317 lines
36 KiB
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317 lines
36 KiB
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<document id="92D90173047015B9612B5EDC561D07DE" ID-DOI="10.11646/zootaxa.5507.1.11" ID-ISSN="1175-5326" ID-Zenodo-Dep="13747567" ID-ZooBank="F39C3608-7943-4E89-9F07-707740CEC61A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726055414319" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03C69063FFFFDD0FFF0C7E0C27FD63EC" docLanguage="en" docName="zootaxa.5507.1.11.pdf" docOrigin="Zootaxa 5507 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.1.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Veraphis shaanxiana Jałoszyński, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="198" masterDocId="FFFFE81BFFFEDD0BFF9B7B3922676169" masterDocTitle="Two new Veraphis species in China (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="200" masterPageNumber="194" pageNumber="195" updateTime="1726056176235" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6555FEF924DE6800D8DD36B142D00F21">Two new Veraphis species in China (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="4812DEDC8A6B88ADDF24B66E4862096C">Jałoszyński, Paweł</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="C9560406EEC76343CBF6EA82A254304D">text</mods:typeOfResource>
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<mods:title id="02CB852903200D06CBEA0699F4E2D95F">Zootaxa</mods:title>
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<mods:part id="A53AA5E96F1C57BC36ACA3461382879E">
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<mods:date id="AAA4E3387EB1DFC74E3C6B36C1967BC6">2024</mods:date>
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<mods:number id="74482A823630432DEBCB3F227133EE5D">2024-09-11</mods:number>
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<mods:number id="930DE4712A0658726A6FD384B42B0319">5507</mods:number>
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<mods:number id="586BBE6E0A57FD097C8A8673EE1ED29D">1</mods:number>
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<mods:url id="75D819D28EC347F6E180089B86B36135">http://dx.doi.org/10.11646/zootaxa.5507.1.11</mods:url>
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</mods:location>
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<mods:classification id="52E1EBE08494E3A49B95D4FAB00C0C25">journal article</mods:classification>
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<mods:identifier id="1894444B8AB55ED9D74BD0A06B960452" type="DOI">10.11646/zootaxa.5507.1.11</mods:identifier>
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<mods:identifier id="94FCA9F958BB12EE618D647BB532845D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5166487551076B1ED942099731B3FCAA" type="Zenodo-Dep">13747567</mods:identifier>
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<mods:identifier id="9562FF09F82852852963114336303192" type="ZooBank">F39C3608-7943-4E89-9F07-707740CEC61A</mods:identifier>
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</mods:mods>
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<treatment id="03C69063FFFFDD0FFF0C7E0C27FD63EC" ID-DOI="http://doi.org/10.5281/zenodo.13747505" ID-Zenodo-Dep="13747505" LSID="urn:lsid:plazi:treatment:03C69063FFFFDD0FFF0C7E0C27FD63EC" httpUri="http://treatment.plazi.org/id/03C69063FFFFDD0FFF0C7E0C27FD63EC" lastPageId="4" lastPageNumber="198" pageId="1" pageNumber="195">
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<subSubSection id="C37572FEFFFFDD0AFF0C7E0C239C6426" box="[151,507,1333,1360]" pageId="1" pageNumber="195" type="nomenclature">
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<paragraph id="8BD02175FFFFDD0AFF0C7E0C239C6426" blockId="1.[151,507,1333,1395]" box="[151,507,1333,1360]" pageId="1" pageNumber="195">
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<heading id="D0989619FFFFDD0AFF0C7E0C239C6426" bold="true" box="[151,507,1333,1360]" fontSize="11" level="1" pageId="1" pageNumber="195" reason="1">
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<emphasis id="B91BFD67FFFFDD0AFF0C7E0C239C6426" bold="true" box="[151,507,1333,1360]" pageId="1" pageNumber="195">
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<taxonomicName id="4C6F5AF6FFFFDD0AFF0C7E0C23F16439" authorityName="Jałoszyński" authorityYear="2024" box="[151,406,1333,1360]" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="195" phylum="Arthropoda" rank="species" species="shaanxiana" status="sp. nov.">
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<emphasis id="B91BFD67FFFFDD0AFF0C7E0C23F16439" bold="true" box="[151,406,1333,1360]" italics="true" pageId="1" pageNumber="195">Veraphis shaanxiana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A228401CFFFFDD0AFE057E0C239C6426" box="[414,507,1333,1359]" pageId="1" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37572FEFFFFDD0AFF0C7E602337641A" box="[151,336,1369,1395]" pageId="1" pageNumber="195" type="description">
