO, lOckIng lObEs;TC, tIbIAl crAck." httpUri="https://zenodo.org/record/1076188/files/figure.png" pageId="3" pageNumber="54">Fig. 3B</figureCitation>
); tegulum C-shaped (
<figureCitation id="13412A10FF87FF85E7CD6A09FD8D624F" box="[489,580,1124,1149]" captionStart="FIGURE 4" captionStartId="8.[151,250,1910,1932]" captionTargetBox="[151,1419,600,1845]" captionTargetId="figure@8.[149,1434,579,1940]" captionTargetPageId="8" captionText="FIGURE 4. Campostichomma manicatum KArsch, 1892. (A–C) mAlE lEFt pAlp; (A) VEntrAl VIEw; (B) prOlAtErAl VIEw; (C) rEtrOlAtErAl VIEw; (D–E) EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw. AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RG, rEtrOlAtErAl grOOVE OF cymbIum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs;S, spErmAthEcAE;ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076192/files/figure.png" pageId="3" pageNumber="54">Fig. 4A</figureCitation>
), simple; firmly attached embolus (E) arising on retrolateral side of tegulum and tapering to apex; median apophysis (MA) concave, attached in middle of tegulum (
<figureCitation id="13412A10FF87FF85E2AE6AEAFB2F6293" box="[1162,1254,1159,1184]" captionStart="FIGURE 3" captionStartId="7.[151,250,1732,1754]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@7.[151,1436,193,1711]" captionTargetPageId="7" captionText="FIGURE 3. Campostichomma manicatum KArsch, 1892. (A–D) mAlE lEFt pAlp; (A) VEntrAl VIEw, (B) rEtrOlAtErAl VIEw, (C) prOlAtErAl VIEw; (D) rIght pAlp, rEtrOlAtErAl VIEw, ExpAndEd; (E) mAlE lEg IV; (F–G) EpIgynum; (F) VEntrAl VIEw; (G) dOrsAl VIEw.AbbrEVIAtIOns:LO, lOckIng lObEs;TC, tIbIAl crAck." httpUri="https://zenodo.org/record/1076188/files/figure.png" pageId="3" pageNumber="54">Fig. 3A</figureCitation>
); conductor (C) hyaline, originating anteriorly of tegulum, fan-shaped and not embracing apex of embolus (
<figureCitation id="13412A10FF87FF85E2856AC1FAD762F7" box="[1185,1310,1196,1221]" captionStart="FIGURE 4" captionStartId="8.[151,250,1910,1932]" captionTargetBox="[151,1419,600,1845]" captionTargetId="figure@8.[149,1434,579,1940]" captionTargetPageId="8" captionText="FIGURE 4. Campostichomma manicatum KArsch, 1892. (A–C) mAlE lEFt pAlp; (A) VEntrAl VIEw; (B) prOlAtErAl VIEw; (C) rEtrOlAtErAl VIEw; (D–E) EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw. AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RG, rEtrOlAtErAl grOOVE OF cymbIum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs;S, spErmAthEcAE;ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076192/files/figure.png" pageId="3" pageNumber="54">Fig. 4A–C</figureCitation>
); fundus in subtegulum, reservoir and ejaculatory duct simple, without loops or switchbacks, spiralling around outer margin of tegulum. Abdomen oval, without scuta (
<figureCitation id="13412A10FF87FF85E47E6A99FD11633F" box="[602,728,1268,1293]" captionStart="FIGURE2" captionStartId="6.[151,250,1960,1982]" captionTargetBox="[235,1353,193,1941]" captionTargetId="figure@6.[235,1353,193,1941]" captionTargetPageId="6" captionText="FIGURE2. Campostichomma manicatumKArsch, 1892. (A) dOrsAl And (B) VEntrAl hAbItus OF mAlE;(C) dOrsAl And (D) FEmAlE hAbItus,VEntrAl VIEw; (E) mAlE And (F) FEmAlE OculAr ArEA, AntErIOrVIEw." httpUri="https://zenodo.org/record/1076184/files/figure.png" pageId="3" pageNumber="54">Fig. 2A–D</figureCitation>
). Respiratory system comprising two anterior book lungs and a posterior tracheal spiracle. Epigynum divided by longitudinal epigynal folds into median lobe and lateral lobes; lateral lobes with or without teeth (