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<paragraph id="8BD02175FFFFDD0AFF0C7E602337641A" blockId="1.[151,507,1333,1395]" box="[151,336,1369,1395]" pageId="1" pageNumber="195">
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(
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<figureCitation id="13543DF0FFFFDD0AFF047E602281641A" box="[159,230,1369,1395]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1959,1983]" captionTargetBox="[158,1430,568,1930]" captionTargetId="figure-163@2.[151,1436,561,1936]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of holotype males. Veraphis shaanxiana sp. nov. (1) and Veraphis dabashana sp. nov. (2)." figureDoi="http://doi.org/10.5281/zenodo.13747569" httpUri="https://zenodo.org/record/13747569/files/figure.png" pageId="1" pageNumber="195">Figs 1</figureCitation>
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,
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<figureCitation id="13543DF0FFFFDD0AFF6F7E602346641A" box="[244,289,1369,1395]" captionStart="FIGURES 3–10" captionStartId="3.[151,264,1930,1954]" captionTargetBox="[158,1429,280,1900]" captionTargetId="figure-39@3.[151,1436,273,1907]" captionTargetPageId="3" captionText="FIGURES 3–10. Aedeagus in ventral (3, 5, 7, 9) and lateral (4, 6, 8, 10) views. Veraphis shaanxiana sp. nov. (3‒6) and Veraphis dabashana sp. nov. (7‒10)." figureDoi="http://doi.org/10.5281/zenodo.13747571" httpUri="https://zenodo.org/record/13747571/files/figure.png" pageId="1" pageNumber="195">3–6</figureCitation>
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,
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<figureCitation id="13543DF0FFFFDD0AFEB77E602320641A" box="[300,327,1369,1395]" captionStart="FIGURE 11" captionStartId="5.[151,250,1368,1392]" captionTargetBox="[151,1410,525,1345]" captionTargetId="figure-146@5.[151,1435,525,1345]" captionTargetPageId="5" captionText="FIGURE 11. Distribution of nominal species of Veraphis in China, with comparison of their aedeagi in ventral view (based on Google Maps). Veraphis calcarifer Jałoszyński (a), V. qinghaiensis Jałoszyński (b), V. gansuana Jałoszyński (c), V. assingi Jałoszyński (d), V. modesta Jałoszyński (e), V. spinosa Jałoszyński (f), V. shaanxiana Jałoszyński (g), and V. dabashana Jałoszyński (h)." figureDoi="http://doi.org/10.5281/zenodo.13747575" httpUri="https://zenodo.org/record/13747575/files/figure.png" pageId="1" pageNumber="195">11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37572FEFFFFDD0AFF0C7E9827236722" pageId="1" pageNumber="195" type="materials_examined">
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<paragraph id="8BD02175FFFFDD0AFF0C7E9827236722" blockId="1.[151,1437,1441,2008]" pageId="1" pageNumber="195">
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<emphasis id="B91BFD67FFFFDD0AFF0C7E98232464D2" bold="true" box="[151,323,1441,1467]" pageId="1" pageNumber="195">Type material.</emphasis>
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<materialsCitation id="3B072B28FFFFDD0AFED27E9821546722" collectingDate="2001-07-25" collectionCode="MNHB" collectorName="M. Schulke & P. Jaloszynski" country="China" county="Qinling Shan" elevation="2675" latitude="33.866665" location="Shagoujie" longLatPrecision="1202" longitude="108.76667" municipality="rd. Xi'an" pageId="1" pageNumber="195" specimenCount="1" specimenCount-male="1" stateProvince="Shaanxi" typeStatus="holotype">
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<emphasis id="B91BFD67FFFFDD0AFED27E9820C564D2" bold="true" box="[329,674,1441,1467]" pageId="1" pageNumber="195">
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<typeStatus id="54D49FD7FFFFDD0AFED27E9823DD64D2" box="[329,442,1441,1467]" pageId="1" pageNumber="195" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F37861E5FFFFDD0AFE5F7E98204764D2" box="[452,544,1441,1467]" name="China" pageId="1" pageNumber="195">CHINA</collectingCountry>
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(
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<collectingRegion id="49ABEF97FFFFDD0AFDB57E9820F164D2" box="[558,662,1441,1467]" country="China" name="Shaanxi" pageId="1" pageNumber="195">Shaanxi</collectingRegion>
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):
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</emphasis>
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<specimenCount id="9D69EAFCFFFFDD0AFD337E9B20D864D5" box="[680,703,1442,1468]" count="1" pageId="1" pageNumber="195" type="male">♂</specimenCount>
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, four labels: “CHINA: S-Shaanxi (
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<collectingCounty id="62B159F9FFFFDD0AFBA57E9826B264D2" box="[1086,1237,1441,1467]" pageId="1" pageNumber="195">Qinling Shan</collectingCounty>
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) / mountain range W pass on
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<collectingMunicipality id="6BB4BB0FFFFFDD0AFE8F7EFC231D64B6" box="[276,378,1477,1503]" pageId="1" pageNumber="195">rd. Xi’an</collectingMunicipality>
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/ -
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<location id="8EB077AEFFFFDD0AFE397EFC207264B6" LSID="urn:lsid:plazi:treatment:03C69063FFFFDD0FFF0C7E0C27FD63EC:8EB077AEFFFFDD0AFE397EFC207264B6" box="[418,533,1477,1503]" country="China" county="Qinling Shan" latitude="33.866665" longLatPrecision="1202" longitude="108.76667" municipality="rd. Xi'an" name="Shagoujie" pageId="1" pageNumber="195" stateProvince="Shaanxi">Shagoujie</location>
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,
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<locationDeviation id="9A623857FFFFDD0AFDBB7EFC209764B6" box="[544,752,1477,1504]" location="Xi'an" pageId="1" pageNumber="195">
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<quantity id="4C978C90FFFFDD0AFDBB7EFC200F64B6" box="[544,616,1477,1504]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="195" unit="km" value="45.0">45 km</quantity>
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SSW Xi’an
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</locationDeviation>
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/
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<geoCoordinate id="EE5B47B2FFFFDD0AFC9C7EFC21176489" box="[775,880,1477,1504]" degrees="33" direction="north" minutes="52" orientation="latitude" pageId="1" pageNumber="195" precision="925" value="33.866665">
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33
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<emphasis id="B91BFD67FFFFDD0AFCB87EFF21496489" box="[803,814,1478,1504]" italics="true" pageId="1" pageNumber="195">°</emphasis>
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52’ N
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</geoCoordinate>
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,
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<geoCoordinate id="EE5B47B2FFFFDD0AFCE77EFC21886489" box="[892,1007,1477,1504]" degrees="108" direction="east" minutes="46" orientation="longitude" pageId="1" pageNumber="195" precision="925" value="108.76667">
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108
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<emphasis id="B91BFD67FFFFDD0AFC3D7EFF21D66489" box="[934,945,1478,1504]" italics="true" pageId="1" pageNumber="195">°</emphasis>
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46’ E
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</geoCoordinate>
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,
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<quantity id="4C978C90FFFFDD0AFC677EFC26346489" box="[1020,1107,1477,1504]" metricMagnitude="3" metricUnit="m" metricValue="2.675" pageId="1" pageNumber="195" unit="m" value="2675.0">
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<elevation id="0042C646FFFFDD0AFC677EFC26346489" box="[1020,1107,1477,1504]" metricMagnitude="3" metricUnit="m" metricValue="2.675" pageId="1" pageNumber="195" unit="m" value="2675.0">2675 m</elevation>
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</quantity>
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, / leg.
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<collectorName id="269A44A3FFFFDD0AFB3B7EFF274664B6" box="[1184,1313,1477,1504]" pageId="1" pageNumber="195">M. Schülke</collectorName>
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[C01-20]” [white, printed], “
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<date id="FFD107B5FFFFDD0AFEF97ED02398676A" box="[354,511,1513,1540]" pageId="1" pageNumber="195" value="2001-07-25">
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<collectingDate id="EF95FE5DFFFFDD0AFEF97ED02398676A" box="[354,511,1513,1540]" pageId="1" pageNumber="195" value="2001-07-25">25. VII. 2001</collectingDate>
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</date>
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. / N- slope, Abies, Betula, Larix, / Rhododendron, / subalpine meadows / (sifted) [C01-20)” [white, printed], “Sammlung /
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<collectorName id="269A44A3FFFFDD0AFDEB7D372095674E" box="[624,754,1549,1576]" pageId="1" pageNumber="195">M. Schülke</collectorName>
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/ Berlin” [white, printed], “
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<taxonomicName id="4C6F5AF6FFFFDD0AFBAA7D3726D56741" authorityName="Casey" authorityYear="1897" box="[1073,1202,1550,1576]" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="195" phylum="Arthropoda" rank="genus">