<figureCitation id="13412A10FF87FF85E4036B51FD4F6367" box="[551,646,1340,1365]" captionStart="FIGURE 4" captionStartId="8.[151,250,1910,1932]" captionTargetBox="[151,1419,600,1845]" captionTargetId="figure@8.[149,1434,579,1940]" captionTargetPageId="8" captionText="FIGURE 4. Campostichomma manicatum KArsch, 1892. (A–C) mAlE lEFt pAlp; (A) VEntrAl VIEw; (B) prOlAtErAl VIEw; (C) rEtrOlAtErAl VIEw; (D–E) EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw. AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RG, rEtrOlAtErAl grOOVE OF cymbIum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs;S, spErmAthEcAE;ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076192/files/figure.png" pageId="3" pageNumber="54">Figs 4D</figureCitation>
,
<figureCitation id="13412A10FF87FF85E4B66B51FD336367" box="[658,762,1340,1364]" captionStart="FIGURE8" captionStartId="12.[151,250,1887,1909]" captionTargetBox="[167,1428,218,1840]" captionTargetId="figure@12.[151,1436,193,1866]" captionTargetPageId="12" captionText="FIGURE8. Campostichomma spp., FEmAlE gEnItAlIA.(A–B) C. alawala sp. nov., EpIgynum;(A) VEntrAl VIEw;(B) dOrsAlVIEw; (C–D) C. harasbedda sp. nov., EpIgynum; (C) VEntrAl VIEw; (D) dOrsAl VIEw;(E–F) C. mudduk sp. nov., EpIgynum; (E)VEntrAl VIEw; (F) dOrsAl VIEw. AbbrEVIAtIOns: CD, cOpulAtOry ducts; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAn lObEOF EpIgynum;S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076206/files/figure.png" pageId="3" pageNumber="54">8A, C, E</figureCitation>
). Vulva with elongated copulatory duct, spermathecae with protuberances and small fertilization ducts (
<figureCitation id="13412A10FF87FF85E4AC6B32FD2A634B" box="[648,739,1375,1400]" captionStart="FIGURE 4" captionStartId="8.[151,250,1910,1932]" captionTargetBox="[151,1419,600,1845]" captionTargetId="figure@8.[149,1434,579,1940]" captionTargetPageId="8" captionText="FIGURE 4. Campostichomma manicatum KArsch, 1892. (A–C) mAlE lEFt pAlp; (A) VEntrAl VIEw; (B) prOlAtErAl VIEw; (C) rEtrOlAtErAl VIEw; (D–E) EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw. AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RG, rEtrOlAtErAl grOOVE OF cymbIum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs;S, spErmAthEcAE;ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076192/files/figure.png" pageId="3" pageNumber="54">Figs 4E</figureCitation>
,
<figureCitation id="13412A10FF87FF85E4D46B0DFC9F634B" box="[752,854,1376,1400]" captionStart="FIGURE8" captionStartId="12.[151,250,1887,1909]" captionTargetBox="[167,1428,218,1840]" captionTargetId="figure@12.[151,1436,193,1866]" captionTargetPageId="12" captionText="FIGURE8. Campostichomma spp., FEmAlE gEnItAlIA.(A–B) C. alawala sp. nov., EpIgynum;(A) VEntrAl VIEw;(B) dOrsAlVIEw; (C–D) C. harasbedda sp. nov., EpIgynum; (C) VEntrAl VIEw; (D) dOrsAl VIEw;(E–F) C. mudduk sp. nov., EpIgynum; (E)VEntrAl VIEw; (F) dOrsAl VIEw. AbbrEVIAtIOns: CD, cOpulAtOry ducts; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAn lObEOF EpIgynum;S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076206/files/figure.png" pageId="3" pageNumber="54">8B, D, F</figureCitation>
). Cribellum absent; colulus a fleshy triangular lobe set with several setae. Six spinnerets, anterior (ALS) and posterior laterals (PLS) two-segmented, posterior medians (PMS) one-segmented; females with ALS having two major ampullate gland spigots and several piriform gland spigots, PMS with multiple minor ampullate, cylindrical and aciniform gland spigots; PLS with at least cylindrical and aciniform gland spigots.
</paragraph>
</subSubSection>
<subSubSection id="C360651EFF87FF85E6E36879FEB56063" pageId="3" pageNumber="54" type="description">
<paragraph id="8BC53695FF87FF85E6E36879FEB56063" blockId="3.[151,1437,151,1869]" pageId="3" pageNumber="54">
<emphasis id="B90EEA87FF87FF85E6E36879FEAF601E" bold="true" box="[199,358,1556,1581]" pageId="3" pageNumber="54">Composition.</emphasis>
Four species:
<taxonomicName id="4C7A4D16FF87FF85E4376879FC93601F" authority="Karsch, 1892" authorityName="Karsch" authorityYear="1892" box="[531,858,1556,1581]" class="Arachnida" family="Zorocratidae" genus="Campostichomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="manicatum">
<emphasis id="B90EEA87FF87FF85E4376879FD7D601F" box="[531,692,1556,1580]" italics="true" pageId="3" pageNumber="54">C. manicatum</emphasis>
Karsch, 1892
</taxonomicName>
,
<taxonomicName id="4C7A4D16FF87FF85E54D6879FC2E601F" authority="Polotow &amp; Griswold, 2017" authorityName="Polotow &amp; Griswold" authorityYear="2017" box="[873,999,1556,1580]" class="Arachnida" family="Zorocratidae" genus="Campostichomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="alawala" status="sp. nov.">
<emphasis id="B90EEA87FF87FF85E54D6879FC2E601F" box="[873,999,1556,1580]" italics="true" pageId="3" pageNumber="54">C. alawala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57FCFF87FF85E5D66879FB87601E" box="[1010,1102,1556,1581]" pageId="3" pageNumber="54" rank="species">
<emphasis id="B90EEA87FF87FF85E5D66879FB87601E" bold="true" box="[1010,1102,1556,1581]" pageId="3" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C7A4D16FF87FF85E2446879FB14601F" authority="Polotow &amp; Griswold, 2017" authorityName="Polotow &amp; Griswold" authorityYear="2017" box="[1120,1245,1556,1580]" class="Arachnida" family="Zorocratidae" genus="Campostichomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="mudduk" status="sp. nov.">
<emphasis id="B90EEA87FF87FF85E2446879FB14601F" box="[1120,1245,1556,1580]" italics="true" pageId="3" pageNumber="54">C. mudduk</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57FCFF87FF85E2CC6879FA8C601E" box="[1256,1349,1556,1581]" pageId="3" pageNumber="54" rank="species">
<emphasis id="B90EEA87FF87FF85E2CC6879FA8C601E" bold="true" box="[1256,1349,1556,1581]" pageId="3" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C7A4D16FF87FF85E3A66879FED56063" authority="Polotow &amp; Griswold, 2017" authorityName="Polotow &amp; Griswold" authorityYear="2017" class="Arachnida" family="Zorocratidae" genus="Campostichomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="harasbedda" status="sp. nov.">
<emphasis id="B90EEA87FF87FF85E3A66879FED56063" italics="true" pageId="3" pageNumber="54">C. harasbedda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57FCFF87FF85E706685AFEB56063" box="[290,380,1591,1616]" pageId="3" pageNumber="54" rank="species">
<emphasis id="B90EEA87FF87FF85E706685AFEB56063" bold="true" box="[290,380,1591,1616]" pageId="3" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C360651EFF87FF85E6E36831FEBA617E" pageId="3" pageNumber="54" type="distribution">
<paragraph id="8BC53695FF87FF85E6E36831FD056047" blockId="3.[151,1437,151,1869]" box="[199,716,1628,1653]" pageId="3" pageNumber="54">
<emphasis id="B90EEA87FF87FF85E6E36831FEA96046" bold="true" box="[199,352,1628,1653]" pageId="3" pageNumber="54">Distribution.</emphasis>
Endemic to
<collectingCountry id="F36D7605FF87FF85E7CA6831FD956046" box="[494,604,1628,1653]" name="Sri Lanka" pageId="3" pageNumber="54">Sri Lanka</collectingCountry>
(