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<emphasis id="B91BFD67FFFFDD0AFBAA7D3726D56741" box="[1073,1202,1550,1576]" italics="true" pageId="1" pageNumber="195">VERAPHIS</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C6F5AF6FFFFDD0AFB517D34272C674E" authorityName="Jałoszyński" authorityYear="2024" box="[1226,1355,1549,1575]" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="195" phylum="Arthropoda" rank="species" species="shaanxiana">
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<emphasis id="B91BFD67FFFFDD0AFB517D34272C674E" box="[1226,1355,1549,1575]" italics="true" pageId="1" pageNumber="195">shaanxiana</emphasis>
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</taxonomicName>
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m. /
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<collectorName id="269A44A3FFFFDD0AFA127D37237C6722" pageId="1" pageNumber="195">P. Jałoszyński</collectorName>
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, 2024 /
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<typeStatus id="54D49FD7FFFFDD0AFEEF7D08207A6722" box="[372,541,1585,1611]" pageId="1" pageNumber="195" type="holotype">HOLOTYPUS</typeStatus>
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” [red, printed] (
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<collectionCode id="ED7EB9B0FFFFDD0AFD4A7D08214C6722" box="[721,811,1585,1611]" pageId="1" pageNumber="195">MNHB</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B072B28FFFFDD0AFCA67D0827586722" box="[829,1343,1585,1612]" collectingDate="2001-07-25" collectionCode="MNHB" collectorName="M. Schulke & P. Jaloszynski" country="China" county="Qinling Shan" elevation="2675" latitude="33.866665" location="Shagoujie" longLatPrecision="1202" longitude="108.76667" municipality="rd. Xi'an" pageId="1" pageNumber="195" specimenCount="1" specimenCount-female="1" stateProvince="Shaanxi" typeStatus="paratype">
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<emphasis id="B91BFD67FFFFDD0AFCA67D0821D36722" bold="true" box="[829,948,1585,1611]" pageId="1" pageNumber="195">
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<typeStatus id="54D49FD7FFFFDD0AFCA67D0821C96722" box="[829,942,1585,1611]" pageId="1" pageNumber="195" type="paratype">Paratype</typeStatus>
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:
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</emphasis>
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<specimenCount id="9D69EAFCFFFFDD0AFC217D0B21A86725" box="[954,975,1586,1612]" count="1" pageId="1" pageNumber="195" type="female">♀</specimenCount>
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, same data as for holotype (cPJ)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37572FEFFFFDD0AFF5C7D6C215567B2" pageId="1" pageNumber="195" type="diagnosis">
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<paragraph id="8BD02175FFFFDD0AFF5C7D6C215567B2" blockId="1.[151,1437,1441,2008]" pageId="1" pageNumber="195">
|
|||
|
|
<emphasis id="B91BFD67FFFFDD0AFF5C7D6C23256706" bold="true" box="[199,322,1621,1647]" pageId="1" pageNumber="195">Diagnosis.</emphasis>
|
|||
|
|
Body only slightly exceeding
|
|||
|
|
<quantity id="4C978C90FFFFDD0AFD287D6C209E6719" box="[691,761,1621,1648]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="195" unit="mm" value="1.0">1 mm</quantity>
|
|||
|
|
in length; males with unmodified trochanters, with deep median impression on metaventrite; aedeagus in ventral view with distinctly asymmetrical apical region, including endophallus shifted laterad midline, in lateral view median lobe curved near base and nearly straight in distal 2/3, with narrow indentation in subapical region of ventral wall.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37572FEFFFFDD0FFF5C7DDC269360E0" lastPageId="4" lastPageNumber="198" pageId="1" pageNumber="195" type="description">
|
|||
|
|
<paragraph id="8BD02175FFFFDD0AFF5C7DDC27ED6796" blockId="1.[151,1437,1441,2008]" box="[199,1418,1765,1792]" pageId="1" pageNumber="195">
|
|||
|
|
<emphasis id="B91BFD67FFFFDD0AFF5C7DDC233E6796" bold="true" box="[199,345,1765,1791]" pageId="1" pageNumber="195">Description.</emphasis>
|
|||
|
|
BL
|
|||
|
|
<quantity id="4C978C90FFFFDD0AFE127DDC23886669" box="[393,495,1765,1792]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="1" pageNumber="195" unit="mm" value="1.11">1.11 mm</quantity>
|
|||
|
|
. Body of male (
|
|||
|
|
<figureCitation id="13543DF0FFFFDD0AFD397DDC20856796" box="[674,738,1765,1791]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1959,1983]" captionTargetBox="[158,1430,568,1930]" captionTargetId="figure-163@2.[151,1436,561,1936]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of holotype males. Veraphis shaanxiana sp. nov. (1) and Veraphis dabashana sp. nov. (2)." figureDoi="http://doi.org/10.5281/zenodo.13747569" httpUri="https://zenodo.org/record/13747569/files/figure.png" pageId="1" pageNumber="195">Fig. 1</figureCitation>
|
|||
|
|