<figureCitation id="13412A10FF87FF85E44F6831FD766047" box="[619,703,1628,1653]" captionStart="FIGURE14" captionStartId="20.[151,250,1476,1498]" captionTargetBox="[150,1436,807,1454]" captionTargetId="figure@20.[150,1436,807,1454]" captionTargetPageId="20" captionText="FIGURE14. (A) PArtIAl wOrld mAp (ExcEptNEArctIc AndNEOtrOpIcAl rEAlms),shOwIng thE lOcAtIOnOF thE IslAnd cOuntry OF SrI LAnkA In SOuth AsIA. (B) ClOsE up OF SrI LAnkA, wIth cOllEctIOn lOcAlItIEs OF Campostichomma And Devendra spEcIEs: whItE squArE, Devendra pardalis (SImOn, 1898), D. pumilus (SImOn, 1898) And D. saama sp. nov.; whItE cIrclE, D seriatus (SImOn, 1898) And Devendra amaiti sp. nov.; blAck squArE, Compostichomma harasbedda sp. nov.; blAck cIrclE, Compostichomma alawala sp. nov.; blAck trIAnglE, Compostichomma mudduk sp. nov. ThE cOllEctIOn dAtA OF Compostichomma manicatum KArsch, 1892 dO nOt prOVIdE A spEcIFIclOcAlIty In SrI LAnkA, thus thE spEcIEs Is nOt shOwn In thIs mAp." httpUri="https://zenodo.org/record/1076226/files/figure.png" pageId="3" pageNumber="54">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8BC53695FF87FF85E6E36812FEBA617E" blockId="3.[151,1437,151,1869]" pageId="3" pageNumber="54">
<emphasis id="B90EEA87FF87FF85E6E36812FEB460AB" bold="true" box="[199,381,1663,1688]" pageId="3" pageNumber="54">Natural history</emphasis>
. The
<typeStatus id="54C18837FF87FF85E79868ECFE2560AA" box="[444,492,1665,1689]" pageId="3" pageNumber="54">type</typeStatus>
specimens of
<taxonomicName id="4C7A4D16FF87FF85E4A868EDFCE060A4" authorityName="Karsch" authorityYear="1892" box="[652,809,1664,1687]" class="Arachnida" family="Zorocratidae" genus="Campostichomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="manicatum">
<emphasis id="B90EEA87FF87FF85E4A868EDFCE060A4" box="[652,809,1664,1687]" italics="true" pageId="3" pageNumber="54">C. manicatum</emphasis>
</taxonomicName>
were collected in the highlands of
<collectingCountry id="F36D7605FF87FF85E2896812FAD760AB" box="[1197,1310,1663,1688]" name="Sri Lanka" pageId="3" pageNumber="54">Sri Lanka</collectingCountry>
. Collection data for the specimens collected by the Lund University
<collectingCountry id="F36D7605FF87FF85E50268C9FCBE608E" box="[806,887,1700,1725]" name="Sri Lanka" pageId="3" pageNumber="54">Ceylon</collectingCountry>
Expedition are detailed in Brinck
<emphasis id="B90EEA87FF87FF85E32268C8FA88608F" box="[1286,1345,1700,1724]" italics="true" pageId="3" pageNumber="54">et al.</emphasis>
(1971). These data suggest that these are running spiders that favor moist, closed canopy forest or woodland. Most were collected in the highland above
<quantity id="4C829B70FF87FF85E42F6881FDAC6137" box="[523,613,1772,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="54" unit="m" value="1000.0">1000 m</quantity>
, in forest, moist ravines or near tea plantations.
<taxonomicName id="4C7A4D16FF87FF85E2BF6881FAD16137" authority="Polotow &amp; Griswold, 2017" authorityName="Polotow &amp; Griswold" authorityYear="2017" box="[1179,1304,1772,1796]" class="Arachnida" family="Zorocratidae" genus="Campostichomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="alawala" status="sp. nov.">
<emphasis id="B90EEA87FF87FF85E2BF6881FAD16137" box="[1179,1304,1772,1796]" italics="true" pageId="3" pageNumber="54">C. alawala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57FCFF87FF85E3076881FAB66136" box="[1315,1407,1772,1797]" pageId="3" pageNumber="54" rank="species">
<emphasis id="B90EEA87FF87FF85E3076881FAB66136" bold="true" box="[1315,1407,1772,1797]" pageId="3" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
is unique in being from lower elevation, from jungle at only
<quantity id="4C829B70FF87FF85E561697DFC59611B" box="[837,912,1808,1832]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="54" unit="m" value="200.0">200 m</quantity>
elevation. More detail is provided with each species description.
</paragraph>
</subSubSection>
</treatment>
</document>
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