) flat, slender, light brown, covered with yellowish vestiture.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD02175FFFFDD09FF5C7C30235161BC" blockId="1.[151,1437,1441,2008]" lastBlockId="2.[151,1437,151,537]" lastPageId="2" lastPageNumber="196" pageId="1" pageNumber="195">
|
|||
|
|
Head broadest at large, moderately convex eyes, HL
|
|||
|
|
<quantity id="4C978C90FFFFDD0AFCDA7C3021CB664D" box="[833,940,1801,1828]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="195" unit="mm" value="0.14">0.14 mm</quantity>
|
|||
|
|
, HW
|
|||
|
|
<quantity id="4C978C90FFFFDD0AFC6C7C302603664D" box="[1015,1124,1801,1828]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="195" unit="mm" value="0.23">0.23 mm</quantity>
|
|||
|
|
; vertex with pair of small posteromedian pits, each prolonged by shallow longitudinal groove extending anteriorly to posteromesal margin of weakly elevated supraantennal tubercle; area between grooves distinctly flattened and impressed in relation to convex sides of vertex and frons. Punctures on head dorsum fine, inconspicuous; setae short, sparse and suberect. Antennae slender, with distinctly demarcated trimerous clubs, AnL
|
|||
|
|
<quantity id="4C978C90FFFFDD0AFC097CA0219E66DD" box="[914,1017,1945,1972]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="195" unit="mm" value="0.48">0.48 mm</quantity>
|
|||
|
|
, antennomeres 1 and 2 each strongly elongate, 3 indistinctly transverse, 4 about as long as broad, 5 slightly elongate, 6 and 7 each about as long as broad, 8 much shorter than 7, distinctly transverse, 9 and 10 each weakly transverse, 11 slightly longer than 10, about as long as broad.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD02175FFFCDD09FF5C7BE620DD60E0" blockId="2.[151,1437,151,537]" pageId="2" pageNumber="196">
|
|||
|
|
Pronotum broadest near middle; PL
|
|||
|
|
<quantity id="4C978C90FFFCDD09FDCD7BE620DD6190" box="[598,698,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="196" unit="mm" value="0.3">0.30 mm</quantity>
|
|||
|
|
, PW
|
|||
|
|
<quantity id="4C978C90FFFCDD09FD6F7BE6213F6190" box="[756,856,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="196" unit="mm" value="0.31">0.31 mm</quantity>
|
|||
|
|
, anterior margin broadly and evenly rounded; lateral margins strongly rounded in anterior half, behind middle nearly straight and barely noticeably converging towards slightly sharp-angled hind corners; posterior margin shallowly bisinuate. Pronotal base with shallow transverse median pit and one pair of distinct lateral pits. Punctures on pronotal disc fine and inconspicuous, shallow and with diffuse margins; setae short and sparse, suberect.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD02175FFFCDD09FF5C7AAA23BA6370" blockId="2.[151,1437,151,537]" pageId="2" pageNumber="196">
|
|||
|
|
Elytra together slightly rhomboidal, broadest slightly anterior to middle; EL
|
|||
|
|
<quantity id="4C978C90FFFCDD09FBB37AAA26E860C4" box="[1064,1167,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="2" pageNumber="196" unit="mm" value="0.68">0.68 mm</quantity>
|
|||
|
|
, EW
|
|||
|
|
<quantity id="4C978C90FFFCDD09FB557AAA275260C4" box="[1230,1333,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="196" unit="mm" value="0.43">0.43 mm</quantity>
|
|||
|
|
, EI 1.59; humeral calli well-marked, elongate. Elytral disc covered with punctures slightly more distinct than those on pronotum but still superficial and inconspicuous, setae similar to those on pronotum. Hind wings well developed, about twice as long as elytra.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF1071FDFFFCDD09FF0C7C9E274F66D6" ID-DOI="http://doi.org/10.5281/zenodo.13747569" ID-Zenodo-Dep="13747569" box="[151,1320,1959,1983]" httpUri="https://zenodo.org/record/13747569/files/figure.png" pageId="2" pageNumber="196" startId="2.[151,264,1959,1983]" targetBox="[158,1430,568,1930]" targetPageId="2" targetType="figure">
|
|||
|
|
<paragraph id="8BD02175FFFCDD09FF0C7C9E274F66D6" blockId="2.[151,1320,1959,1983]" box="[151,1320,1959,1983]" pageId="2" pageNumber="196">
|
|||
|
|
<emphasis id="B91BFD67FFFCDD09FF0C7C9E235D66D6" bold="true" box="[151,314,1959,1983]" pageId="2" pageNumber="196">FIGURES 1–2.</emphasis>
|
|||
|
|
Habitus of holotype males.
|
|||
|
|
<taxonomicName id="4C6F5AF6FFFCDD09FDC37C9E214B66D6" authorityName="Jałoszyński" authorityYear="2024" box="[600,812,1959,1983]" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="196" phylum="Arthropoda" rank="species" species="shaanxiana" status="sp. nov.">
|
|||
|
|
<emphasis id="B91BFD67FFFCDD09FDC37C9E214B66D6" box="[600,812,1959,1983]" italics="true" pageId="2" pageNumber="196">Veraphis shaanxiana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B91BFD67FFFCDD09FCA97C9E21E466D6" bold="true" box="[818,899,1959,1983]" pageId="2" pageNumber="196">
|
|||
|
|
<taxonomicNameLabel id="A228401CFFFCDD09FCA97C9E21E466D6" box="[818,899,1959,1983]" pageId="2" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B91BFD67FFFCDD09FC097C9E21F866D6" bold="true" box="[914,927,1959,1983]" pageId="2" pageNumber="196">1</emphasis>
|
|||
|
|
) and
|
|||
|
|
<taxonomicName id="4C6F5AF6FFFCDD09FC437C9E26C166D6" authorityName="Jałoszyński" authorityYear="2024" box="[984,1190,1959,1983]" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="196" phylum="Arthropoda" rank="species" species="dabashana" status="sp. nov.">
|
|||
|
|
<emphasis id="B91BFD67FFFCDD09FC437C9E26C166D6" box="[984,1190,1959,1983]" italics="true" pageId="2" pageNumber="196">Veraphis dabashana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B91BFD67FFFCDD09FB367C9E269A66D6" bold="true" box="[1197,1277,1959,1983]" pageId="2" pageNumber="196">
|
|||
|
|
<taxonomicNameLabel id="A228401CFFFCDD09FB367C9E269A66D6" box="[1197,1277,1959,1983]" pageId="2" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B91BFD67FFFCDD09FA967C9E277D66D6" bold="true" box="[1293,1306,1959,1983]" pageId="2" pageNumber="196">2</emphasis>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BD02175FFFDDD08FF5C7BAE237661BC" blockId="3.[151,1436,151,249]" pageId="3" pageNumber="197">Legs moderately long and slender; protrochanters unmodified, protibia with small subapical fin-like projection.</paragraph>
|
|||
|
|
<paragraph id="8BD02175FFFDDD08FF5C7BE621566190" blockId="3.[151,1436,151,249]" box="[199,817,223,249]" pageId="3" pageNumber="197">Metaventrite with distinct elongate median impression.</paragraph>
|
|||
|
|
<caption id="DF1071FDFFFDDD08FF0C7CB323CA66AF" ID-DOI="http://doi.org/10.5281/zenodo.13747571" ID-Zenodo-Dep="13747571" httpUri="https://zenodo.org/record/13747571/files/figure.png" pageId="3" pageNumber="197" startId="3.[151,264,1930,1954]" targetBox="[158,1429,280,1900]" targetPageId="3" targetType="figure">
|
|||
|
|
<paragraph id="8BD02175FFFDDD08FF0C7CB323CA66AF" blockId="3.[151,1437,1930,1990]" pageId="3" pageNumber="197">
|
|||
|
|
<emphasis id="B91BFD67FFFDDD08FF0C7CB3232166CB" bold="true" box="[151,326,1930,1954]" pageId="3" pageNumber="197">FIGURES 3–10.</emphasis>
|
|||
|
|
Aedeagus in ventral (
|
|||
|
|
<emphasis id="B91BFD67FFFDDD08FDBB7CB3201166CB" bold="true" box="[544,630,1930,1954]" pageId="3" pageNumber="197">3, 5, 7, 9</emphasis>
|
|||
|
|
) and lateral (
|
|||
|
|
<emphasis id="B91BFD67FFFDDD08FD617CB3213B66CB" bold="true" box="[762,860,1930,1954]" pageId="3" pageNumber="197">4, 6, 8, 10</emphasis>
|
|||
|
|
) views.
|
|||
|
|
<taxonomicName id="4C6F5AF6FFFDDD08FC2B7CB326E466CB" authorityName="Jałoszyński" authorityYear="2024" box="[944,1155,1930,1954]" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="species" species="shaanxiana" status="sp. nov.">
|
|||
|
|
<emphasis id="B91BFD67FFFDDD08FC2B7CB326E466CB" box="[944,1155,1930,1954]" italics="true" pageId="3" pageNumber="197">Veraphis shaanxiana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B91BFD67FFFDDD08FB137CB326BF66CB" bold="true" box="[1160,1240,1930,1954]" pageId="3" pageNumber="197">
|
|||
|
|
<taxonomicNameLabel id="A228401CFFFDDD08FB137CB326BF66CB" box="[1160,1240,1930,1954]" pageId="3" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B91BFD67FFFDDD08FB7D7CB3276B66CB" bold="true" box="[1254,1292,1930,1954]" pageId="3" pageNumber="197">3‒6</emphasis>
|
|||
|
|
) and
|
|||
|
|
<taxonomicName id="4C6F5AF6FFFDDD08FADF7CB3236166AF" authorityName="Jałoszyński" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="197" phylum="Arthropoda" rank="species" species="dabashana" status="sp. nov.">
|
|||
|
|
<emphasis id="B91BFD67FFFDDD08FADF7CB3236166AF" italics="true" pageId="3" pageNumber="197">Veraphis dabashana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B91BFD67FFFDDD08FE967C97233A66AF" bold="true" box="[269,349,1966,1990]" pageId="3" pageNumber="197">
|
|||
|
|
<taxonomicNameLabel id="A228401CFFFDDD08FE967C97233A66AF" box="[269,349,1966,1990]" pageId="3" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B91BFD67FFFDDD08FEF77C9723F866AF" bold="true" box="[364,415,1966,1990]" pageId="3" pageNumber="197">7‒10</emphasis>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BD02175FFFADD0FFF5C7BAE27376028" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="198">
|
|||
|
|
Aedeagus (
|
|||
|
|
<figureCitation id="13543DF0FFFADD0FFED37BAE23C861D8" box="[328,431,151,177]" captionStart="FIGURES 3–10" captionStartId="3.[151,264,1930,1954]" captionTargetBox="[158,1429,280,1900]" captionTargetId="figure-39@3.[151,1436,273,1907]" captionTargetPageId="3" captionText="FIGURES 3–10. Aedeagus in ventral (3, 5, 7, 9) and lateral (4, 6, 8, 10) views. Veraphis shaanxiana sp. nov. (3‒6) and Veraphis dabashana sp. nov. (7‒10)." figureDoi="http://doi.org/10.5281/zenodo.13747571" httpUri="https://zenodo.org/record/13747571/files/figure.png" pageId="4" pageNumber="198">Figs 3–6</figureCitation>
|
|||
|
|
) strongly elongate; AeL
|
|||
|
|
<quantity id="4C978C90FFFADD0FFD487BAE215A61D8" box="[723,829,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="198" unit="mm" value="0.28">0.28 mm</quantity>
|
|||
|
|
; in ventral view median lobe broadly and shallowly constricted proximally to middle, broadening distally and in subapical region narrowing towards broadly rounded apex, apical region clearly asymmetrical, in lateral view median lobe curved near base and nearly straight in distal 2/3, with shallow ventral indentation in subapical region. Endophallus strongly asymmetrical, shifted laterad midline. Parameres slender, exceeding half length of median lobe, each with several short apical and subapical setae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD02175FFFADD0FFF5C7A72269360E0" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="198">
|
|||
|
|
Female. Externally similar to male except for metaventrite lacking median impression. BL
|
|||
|
|
<quantity id="4C978C90FFFADD0FFB507A722754600C" box="[1227,1331,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="198" unit="mm" value="1.1">1.10 mm</quantity>
|
|||
|
|
; HL
|
|||
|
|
<quantity id="4C978C90FFFADD0FFAF07A7222A260E0" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="198" unit="mm" value="0.14">0.14 mm</quantity>
|
|||
|
|
, HW
|
|||
|
|
<quantity id="4C978C90FFFADD0FFE9D7A56230B60E0" box="[262,364,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="198" unit="mm" value="0.23">0.23 mm</quantity>
|
|||
|
|
, AnL
|
|||
|
|
<quantity id="4C978C90FFFADD0FFE347A56207160E0" box="[431,534,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="4" pageNumber="198" unit="mm" value="0.48">0.48 mm</quantity>
|
|||
|
|
; PL
|
|||
|
|
<quantity id="4C978C90FFFADD0FFDD37A5620C960E0" box="[584,686,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="198" unit="mm" value="0.3">0.30 mm</quantity>
|
|||
|
|
, PW
|
|||
|
|
<quantity id="4C978C90FFFADD0FFD717A56213660E0" box="[746,849,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="198" unit="mm" value="0.31">0.31 mm</quantity>
|
|||
|
|
; EL
|
|||
|
|
<quantity id="4C978C90FFFADD0FFC1E7A56218C60E0" box="[901,1003,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="198" unit="mm" value="0.68">0.68 mm</quantity>
|
|||
|
|
, EW
|
|||
|
|
<quantity id="4C978C90FFFADD0FFBB37A5726E960E0" box="[1064,1166,366,393]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="198" unit="mm" value="0.45">0.45 mm</quantity>
|
|||
|
|
, EI 1.50.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37572FEFFFADD0FFF5C7AAB215B60C4" box="[199,828,402,429]" pageId="4" pageNumber="198" type="distribution">
|
|||
|
|
<paragraph id="8BD02175FFFADD0FFF5C7AAB215B60C4" blockId="4.[151,1437,151,645]" box="[199,828,402,429]" pageId="4" pageNumber="198">
|
|||
|
|
<emphasis id="B91BFD67FFFADD0FFF5C7AAB230660C5" bold="true" box="[199,353,402,428]" pageId="4" pageNumber="198">Distribution.</emphasis>
|
|||
|
|
Qinling Mountains in CE
|
|||
|
|
<collectingCountry id="F37861E5FFFADD0FFD117AAA20AA60C4" box="[650,717,403,429]" name="China" pageId="4" pageNumber="198">China</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13543DF0FFFADD0FFD467AAA214960C4" box="[733,814,403,429]" captionStart="FIGURE 11" captionStartId="5.[151,250,1368,1392]" captionTargetBox="[151,1410,525,1345]" captionTargetId="figure-146@5.[151,1435,525,1345]" captionTargetPageId="5" captionText="FIGURE 11. Distribution of nominal species of Veraphis in China, with comparison of their aedeagi in ventral view (based on Google Maps). Veraphis calcarifer Jałoszyński (a), V. qinghaiensis Jałoszyński (b), V. gansuana Jałoszyński (c), V. assingi Jałoszyński (d), V. modesta Jałoszyński (e), V. spinosa Jałoszyński (f), V. shaanxiana Jałoszyński (g), and V. dabashana Jałoszyński (h)." figureDoi="http://doi.org/10.5281/zenodo.13747575" httpUri="https://zenodo.org/record/13747575/files/figure.png" pageId="4" pageNumber="198">Fig. 11</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37572FEFFFADD0FFF5C7A8E20A060B8" box="[199,711,439,465]" pageId="4" pageNumber="198" type="etymology">
|
|||
|
|
<paragraph id="8BD02175FFFADD0FFF5C7A8E20A060B8" blockId="4.[151,1437,151,645]" box="[199,711,439,465]" pageId="4" pageNumber="198">
|
|||
|
|
<emphasis id="B91BFD67FFFADD0FFF5C7A8E232960B8" bold="true" box="[199,334,439,465]" pageId="4" pageNumber="198">Etymology.</emphasis>
|
|||
|
|
After
|
|||
|
|
<collectingRegion id="49ABEF97FFFADD0FFE0E7A8E203F60B8" box="[405,600,439,465]" country="China" name="Shaanxi" pageId="4" pageNumber="198">Shaanxi province</collectingRegion>
|
|||
|
|
in
|
|||
|
|
<collectingCountry id="F37861E5FFFADD0FFDE77A8E20A460B8" box="[636,707,439,465]" name="China" pageId="4" pageNumber="198">China</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
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<subSubSection id="C37572FEFFFADD0FFF5C7AE227FD63EC" pageId="4" pageNumber="198" type="discussion">
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<paragraph id="8BD02175FFFADD0FFF5C7AE227FD63EC" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="198">
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<emphasis id="B91BFD67FFFADD0FFF5C7AE2235C609C" bold="true" box="[199,315,475,501]" pageId="4" pageNumber="198">Remarks.</emphasis>
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<taxonomicName id="4C6F5AF6FFFADD0FFEDC7AE22051609C" authorityName="Jałoszyński" authorityYear="2024" box="[327,566,475,501]" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="shaanxiana">
|
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<emphasis id="B91BFD67FFFADD0FFEDC7AE22051609C" box="[327,566,475,501]" italics="true" pageId="4" pageNumber="198">Veraphis shaanxiana</emphasis>
|
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</taxonomicName>
|
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|
|
is one of a few species with a clearly asymmetrical apical region of the aedeagus. Among Chinese species, a strongly asymmetrical aedeagal apex can be found in
|
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|
<taxonomicName id="4C6F5AF6FFFADD0FFB1E7AC6276A6371" authorityName="Jaloszynski" authorityYear="2013" box="[1157,1293,511,536]" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="gansuana">
|
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|
|
<emphasis id="B91BFD67FFFADD0FFB1E7AC6276A6371" box="[1157,1293,511,536]" italics="true" pageId="4" pageNumber="198">V. gansuana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and slightly asymmetrical in
|
|||
|
|
<taxonomicName id="4C6F5AF6FFFADD0FFEC9791A206E6354" authorityName="Jałoszyński" authorityYear="2012" box="[338,521,547,573]" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="qinghaienasis">
|
|||
|
|
<emphasis id="B91BFD67FFFADD0FFEC9791A206E6354" box="[338,521,547,573]" italics="true" pageId="4" pageNumber="198">V. qinghaienasis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C6F5AF6FFFADD0FFDA4791A20CD6355" authorityName="Jaloszynski" authorityYear="2013" box="[575,682,547,572]" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="assingi">
|
|||
|
|
<emphasis id="B91BFD67FFFADD0FFDA4791A20CD6355" box="[575,682,547,572]" italics="true" pageId="4" pageNumber="198">V. assingi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Comparison of the aedeagi (
|
|||
|
|
<figureCitation id="13543DF0FFFADD0FFC6F791A26236354" box="[1012,1092,547,573]" captionStart="FIGURE 11" captionStartId="5.[151,250,1368,1392]" captionTargetBox="[151,1410,525,1345]" captionTargetId="figure-146@5.[151,1435,525,1345]" captionTargetPageId="5" captionText="FIGURE 11. Distribution of nominal species of Veraphis in China, with comparison of their aedeagi in ventral view (based on Google Maps). Veraphis calcarifer Jałoszyński (a), V. qinghaiensis Jałoszyński (b), V. gansuana Jałoszyński (c), V. assingi Jałoszyński (d), V. modesta Jałoszyński (e), V. spinosa Jałoszyński (f), V. shaanxiana Jałoszyński (g), and V. dabashana Jałoszyński (h)." figureDoi="http://doi.org/10.5281/zenodo.13747575" httpUri="https://zenodo.org/record/13747575/files/figure.png" pageId="4" pageNumber="198">Fig. 11</figureCitation>
|
|||
|
|
) demonstrates that they all are clearly different. An asymmetrical aedeagal apex can be also found in the Japanese
|
|||
|
|
<taxonomicName id="4C6F5AF6FFFADD0FFBA3797E27FF6308" authority="Jaloszynski, 2014" authorityName="Jaloszynski" authorityYear="2014" box="[1080,1432,583,609]" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="yoshitomii">
|
|||
|
|
<emphasis id="B91BFD67FFFADD0FFBA3797E26AF6308" box="[1080,1224,583,609]" italics="true" pageId="4" pageNumber="198">V. yoshitomii</emphasis>
|
|||
|
|
<bibRefCitation id="EFFE5C84FFFADD0FFB54797E27FF6308" author="Jaloszynski, P." box="[1231,1432,583,609]" pageId="4" pageNumber="198" pagination="589 - 595" refId="ref3095" refString="Jaloszynski, P. (2014) Veraphis yoshitomii sp. n. (Coleoptera, Staphylinidae, Scydmaeninae) from Japan. Zootaxa, 3860 (6), 589 - 595. https: // doi. org / 10.11646 / zootaxa. 3860.6.7" type="journal article" year="2014">Jałoszyński, 2014</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, but the general shapes of the aedeagi are different, and
|
|||
|
|
<taxonomicName id="4C6F5AF6FFFADD0FFC9A795221F663EC" authorityName="Jaloszynski" authorityYear="2014" box="[769,913,619,645]" class="Insecta" family="Staphylinidae" genus="Veraphis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="198" phylum="Arthropoda" rank="species" species="yoshitomii">
|
|||
|
|
<emphasis id="B91BFD67FFFADD0FFC9A795221F663EC" box="[769,913,619,645]" italics="true" pageId="4" pageNumber="198">V. yoshitomii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is much longer, stouter and darkly pigmented.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
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</document>
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